treatments-xml/data/03/B0/87/03B087B8FFB5A224DCDAFB82D5D618C2.xml
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<document id="D99D8D5AF1BCEB0E2C1718FD5175301B" ID-CLB-Dataset="26954" ID-DOI="10.5252/adansonia2019v41a4" ID-GBIF-Dataset="9dfbe7ab-f1ae-49ff-af7b-14dff6879995" ID-Zenodo-Dep="2653394" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556520005505" checkinUser="plazi" docAuthor="Salvator NTORE &amp; Olivier LACHENAUD" docDate="2019" docId="03B087B8FFB5A224DCDAFB82D5D618C2" docLanguage="en" docName="a2019v41a4.pdf" docOrigin="Adansonia 41 (4)" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Pauridiantha gracilipes Salvator NTORE &amp; Olivier LACHENAUD 2019, sp. nov." docType="treatment" docVersion="14" lastPageNumber="38" masterDocId="FF89FFC0FFBDA22FDC4FFFBED4401E0D" masterDocTitle="Two new species and a new combination in the genus Pauridiantha Hook. f. (Rubiaceae) from tropical Africa" masterLastPageNumber="40" masterPageNumber="29" pageNumber="35" updateTime="1698724171376" updateUser="ExternalLinkService">
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<mods:title id="7CFA63334EE2366CADCCDEE447E05026">Two new species and a new combination in the genus Pauridiantha Hook. f. (Rubiaceae) from tropical Africa</mods:title>
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<mods:name id="42DF4F8EED1E8204A4D440D22E7CA825" type="personal">
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<mods:namePart id="31AD0BAEE4B534CD85E887D198A2ACCE">Salvator NTORE</mods:namePart>
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<mods:namePart id="85228932728B747522EC935CDC328AA9">Olivier LACHENAUD</mods:namePart>
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<mods:title id="1C4FC74B2ABFDFD0CC0044A1B46A709A">Adansonia</mods:title>
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<mods:date id="169B653DE94B0573065818A324DBBF4A">2019</mods:date>
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<mods:number id="D32A3B5650CA743C729DB4C73841AC83">2019-04-08</mods:number>
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<heading id="D0EE81C2FFB5A227DCDAFB82D6B71A5A" box="[149,759,1084,1111]" centered="true" fontSize="11" level="3" pageId="8" pageNumber="35" reason="4">
<taxonomicName id="4C194D2DFFB5A227DCDAFB82D6D61A5A" ID-CoL="9V3DS" authority="O.Lachenaud &amp; Ntore" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[149,662,1084,1111]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="B96DEABCFFB5A227DCDAFB82D5D41A5A" bold="true" box="[149,404,1084,1111]" italics="true" pageId="8" pageNumber="35">Pauridiantha gracilipes</emphasis>
O.Lachenaud &amp; Ntore
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,
<taxonomicNameLabel id="A25E57C7FFB5A227DEEDFB83D6B71A5A" box="[674,759,1085,1111]" pageId="8" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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<heading id="D0EE81C2FFB5A227DD3CFBE2D6591A7A" box="[371,537,1116,1143]" centered="true" fontSize="11" level="3" pageId="8" pageNumber="35" reason="1">
(
<figureCitation id="13222A2BFFB5A227DD35FBE2D5B61A7A" box="[378,502,1116,1143]" captionStart="FIG" captionStartId="4.[132,143,1134,1152]" captionTargetBox="[151,1437,215,1096]" captionTargetId="figure@4.[793,1437,203,665]" captionTargetPageId="4" captionText="FIG. 1. — Seeds of Pauridiantha chlorantha (K.Schum.) Ntore &amp; O.Lachenaud, comb. nov., from Jannerup &amp; Mhoro 129 (A, B) and P. gracilipes O.Lachenaud &amp; Ntore, sp. nov., from Lachenaud et al. 1981 (C, D) in electron microscopy: A, C, entire seed; B, detail of exotesta cells with outer tangential wall removed, showing thickened radial walls with small pits, and large pits in the inner tangential walls; D, detail of exotesta cells with outer tangential wall removed, showing thin radial walls with small pits, and large pits in the inner tangential walls. Scale bars: A, C, 100 µm; B, D, 10 µm." httpUri="https://zenodo.org/record/2653396/files/figure.png" pageId="8" pageNumber="35">Figs 1C, D</figureCitation>
;
<figureCitation id="13222A2BFFB5A227DE4DFBE3D6501A7A" box="[514,528,1117,1143]" captionStart="FIG" captionStartId="6.[132,143,1937,1955]" captionTargetBox="[162,1432,215,1889]" captionTargetId="figure@6.[139,1447,207,1908]" captionTargetPageId="6" captionText="FIG. 3. — Pauridiantha gracilipes O.Lachenaud &amp; Ntore,sp.nov.: A, fruiting stem; B, portion of old leafless stem, with spines formed by the persistent basal part of old twigs;C, detail of lower leaf surface,showing reticulate venation;D, detail of lower surface showing a domatia;E, node with paired inflorescences and stipule;F, calyx and disk,seen from above; G, flower; H, infructescence with one fruit and two undeveloped persistent ovaries; I, cross-section of fruit,basal part; J, cross-section of fruit, apical part. All details from Lachenaud et al. 1981. Scale bars: A, 4 cm; B, 2 cm; C, E, I, J, 5 mm; D, 1 mm; F, G, 2 mm; H, 1 cm. Illustration: Antonio Fernandez." httpUri="https://zenodo.org/record/2653398/files/figure.png" pageId="8" pageNumber="35">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB5A227DCCBFB22D5EC1BAF" pageId="8" pageNumber="35" type="diagnosis">
<paragraph id="8BA636AEFFB5A227DCCBFB22D5EC1BAF" blockId="8.[132,776,1180,1442]" pageId="8" pageNumber="35">
<emphasis id="B96DEABCFFB5A227DCCBFB22D6AC1B0E" italics="true" pageId="8" pageNumber="35">Stipulis breviter triangularibus quam ramis angustioribus, ramis gracilibus glabris vel subglabris, foliis domatiis pubescentibus exceptis glaberrimis subtus dense et conspicue reticulatis, seminibus sublaevibus (nec prominente reticulatis ut in omnibus alteribus congeneribus)</emphasis>
P. liebrechtsianae
<emphasis id="B96DEABCFFB5A227DD50FAB8D5EC1BAF" italics="true" pageId="8" pageNumber="35">
(De Wild. &amp; T.Durand) Ntore &amp; Dessein affinis, sed pedicellis fructiferis
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longis (nec
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), ovario basi 2-loculare et apice 4-loculare (nec basi 3-4-loculare et apice 6-8-loculare), inflorescentiis 2(-3)-floris et umbelliformibus (nec 3-9-floris et saepe racemosis) caulibusque nodis inferioribus saepe spinosis (nec inermibus) differt.
</emphasis>
</paragraph>
</subSubSection>
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TYPUS. —
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.
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,
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Région du Lac Alombié, ±
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au nord de Mpaga
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,
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,
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<collectorName id="26EC5378FFB5A227DE39FA64D69A1BFF" box="[630,730,1498,1522]" pageId="8" pageNumber="35">Lachenaud</collectorName>
,
<collectorName id="26EC5378FFB5A227DEADFA64D4FA1800" pageId="8" pageNumber="35">Ikabanga</collectorName>
&amp;
<collectorName id="26EC5378FFB5A227DC99FA4BD55B1800" box="[214,283,1525,1549]" pageId="8" pageNumber="35">Lafferty</collectorName>
<collectingDate id="EFE3E986FFB5A227DD50FA4BD5101800" box="[287,336,1525,1549]" pageId="8" pageNumber="35">1981</collectingDate>
</emphasis>
(
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,
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!; iso-,
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!; P!;
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!)
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.
</paragraph>
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. —
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<emphasis id="B96DEABCFFB5A227DD5EF9FBD5171850" bold="true" box="[273,343,1605,1629]" pageId="8" pageNumber="35">Gabon</emphasis>
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Préfecture
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, à environ
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au nord
<collectorName id="26EC5378FFB5A227DCF1F9DED5291875" box="[190,361,1632,1656]" pageId="8" pageNumber="35">du Lac Alombié</collectorName>
,
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<collectorName id="26EC5378FFB5A227DEA7F9DFD4F5189F" pageId="8" pageNumber="35">Bidault</collectorName>
&amp;
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1726
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(
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[
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!;
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n.v.)
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;
<emphasis id="B96DEABCFFB5A227DC88F92BD4B518A0" box="[199,245,1685,1709]" italics="true" pageId="8" pageNumber="35">ibid.</emphasis>
,
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,
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</collectingDate>
,
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<collectorName id="26EC5378FFB5A227DE28F92BD6F218A0" box="[615,690,1685,1709]" pageId="8" pageNumber="35">Bidault</collectorName>
,
<collectorName id="26EC5378FFB5A227DEF1F92BD74318A1" box="[702,771,1685,1708]" pageId="8" pageNumber="35">Stévart</collectorName>
,
<collectorName id="26EC5378FFB5A227DCCBF90ED49518CA" box="[132,213,1712,1735]" pageId="8" pageNumber="35">Nguema</collectorName>
&amp;
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1735
</emphasis>
(
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[
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!];
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!;
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n.v.);
</materialsCitation>
<materialsCitation id="3B713CF3FFB5A227DCA0F974D633191A" ID-GBIF-Occurrence="2237036577" collectingDate="2014-10-25" collectingDateMax="2014-10-25" collectingDateMin="2014-10-15" collectionCode="BR, BRLU, LBV" collectorName="Zebaze &amp; Barbier &amp; Niangadouma &amp; Stevart" country="Gabon" latitude="-0.8183333" location="nord du Lac Alombie" longLatPrecision="21" longitude="9.456111" municipality="Prefecture de Mpaga" pageId="8" pageNumber="35" specimenCode="BR0000016173910" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BC2ACD4FFB5A227DCA0F974D5F918EF" box="[239,441,1738,1762]" pageId="8" pageNumber="35">Préfecture de Mpaga</collectingMunicipality>
,
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,
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,
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<date id="FFA7106EFFB5A227DCB5F95AD52718F1" box="[250,359,1764,1788]" pageId="8" pageNumber="35" value="2014-10-25">25.X.2014</date>
</collectingDate>
,
<emphasis id="B96DEABCFFB5A227DD3CF95AD4EA191A" italics="true" pageId="8" pageNumber="35">
<collectorName id="26EC5378FFB5A227DD3CF95AD5F918F1" box="[371,441,1764,1788]" pageId="8" pageNumber="35">Zébazé</collectorName>
,
<collectorName id="26EC5378FFB5A227DD8BF95AD64D18F1" box="[452,525,1764,1788]" pageId="8" pageNumber="35">Barbier</collectorName>
,
<collectorName id="26EC5378FFB5A227DE56F95AD6E118F1" box="[537,673,1764,1788]" pageId="8" pageNumber="35">Niangadouma</collectorName>
&amp;
<collectorName id="26EC5378FFB5A227DE8AF95BD74718F1" box="[709,775,1765,1788]" pageId="8" pageNumber="35">Stévart</collectorName>
419
</emphasis>
(
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[
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!];
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!;
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n.v.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB5A227DCCBF88AD746196A" pageId="8" pageNumber="35" type="etymology">
<paragraph id="8BA636AEFFB5A227DCCBF88AD746196A" blockId="8.[132,775,1844,1895]" pageId="8" pageNumber="35">ETYMOLOGY. — The specific name refers to the particularly long and slender fruiting pedicels, a diagnostic character of the species.</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB5A227DCCBF83BD52519E0" pageId="8" pageNumber="35" type="distribution">
<paragraph id="8BA636AEFFB5A227DCCBF83BD52519E0" blockId="8.[132,775,1925,2029]" pageId="8" pageNumber="35">
DISTRIBUTION. — Lower
<collectingCountry id="F30E763EFFB5A227DD34F83BD5821990" box="[379,450,1925,1949]" name="Guinea-Bissau" pageId="8" pageNumber="35">Guinea</collectingCountry>
subcentre of endemism. Endemic to the lower Ogooué basin in
<collectingCountry id="F30E763EFFB5A227DDD5F81ED59C19B5" box="[410,476,1952,1976]" name="Gabon" pageId="8" pageNumber="35">Gabon</collectingCountry>
(
<figureCitation id="13222A2BFFB5A227DDA6F81ED65E19B5" box="[489,542,1952,1976]" captionStart="FIG" captionStartId="9.[225,236,751,769]" captionTargetBox="[291,1321,203,721]" captionTargetId="figure@9.[346,973,213,701]" captionTargetPageId="9" captionText="FIG. 4. — Distribution of Pauridiantha gracilipes O.Lachenaud &amp; Ntore, sp. nov. () and P. principensis Ntore &amp; O.Lachenaud, sp. nov. ()." httpUri="https://zenodo.org/record/2653647/files/figure.png" pageId="8" pageNumber="35">Fig. 4</figureCitation>
), where it is only known from a small area north of Lake Alombié; locally abundant in its highly restricted range.
</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB5A227DF62FC6BD0C81B7A" pageId="8" pageNumber="35" type="biology_ecology">
<paragraph id="8BA636AEFFB5A227DF62FC6BD0D21B26" blockId="8.[813,1456,981,1323]" pageId="8" pageNumber="35">
HABITAT. — Restricted to seasonally flooded swamp forests dominated by
<taxonomicName id="4C194D2DFFB5A227DFCDFC4ED0F31A05" authority="Mildbr." authorityName="Mildbr." box="[898,1203,1008,1032]" class="Magnoliopsida" family="Ctenolophonaceae" genus="Ctenolophon" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="englerianus">
<emphasis id="B96DEABCFFB5A227DFCDFC4ED0221A05" box="[898,1122,1008,1032]" italics="true" pageId="8" pageNumber="35">Ctenolophon englerianus</emphasis>
Mildbr.
</taxonomicName>
,
<taxonomicName id="4C194D2DFFB5A227D8F5FC4ED0621A2E" authority="(De Wild.) Prance &amp; F. White" authorityName="Prance &amp; F. White" baseAuthorityName="De Wild." class="Magnoliopsida" family="Chrysobalanaceae" genus="Dactyladenia" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="eketensis">
<emphasis id="B96DEABCFFB5A227D8F5FC4ED1C51A05" box="[1210,1413,1008,1032]" italics="true" pageId="8" pageNumber="35">Dactyladenia eketensis</emphasis>
(De Wild.) Prance &amp; F.White
</taxonomicName>
and
<taxonomicName id="4C194D2DFFB5A227D81EFBB5D12D1A2E" authority="Baill." authorityName="Baill." box="[1105,1389,1035,1059]" class="Magnoliopsida" family="Euphorbiaceae" genus="Anthostema" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="aubryanum">
<emphasis id="B96DEABCFFB5A227D81EFBB5D1711A2E" box="[1105,1329,1035,1059]" italics="true" pageId="8" pageNumber="35">Anthostema aubryanum</emphasis>
Baill.
</taxonomicName>
, where it is locally dominant in the undergrowth and forms dense thickets. This peculiar habitat is described by
<bibRefCitation id="EF884B5FFFB5A227D8C4FBFED1E91A55" author="VANDEWEGHE J. - P. &amp; STEVART T." box="[1163,1449,1088,1112]" journalOrPublisher="Agence nationale des Parcs nationaux, Libreville" pageId="8" pageNumber="35" refId="ref9434" refString="VANDEWEGHE J. - P. &amp; STEVART T. 2017. - Le delta de l'Ogooue. Agence nationale des Parcs nationaux, Libreville." title="Le delta de l'Ogooue" type="book" year="2017">Vandeweghe &amp; Stévart (2017</bibRefCitation>
: 207-209) as “forêt inondable monodominante” (although, in fact,
<taxonomicName id="4C194D2DFFB5A227DF62FBCAD7E51A81" box="[813,933,1140,1164]" class="Magnoliopsida" family="Ctenolophonaceae" genus="Ctenolophon" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEABCFFB5A227DF62FBCAD7E51A81" box="[813,933,1140,1164]" italics="true" pageId="8" pageNumber="35">Ctenolophon</emphasis>
</taxonomicName>
is not always strictly monodominant) and, from an analysis of satellite images, is estimated to cover around
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a restricted area north of Lake Alombié.
<taxonomicName id="4C194D2DFFB5A227D8FAFB17D16B1ACC" box="[1205,1323,1193,1217]" class="Magnoliopsida" family="Ctenolophonaceae" genus="Ctenolophon" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEABCFFB5A227D8FAFB17D16B1ACC" box="[1205,1323,1193,1217]" italics="true" pageId="8" pageNumber="35">Ctenolophon</emphasis>
</taxonomicName>
is believed to have been a key species for the maintenance of forest micro-refugia in the Ogooué delta during dry periods, due to its unusual ability to grow both on well drained and seasonally flooded soils (
<bibRefCitation id="EF884B5FFFB5A227D9CFFB47D05E1B26" author="VANDEWEGHE J. - P. &amp; STEVART T." journalOrPublisher="Agence nationale des Parcs nationaux, Libreville" pageId="8" pageNumber="35" refId="ref9434" refString="VANDEWEGHE J. - P. &amp; STEVART T. 2017. - Le delta de l'Ogooue. Agence nationale des Parcs nationaux, Libreville." title="Le delta de l'Ogooue" type="book" year="2017">Vandeweghe &amp; Stévart 2017</bibRefCitation>
: 159, 177).
</paragraph>
<paragraph id="8BA636AEFFB5A227DF62FAFBD0C81B7A" blockId="8.[813,1455,1349,1400]" pageId="8" pageNumber="35">PHENOLOGY. — Fruits (not fully mature) in October, corresponding to the onset of the rainy season.</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB5A226DF62FA2FD66E1A7A" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="conservation">
<paragraph id="8BA636AEFFB5A227DF62FA2FD1F01966" blockId="8.[813,1456,1425,1899]" pageId="8" pageNumber="35">
PRELIMINARY CONSERVATION ASSESSMENT. — Critically Endangered [CR B1ab(iii)+B2ab(iii)]. The extent of occurrence (EOO) of
<taxonomicName id="4C194D2DFFB5A227D9CCFA13D08A1BD3" authority="O.Lachenaud &amp; Ntore" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="B96DEABCFFB5A227D9CCFA13D7A01BD3" italics="true" pageId="8" pageNumber="35">Pauridiantha gracilipes</emphasis>
O.Lachenaud &amp; Ntore
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB5A227D89CFA79D15E1BD3" box="[1235,1310,1479,1502]" pageId="8" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
is estimated to be
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2, and its area of occupancy (AOO) to be
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², both values being within the limit for Critically Endangered status under subcriteria B1 and B2. The species is endemic to the coastal part of the Ogooué delta in
<collectingCountry id="F30E763EFFB5A227DFBBF98ED07A1845" box="[1012,1082,1584,1608]" name="Gabon" pageId="8" pageNumber="35">Gabon</collectingCountry>
, and occurs in
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<emphasis id="B96DEABCFFB5A227D885F98ED1EF1845" box="[1226,1455,1584,1608]" italics="true" pageId="8" pageNumber="35">Ctenolophon englerianus</emphasis>
</taxonomicName>
seasonally flooded forests, a highly localized habitat, which is estimated to cover around
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the delta (
<bibRefCitation id="EF884B5FFFB5A227D89CF9DBD7201895" author="VANDEWEGHE J. - P. &amp; STEVART T." journalOrPublisher="Agence nationale des Parcs nationaux, Libreville" pageId="8" pageNumber="35" refId="ref9434" refString="VANDEWEGHE J. - P. &amp; STEVART T. 2017. - Le delta de l'Ogooue. Agence nationale des Parcs nationaux, Libreville." title="Le delta de l'Ogooue" type="book" year="2017">Vandeweghe &amp; Stévart 2017</bibRefCitation>
: 207-209). It is known from four specimens representing four subpopulations, all situated outside protected areas (although its presence in the adjacent Wonga-Wongué National Park is highly probable). Though its habitat currently suffers little from human activities, forest exploitation has impacted it in the past and may do so again in the future; a decline in the quality of the habitat may therefore be expected. The four subpopulations represent one location according to
<bibRefCitation id="EF884B5FFFB5A227D84AF887D0CF195C" author="IUCN" box="[1029,1167,1849,1873]" journalOrPublisher="Gland, Switzerland and Cambridge, UK" pageId="8" pageNumber="35" refId="ref8981" refString="IUCN. 2012. - IUCN Red List Categories and Criteria: Version 3.1. Second edition. Gland, Switzerland and Cambridge, UK." title="IUCN Red List Categories and Criteria: Version 3.1. Second edition" type="book" year="2012">IUCN (2012)</bibRefCitation>
, and the species is evaluated as Critically Endangered according to the conditions B1ab(iii)+B2ab(iii).
</paragraph>
<paragraph id="8BA636AEFFB5A226DF62F83BD66E1A7A" blockId="8.[813,1455,1925,2029]" lastBlockId="9.[132,775,855,1143]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
AFFINITIES.
<emphasis id="B96DEABCFFB5A227DFE6F838D7831990" box="[937,963,1926,1949]" italics="true" pageId="8" pageNumber="35"></emphasis>
<taxonomicName id="4C194D2DFFB5A227DF84F83BD1EB1990" authority="O.Lachenaud &amp; Ntore" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[971,1451,1925,1949]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="B96DEABCFFB5A227DF84F83BD0F21990" box="[971,1202,1925,1949]" italics="true" pageId="8" pageNumber="35">Pauridiantha gracilipes</emphasis>
O.Lachenaud &amp; Ntore
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB5A227DF62F81ED73919BA" box="[813,889,1952,1975]" pageId="8" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
appears to be most closely related to
<taxonomicName id="4C194D2DFFB5A227D8A3F81ED08719DF" authority="(De Wild. &amp; T. Durand) Ntore &amp; Dessein. The" authorityName="Ntore &amp; Dessein. The" baseAuthorityName="De Wild. &amp; T. Durand" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="35" phylum="Tracheophyta" rank="species" species="liebrechtsiana">
<emphasis id="B96DEABCFFB5A227D8A3F81ED1C219B5" box="[1260,1410,1952,1976]" italics="true" pageId="8" pageNumber="35">P. liebrechtsiana</emphasis>
(De Wild. &amp; T.Durand) Ntore &amp; Dessein. The
</taxonomicName>
two species share nearly glabrous vegetative parts, stipules narrower than the twigs, leaves with hairy domatia and densely reticulate tertiary veins, and, most importantly, a nearly smooth seed surface (in all other species of the genus the margins of the exotestal cells are markedly raised). It is also related, though more distantly, to
<taxonomicName id="4C194D2DFFB4A226DE62FC19D5681DD5" authority="(N. Halle) Ntore &amp; Dessein" authorityName="Ntore &amp; Dessein" baseAuthorityName="N. Halle" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="letestuana">
<emphasis id="B96DEABCFFB4A226DE62FC19D6E11DB3" box="[557,673,934,958]" italics="true" pageId="9" pageNumber="36">P. letestuana</emphasis>
(N.Hallé) Ntore &amp; Dessein
</taxonomicName>
and
<taxonomicName id="4C194D2DFFB4A226DD1AFC7FD6DB1DD5" authority="Ntore &amp; Dessein. The" authorityName="Ntore &amp; Dessein. The" authorityYear="2003" box="[341,667,960,985]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="smetsiana">
<emphasis id="B96DEABCFFB4A226DD1AFC7FD5841DD5" box="[341,452,961,984]" italics="true" pageId="9" pageNumber="36">P. smetsiana</emphasis>
Ntore &amp; Dessein. The
</taxonomicName>
differences between these four species are shown in
<tableCitation id="C69B0315FFB4A226DE4BFC65D60D1DFE" box="[516,589,987,1011]" captionStart="TABLE" captionStartId="8.[347,358,221,239]" captionTargetBox="[134,1455,297,920]" captionTargetId="graphics@8.[132,1455,596,620]" captionTargetPageId="8" captionText="TABLE 1. — Differences between Pauridiantha gracilipes O.Lachenaud &amp; Ntore, sp. nov. and related species." httpUri="http://table.plazi.org/id/DF666626FFB5A227DD14FF63D09A1EE2" pageId="9" pageNumber="36" tableUuid="DF666626FFB5A227DD14FF63D09A1EE2">Table 1</tableCitation>
. Remarkably, three of them are endemic to west-central
<collectingCountry id="F30E763EFFB4A226DDA1FC4BD6741A00" box="[494,564,1013,1037]" name="Gabon" pageId="9" pageNumber="36">Gabon</collectingCountry>
; only
<taxonomicName id="4C194D2DFFB4A226DE3EFC48D7461A00" box="[625,774,1013,1037]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="liebrechtsiana">
<emphasis id="B96DEABCFFB4A226DE3EFC48D7461A00" box="[625,774,1013,1037]" italics="true" pageId="9" pageNumber="36">P. liebrechtsiana</emphasis>
</taxonomicName>
is more widespread, occurring in seasonally flooded forests both in the
<collectingCountry id="F30E763EFFB4A226DCE6FB94D4AC1A4F" box="[169,236,1066,1090]" name="Democratic Republic of the Congo" pageId="9" pageNumber="36">Congo</collectingCountry>
and the Ogooué basins, its westernmost known locality being Lake Gomé in
<collectingCountry id="F30E763EFFB4A226DD18FBFBD5DD1A50" box="[343,413,1093,1117]" name="Gabon" pageId="9" pageNumber="36">Gabon</collectingCountry>
,
<emphasis id="B96DEABCFFB4A226DDE7FBFBD5F71A51" box="[424,439,1093,1116]" italics="true" pageId="9" pageNumber="36">c.</emphasis>
<quantity id="4CE19B4BFFB4A226DDF1FBFBD64B1A50" box="[446,523,1093,1117]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="36" unit="km" value="120.0">120 km</quantity>
northeast of the range of
<taxonomicName id="4C194D2DFFB4A226DCCBFBDED5991A7A" authority="O.Lachenaud &amp; Ntore" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[132,473,1119,1143]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="B96DEABCFFB4A226DCCBFBDED4AE1A7A" box="[132,238,1119,1143]" italics="true" pageId="9" pageNumber="36">P. gracilipes</emphasis>
O.Lachenaud &amp; Ntore
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB4A226DDACFBDED66E1A7A" box="[483,558,1120,1143]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF666626FFB4A226DCAEFD51D1141D0C" box="[225,1364,751,769]" httpUri="https://zenodo.org/record/2653647/files/figure.png" pageId="9" pageNumber="36" startId="9.[225,236,751,769]" targetBox="[291,1321,203,721]" targetPageId="9">
<paragraph id="8BA636AEFFB4A226DCAEFD51D1141D0C" blockId="9.[225,1364,751,769]" box="[225,1364,751,769]" pageId="9" pageNumber="36">
FIG. 4. — Distribution of
<taxonomicName id="4C194D2DFFB4A226DDE4FD51D7631D0C" authority="O.Lachenaud &amp; Ntore" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[427,803,751,769]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="B96DEABCFFB4A226DDE4FD51D6261D0C" box="[427,614,751,769]" italics="true" pageId="9" pageNumber="36">Pauridiantha gracilipes</emphasis>
O.Lachenaud &amp; Ntore
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB4A226DF65FD51D72D1D0C" box="[810,877,751,769]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
() and
<taxonomicName id="4C194D2DFFB4A226DFFCFD51D0A51D0C" authority="Ntore &amp; O.Lachenaud" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[947,1253,751,769]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="principensis" status="sp. nov.">
<emphasis id="B96DEABCFFB4A226DFFCFD51D06A1D0C" box="[947,1066,751,769]" italics="true" pageId="9" pageNumber="36">P. principensis</emphasis>
Ntore &amp; O.Lachenaud
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB4A226D8A2FD51D1701D0C" box="[1261,1328,751,769]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
().
</paragraph>
</caption>
<subSubSection id="C3036525FFB4A224DCCBFB2BD5BB1ABD" lastPageId="11" lastPageNumber="38" pageId="9" pageNumber="36" type="description">
<paragraph id="8BA636AEFFB4A226DCCBFB2BD5551AA3" blockId="9.[132,775,1173,1392]" box="[132,277,1173,1199]" pageId="9" pageNumber="36">DESCRIPTION</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBFB08D5701ADD" blockId="9.[132,775,1173,1392]" box="[132,304,1206,1232]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DCCBFB08D5701ADD" box="[132,304,1206,1232]" italics="true" pageId="9" pageNumber="36">Lianescent shrub</emphasis>
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBFB68D69B1B7D" blockId="9.[132,775,1173,1392]" pageId="9" pageNumber="36">
To
<quantity id="4CE19B4BFFB4A226DCE5FB68D4AF1AFD" box="[170,239,1238,1264]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="9" pageNumber="36" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3-4 m</quantity>
tall, with drooping branches bearing divaricate, ± horizontal flowering twigs; older stems sometimes bearing opposite spines,
<quantity id="4CE19B4BFFB4A226DD75FAA8D5F21B3D" box="[314,434,1302,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="9" pageNumber="36" unit="cm" value="1.25" valueMax="1.7" valueMin="0.8">0.8-1.7 cm</quantity>
long, formed by the persistent base of twigs; twigs cylindrical,
<quantity id="4CE19B4BFFB4A226DD93FA8BD61F1B5D" box="[476,607,1333,1360]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="9" pageNumber="36" unit="mm" value="1.1" valueMax="1.5" valueMin="0.7">0.7-1.5 mm</quantity>
thick, glabrous or with sparse minute appressed hairs towards the apex.
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBFA2BD4931BA2" blockId="9.[132,775,1429,1551]" box="[132,211,1429,1455]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DCCBFA2BD4931BA2" box="[132,211,1429,1455]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DCCBFA2BD4931BA2" box="[132,211,1429,1455]" italics="true" pageId="9" pageNumber="36">Stipules</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBFA0BD66A1802" blockId="9.[132,775,1429,1551]" pageId="9" pageNumber="36">
Tardily caducous, narrowly triangular, 1.3-2 ×
<quantity id="4CE19B4BFFB4A226DE32FA0BD7431BC2" box="[637,771,1461,1488]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="9" pageNumber="36" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5-0.7 mm</quantity>
, ± thickened, narrower than the twig and slightly diverging from it, sparsely appressed-puberulous.
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBF98BD4861843" blockId="9.[132,774,1589,1679]" box="[132,198,1589,1614]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DCCBF98BD4861843" box="[132,198,1589,1614]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DCCBF98BD4861843" box="[132,198,1589,1614]" italics="true" pageId="9" pageNumber="36">Leaves</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBF9EAD5651883" blockId="9.[132,774,1589,1679]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DCCBF9EAD48F1863" bold="true" box="[132,207,1620,1646]" pageId="9" pageNumber="36">Petiole</emphasis>
.
<quantity id="4CE19B4BFFB4A226DC91F9EAD5151862" box="[222,341,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="9" pageNumber="36" unit="cm" value="0.55" valueMax="0.7" valueMin="0.4">0.4-0.7 cm</quantity>
long, canaliculate, shortly pubescent on the upper side.
</paragraph>
<paragraph id="8BA636AEFFB4A226DCCBF90DD5A119E0" blockId="9.[132,776,1715,2030]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DCCBF90DD4BE18C0" bold="true" box="[132,254,1715,1741]" pageId="9" pageNumber="36">Leaf-blade.</emphasis>
Elliptic, 5.5-10.8 ×
<quantity id="4CE19B4BFFB4A226DD95F90AD61418C2" box="[474,596,1716,1743]" metricMagnitude="-2" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="4.1" metricValueMin="2.1" pageId="9" pageNumber="36" unit="cm" value="3.0999999999999996" valueMax="4.1" valueMin="2.1">2.1-4.1 cm</quantity>
, acute to rounded at base, narrowly acuminate at apex with acumen
<quantity id="4CE19B4BFFB4A226DEF5F96AD74718E3" box="[698,775,1748,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="36" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 cm</quantity>
long, papyraceous, glabrous except the underside of the nerves very sparsely appressed-puberulous, dark green above and pale green below, drying reddish on both sides; midrib concave above; secondary veins 5-7 pairs, strongly curved and brochidodromous, forming loops
<quantity id="4CE19B4BFFB4A226DDA9F8CDD60F1983" box="[486,591,1907,1934]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="9" pageNumber="36" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
from the margin; tertiary veins reticulate, conspicuous and dark green in life on the lower side of the leaf, forming a dense network with areolae
<quantity id="4CE19B4BFFB4A226DC99F86DD51819E3" box="[214,344,2003,2030]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="9" pageNumber="36" unit="mm" value="1.1" valueMax="1.5" valueMin="0.7">0.7-1.5 mm</quantity>
in diameter.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FCE8D06B1D9C" blockId="9.[813,1455,854,913]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62FCE8D7D51D7D" bold="true" box="[813,917,854,880]" pageId="9" pageNumber="36">Domatia.</emphasis>
Present in the main axils but not very conspicuous, crypt-like, ± pubescent.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FC09D7F31DDC" blockId="9.[813,1455,951,1072]" box="[813,947,951,977]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DF62FC09D7F31DDC" box="[813,947,951,977]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DF62FC09D7F31DDC" box="[813,947,951,977]" italics="true" pageId="9" pageNumber="36">Inflorescences</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FC68D14A1A3D" blockId="9.[813,1455,951,1072]" pageId="9" pageNumber="36">
Axillary, solitary in each axil, in contracted cymes, (2-)3-flowered,
<emphasis id="B96DEABCFFB4A226DF25FC49D73B1A1D" box="[874,891,1015,1040]" italics="true" pageId="9" pageNumber="36">c.</emphasis>
<quantity id="4CE19B4BFFB4A226DFCCFC49D7901A1C" box="[899,976,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="36" unit="cm" value="0.8">0.8 cm</quantity>
long in the flowering stage and
<quantity id="4CE19B4BFFB4A226D979FC48D1EF1A1C" box="[1334,1455,1014,1041]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="9" pageNumber="36" unit="cm" value="2.15" valueMax="2.5" valueMin="1.8">1.8-2.5 cm</quantity>
long in fruit, sparsely appressed-puberulous.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FBE8D0601A9D" blockId="9.[813,1455,1110,1169]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62FBE8D7D61A7D" bold="true" box="[813,918,1110,1136]" pageId="9" pageNumber="36">Peduncle.</emphasis>
<quantity id="4CE19B4BFFB4A226DFD3FBE8D0501A7C" box="[924,1040,1110,1137]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="9" pageNumber="36" unit="cm" value="0.4" valueMax="0.6" valueMin="0.2">0.2-0.6 cm</quantity>
long, with a pair of prophylls
<emphasis id="B96DEABCFFB4A226D909FBE9D1171A7D" box="[1350,1367,1111,1136]" italics="true" pageId="9" pageNumber="36">c.</emphasis>
<quantity id="4CE19B4BFFB4A226D913FBE8D1EF1A7C" box="[1372,1455,1110,1137]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="36" unit="mm" value="0.5">0.5 mm</quantity>
long in the upper half.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FB0BD0561ADD" blockId="9.[813,1046,1205,1232]" box="[813,1046,1205,1232]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DF62FB0BD0561ADD" box="[813,1046,1205,1232]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DF62FB0BD7FC1AC2" bold="true" box="[813,956,1205,1231]" pageId="9" pageNumber="36">Floral bracts.</emphasis>
Absent.
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FB4BD73A1B02" blockId="9.[813,922,1269,1328]" box="[813,890,1269,1295]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DF62FB4BD73A1B02" box="[813,890,1269,1295]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DF62FB4BD73A1B02" box="[813,890,1269,1295]" italics="true" pageId="9" pageNumber="36">Flowers</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FAABD7DA1B3D" blockId="9.[813,922,1269,1328]" box="[813,922,1301,1328]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DF62FAABD7DA1B3D" box="[813,922,1301,1328]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">5-merous.</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FAEBD1311B62" blockId="9.[813,1393,1365,1392]" box="[813,1393,1365,1392]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62FAEBD7C11B62" bold="true" box="[813,897,1365,1391]" pageId="9" pageNumber="36">Pedicel.</emphasis>
<quantity id="4CE19B4BFFB4A226DFC6FAEBD7A11B7D" box="[905,993,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="9" pageNumber="36" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
long, sparsely appressed-puberulous.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62FA2AD0DE1BE2" blockId="9.[813,1455,1428,1519]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62FA2AD7331BA3" bold="true" box="[813,883,1428,1454]" pageId="9" pageNumber="36">Calyx.</emphasis>
Pale green, cupuliform,
<emphasis id="B96DEABCFFB4A226D8CEFA28D0D11BA2" box="[1153,1169,1430,1455]" italics="true" pageId="9" pageNumber="36">c.</emphasis>
<quantity id="4CE19B4BFFB4A226D8D6FA2BD0B01BBD" box="[1177,1264,1429,1456]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="36" unit="mm" value="0.7">0.7 mm</quantity>
long, divided for 1/3 to 1/2 of its length, the lobes broadly triangular and acute at apex, puberulous on both sides.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62F9AAD7841883" blockId="9.[813,1455,1556,1679]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62F9AAD02B1823" bold="true" box="[813,1131,1556,1582]" pageId="9" pageNumber="36">Corolla (only one seen, dry).</emphasis>
With tube 1.5 ×
<quantity id="4CE19B4BFFB4A226D964F9AAD1C51822" box="[1323,1413,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="36" unit="mm" value="1.5">1.5 mm</quantity>
, inflated near the base, glabrous outside and densely hairy in the throat, and lobes triangular, 1.5
<emphasis id="B96DEABCFFB4A226D8C3F9EAD0DA1863" bold="true" box="[1164,1178,1620,1646]" pageId="9" pageNumber="36">×</emphasis>
<quantity id="4CE19B4BFFB4A226D8EDF9EAD0A71862" box="[1186,1255,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="36" unit="mm" value="1.0">1 mm</quantity>
, reflexed, glabrous on both sides.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62F90DD05518C2" blockId="9.[813,1045,1715,1743]" box="[813,1045,1715,1743]" pageId="9" pageNumber="36">
<heading id="D0EE81C2FFB4A226DF62F90DD05518C2" box="[813,1045,1715,1743]" fontSize="11" level="4" pageId="9" pageNumber="36" reason="1">
<emphasis id="B96DEABCFFB4A226DF62F90DD7E618C0" bold="true" box="[813,934,1715,1741]" pageId="9" pageNumber="36">Floral bud.</emphasis>
Not seen.
</heading>
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62F94DD10C1903" blockId="9.[813,1356,1779,1806]" box="[813,1356,1779,1806]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62F94DD7B21900" bold="true" box="[813,1010,1779,1805]" pageId="9" pageNumber="36">Stamens and style</emphasis>
. Not seen, apparently included.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62F88DD0C51980" blockId="9.[813,1455,1843,1934]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62F88DD7281940" bold="true" box="[813,872,1843,1869]" pageId="9" pageNumber="36">Disk.</emphasis>
<emphasis id="B96DEABCFFB4A226DF20F88AD7C91943" box="[879,905,1844,1870]" italics="true" pageId="9" pageNumber="36">C.</emphasis>
<quantity id="4CE19B4BFFB4A226DFDFF88DD7A61943" box="[912,998,1843,1870]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="36" unit="mm" value="0.5">0.5 mm</quantity>
high, hemispherical with 5 lateral grooves and 10 apical fossulae (formed by the imprints of the filaments and anthers), whitish, papillose.
</paragraph>
<paragraph id="8BA636AEFFB4A226DF62F80DD0F219E0" blockId="9.[813,1455,1971,2030]" pageId="9" pageNumber="36">
<emphasis id="B96DEABCFFB4A226DF62F80DD73619C0" bold="true" box="[813,886,1971,1997]" pageId="9" pageNumber="36">Ovary.</emphasis>
2-locular below, 4-locular by false septa in the upper part, sparsely appressed-puberulous.
</paragraph>
<caption id="DF666626FFB7A225DCCBF812D0D919E6" httpUri="https://zenodo.org/record/2653400/files/figure.png" pageId="10" pageNumber="37" startId="10.[132,143,1964,1982]" targetBox="[279,1312,215,1900]" targetPageId="10">
<paragraph id="8BA636AEFFB7A225DCCBF812D0D919E6" blockId="10.[132,1455,1964,2027]" pageId="10" pageNumber="37">
FIG. 5. —
<taxonomicName id="4C194D2DFFB7A225DC9AF812D62219B3" authority="Ntore &amp; O.Lachenaud" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[213,610,1964,1982]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="37" phylum="Tracheophyta" rank="species" species="principensis" status="sp. nov.">
<emphasis id="B96DEABCFFB7A225DC9AF812D5E619B3" box="[213,422,1964,1982]" italics="true" pageId="10" pageNumber="37">Pauridiantha principensis</emphasis>
Ntore &amp; O.Lachenaud
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB7A225DE25F812D6EF19B3" box="[618,687,1964,1982]" pageId="10" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B96DEABCFFB7A225DEFAF812D68219B3" bold="true" box="[693,706,1964,1982]" pageId="10" pageNumber="37">A</emphasis>
, leafy stem;
<emphasis id="B96DEABCFFB7A225DF62F812D77A19B3" bold="true" box="[813,826,1964,1982]" pageId="10" pageNumber="37">B</emphasis>
, node with stipules;
<emphasis id="B96DEABCFFB7A225DFA8F812D7B519B3" bold="true" box="[999,1013,1964,1982]" pageId="10" pageNumber="37">C</emphasis>
, inflorescence;
<emphasis id="B96DEABCFFB7A225D837F812D0C619B3" bold="true" box="[1144,1158,1964,1982]" pageId="10" pageNumber="37">D</emphasis>
, disk, viewed from above;
<emphasis id="B96DEABCFFB7A225D928F812D13319B3" bold="true" box="[1383,1395,1964,1982]" pageId="10" pageNumber="37">E</emphasis>
, shortstyled flower;
<emphasis id="B96DEABCFFB7A225DCB6F87DD54419D8" bold="true" box="[249,260,1987,2005]" pageId="10" pageNumber="37">F</emphasis>
, same, with calyx and ovary cut longitudinally and corolla laid open;
<emphasis id="B96DEABCFFB7A225DF0CF87DD71119D8" bold="true" box="[835,849,1987,2005]" pageId="10" pageNumber="37">G</emphasis>
, long-styled flower;
<emphasis id="B96DEABCFFB7A225DFB3F87DD04A19D8" bold="true" box="[1020,1034,1987,2005]" pageId="10" pageNumber="37">H</emphasis>
, same, with calyx and ovary cut longitudinally and corolla laid open. All details from
<emphasis id="B96DEABCFFB7A225DDD6F867D65619E6" box="[409,534,2009,2027]" italics="true" pageId="10" pageNumber="37">Oliveira 192/98</emphasis>
. Scale bars: A-C, 1 cm; D, 1 mm; E-H, 5 mm. Illustration: Marijke Meersman.
</paragraph>
</caption>
<caption id="DF666626FFB6A224DCCBFF63D6E21F11" ID-Table-UUID="DF666626FFB6A224DCCBFF63D6E21F11" httpUri="http://table.plazi.org/id/DF666626FFB6A224DCCBFF63D6E21F11" pageId="11" pageNumber="38" startId="11.[132,143,221,239]" targetBox="[134,1362,342,616]" targetIsTable="true" targetPageId="11">
<paragraph id="8BA636AEFFB6A224DCCBFF63D6E21F11" blockId="11.[132,1455,221,284]" pageId="11" pageNumber="38">
TABLE 2. — Differences between
<taxonomicName id="4C194D2DFFB6A224DDD9FF63D73A1EE2" authority="(K. Schum.) Ntore &amp; O. Lachenaud" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" baseAuthorityName="K. Schum." box="[406,890,221,239]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="chlorantha" status="comb. nov.">
<emphasis id="B96DEABCFFB6A224DDD9FF63D5BE1EE2" box="[406,510,221,239]" italics="true" pageId="11" pageNumber="38">Pauridiantha</emphasis>
<emphasis id="B96DEABCFFB6A224DE4CFF63D61A1EE2" box="[515,602,221,239]" italics="true" pageId="11" pageNumber="38">chlorantha</emphasis>
(K.Schum.) Ntore &amp; O.Lachenaud
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB6A224DFCDFF63D7A01EE2" box="[898,992,221,239]" pageId="11" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C194D2DFFB6A224DFA5FF63D0871EE2" authority="(Hiern) Bremek." authorityName="Bremek." baseAuthorityName="Hiern" box="[1002,1223,221,239]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="B96DEABCFFB6A224DFA5FF63D7B91EE2" box="[1002,1017,221,239]" italics="true" pageId="11" pageNumber="38">P.</emphasis>
<emphasis id="B96DEABCFFB6A224DFB2FF63D0001EE2" box="[1021,1088,221,239]" italics="true" pageId="11" pageNumber="38">insularis</emphasis>
(Hiern) Bremek.
</taxonomicName>
and
<taxonomicName id="4C194D2DFFB6A224D8BCFF63D4B71F08" authority="Ntore &amp; O.Lachenaud" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="principensis" status="sp. nov.">
<emphasis id="B96DEABCFFB6A224D8BCFF63D12A1EE2" box="[1267,1386,221,239]" italics="true" pageId="11" pageNumber="38">P. principensis</emphasis>
Ntore &amp; O.Lachenaud
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB6A224DCB0FF4AD5021F0B" box="[255,322,244,262]" pageId="11" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
Note: The ovary of
<taxonomicName id="4C194D2DFFB6A224DDA7FF4AD67F1F0B" box="[488,575,244,262]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="B96DEABCFFB6A224DDA7FF4AD67F1F0B" box="[488,575,244,262]" italics="true" pageId="11" pageNumber="38">P. insularis</emphasis>
</taxonomicName>
is described by Hiern (1877: 73) and Ntore (2008) as 4-5-locular, but is actually 2-locular at the base and 4-locular by false septa at the apex in the material we have seen.
</paragraph>
</caption>
<paragraph id="8BA636AEFFB6A224DCC9FEE8D0DB1C65" pageId="11" pageNumber="38">
<table id="F919C40EFFB65DD0DCC9FEE8D1121C65" box="[134,1362,342,616]" gridcols="4" gridrows="10" pageId="11" pageNumber="38">
<tableNote id="76FF3720FFB6A224DCC9FEE8D0DB1C65" pageId="11" pageNumber="38" targetBox="[134,1362,377,616]" targetPageId="11">
<tr id="352934ECFFB65DD0DCC9FEE8D1121F66" box="[134,1362,342,363]" gridrow="0" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FEE8D50B1F66" box="[134,331,342,363]" gridcol="0" gridrow="0" pageId="11" pageNumber="38">Characters</th>
<th id="76F85D90FFB65DD0DDCAFEE8D6991F66" box="[389,729,342,363]" gridcol="1" gridrow="0" pageId="11" pageNumber="38">
<taxonomicName id="4C194D2DFFB6A224DDCAFEE9D6491F66" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" baseAuthorityName="K. Schum." box="[389,521,343,363]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="chlorantha" status="comb. nov.">
<emphasis id="B96DEABCFFB6A224DDCAFEE9D6491F66" bold="true" box="[389,521,343,363]" italics="true" pageId="11" pageNumber="38">P. chlorantha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB6A224DE5AFEE9D6C41F66" box="[533,644,343,363]" pageId="11" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
</th>
<th id="76F85D90FFB65DD0DEA6FEE8D7B91F66" box="[745,1017,342,363]" gridcol="2" gridrow="0" pageId="11" pageNumber="38">
<taxonomicName id="4C194D2DFFB6A224DEA6FEE9D7B91F66" authority="(Hiern) Bremek." authorityName="Bremek." baseAuthorityName="Hiern" box="[745,1017,343,363]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="B96DEABCFFB6A224DEA6FEE9D7141F66" bold="true" box="[745,852,343,363]" italics="true" pageId="11" pageNumber="38">P. insularis</emphasis>
(Hiern) Bremek.
</taxonomicName>
</th>
<th id="76F85D90FFB65DD0D802FEE8D1121F66" box="[1101,1362,342,363]" gridcol="3" gridrow="0" pageId="11" pageNumber="38">
<taxonomicName id="4C194D2DFFB6A224D802FEE9D09E1F66" authority="Salvator NTORE &amp; Olivier LACHENAUD, 2019" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[1101,1246,343,363]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="principensis" status="sp. nov.">
<emphasis id="B96DEABCFFB6A224D802FEE9D09E1F66" bold="true" box="[1101,1246,343,363]" italics="true" pageId="11" pageNumber="38">P. principensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C7FFB6A224D8A5FEE9D1781F66" box="[1258,1336,343,363]" pageId="11" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</th>
</tr>
<tr id="352934ECFFB65DD0DCC9FEC7D1121F83" box="[134,1362,377,398]" gridrow="1" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FEC7D50B1F83" box="[134,331,377,398]" gridcol="0" gridrow="1" pageId="11" pageNumber="38">Stipules</th>
<td id="76F85D90FFB65DD0DDCAFEC7D6991F83" box="[389,729,377,398]" gridcol="1" gridrow="1" pageId="11" pageNumber="38">glabrous to sparsely pubescent</td>
<td id="76F85D90FFB65DD0DEA6FEC7D7B91F83" box="[745,1017,377,398]" gridcol="2" gridrow="1" pageId="11" pageNumber="38">sparsely pubescent</td>
<td id="76F85D90FFB65DD0D802FEC7D1121F83" box="[1101,1362,377,398]" gridcol="3" gridrow="1" pageId="11" pageNumber="38">glabrous</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FE2CD1121FAB" box="[134,1362,402,422]" gridrow="2" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FE2CD50B1FAB" box="[134,331,402,422]" gridcol="0" gridrow="2" pageId="11" pageNumber="38">Leaf domatia</th>
<td id="76F85D90FFB65DD0DDCAFE2CD6991FAB" box="[389,729,402,422]" gridcol="1" gridrow="2" pageId="11" pageNumber="38">in primary and secondary axils</td>
<td id="76F85D90FFB65DD0DEA6FE2CD7B91FAB" box="[745,1017,402,422]" gridcol="2" gridrow="2" pageId="11" pageNumber="38">in primary axils only</td>
<td id="76F85D90FFB65DD0D802FE2CD1121FAB" box="[1101,1362,402,422]" gridcol="3" gridrow="2" pageId="11" pageNumber="38">absent</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FE14D1121FB3" box="[134,1362,426,446]" gridrow="3" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FE14D50B1FB3" box="[134,331,426,446]" gridcol="0" gridrow="3" pageId="11" pageNumber="38">Secondary leaf veins</th>
<td id="76F85D90FFB65DD0DDCAFE14D6991FB3" box="[389,729,426,446]" gridcol="1" gridrow="3" pageId="11" pageNumber="38">10-16, weakly ascending</td>
<td id="76F85D90FFB65DD0DEA6FE14D7B91FB3" box="[745,1017,426,446]" gridcol="2" gridrow="3" pageId="11" pageNumber="38">10-17, weakly ascending</td>
<td id="76F85D90FFB65DD0D802FE14D1121FB3" box="[1101,1362,426,446]" gridcol="3" gridrow="3" pageId="11" pageNumber="38">8-11, strongly ascending</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FE7CD1121FDB" box="[134,1362,450,470]" gridrow="4" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FE7CD50B1FDB" box="[134,331,450,470]" gridcol="0" gridrow="4" pageId="11" pageNumber="38">Prophylls</th>
<td id="76F85D90FFB65DD0DDCAFE7CD6991FDB" box="[389,729,450,470]" gridcol="1" gridrow="4" pageId="11" pageNumber="38">± halfway on the peduncle or below</td>
<td id="76F85D90FFB65DD0DEA6FE7CD7B91FDB" box="[745,1017,450,470]" gridcol="2" gridrow="4" pageId="11" pageNumber="38">at the apex of the peduncle</td>
<td id="76F85D90FFB65DD0D802FE7CD1121FDB" box="[1101,1362,450,470]" gridcol="3" gridrow="4" pageId="11" pageNumber="38">at the apex of the peduncle</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FE64D1121FE3" box="[134,1362,474,494]" gridrow="5" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FE64D50B1FE3" box="[134,331,474,494]" gridcol="0" gridrow="5" pageId="11" pageNumber="38">Flowering pedicels</th>
<td id="76F85D90FFB65DD0DDCAFE64D6991FE3" box="[389,729,474,494]" gridcol="1" gridrow="5" pageId="11" pageNumber="38">1.5-4 mm</td>
<td id="76F85D90FFB65DD0DEA6FE64D7B91FE3" box="[745,1017,474,494]" gridcol="2" gridrow="5" pageId="11" pageNumber="38">2-3 mm</td>
<td id="76F85D90FFB65DD0D802FE64D1121FE3" box="[1101,1362,474,494]" gridcol="3" gridrow="5" pageId="11" pageNumber="38">3.5-5 mm</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FE4CD1121C0B" box="[134,1362,498,518]" gridrow="6" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FE4CD50B1C0B" box="[134,331,498,518]" gridcol="0" gridrow="6" pageId="11" pageNumber="38">Fruiting pedicels</th>
<td id="76F85D90FFB65DD0DDCAFE4CD6991C0B" box="[389,729,498,518]" gridcol="1" gridrow="6" pageId="11" pageNumber="38">4-6 mm</td>
<td id="76F85D90FFB65DD0DEA6FE4CD7B91C0B" box="[745,1017,498,518]" gridcol="2" gridrow="6" pageId="11" pageNumber="38">4-7.5 mm</td>
<td id="76F85D90FFB65DD0D802FE4CD1121C0B" box="[1101,1362,498,518]" gridcol="3" gridrow="6" pageId="11" pageNumber="38">7.5-10 mm</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FDB4D1121C13" box="[134,1362,522,542]" gridrow="7" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FDB4D50B1C13" box="[134,331,522,542]" gridcol="0" gridrow="7" pageId="11" pageNumber="38">Calyx</th>
<td id="76F85D90FFB65DD0DDCAFDB4D6991C13" box="[389,729,522,542]" gridcol="1" gridrow="7" pageId="11" pageNumber="38">1.5-2 mm wide</td>
<td id="76F85D90FFB65DD0DEA6FDB4D7B91C13" box="[745,1017,522,542]" gridcol="2" gridrow="7" pageId="11" pageNumber="38">2.5-4.5 mm wide</td>
<td id="76F85D90FFB65DD0D802FDB4D1121C13" box="[1101,1362,522,542]" gridcol="3" gridrow="7" pageId="11" pageNumber="38">2-3.5 mm wide</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FD9CD1121C5D" box="[134,1362,546,592]" gridrow="8" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FD9CD50B1C5D" box="[134,331,546,592]" gridcol="0" gridrow="8" pageId="11" pageNumber="38">Ovary</th>
<td id="76F85D90FFB65DD0DDCAFD9CD6991C5D" box="[389,729,546,592]" gridcol="1" gridrow="8" pageId="11" pageNumber="38">4-5-locular at base (8-10-locular by false septa at apex)</td>
<td id="76F85D90FFB65DD0DEA6FD9CD7B91C5D" box="[745,1017,546,592]" gridcol="2" gridrow="8" pageId="11" pageNumber="38">2-locular at base (4-locular by false septa at apex)</td>
<td id="76F85D90FFB65DD0D802FD9CD1121C5D" box="[1101,1362,546,592]" gridcol="3" gridrow="8" pageId="11" pageNumber="38">2-locular at base (4-locular by false septa at apex)</td>
</tr>
<tr id="352934ECFFB65DD0DCC9FDEAD1121C65" box="[134,1362,596,616]" gridrow="9" pageId="11" pageNumber="38">
<th id="76F85D90FFB65DD0DCC9FDEAD50B1C65" box="[134,331,596,616]" gridcol="0" gridrow="9" pageId="11" pageNumber="38">Distribution</th>
<td id="76F85D90FFB65DD0DDCAFDEAD6991C65" box="[389,729,596,616]" gridcol="1" gridrow="9" pageId="11" pageNumber="38">Tanzania</td>
<td id="76F85D90FFB65DD0DEA6FDEAD7B91C65" box="[745,1017,596,616]" gridcol="2" gridrow="9" pageId="11" pageNumber="38">São Tomé</td>
<td id="76F85D90FFB65DD0D802FDEAD1121C65" box="[1101,1362,596,616]" gridcol="3" gridrow="9" pageId="11" pageNumber="38">Príncipe</td>
</tr>
</tableNote>
</table>
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFD06D4811CDC" blockId="11.[132,775,696,850]" box="[132,193,696,721]" pageId="11" pageNumber="38">
<heading id="D0EE81C2FFB6A224DCCBFD06D4811CDC" box="[132,193,696,721]" fontSize="11" level="4" pageId="11" pageNumber="38" reason="1">
<emphasis id="B96DEABCFFB6A224DCCBFD06D4811CDC" box="[132,193,696,721]" italics="true" pageId="11" pageNumber="38">Fruits</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFD69D5601D5F" blockId="11.[132,775,696,850]" pageId="11" pageNumber="38">
Dark green (not fully mature), subglobose, 5.5-7.5 ×
<quantity id="4CE19B4BFFB6A224DE95FD69D4991D1F" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="11" pageNumber="38" unit="mm" value="7.5" valueMax="8.5" valueMin="6.5">6.5- 8.5 mm</quantity>
in life, 4-5 ×
<quantity id="4CE19B4BFFB6A224DD29FD49D5911D1F" box="[358,465,759,786]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="7.0" metricValueMin="5.3" pageId="11" pageNumber="38" unit="mm" value="6.15" valueMax="7.0" valueMin="5.3">5.3-7 mm</quantity>
when dry, smooth, glabrous, crowned with persistent calyx slightly enlarged in diameter (
<quantity id="4CE19B4BFFB6A224DCC3FC89D5531D5F" box="[140,275,823,850]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="4.5" metricValueMin="2.3" pageId="11" pageNumber="38" unit="mm" value="3.4" valueMax="4.5" valueMin="2.3">2.3-4.5 mm</quantity>
).
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFCC8D5391DBC" blockId="11.[132,775,886,945]" pageId="11" pageNumber="38">
<emphasis id="B96DEABCFFB6A224DCCBFCC8D4A41D9D" bold="true" box="[132,228,886,912]" pageId="11" pageNumber="38">Pedicels.</emphasis>
Drooping and strongly accrescent in the fruiting stage,
<quantity id="4CE19B4BFFB6A224DC88FC29D57B1DBC" box="[199,315,919,945]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="11" pageNumber="38" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12-18 mm</quantity>
long.
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFC68D4F91DFD" blockId="11.[132,546,982,1041]" box="[132,185,982,1008]" pageId="11" pageNumber="38">
<heading id="D0EE81C2FFB6A224DCCBFC68D4F91DFD" box="[132,185,982,1008]" fontSize="11" level="4" pageId="11" pageNumber="38" reason="1">
<emphasis id="B96DEABCFFB6A224DCCBFC68D4F91DFD" box="[132,185,982,1008]" italics="true" pageId="11" pageNumber="38">Seeds</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFC49D6621A1C" blockId="11.[132,546,982,1041]" box="[132,546,1014,1041]" pageId="11" pageNumber="38">
Numerous, ellipsoid,
<emphasis id="B96DEABCFFB6A224DD22FC49D53E1A1D" box="[365,382,1015,1040]" italics="true" pageId="11" pageNumber="38">c.</emphasis>
0.7
<emphasis id="B96DEABCFFB6A224DDE0FC48D5FD1A1D" bold="true" box="[431,445,1014,1040]" pageId="11" pageNumber="38">×</emphasis>
<quantity id="4CE19B4BFFB6A224DD8BFC48D65E1A1C" box="[452,542,1014,1041]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="38" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFB89D5471A5D" blockId="11.[132,775,1078,1200]" box="[132,263,1078,1104]" pageId="11" pageNumber="38">
<heading id="D0EE81C2FFB6A224DCCBFB89D5471A5D" box="[132,263,1078,1104]" fontSize="11" level="4" pageId="11" pageNumber="38" reason="1">
<emphasis id="B96DEABCFFB6A224DCCBFB89D5471A5D" box="[132,263,1078,1104]" italics="true" pageId="11" pageNumber="38">Exotesta cells</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFBE8D5BB1ABD" blockId="11.[132,775,1078,1200]" pageId="11" pageNumber="38">Polygonal without raised margins, forming a very faint reticulum; radial walls pitted, not thickened; inner tangential wall broadly pitted, not thickened.</paragraph>
</subSubSection>
<subSubSection id="C3036525FFB6A224DCCBFB6BD5D618C2" pageId="11" pageNumber="38" type="discussion">
<paragraph id="8BA636AEFFB6A224DCCBFB6BD4A81AE3" blockId="11.[132,775,1237,1743]" box="[132,232,1237,1263]" pageId="11" pageNumber="38">REMARKS</paragraph>
<paragraph id="8BA636AEFFB6A224DCCBFB4BD5D618C2" blockId="11.[132,775,1237,1743]" pageId="11" pageNumber="38">
This very peculiar species was discovered in 2014 during the botanical exploration of the Lower Ogooué, and had apparently never been collected before. Although the flowering material is incomplete (only one dried flower was found, with both the style and stamens apparently included), the plant is so distinctive that we have no hesitation in describing it. Its remarkably long fruiting pedicels are unique in the genus, as is the presence of spines on the older stems, which are formed by the persistent base of broken twigs (these spines are only seen on the older individuals). The lianescent habit, with drooping branches somewhat reminiscent of
<taxonomicName id="4C194D2DFFB6A224DCB0F9EAD5021863" box="[255,322,1620,1646]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEABCFFB6A224DCB0F9EAD5021863" box="[255,322,1620,1646]" italics="true" pageId="11" pageNumber="38">Rubus</emphasis>
</taxonomicName>
L. spp., is further unusual, most other
<taxonomicName id="4C194D2DFFB6A224DCCBF9CAD5541883" authorityName="Salvator NTORE &amp; Olivier LACHENAUD" authorityYear="2019" box="[132,276,1652,1678]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="genus">
<emphasis id="B96DEABCFFB6A224DCCBF9CAD5541883" box="[132,276,1652,1678]" italics="true" pageId="11" pageNumber="38">Pauridiantha</emphasis>
</taxonomicName>
species being erect shrubs or trees, although a few, such as
<taxonomicName id="4C194D2DFFB6A224DD65F92BD6F018A3" authority="(K. Krause) Bremek." authorityName="Bremek." baseAuthorityName="K. Krause" box="[298,688,1684,1710]" class="Magnoliopsida" family="Rubiaceae" genus="Pauridiantha" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="38" phylum="Tracheophyta" rank="species" species="pyramidata">
<emphasis id="B96DEABCFFB6A224DD65F92BD58118A3" box="[298,449,1684,1710]" italics="true" pageId="11" pageNumber="38">P. pyramidata</emphasis>
(K.Krause) Bremek.
</taxonomicName>
, can be more or less sarmentose.
</paragraph>
</subSubSection>
</treatment>
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