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<document id="025D1FF2AA37AF8CDD734D44A1552787" ID-DOI="10.5852/ejt.2024.926.2469" ID-GBIF-Dataset="4511b4bd-f026-49d3-9d4d-a39875e75cc5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFF94FFEFFE2FD609FC54FADF" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum njegusiense Jochum &amp; Ruthensteiner 2024, sp. nov." docType="treatment" docUuid="882B9D99-6028-4DC7-B9A0-82033E474B39" docUuidSource="ZooBank" docVersion="11" lastPageNumber="48" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="47" updateTime="1722640713706" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="27CA06DB9E5C183452CD043A090123E9">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
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<mods:namePart id="45043C06F4FFDC45C3025AE47CD7EE15">Jochum, Adrienne</mods:namePart>
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<mods:affiliation id="3A950CC4CB6697DD6FB4AAFE5E5900BA">Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, D- 60325 Frankfurt / M., Germany.</mods:affiliation>
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<mods:namePart id="424617EC878B6835E8DA2008BB4D8C47">Michalik, Peter</mods:namePart>
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<mods:affiliation id="5DD7D798FE7D2525960727D571DC062D">Zoological Institute and Museum, University of Greifswald, Loitzer Strasse 26, D- 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="4C9DE94304933EF37795771A20FC724C" type="email">michalik@uni-greifswald.de</mods:nameIdentifier>
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<mods:namePart id="B8D645539139294C3E5EB1F94C668CC4">Inäbnit, Thomas</mods:namePart>
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<mods:affiliation id="C16AF841E1BC5C3F5B843664C4F3A1F8">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="8E208CC3E24F7249B123867E72616A82" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="62CB735B748716596BEB24B99C60A7EA">Kneubühler, Jeannette</mods:namePart>
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<mods:affiliation id="F95454E41353C27FE0DF3B926AE68E96">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="95AFEC0281F1EF5DDB2710A05D50EE33" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
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<mods:namePart id="36DC2DCF9982AF9F0A2A78D51E6CEFC4">Slapnik, Rajko</mods:namePart>
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<mods:affiliation id="E4E8FECC3F1B4DC5D5C04F143E35BB98">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="26529D7328E57E727B92B5375D5AF7FB" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="77C2AB5C3ECE0C44D778CA15DD0C782F">Vrabec, Marko</mods:namePart>
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<mods:affiliation id="79411D30CF3398E0522B8803262304F2">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="DE70A9FC7C4BE31953609A3AC4665BAF" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
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<mods:namePart id="B858ECC242E52944F215EC6D4393A072">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="7521AABB5418E158CFA302BC84FA1EFE">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="671C8E4934561B8375B1E6D3CE8C4573">Ruthensteiner, Bernhard</mods:namePart>
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<mods:affiliation id="745D44BE428D49D23C0B4540442BEFCA">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="793D857183245FE667321E6F4F0FF4F6" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
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<mods:title id="03EA980BA21FC4032EFF1AD45EF2B4CB">European Journal of Taxonomy</mods:title>
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<mods:part id="8B63C8AE196FEDB25C608D6835DDD918">
<mods:date id="F91C85FD5FCFA994046C567011C5EF71">2024</mods:date>
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<mods:number id="96D502F498A57B90F20535B3D69B077C">2024-03-20</mods:number>
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<mods:number id="8E87AAF6336477AA0DF722BBB4A51232">926</mods:number>
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<treatment id="03B0879DFF94FFEFFE2FD609FC54FADF" ID-DOI="http://doi.org/10.5281/zenodo.10847868" ID-GBIF-Taxon="224067518" ID-Zenodo-Dep="10847868" LSID="urn:lsid:zoobank.org:act:882B9D99-6028-4DC7-B9A0-82033E474B39" httpUri="http://treatment.plazi.org/id/03B0879DFF94FFEFFE2FD609FC54FADF" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C3036500FF94FFEEFE2FD609FBB0FEF7" box="[464,1124,264,291]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA6368BFF94FFEEFE2FD609FBB0FEF7" blockId="46.[372,1216,264,361]" box="[464,1124,264,291]" pageId="46" pageNumber="47">
<heading id="D0EE81E7FF94FFEEFE2FD609FBB0FEF7" box="[464,1124,264,291]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="4C194D08FF94FFEEFE2FD609FBD6FEF6" ID-CoL="C8NTC" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" box="[464,1026,264,291]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="njegusiense" status="sp. nov.">
<emphasis id="B96DEA99FF94FFEEFE2FD609FD02FEF6" bold="true" box="[464,726,264,291]" italics="true" pageId="46" pageNumber="47">Zospeum njegusiense</emphasis>
Jochum &amp; Ruthensteiner
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<taxonomicNameLabel id="A25E57E2FF94FFEEFBF6D609FBB0FEF7" box="[1033,1124,265,291]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA6368BFF94FFEEFD0FD64FFC97FEBD" blockId="46.[372,1216,264,361]" box="[752,835,335,361]" pageId="46" pageNumber="47">
<figureCitation id="13222A0EFF94FFEEFD0FD64FFC97FEBD" box="[752,835,335,361]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[189,1398,1058,1782]" captionTargetId="figure-216@46.[189,1398,1058,1782]" captionTargetPageId="46" captionText="Fig. 17. Z. njegusiense Jochum &amp; Ruthensteiner sp. nov., Scanning Electron Microscopy images. A. Holotype (NMBE 572639), aperture view showing subfossil shell with superficial erosion of shell layer. B. Paratype (NMBE 578378), aperture view showing close up of lamellar band on columella (Fig. C). C. Paratype (NMBE 578378), aperture view showing damaged shell and columella. D. Close up view of holotype (NMBE 572639), showing body whorl with low, irregular growth lines. E. Paratype (NMBE 578378), protoconch." figureDoi="http://doi.org/10.5281/zenodo.10847854" httpUri="https://zenodo.org/record/10847854/files/figure.png" pageId="46" pageNumber="47">Fig. 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF94FFEEFF42D690FDC9FD96" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BA6368BFF94FFEEFF42D690FEE2FE7E" blockId="46.[189,310,400,426]" box="[189,310,400,426]" pageId="46" pageNumber="47">
<heading id="D0EE81E7FF94FFEEFF42D690FEE2FE7E" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96DEA99FF94FFEEFF42D690FEE2FE7E" bold="true" box="[189,310,400,426]" pageId="46" pageNumber="47">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF94FFEEFF42D6BFFDC9FD96" blockId="46.[188,1399,445,579]" pageId="46" pageNumber="47">Shell conical with 5¼ flatly convex whorls; aperture reniform; penultimate whorl over ½ height of body whorl; columella slender with a distinctly sculpted and broad lamellar band obliquely projecting ca ⅓ beyond the width of the slender columella; a second, barely visible low lamella is present at mid-section of the columella. Suture deep.</paragraph>
</subSubSection>
<subSubSection id="C3036500FF94FFEEFF42D56AFC6BFD67" box="[189,959,618,691]" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8BA6368BFF94FFEEFF42D56AFE90FD50" blockId="46.[189,324,618,644]" box="[189,324,618,644]" pageId="46" pageNumber="47">
<heading id="D0EE81E7FF94FFEEFF42D56AFE90FD50" bold="true" box="[189,324,618,644]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96DEA99FF94FFEEFF42D56AFE90FD50" bold="true" box="[189,324,618,644]" pageId="46" pageNumber="47">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF94FFEEFF42D598FC6BFD67" blockId="46.[189,959,664,691]" box="[189,959,664,691]" pageId="46" pageNumber="47">
This species is named after its
<typeStatus id="54A28829FF94FFEEFDDAD599FD8CFD67" box="[549,600,665,691]" pageId="46" pageNumber="47">type</typeStatus>
locality Njeguši,
<collectingCountry id="F30E761BFF94FFEEFCD7D599FC6FFD67" box="[808,955,665,691]" name="Serbia" pageId="46" pageNumber="47">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF94FFEFFF42D5DAFDFFFEBD" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BA6368BFF94FFEEFF42D5DAFEB8FD20" blockId="46.[189,364,730,756]" box="[189,364,730,756]" pageId="46" pageNumber="47">
<heading id="D0EE81E7FF94FFEEFF42D5DAFEB8FD20" bold="true" box="[189,364,730,756]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96DEA99FF94FFEEFF42D5DAFEB8FD20" bold="true" box="[189,364,730,756]" pageId="46" pageNumber="47">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B713CD6FF94FFEEFF15D409FDFFFC58" ID-GBIF-Occurrence="4598261308" collectingDate="2021-08-05" collectionCode="NMBE, AJC" collectorName="Laszlo Danyi &amp; Nikolett Ujhegyi" country="Serbia" county="Njegusi" elevation="915" latitude="42.4307" location="St John's cave" longLatPrecision="6" longitude="18.8115" municipality="St John's cave" pageId="46" pageNumber="47" specimenCode="NMBE 572639" specimenCount="1" typeStatus="holotype">
<paragraph id="8BA6368BFF94FFEEFF15D409FDDAFCF6" blockId="46.[189,1398,776,909]" box="[234,526,776,803]" pageId="46" pageNumber="47">
<emphasis id="B96DEA99FF94FFEEFF15D409FE88FCF7" bold="true" box="[234,348,777,803]" pageId="46" pageNumber="47">
<typeStatus id="54A28829FF94FFEEFF15D409FE88FCF7" box="[234,348,777,803]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
</emphasis>
(SEM sample)
</paragraph>
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<collectingCountry id="F30E761BFF94FFEEFF42D42CFE5EFC92" box="[189,394,812,838]" name="Serbia" pageId="46" pageNumber="47">MONTENEGRO</collectingCountry>
• (
<figureCitation id="13222A0EFF94FFEEFE4ED42CFD93FC92" box="[433,583,812,838]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[189,1398,1058,1782]" captionTargetId="figure-216@46.[189,1398,1058,1782]" captionTargetPageId="46" captionText="Fig. 17. Z. njegusiense Jochum &amp; Ruthensteiner sp. nov., Scanning Electron Microscopy images. A. Holotype (NMBE 572639), aperture view showing subfossil shell with superficial erosion of shell layer. B. Paratype (NMBE 578378), aperture view showing close up of lamellar band on columella (Fig. C). C. Paratype (NMBE 578378), aperture view showing damaged shell and columella. D. Close up view of holotype (NMBE 572639), showing body whorl with low, irregular growth lines. E. Paratype (NMBE 578378), protoconch." figureDoi="http://doi.org/10.5281/zenodo.10847854" httpUri="https://zenodo.org/record/10847854/files/figure.png" pageId="46" pageNumber="47">Fig. 17A, D</figureCitation>
);
<collectingCounty id="62C74E07FF94FFEEFDA0D42CFD69FC92" box="[607,701,812,838]" pageId="46" pageNumber="47">Njeguši</collectingCounty>
,
<location id="8EC66050FF94FFEEFD33D42CFCA8FC92" LSID="urn:lsid:plazi:treatment:03B0879DFF94FFEFFE2FD609FC54FADF:8EC66050FF94FFEEFD33D42CFCA8FC92" box="[716,892,812,838]" country="Serbia" county="Njegusi" latitude="42.4307" longLatPrecision="6" longitude="18.8115" municipality="St John's cave" name="St John's cave" pageId="46" pageNumber="47">St John’s cave</location>
;
<geoCoordinate id="EE2D504CFF94FFEEFC74D42CFBCCFC92" box="[907,1048,812,838]" degrees="42.4307" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="5" value="42.4307">42.4307° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFF94FFEEFBD9D42BFB7AFC92" box="[1062,1198,811,838]" degrees="18.8115" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="5" value="18.8115">18.8115° E</geoCoordinate>
,
<elevation id="0034D1B8FF94FFEEFB42D42BFA98FC92" box="[1213,1356,811,838]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="46" pageNumber="47" unit="m" value="915.0">
<quantity id="4CE19B6EFF94FFEEFB42D42BFADEFC92" box="[1213,1290,811,838]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="46" pageNumber="47" unit="m" value="915.0">915 m</quantity>
a.s.l.
</elevation>
; ca -
<quantity id="4CE19B6EFF94FFEEFF39D44FFED2FCBD" box="[198,262,847,874]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="46" pageNumber="47" unit="m" value="85.0">85 m</quantity>
vertical depth below the cave entrance;
<date id="FFA7104BFF94FFEEFD08D44FFC46FCBD" box="[759,914,847,874]" pageId="46" pageNumber="47" value="2021-08-05">
<collectingDate id="EFE3E9A3FF94FFEEFD08D44FFC46FCBD" box="[759,914,847,874]" pageId="46" pageNumber="47" value="2021-08-05">5 Aug. 2021</collectingDate>
</date>
;
<collectorName id="26EC535DFF94FFEEFC5DD44FFB97FCBD" box="[930,1091,847,873]" pageId="46" pageNumber="47">László Dányi</collectorName>
&amp;
<collectorName id="26EC535DFF94FFEEFB90D44FFAEDFCBD" box="[1135,1337,847,873]" pageId="46" pageNumber="47">Nikolett Ujhegyi</collectorName>
leg.;
<specimenCode id="DBBF9EF0FF94FFEEFF42D473FEA5FC59" box="[189,369,882,909]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="46" pageNumber="47">NMBE 572639</specimenCode>
(ex AJC 2483).
</paragraph>
</materialsCitation>
<caption id="DF666603FF94FFEEFF42D01EFDF5F83C" ID-DOI="http://doi.org/10.5281/zenodo.10847854" ID-Zenodo-Dep="10847854" httpUri="https://zenodo.org/record/10847854/files/figure.png" pageId="46" pageNumber="47" startId="46.[189,232,1822,1848]" targetBox="[189,1398,1058,1782]" targetPageId="46">
<paragraph id="8BA6368BFF94FFEEFF42D01EFDF5F83C" blockId="46.[188,1399,1822,2025]" pageId="46" pageNumber="47">
<emphasis id="B96DEA99FF94FFEEFF42D01EFEF4F8EC" bold="true" box="[189,288,1822,1848]" pageId="46" pageNumber="47">Fig. 17.</emphasis>
<taxonomicName id="4C194D08FF94FFEEFED2D01FFCCCF8EC" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" box="[301,792,1822,1848]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="njegusiense" status="sp. nov.">
<emphasis id="B96DEA99FF94FFEEFED2D01FFE08F8EC" box="[301,476,1823,1848]" italics="true" pageId="46" pageNumber="47">Z. njegusiense</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF94FFEEFCDAD01FFC53F8ED" box="[805,903,1823,1849]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, Scanning Electron Microscopy images.
<emphasis id="B96DEA99FF94FFEEFF42D041FF06F88F" bold="true" box="[189,210,1857,1883]" pageId="46" pageNumber="47">A</emphasis>
. Holotype (NMBE 572639), aperture view showing subfossil shell with superficial erosion of shell layer.
<emphasis id="B96DEA99FF94FFEEFEFBD065FECCF8AB" bold="true" box="[260,280,1893,1919]" pageId="46" pageNumber="47">B</emphasis>
. Paratype (NMBE 578378), aperture view showing close up of lamellar band on columella (Fig. C).
<emphasis id="B96DEA99FF94FFEEFF13D088FED5F876" bold="true" box="[236,257,1928,1954]" pageId="46" pageNumber="47">C</emphasis>
. Paratype (NMBE 578378), aperture view showing damaged shell and columella.
<emphasis id="B96DEA99FF94FFEEFB19D088FB2FF876" bold="true" box="[1254,1275,1928,1954]" pageId="46" pageNumber="47">D</emphasis>
. Close up view of holotype (NMBE 572639), showing body whorl with low, irregular growth lines.
<emphasis id="B96DEA99FF94FFEEFB11D0ABFAD4F811" bold="true" box="[1262,1280,1963,1989]" pageId="46" pageNumber="47">E</emphasis>
. Paratype (NMBE 578378), protoconch.
</paragraph>
</caption>
<paragraph id="8BA6368BFF95FFEFFF15D609FDCDFEF7" blockId="47.[189,1398,265,362]" box="[234,537,265,291]" pageId="47" pageNumber="48">
<emphasis id="B96DEA99FF95FFEFFF15D609FEB3FEF7" bold="true" box="[234,359,265,291]" pageId="47" pageNumber="48">
<typeStatus id="54A28829FF95FFEFFF15D609FEB3FEF7" box="[234,359,265,291]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
(SEM sample)
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D62CFDFFFEBD" blockId="47.[189,1398,265,362]" pageId="47" pageNumber="48">
<materialsCitation id="3B713CD6FF95FFEFFF42D62CFDFCFEBD" ID-GBIF-Occurrence="4598261303" collectingDate="2021-08-05" collectionCode="NMBE, AJC" collectorName="Laszlo Danyi &amp; Nikolett Ujhegyi" country="Serbia" county="Njegusi" elevation="915" latitude="42.4307" location="MONTENEGRO" longLatPrecision="6" longitude="18.8115" municipality="St John's cave" pageId="47" pageNumber="48" specimenCode="NMBE 578378" specimenCount="2" typeStatus="paratype">
<collectingCountry id="F30E761BFF95FFEFFF42D62CFE5EFE92" box="[189,394,300,326]" name="Serbia" pageId="47" pageNumber="48">MONTENEGRO</collectingCountry>
• (
<figureCitation id="13222A0EFF95FFEFFE4AD62CFDBFFE92" box="[437,619,300,326]" captionStart="Fig" captionStartId="46.[189,232,1822,1848]" captionTargetBox="[189,1398,1058,1782]" captionTargetId="figure-216@46.[189,1398,1058,1782]" captionTargetPageId="46" captionText="Fig. 17. Z. njegusiense Jochum &amp; Ruthensteiner sp. nov., Scanning Electron Microscopy images. A. Holotype (NMBE 572639), aperture view showing subfossil shell with superficial erosion of shell layer. B. Paratype (NMBE 578378), aperture view showing close up of lamellar band on columella (Fig. C). C. Paratype (NMBE 578378), aperture view showing damaged shell and columella. D. Close up view of holotype (NMBE 572639), showing body whorl with low, irregular growth lines. E. Paratype (NMBE 578378), protoconch." figureDoi="http://doi.org/10.5281/zenodo.10847854" httpUri="https://zenodo.org/record/10847854/files/figure.png" pageId="47" pageNumber="48">Fig. 17B–C, E</figureCitation>
);
<specimenCount id="9D1FFD02FF95FFEFFD7AD62CFD34FE93" box="[645,736,300,327]" count="2" pageId="47" pageNumber="48" type="generic">2 specs</specimenCount>
(broken shells); same collection data as for holotype;
<specimenCode id="DBBF9EF0FF95FFEFFF42D650FEA5FEBE" box="[189,369,335,362]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="47" pageNumber="48">NMBE 578378</specimenCode>
(ex AJC 2483)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF95FFEFFF42D690FDE6FCC5" pageId="47" pageNumber="48" type="description">
<paragraph id="8BA6368BFF95FFEFFF42D690FE9AFE7E" blockId="47.[189,334,400,426]" box="[189,334,400,426]" pageId="47" pageNumber="48">
<heading id="D0EE81E7FF95FFEFFF42D690FE9AFE7E" bold="true" box="[189,334,400,426]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96DEA99FF95FFEFFF42D690FE9AFE7E" bold="true" box="[189,334,400,426]" pageId="47" pageNumber="48">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D6C0FEB9FE28" blockId="47.[189,1398,447,509]" pageId="47" pageNumber="48">
<smallCapsWord id="8D40A057FF95FFEFFF42D6C0FEBAFE0C" baselines="467,467" box="[189,366,448,474]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="47" pageNumber="48">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="54A28829FF95FFEFFE82D6BFFE3BFE0D" box="[381,495,447,473]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
: sh:
<quantity id="4CE19B6EFF95FFEFFDDDD6BFFD59FE0E" box="[546,653,447,474]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="47" pageNumber="48" unit="mm" value="1.37">1.37 mm</quantity>
; sw:
<quantity id="4CE19B6EFF95FFEFFD37D6BFFCE0FE0E" box="[712,820,447,474]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="47" pageNumber="48" unit="mm" value="0.96">0.96 mm</quantity>
; ah:
<quantity id="4CE19B6EFF95FFEFFC95D6BFFC01FE0E" box="[874,981,447,474]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="47" pageNumber="48" unit="mm" value="0.61">0.61 mm</quantity>
; aw:
<quantity id="4CE19B6EFF95FFEFFBEDD6BFFBA9FE0E" box="[1042,1149,447,474]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="47" pageNumber="48" unit="mm" value="0.61">0.61 mm</quantity>
; 0.63; hlw:
<quantity id="4CE19B6EFF95FFEFFAF9D6BFFAA6FE0E" box="[1286,1394,447,474]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="47" pageNumber="48" unit="mm" value="0.88">0.88 mm</quantity>
; SA: 68.04 deg.
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D524FDE6FCC5" blockId="47.[189,1400,547,786]" pageId="47" pageNumber="48">Shell subfossil, eroded; conical with 5¼ flatly convex whorls; aperture reniform; penultimate whorl over ½ height of body whorl; teleoconch smooth with intermittent impressed striations on the body whorl; suture deep; parietal shield well defined, thick, long, and angular, curved at max. whorl convexity; peristome notch prominent at upper junction with parietal shield; smooth, impressed irregular ribbing behind peristome; columella slender with a distinctly sculpted and broad lamellar band obliquely projecting ca ⅓ beyond the width of the slender columella; a second, barely visible low ripple is present at mid-section of the columella.</paragraph>
</subSubSection>
<subSubSection id="C3036500FF95FFEFFF42D438FB0FFC56" box="[189,1243,824,898]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BA6368BFF95FFEFFF42D438FE82FC86" blockId="47.[189,342,824,850]" box="[189,342,824,850]" pageId="47" pageNumber="48">
<heading id="D0EE81E7FF95FFEFFF42D438FE82FC86" bold="true" box="[189,342,824,850]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96DEA99FF95FFEFFF42D438FE82FC86" bold="true" box="[189,342,824,850]" pageId="47" pageNumber="48">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D467FB0FFC56" blockId="47.[189,1243,871,898]" box="[189,1243,871,898]" pageId="47" pageNumber="48">
This species is only known from the
<typeStatus id="54A28829FF95FFEFFD91D468FD75FC56" box="[622,673,872,898]" pageId="47" pageNumber="48">type</typeStatus>
locality, St John’s cave in Njeguši,
<collectingCountry id="F30E761BFF95FFEFFBBCD468FB02FC56" box="[1091,1238,872,898]" name="Serbia" pageId="47" pageNumber="48">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF95FFEFFF42D4A9FC54FADF" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8BA6368BFF95FFEFFF42D4A9FEFBFC17" blockId="47.[189,303,937,963]" box="[189,303,937,963]" pageId="47" pageNumber="48">
<heading id="D0EE81E7FF95FFEFFF42D4A9FEFBFC17" bold="true" box="[189,303,937,963]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96DEA99FF95FFEFFF42D4A9FEFBFC17" bold="true" box="[189,303,937,963]" pageId="47" pageNumber="48">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D4D7FC54FADF" blockId="47.[187,1400,983,1292]" pageId="47" pageNumber="48">
The species differs from
<taxonomicName id="4C194D08FF95FFEFFE28D4D8FBB5FC25" authority="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityName="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityYear="2024" box="[471,1121,983,1009]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="kolbae" status="sp. nov.">
<emphasis id="B96DEA99FF95FFEFFE28D4D8FD90FC25" box="[471,580,983,1009]" italics="true" pageId="47" pageNumber="48">Z. kolbae</emphasis>
Jochum, Inäbnit, Kneubühler &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF95FFEFFB98D4D8FB6BFC26" box="[1127,1215,984,1010]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
in its uniformly conical shell form and its narrow, reniform apertural shape. Though the straight and robust columella is reminiscent of that in
<taxonomicName id="4C194D08FF95FFEFFE19D31EFB9CFBE3" authority="Jochum, Schilthuizen &amp; Ruthensteiner" authorityName="Jochum, Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[486,1096,1053,1080]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="njunjicae" status="sp. nov.">
<emphasis id="B96DEA99FF95FFEFFE19D31EFDA1FBE3" box="[486,629,1054,1079]" italics="true" pageId="47" pageNumber="48">Z. njunjicae</emphasis>
Jochum, Schilthuizen &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF95FFEFFBADD31EFB65FBEC" box="[1106,1201,1054,1080]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, the pronounced configuration of the lamella differentiates significantly from the weak low lamella of that species (as seen in all its Micro-CT perspectives) except perhaps the dorsal view, which is different altogether in bearing a distinctive lamellar bulge and a secondary low lamella above the tortional formation at the base of the columella. Though the shell is eroded, it shares no whorl size dimensions nor body whorl or penultimate whorl shape with the unidentified and unassessed morphospecies,
<taxonomicName id="4C194D08FF95FFEFFB9CD3CEFAC4FB3C" box="[1123,1296,1230,1256]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="undefined-1">
<emphasis id="B96DEA99FF95FFEFFB9CD3CEFB1AFB33" box="[1123,1230,1230,1255]" italics="true" pageId="47" pageNumber="48">Zospeum</emphasis>
sp. 1
</taxonomicName>
(NMBE 577052–577053) (
<figureCitation id="13222A0EFF95FFEFFE68D3F1FDF5FADF" box="[407,545,1265,1291]" captionStart="Fig" captionStartId="45.[189,232,1787,1813]" captionTargetBox="[220,1366,561,1748]" captionTargetId="figure-212@45.[220,1366,561,1748]" captionTargetPageId="45" captionText="Fig. 16. Light microscopic images of full-bodied Zospeum Bourguignat, 1856 from Njeguši, St John’s cave. A–B. Zospeum kolbae Jochum, Inäbnit, Kneubühler &amp; Ruthensteiner sp. nov. (NMBE 571122– 571123), individuals assessed by DNA sequencing with sigmoid intestine showing through shells (shells were destroyed post imaging for tissue extraction). A. Holotype (NMBE 571122), shell of aliquot, aperture, and aperture facing left view. B. Paratype (NMBE 571123), shell of aliquot, aperture, and aperture facing left view. C. Undescribed Zospeum sp. 1 (NMBE 577052) showing all perspectives. D–E. Undescribed Zospeum sp. 1 (NMBE 577053/2) showing all perspectives." figureDoi="http://doi.org/10.5281/zenodo.10847850" httpUri="https://zenodo.org/record/10847850/files/figure.png" pageId="47" pageNumber="48">Fig. 16C–E</figureCitation>
) also found in St John’s cave.
</paragraph>
</subSubSection>
</treatment>
</document>
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