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<document id="EAB508DF850216F2FC8E9CB74A6F041C" ID-CLB-Dataset="8958" ID-DOI="10.11646/zootaxa.4820.3.1" ID-GBIF-Dataset="fd413580-2960-463f-a924-8c27eaeafc6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4397861" ID-ZooBank="44BD1E07-C9F3-4488-936E-819C2FCA18C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609210733456" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="03AF3A058D69FF8BFAA9FA02A172FA05" docLanguage="en" docName="zootaxa.4820.3.1.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Oculogaster) malabarensis Kluge 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="433" masterDocId="FF96427D8D72FFABFA3EFFACA367FFC4" masterDocTitle="Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="437" masterPageNumber="401" pageNumber="428" updateTime="1698848343517" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9657B79ED2B52E1C898E2DD3ECFA24E8">Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<treatment id="03AF3A058D69FF8BFAA9FA02A172FA05" ID-DOI="http://doi.org/10.5281/zenodo.4436259" ID-GBIF-Taxon="176573418" ID-Zenodo-Dep="4436259" LSID="urn:lsid:plazi:treatment:03AF3A058D69FF8BFAA9FA02A172FA05" httpUri="http://treatment.plazi.org/id/03AF3A058D69FF8BFAA9FA02A172FA05" lastPageId="32" lastPageNumber="433" pageId="27" pageNumber="428">
<subSubSection id="C31CD8988D69FFB0FAA9FA02A1D4FA0C" box="[151,691,1454,1481]" pageId="27" pageNumber="428" type="nomenclature">
<paragraph id="8BB98B138D69FFB0FAA9FA02A1D4FA0C" blockId="27.[151,691,1454,1516]" box="[151,691,1454,1481]" pageId="27" pageNumber="428">
<heading id="D0F13C7F8D69FFB0FAA9FA02A1D4FA0C" bold="true" box="[151,691,1454,1481]" fontSize="11" level="1" pageId="27" pageNumber="428" reason="1">
<taxonomicName id="4C06F0908D69FFB0FAA9FA02A1D4FA0C" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[151,691,1454,1481]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D69FFB0FAA9FA02A1D4FA0C" bold="true" box="[151,691,1454,1481]" pageId="27" pageNumber="428">
<emphasis id="B97257018D69FFB0FAA9FA02A274FA0D" bold="true" box="[151,275,1454,1481]" italics="true" pageId="27" pageNumber="428">Procloeon</emphasis>
(
<emphasis id="B97257018D69FFB0FB1AFA02A2D0FA0D" bold="true" box="[292,439,1454,1481]" italics="true" pageId="27" pageNumber="428">Oculogaster</emphasis>
)
<emphasis id="B97257018D69FFB0FBF6FA02A10EFA0D" bold="true" box="[456,617,1454,1481]" italics="true" pageId="27" pageNumber="428">malabarensis</emphasis>
<taxonomicNameLabel id="A241EA7A8D69FFB0F84FFA02A1D4FA0C" box="[625,691,1454,1480]" pageId="27" pageNumber="428" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C31CD8988D69FFB0FAA9FA7EA258FA28" box="[151,319,1490,1516]" pageId="27" pageNumber="428" type="description">
<paragraph id="8BB98B138D69FFB0FAA9FA7EA258FA28" blockId="27.[151,691,1454,1516]" box="[151,319,1490,1516]" pageId="27" pageNumber="428">
(
<figureCitation id="133D97968D69FFB0FA9EFA7EA250FA28" box="[160,311,1490,1516]" captionStart-0="FIGURES 109112" captionStart-1="FIGURES 113117" captionStart-2="FIGURES 118126" captionStart-3="FIGURES 127133" captionStartId-0="28.[151,264,1917,1941]" captionStartId-1="29.[151,264,1524,1548]" captionStartId-2="30.[151,264,1969,1993]" captionStartId-3="31.[151,264,1707,1731]" captionTargetBox-0="[151,1436,181,1894]" captionTargetBox-1="[151,1436,181,1501]" captionTargetBox-2="[151,1430,424,1938]" captionTargetBox-3="[151,1436,309,1684]" captionTargetId-0="figure-27@28.[151,1436,181,1894]" captionTargetId-1="figure-16@29.[151,1436,181,1501]" captionTargetId-2="figure-126@30.[151,1436,417,1946]" captionTargetId-3="figure-69@31.[151,1436,309,1684]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionTargetPageId-3="31" captionText-0="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." captionText-1="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." captionText-2="FIGURES 118126. Procloeon (Oculogaster) malabarensis sp. n. 118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae." captionText-3="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.4397915" figureDoi-1="http://doi.org/10.5281/zenodo.4397917" figureDoi-2="http://doi.org/10.5281/zenodo.4397919" figureDoi-3="http://doi.org/10.5281/zenodo.4397921" httpUri-0="https://zenodo.org/record/4397915/files/figure.png" httpUri-1="https://zenodo.org/record/4397917/files/figure.png" httpUri-2="https://zenodo.org/record/4397919/files/figure.png" httpUri-3="https://zenodo.org/record/4397921/files/figure.png" pageId="27" pageNumber="428">Fig. 109133</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D69FFB0FAA9F9BEA00EF9EE" box="[151,873,1553,1578]" pageId="27" pageNumber="428" type="reference_group">
<paragraph id="8BB98B138D69FFB0FAA9F9BEA00EF9EE" blockId="27.[151,873,1553,1578]" box="[151,873,1553,1578]" pageId="27" pageNumber="428">
<treatmentCitationGroup id="AB16AC3D8D69FFB0FAA9F9BEA00EF9EE" box="[151,873,1553,1578]" pageId="27" pageNumber="428">
?
<treatmentCitation id="0AA7AD028D69FFB0FA97F9BDA002F9EE" author="Sivaramakrishnan, K. G. &amp; Venkataraman, K." box="[169,869,1553,1578]" page="211" pageId="27" pageNumber="428" year="1990">
<taxonomicName id="4C06F0908D69FFB0FA97F9BDA002F9EE" ID-CoL="4MNWG" authority=": Sivaramakrishnan &amp; Venkataraman 1990: 211" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityPageNumber="211" authorityYear="1990" box="[169,869,1553,1578]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="regularum">
<emphasis id="B97257018D69FFB0FA97F9BDA2E5F9ED" box="[169,386,1553,1577]" italics="true" pageId="27" pageNumber="428">Procloeon regularum</emphasis>
:
<bibRefCitation id="EF97F6E28D69FFB0FBB1F9BDA002F9EE" author="Sivaramakrishnan, K. G. &amp; Venkataraman, K." box="[399,869,1553,1578]" pageId="27" pageNumber="428" pagination="209 - 213" refId="ref18995" refString="Sivaramakrishnan, K. G. &amp; Venkataraman, K. (1990) Abundance, altitudinal distribution and swarming of Ephemeroptera in Palni Hills, South India. In: Campbell, I. C. (Ed.), Mayflies and Stoneflies: Life Histories and Biology. Proceeding of the 5 th International Conference on Ephemeroptera &amp; 9 th Intfernational Symposium on Plecoptera, Marysville, Australia, 18 - 24 February 1987. Kluwer Acad. Publ., Dordrecht, pp. 209 - 213. https: // doi. org / 10.1007 / 978 - 94 - 009 - 2397 - 3 _ 24" type="book chapter" year="1990">Sivaramakrishnan &amp; Venkataraman 1990: 211</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D69FFB0FAA9F9FEA0E9F9A8" box="[151,910,1618,1645]" pageId="27" pageNumber="428" type="etymology">
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<emphasis id="B97257018D69FFB0FAA9F9FEA279F9A8" bold="true" box="[151,286,1618,1644]" pageId="27" pageNumber="428">Etymology.</emphasis>
The name refers to the historical area Malabar in
<collectingCountry id="F311CB838D69FFB0F973F9FEA0EDF9A8" box="[845,906,1618,1644]" name="India" pageId="27" pageNumber="428">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D69FF8BFAF9F9DAA269FA59" lastPageId="32" lastPageNumber="433" pageId="27" pageNumber="428" type="materials_examined">
<paragraph id="8BB98B138D69FFB0FAF9F9DAA055F93F" blockId="27.[151,1437,1618,2038]" pageId="27" pageNumber="428">
<emphasis id="B97257018D69FFB0FAF9F9DAA2CAF954" bold="true" box="[199,429,1654,1680]" pageId="27" pageNumber="428">Material examined</emphasis>
(
<collectionCode id="ED1713D68D69FFB0FBFFF9DAA292F954" box="[449,501,1654,1680]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="428" type="Museum">ZIN</collectionCode>
).
<emphasis id="B97257018D69FFB0F832F9DAA1E5F954" bold="true" box="[524,642,1654,1680]" pageId="27" pageNumber="428">
<typeStatus id="54BD35B18D69FFB0F832F9DAA11AF954" box="[524,637,1654,1680]" pageId="27" pageNumber="428" type="holotype">Holotype</typeStatus>
:
</emphasis>
L-S-I
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{
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specimen [IX](11)2013},
<collectingCountry id="F311CB838D69FFB0FE1CF9DAA713F954" box="[1058,1140,1654,1680]" name="India" pageId="27" pageNumber="428">INDIA</collectingCountry>
,
<collectingRegion id="49C245F18D69FFB0FEBDF9DAA79EF954" box="[1155,1273,1654,1680]" country="India" name="Karnataka" pageId="27" pageNumber="428">Karnataka</collectingRegion>
,
<collectingCounty id="62D8F39F8D69FFB0FF36F9DAA6F4F954" box="[1288,1427,1654,1680]" pageId="27" pageNumber="428">Shivamogga</collectingCounty>
/
<collectingMunicipality id="6BDD11698D69FFB0FAA9F935A3BAF977" box="[151,221,1689,1715]" pageId="27" pageNumber="428">Udupi</collectingMunicipality>
districts near
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,
<date id="FFB8ADD38D69FFB0FBCBF935A104F970" box="[501,611,1689,1716]" pageId="27" pageNumber="428" value="2013-01-25">
<collectingDate id="EFFC543B8D69FFB0FBCBF935A104F970" box="[501,611,1689,1716]" pageId="27" pageNumber="428" value="2013-01-25">25.I.2013</collectingDate>
</date>
, coll.
<collectorName id="26F3EEC58D69FFB0F893F936A071F977" box="[685,790,1689,1716]" pageId="27" pageNumber="428">N. Kluge</collectorName>
&amp;
<collectorName id="26F3EEC58D69FFB0F97FF936A0D8F977" box="[833,959,1689,1716]" pageId="27" pageNumber="428">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation id="3B6E814E8D69FFB0F9F6F936A2F7F913" ID-GBIF-Occurrence="3014991302" collectingDate="2013-01-11" collectingDateMax="2013-01-31" collectingDateMin="2013-01-11" collectorName="N. Kluge &amp; L. Sheyko." country="India" county="Shivamogga" location="Agumbe" municipality="Udupi" pageId="27" pageNumber="428" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="paratype">
<emphasis id="B97257018D69FFB0F9F6F936A72DF970" bold="true" box="[968,1098,1690,1716]" pageId="27" pageNumber="428">
<typeStatus id="54BD35B18D69FFB0F9F6F936A723F970" box="[968,1092,1690,1716]" pageId="27" pageNumber="428" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same locality and collectors,
<date id="FFB8ADD38D69FFB0FAA9F911A257F913" box="[151,304,1725,1751]" pageId="27" pageNumber="428" value="2013-01-11" valueMax="2013-01-31" valueMin="2013-01-11">
<collectingDate id="EFFC543B8D69FFB0FAA9F911A257F913" box="[151,304,1725,1751]" pageId="27" pageNumber="428" value="2013-01-11" valueMax="2013-01-31" valueMin="2013-01-11">1131.I.2013</collectingDate>
</date>
: 2 L-S
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, 1 L/S
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, 3 L-S
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,
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;
<materialsCitation id="3B6E814E8D69FFB0F8FAF911A2E5F93F" ID-GBIF-Occurrence="3014991311" collectingDate="2016-01-18" collectingDateMax="2016-01-21" collectingDateMin="2016-01-18" collectorName="N. Kluge &amp; L. Sheyko" country="India" location="Erumeli" municipality="Kottayam district" pageId="27" pageNumber="428" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="paratype">
<collectingRegion id="49C245F18D69FFB0F8FAF911A076F913" box="[708,785,1725,1751]" country="India" name="Kerala" pageId="27" pageNumber="428">Kerala</collectingRegion>
,
<collectingMunicipality id="6BDD11698D69FFB0F922F912A085F913" box="[796,994,1725,1752]" pageId="27" pageNumber="428">Kottayam district</collectingMunicipality>
,
<location id="8ED9DDC88D69FFB0F9D2F911A72EF913" LSID="urn:lsid:plazi:treatment:03AF3A058D69FF8BFAA9FA02A172FA05:8ED9DDC88D69FFB0F9D2F911A72EF913" box="[1004,1097,1725,1751]" country="India" municipality="Kottayam district" name="Erumeli" pageId="27" pageNumber="428" stateProvince="Kerala">Erumeli</location>
,
<date id="FFB8ADD38D69FFB0FE6AF911A78BF913" box="[1108,1260,1725,1751]" pageId="27" pageNumber="428" value="2016-01-18" valueMax="2016-01-21" valueMin="2016-01-18">
<collectingDate id="EFFC543B8D69FFB0FE6AF911A78BF913" box="[1108,1260,1725,1751]" pageId="27" pageNumber="428" value="2016-01-18" valueMax="2016-01-21" valueMin="2016-01-18">1821.I.2016</collectingDate>
</date>
, coll.
<collectorName id="26F3EEC58D69FFB0FF11F912A6FFF913" box="[1327,1432,1725,1752]" pageId="27" pageNumber="428">N. Kluge</collectorName>
,
<collectorName id="26F3EEC58D69FFB0FAA9F94EA269F93F" box="[151,270,1761,1788]" pageId="27" pageNumber="428">L. Sheyko</collectorName>
: 3 L-S-I
<specimenCount id="9D00409A8D69FFB0FB51F94DA2E5F93F" box="[367,386,1761,1787]" pageId="27" pageNumber="428" type="male"></specimenCount>
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, 4 S-I
<specimenCount id="9D00409A8D69FFB0FBF6F94DA2BBF93F" box="[456,476,1761,1787]" pageId="27" pageNumber="428" type="male"></specimenCount>
, 4 S
<specimenCount id="9D00409A8D69FFB0F833F94DA144F93F" box="[525,547,1761,1787]" pageId="27" pageNumber="428" type="male"></specimenCount>
, 2 S-I
<specimenCount id="9D00409A8D69FFB0F850F94DA119F93F" box="[622,638,1761,1787]" pageId="27" pageNumber="428" type="female"></specimenCount>
, 2 S
<specimenCount id="9D00409A8D69FFB0F88CF94DA1A3F93F" box="[690,708,1761,1787]" pageId="27" pageNumber="428" type="female"></specimenCount>
,
<specimenCount id="9D00409A8D69FFB0F8EFF94DA049F93F" box="[721,814,1761,1787]" pageId="27" pageNumber="428" type="larva">2 larvae</specimenCount>
.
</paragraph>
<paragraph id="8BB98B138D69FFB0FAF9F8A9A66FF832" blockId="27.[151,1437,1618,2038]" pageId="27" pageNumber="428">
<emphasis id="B97257018D69FFB0FAF9F8A9A274F8DB" bold="true" box="[199,275,1797,1823]" pageId="27" pageNumber="428">
<emphasis id="B97257018D69FFB0FAF9F8A9A26CF8DB" bold="true" box="[199,267,1797,1823]" italics="true" pageId="27" pageNumber="428">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION. Head with brownish and colorless areas. Pronotum and mesonotum ocher or brownish with lighter and darker areas. Metanotum and thoracic pleura with colorless and brown areas, thoracic sterna colorless. Legs colorless or brownish, either unicolor, or with diffuse brown band in distal part of femur; tarsus darkened. Abdominal terga with maculation different on different segments; most or all terga with submedian pair of blanks, either corresponding to sigilla, or larger than sigilla; terga IV and VII more-or-less lighter than others; tergum
<collectionCode id="ED1713D68D69FFB0FB45F814A2FEF816" box="[379,409,1976,2002]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
darker than others (
<figureCitation id="133D97968D69FFB0F840F81BA185F816" box="[638,738,1975,2002]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="27" pageNumber="428">Fig. 109</figureCitation>
). Abdominal sterna lighter than terga, either unicolor, or with lighter and darker areas. Caudalii ocher with posterior margin of every 4th segment brown (
<figureCitation id="133D97968D69FFB0FEA4F877A79BF832" box="[1178,1276,2011,2038]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="27" pageNumber="428">Fig. 112</figureCitation>
).
</paragraph>
<caption id="DF79DB9B8D6EFFB7FAA9F8D1A20CF81A" ID-DOI="http://doi.org/10.5281/zenodo.4397915" ID-Zenodo-Dep="4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="28" pageNumber="429" startId="28.[151,264,1917,1941]" targetBox="[151,1436,181,1894]" targetPageId="28">
<paragraph id="8BB98B138D6EFFB7FAA9F8D1A20CF81A" blockId="28.[151,1437,1917,2014]" pageId="28" pageNumber="429">
<emphasis id="B97257018D6EFFB7FAA9F8D1A208F852" bold="true" box="[151,367,1917,1942]" pageId="28" pageNumber="429">FIGURES 109112.</emphasis>
Larvae of
<taxonomicName id="4C06F0908D6EFFB7FBDBF8D2A012F852" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[485,885,1918,1942]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D6EFFB7FBDBF8D2A129F852" box="[485,590,1918,1942]" italics="true" pageId="28" pageNumber="429">Procloeon</emphasis>
(
<emphasis id="B97257018D6EFFB7F861F8D2A1BBF852" box="[607,732,1918,1942]" italics="true" pageId="28" pageNumber="429">Oculogaster</emphasis>
)
<emphasis id="B97257018D6EFFB7F8D3F8D2A012F852" box="[749,885,1918,1942]" italics="true" pageId="28" pageNumber="429">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D6EFFB7F940F8D2A0DEF852" bold="true" box="[894,953,1918,1942]" pageId="28" pageNumber="429">
<taxonomicNameLabel id="A241EA7A8D6EFFB7F940F8D2A0DEF852" box="[894,953,1918,1942]" pageId="28" pageNumber="429" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype).
</paragraph>
</caption>
<caption id="DF79DB9B8D6FFFB6FAA9FA58A634F9F5" ID-DOI="http://doi.org/10.5281/zenodo.4397917" ID-Zenodo-Dep="4397917" httpUri="https://zenodo.org/record/4397917/files/figure.png" pageId="29" pageNumber="430" startId="29.[151,264,1524,1548]" targetBox="[151,1436,181,1501]" targetPageId="29">
<paragraph id="8BB98B138D6FFFB6FAA9FA58A634F9F5" blockId="29.[151,1436,1524,1585]" pageId="29" pageNumber="430">
<emphasis id="B97257018D6FFFB6FAA9FA58A20BF9C8" bold="true" box="[151,364,1524,1548]" pageId="29" pageNumber="430">FIGURES 113117.</emphasis>
Larva of
<taxonomicName id="4C06F0908D6FFFB6FBE6FA59A00FF9C9" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[472,872,1525,1549]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D6FFFB6FBE6FA59A126F9C9" box="[472,577,1525,1549]" italics="true" pageId="29" pageNumber="430">Procloeon</emphasis>
(
<emphasis id="B97257018D6FFFB6F86CFA59A1A8F9C9" box="[594,719,1525,1549]" italics="true" pageId="29" pageNumber="430">Oculogaster</emphasis>
)
<emphasis id="B97257018D6FFFB6F8DEFA59A00FF9C9" box="[736,872,1525,1549]" italics="true" pageId="29" pageNumber="430">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D6FFFB6F94FFA59A0CBF9C9" bold="true" box="[881,940,1525,1549]" pageId="29" pageNumber="430">
<taxonomicNameLabel id="A241EA7A8D6FFFB6F94FFA59A0CBF9C9" box="[881,940,1525,1549]" pageId="29" pageNumber="430" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side).
</paragraph>
</caption>
<paragraph id="8BB98B138D6FFFB6FAF9F9F3A73DF959" blockId="29.[151,1437,1631,2017]" pageId="29" pageNumber="430">
HYPODERMAL COLORATION. Either absent or (just before moult to subimago) as in respective winged stages; male (
<figureCitation id="133D97968D6FFFB6FB0EF92FA2F5F959" box="[304,402,1667,1693]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="29" pageNumber="430">Fig. 109</figureCitation>
; as in
<figureCitation id="133D97968D6FFFB6FBE6F92FA114F959" box="[472,627,1667,1693]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="29" pageNumber="430">Figs 127129</figureCitation>
) and female (as in
<figureCitation id="133D97968D6FFFB6F977F92FA084F959" box="[841,995,1667,1693]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="29" pageNumber="430">Figs 132133</figureCitation>
) different.
</paragraph>
<paragraph id="8BB98B138D6FFFB5FAF9F90BA2D3FE85" blockId="29.[151,1437,1631,2017]" lastBlockId="30.[151,1437,151,393]" lastPageId="30" lastPageNumber="431" pageId="29" pageNumber="430">
<collectionCode id="ED1713D68D6FFFB6FAF9F90BA288F905" box="[199,495,1703,1729]" pageId="29" pageNumber="430">SHAPE AND SETATION</collectionCode>
. Mouth parts as in
<figureCitation id="133D97968D6FFFB6F8FFF90BA03CF905" box="[705,859,1703,1729]" captionStart="FIGURES 113117" captionStartId="29.[151,264,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-16@29.[151,1436,181,1501]" captionTargetPageId="29" captionText="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.4397917" httpUri="https://zenodo.org/record/4397917/files/figure.png" pageId="29" pageNumber="430">Figs 113117</figureCitation>
; maxillary palp 3-segmented, with 2nd and 3rd segments subequal (
<figureCitation id="133D97968D6FFFB6FB6CF967A2D5F922" box="[338,434,1739,1766]" captionStart="FIGURES 113117" captionStartId="29.[151,264,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-16@29.[151,1436,181,1501]" captionTargetPageId="29" captionText="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.4397917" httpUri="https://zenodo.org/record/4397917/files/figure.png" pageId="29" pageNumber="430">Fig. 116</figureCitation>
). Femora with very small spine-like setae on outer and inner sides, with 2 spine-like apical setae located in plane of leg (
<figureCitation id="133D97968D6FFFB6FBE0F943A158F8CD" box="[478,575,1775,1801]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="29" pageNumber="430">Fig. 110</figureCitation>
). Claws moderately long, in proximal part with two rows of minute denticles increasing distally (
<figureCitation id="133D97968D6FFFB6FB49F8BFA2BEF8E9" box="[375,473,1811,1837]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="29" pageNumber="430">Fig. 111</figureCitation>
). Hind protoptera completely absent. Abdominal segment IX with 37 lateral spines on each side, other segments without lateral spines; postero-lateral spines present on segments IIIX or IIIIX, being minute on anterior segments. Posterior margin of abdominal tergum I with few sparse small spine-like pointed denticles; posterior margins of terga IIX with spine-like pointed denticles of unequal length; on all terga, including tergum IX, row of denticles not interrupted medially. Abdominal terga with short scales in wide semilunar or W-shaped non-opercula-bearing sockets, with sparse fine setae. Posterior margins of abdominal sterna IIV or IIII smooth, posterior margin of sternum
<collectionCode id="ED1713D68D6CFFB5F80CFF3BA121FF75" box="[562,582,151,177]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="431" type="Museum">V</collectionCode>
or sterna IVV with few minute denticles, posterior margins of sterna VIIX and paraprocts with spine-like pointed denticles of unequal length. Abdominal sterna with sparse scales and sparse fine setae not forming regular rows. Tergalii widest near base, without dorsal lamellae; costal ribs without denticles (as in
<figureCitation id="133D97968D6CFFB5FAE3FEAFA21DFED9" box="[221,378,259,285]" captionStart="FIGURES 118126" captionStartId="30.[151,264,1969,1993]" captionTargetBox="[151,1430,424,1938]" captionTargetId="figure-126@30.[151,1436,417,1946]" captionTargetPageId="30" captionText="FIGURES 118126. Procloeon (Oculogaster) malabarensis sp. n. 118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397919" httpUri="https://zenodo.org/record/4397919/files/figure.png" pageId="30" pageNumber="431">Figs 118124</figureCitation>
). In distal part of cercus, spine on outer side of each segment as long as combined length of 2 next segments (
<figureCitation id="133D97968D6CFFB5FB78FE8BA2C0FE85" box="[326,423,295,321]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="30" pageNumber="431">Fig. 112</figureCitation>
).
</paragraph>
<paragraph id="8BB98B138D6CFFB5FAF9FEE6A23AFE4D" blockId="30.[151,1437,151,393]" pageId="30" pageNumber="431">
<emphasis id="B97257018D6CFFB5FAF9FEE6A224FEA1" bold="true" box="[199,323,330,357]" italics="true" pageId="30" pageNumber="431">Subimago.</emphasis>
CUTICULAR COLORATION. Cuticle either entirely colorless, or with postsubalar sclerite partly brown (
<figureCitation id="133D97968D6CFFB5FAD0FEC3A237FE4D" box="[238,336,367,393]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="30" pageNumber="431">Fig. 130</figureCitation>
).
</paragraph>
<caption id="DF79DB9B8D6CFFB5FAA9F81DA19BF829" ID-DOI="http://doi.org/10.5281/zenodo.4397919" ID-Zenodo-Dep="4397919" httpUri="https://zenodo.org/record/4397919/files/figure.png" pageId="30" pageNumber="431" startId="30.[151,264,1969,1993]" targetBox="[151,1430,424,1938]" targetPageId="30">
<paragraph id="8BB98B138D6CFFB5FAA9F81DA19BF829" blockId="30.[151,1436,1969,2029]" pageId="30" pageNumber="431">
<emphasis id="B97257018D6CFFB5FAA9F81DA209F80D" bold="true" box="[151,366,1969,1993]" pageId="30" pageNumber="431">FIGURES 118126.</emphasis>
<taxonomicName id="4C06F0908D6CFFB5FB4BF81DA064F80D" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[373,771,1969,1994]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D6CFFB5FB4BF81DA2B9F80D" box="[373,478,1969,1993]" italics="true" pageId="30" pageNumber="431">Procloeon</emphasis>
(
<emphasis id="B97257018D6CFFB5FBD0F81DA10CF80D" box="[494,619,1969,1993]" italics="true" pageId="30" pageNumber="431">Oculogaster</emphasis>
)
<emphasis id="B97257018D6CFFB5F845F81DA064F80D" box="[635,771,1969,1993]" italics="true" pageId="30" pageNumber="431">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D6CFFB5F935F81EA022F80E" bold="true" box="[779,837,1970,1994]" pageId="30" pageNumber="431">
<taxonomicNameLabel id="A241EA7A8D6CFFB5F935F81EA022F80E" box="[779,837,1970,1994]" pageId="30" pageNumber="431" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae.
</paragraph>
</caption>
<paragraph id="8BB98B138D6DFFB4FAF9FF3BA749FF76" blockId="31.[151,1436,151,285]" box="[199,1070,151,178]" pageId="31" pageNumber="432">HYPODERMAL COLORATION. As in imago (see below, male and female).</paragraph>
<paragraph id="8BB98B138D6DFFB4FAF9FF17A26DFED9" blockId="31.[151,1436,151,285]" pageId="31" pageNumber="432">
TEXTURE. On fore leg of male and female 1st tarsal segment covered partly with microtrichia (as tibia), partly with pointed microlepides; all other tarsal segments entirely covered with pointed microlepides (as in
<figureCitation id="133D97968D6DFFB4FF52FF73A399FED9" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="31" pageNumber="432">Figs 105106</figureCitation>
).
</paragraph>
<caption id="DF79DB9B8D6DFFB4FAA9F907A267F8CF" ID-DOI="http://doi.org/10.5281/zenodo.4397921" ID-Zenodo-Dep="4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="31" pageNumber="432" startId="31.[151,264,1707,1731]" targetBox="[151,1436,309,1684]" targetPageId="31">
<paragraph id="8BB98B138D6DFFB4FAA9F907A267F8CF" blockId="31.[151,1436,1707,1803]" pageId="31" pageNumber="432">
<emphasis id="B97257018D6DFFB4FAA9F907A209F907" bold="true" box="[151,366,1707,1731]" pageId="31" pageNumber="432">FIGURES 127133.</emphasis>
<taxonomicName id="4C06F0908D6DFFB4FB4AF907A066F907" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[372,769,1707,1732]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D6DFFB4FB4AF907A2BAF907" box="[372,477,1707,1731]" italics="true" pageId="31" pageNumber="432">Procloeon</emphasis>
(
<emphasis id="B97257018D6DFFB4FBD2F907A10EF907" box="[492,617,1707,1731]" italics="true" pageId="31" pageNumber="432">Oculogaster</emphasis>
)
<emphasis id="B97257018D6DFFB4F847F907A066F907" box="[633,769,1707,1731]" italics="true" pageId="31" pageNumber="432">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D6DFFB4F939F900A026F900" bold="true" box="[775,833,1708,1732]" pageId="31" pageNumber="432">
<taxonomicNameLabel id="A241EA7A8D6DFFB4F939F900A026F900" box="[775,833,1708,1732]" pageId="31" pageNumber="432" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype).
</paragraph>
</caption>
<paragraph id="8BB98B138D6DFF8BFAF9F896A116FEA1" blockId="31.[151,1436,1849,2020]" lastBlockId="32.[151,1437,150,1473]" lastPageId="32" lastPageNumber="433" pageId="31" pageNumber="432">
<emphasis id="B97257018D6DFFB4FAF9F896A23FF890" bold="true" box="[199,344,1849,1876]" italics="true" pageId="31" pageNumber="432">Imago, male</emphasis>
(
<figureCitation id="133D97968D6DFFB4FB56F896A160F890" box="[360,519,1850,1876]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="31" pageNumber="432">Figs 128129</figureCitation>
). Head ocher or whitish. Antenna with scapus and pedicellus non-pigmented, flagellum either non-pigmented, or brown. Turbinate eyes widened apically; either uniformly orange, or orange with yellow ring bordering facetted surface. Thorax ocher or whitish. Legs pale ocher. In
<typeStatus id="54BD35B18D6DFFB4FE6BF82EA7DFF858" box="[1109,1208,1922,1948]" pageId="31" pageNumber="432" type="paratype">paratype</typeStatus>
, ratio femur / tibia / tarsal segments (mm) on fore leg 0.80: 1.02: 0.03: 0.47: 0.30: 0.16: 0.12; on middle and hind leg 0.68: 0.68: 0.29: 0.10: 0.03: 0.11. Middle and hind legs without apical spine on 1st+2nd tarsomere, with single apical spine on primary 3rd tarsomere (as in
<figureCitation id="133D97968D52FF8BF834FF3BA10AFF75" box="[522,621,150,177]" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="32" pageNumber="433">Fig. 105</figureCitation>
). Wing with membrane colorless, veins ocher or brownish. Pterostigma with 13 oblique veins; marginal intercalary present in each interspace. Hind wing absent. Abdominal terga and sterna IVI translucent, whitish, either without hypodermal markings, or with pair of small orange or red-brown submedian spots on tergum IV (
<figureCitation id="133D97968D52FF8BF83FFEAFA1FBFED9" box="[513,668,259,285]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Fig. 127129</figureCitation>
); lateral tracheae black. Most part of tergum VII and smaller part of tergum VIII red-brown; terga IXX entirely white or ocher. Genitalia non-pigmented. Median projection of penial bridge truncated (
<figureCitation id="133D97968D52FF8BFB63FEE7A2A7FEA1" box="[349,448,330,357]" captionStart="FIGURES 118126" captionStartId="30.[151,264,1969,1993]" captionTargetBox="[151,1430,424,1938]" captionTargetId="figure-126@30.[151,1436,417,1946]" captionTargetPageId="30" captionText="FIGURES 118126. Procloeon (Oculogaster) malabarensis sp. n. 118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397919" httpUri="https://zenodo.org/record/4397919/files/figure.png" pageId="32" pageNumber="433">Fig. 125</figureCitation>
). Cerci whitish.
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FEC3A6FAFD6D" blockId="32.[151,1437,150,1473]" pageId="32" pageNumber="433">
<emphasis id="B97257018D52FF8BFAF9FEC3A209FE4D" bold="true" box="[199,366,366,393]" italics="true" pageId="32" pageNumber="433">Imago, female</emphasis>
(
<figureCitation id="133D97968D52FF8BFB43FEC3A283FE4D" box="[381,484,367,393]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Figs 131</figureCitation>
). Head moderately wide, distance between eyes subequal to eye length; eyes significantly elevated above head surface (as in
<figureCitation id="133D97968D52FF8BF859FE3FA1AFFE69" box="[615,712,403,429]" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="32" pageNumber="433">Fig. 103</figureCitation>
). Head and thorax ocher, without brown spots, thorax sometimes with white markings. Legs as in male (see above); fore leg without apical spine on 2nd tarsomere, with single apical spine on primary 3rd tarsomere (as in
<figureCitation id="133D97968D52FF8BF87DFE77A1C2FE32" box="[579,677,475,502]" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="32" pageNumber="433">Fig. 106</figureCitation>
); same on middle and hind legs (as in
<figureCitation id="133D97968D52FF8BFE6AFE77A7D5FE31" box="[1108,1202,474,501]" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="32" pageNumber="433">Fig. 105</figureCitation>
) (as in male). Wings as in male (see above). Abdomen ocher; terga IIVII laterally with contrastingly brown or black markings forming oblique stripes, often interrupted; markings on segments II, IV and VII larger than others (
<figureCitation id="133D97968D52FF8BFE42FD8FA67CFDF9" box="[1148,1307,547,573]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Figs 131133</figureCitation>
). Orange or red spots never present; white pigmentation between black markings either present (
<figureCitation id="133D97968D52FF8BFE72FDEBA78EFDA5" box="[1100,1257,583,609]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Figs 131132</figureCitation>
) or absent (
<figureCitation id="133D97968D52FF8BFF51FDEBA3A2FD41" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Fig. 133</figureCitation>
). Sterna IIVII or IIVIII laterally with dark brown or black markings in form of longitudinal interrupted lines or small spots (absent in young subimagines). Lateral tracheae colored with dark brown or black. Cerci whitish.
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FD1FA1E2FD0A" blockId="32.[151,1437,150,1473]" box="[199,645,691,718]" pageId="32" pageNumber="433">
<emphasis id="B97257018D52FF8BFAF9FD1FA39BFD09" bold="true" box="[199,252,691,717]" italics="true" pageId="32" pageNumber="433">Egg.</emphasis>
Without rigid chorion (viviparity).
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FD7AA1F3FD35" blockId="32.[151,1437,150,1473]" box="[199,660,726,753]" pageId="32" pageNumber="433">
<emphasis id="B97257018D52FF8BFAF9FD7AA229FD34" bold="true" box="[199,334,726,752]" pageId="32" pageNumber="433">Dimension.</emphasis>
Fore wing length
<quantity id="4CFE26F68D52FF8BF825FD7AA1F7FD35" box="[539,656,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="32" pageNumber="433" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.54 mm</quantity>
.
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FD56A629FCFD" blockId="32.[151,1437,150,1473]" pageId="32" pageNumber="433">
<emphasis id="B97257018D52FF8BFAF9FD56A205FCD0" bold="true" box="[199,354,762,788]" pageId="32" pageNumber="433">Comparison.</emphasis>
The new species
<taxonomicName id="4C06F0908D52FF8BF817FD57A077FCD1" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[553,784,763,789]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BF817FD57A158FCD0" box="[553,575,763,788]" italics="true" pageId="32" pageNumber="433">P.</emphasis>
(
<emphasis id="B97257018D52FF8BF870FD57A10EFCD1" box="[590,617,763,789]" italics="true" pageId="32" pageNumber="433">O.</emphasis>
)
<emphasis id="B97257018D52FF8BF844FD57A077FCD1" box="[634,784,763,789]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BF929FD57A030FCD1" bold="true" box="[791,855,763,789]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BF929FD57A030FCD1" box="[791,855,763,789]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is closely related to
<taxonomicName id="4C06F0908D52FF8BFE00FD57A663FCD1" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" box="[1086,1284,763,789]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFE00FD57A734FCD0" box="[1086,1107,763,788]" italics="true" pageId="32" pageNumber="433">P.</emphasis>
(
<emphasis id="B97257018D52FF8BFE5DFD57A719FCD1" box="[1123,1150,763,789]" italics="true" pageId="32" pageNumber="433">O.</emphasis>
)
<emphasis id="B97257018D52FF8BFEB1FD57A663FCD1" box="[1167,1284,763,789]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
, but it differs by smaller size, by truncated median projection of penial bridge and by hypodermal coloration of abdomen.
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FCEFA273FB65" blockId="32.[151,1437,150,1473]" pageId="32" pageNumber="433">
The difference in hypodermal coloration is better expressed in the female (thats why the reared female imago is designated as the
<typeStatus id="54BD35B18D52FF8BFB41FCCBA281FC46" box="[383,486,871,898]" pageId="32" pageNumber="433" type="holotype">holotype</typeStatus>
). Females differ as follows (compare
<figureCitation id="133D97968D52FF8BF9A8FCCBA756FC45" box="[918,1073,871,897]" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="32" pageNumber="433">Figs 103104</figureCitation>
and
<figureCitation id="133D97968D52FF8BFE55FCCBA7B4FC45" box="[1131,1235,871,897]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">131133</figureCitation>
): In contrast to
<taxonomicName id="4C06F0908D52FF8BFFB6FCCBA225FC61" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFFB6FCCBA6FAFC44" box="[1416,1437,871,896]" italics="true" pageId="32" pageNumber="433">P.</emphasis>
(
<emphasis id="B97257018D52FF8BFA9FFC27A3DBFC61" box="[161,188,907,933]" italics="true" pageId="32" pageNumber="433">O.</emphasis>
)
<emphasis id="B97257018D52FF8BFAF3FC27A225FC61" box="[205,322,907,933]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
, whose winged females have paired reddish spots on terga IV and VII, females of
<taxonomicName id="4C06F0908D52FF8BFED3FC27A3BEFC0C" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFED3FC27A665FC60" box="[1261,1282,907,932]" italics="true" pageId="32" pageNumber="433">P.</emphasis>
(
<emphasis id="B97257018D52FF8BFF2CFC27A649FC61" box="[1298,1326,907,933]" italics="true" pageId="32" pageNumber="433">O.</emphasis>
)
<emphasis id="B97257018D52FF8BFF01FC27A3BEFC0C" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BFADFFC03A246FC0D" bold="true" box="[225,289,943,969]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BFADFFC03A246FC0D" box="[225,289,943,969]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
have no such spots, but only brown or black lateral markings. The number and shape of the brown or black lateral markings also differs in these two species: In
<taxonomicName id="4C06F0908D52FF8BF913FC7FA090FC29" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" box="[813,1015,979,1005]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BF913FC7FA059FC28" box="[813,830,979,1004]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BF968FC7FA00DFC29" box="[854,874,979,1005]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF9BCFC7FA090FC29" box="[898,1015,979,1005]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
such markings are present beginning from tergum I, but in
<taxonomicName id="4C06F0908D52FF8BFBB0FC5BA11DFBD5" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[398,634,1015,1041]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFBB0FC5BA2F8FBD4" box="[398,415,1015,1040]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFB89FC5BA2ACFBD5" box="[439,459,1015,1041]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BFBDAFC5BA11DFBD5" box="[484,634,1015,1041]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BF8BFFC5BA1A5FBD5" bold="true" box="[641,706,1015,1041]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BF8BFFC5BA1A5FBD5" box="[641,706,1015,1041]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
they are present beginning from tergum II; in
<taxonomicName id="4C06F0908D52FF8BFEEFFC5BA6FBFBD5" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" box="[1233,1436,1015,1041]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFEEFFC5BA785FBD4" box="[1233,1250,1015,1040]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFEC5FC5BA668FBD5" box="[1275,1295,1015,1041]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BFF19FC5BA6FBFBD5" box="[1319,1436,1015,1041]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
markings on terga IIIII and VVI are integral and roundish, but in
<taxonomicName id="4C06F0908D52FF8BF9ABFBB7A7E7FBF1" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[917,1152,1051,1077]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BF9ABFBB7A0C1FBF0" box="[917,934,1051,1076]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BF980FBB7A0B5FBF1" box="[958,978,1051,1077]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF9D4FBB7A7E7FBF1" box="[1002,1152,1051,1077]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BFEB6FBB7A7AFFBF1" bold="true" box="[1160,1224,1051,1077]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BFEB6FBB7A7AFFBF1" box="[1160,1224,1051,1077]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
markings on these terga have form of oblique, narrow, interrupted lines. Dark brown or black markings on abdominal sterna are present in female imagines of
<taxonomicName id="4C06F0908D52FF8BFB85FBCFA1C2FBB9" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[443,677,1123,1149]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFB85FBCFA2ABFBB8" box="[443,460,1123,1148]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFBDDFBCFA290FBB9" box="[483,503,1123,1149]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF831FBCFA1C2FBB9" box="[527,677,1123,1149]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BF892FBCFA18CFBB9" bold="true" box="[684,747,1123,1149]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BF892FBCFA18CFBB9" box="[684,747,1123,1149]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but are absent in all 71 examined female imagines of
<taxonomicName id="4C06F0908D52FF8BFF71FBCFA277FB65" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFF71FBCFA607FBB8" box="[1359,1376,1123,1148]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFF46FBCFA6EBFBB9" box="[1400,1420,1123,1149]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BFAA9FB2BA277FB65" box="[151,272,1159,1185]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BFB2EFB2BA273FB65" box="[272,276,1159,1185]" italics="true" pageId="32" pageNumber="433">.</emphasis>
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FB07A175FAC9" blockId="32.[151,1437,150,1473]" pageId="32" pageNumber="433">
In winged males of
<taxonomicName id="4C06F0908D52FF8BFB97FB07A1F2FB01" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[425,661,1195,1221]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFB97FB07A2DDFB00" box="[425,442,1195,1220]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFBECFB07A281FB01" box="[466,486,1195,1221]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BFBC1FB07A1F2FB01" box="[511,661,1195,1221]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BF8A2FB07A1BAFB01" bold="true" box="[668,733,1195,1221]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BF8A2FB07A1BAFB01" box="[668,733,1195,1221]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the pair of reddish markings of abdominal tergum IV are limited to the median area of the tergum (
<figureCitation id="133D97968D52FF8BF877FB63A1CCFB2D" box="[585,683,1231,1257]" captionStart="FIGURES 127133" captionStartId="31.[151,264,1707,1731]" captionTargetBox="[151,1436,309,1684]" captionTargetId="figure-69@31.[151,1436,309,1684]" captionTargetPageId="31" captionText="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397921" httpUri="https://zenodo.org/record/4397921/files/figure.png" pageId="32" pageNumber="433">Fig. 127</figureCitation>
), while in
<taxonomicName id="4C06F0908D52FF8BF91DFB63A089FB2D" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" box="[803,1006,1231,1257]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BF91DFB63A053FB2C" box="[803,820,1231,1256]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BF972FB63A007FB2D" box="[844,864,1231,1257]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF947FB63A089FB2D" box="[889,1006,1231,1257]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
such markings are more or less wider expanded laterally (
<figureCitation id="133D97968D52FF8BFB4AFB5FA162FAC9" box="[372,517,1267,1293]" captionStart="FIGURES 99102" captionStartId="25.[151,264,2003,2027]" captionTargetBox="[151,1436,669,1980]" captionTargetId="figure-242@25.[151,1436,669,1980]" captionTargetPageId="25" captionText="FIGURES 99102. Male imagines of Procloeon (Oculogaster) regularum. 99101, general view; 102, abdominal tergum IV." figureDoi="http://doi.org/10.5281/zenodo.4397909" httpUri="https://zenodo.org/record/4397909/files/figure.png" pageId="32" pageNumber="433">Figs 99102</figureCitation>
).
</paragraph>
<paragraph id="8BB98B138D52FF8BFAF9FABBA269FA59" blockId="32.[151,1437,150,1473]" pageId="32" pageNumber="433">
The last instar larva of
<taxonomicName id="4C06F0908D52FF8BFBEDFABBA1F8FAF5" authorityName=": Sivaramakrishnan &amp; Venkataraman" authorityYear="1990" box="[467,671,1303,1329]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BFBEDFABBA283FAF4" box="[467,484,1303,1328]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BFBC3FABBA176FAF5" box="[509,529,1303,1329]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF814FABBA1F8FAF5" box="[554,671,1303,1329]" italics="true" pageId="32" pageNumber="433">regularum</emphasis>
</taxonomicName>
has paired reddish markings similar on terga IV and VII (
<figureCitation id="133D97968D52FF8BFF04FABBA6E8FAF5" box="[1338,1423,1303,1329]" captionStart="FIGURES 8489" captionStartId="23.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-16@23.[151,1436,181,1894]" captionTargetPageId="23" captionText="FIGURES 8489. Larvae of Procloeon (Oculogaster) regularum. 8488, exuviae: 84, abdomen; 8586, its caudalii; 8788, fragment of middle part of cercus, with focus on lateral and median sides; 89, male larva." figureDoi="http://doi.org/10.5281/zenodo.4397903" httpUri="https://zenodo.org/record/4397903/files/figure.png" pageId="32" pageNumber="433">Fig. 89</figureCitation>
), that differs from the coloration of winged stages; the larva of
<taxonomicName id="4C06F0908D52FF8BF96EFA97A75BFA91" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[848,1084,1339,1365]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D52FF8BF96EFA97A006FA90" box="[848,865,1339,1364]" italics="true" pageId="32" pageNumber="433">P</emphasis>
. (
<emphasis id="B97257018D52FF8BF947FA97A0EAFA91" box="[889,909,1339,1365]" italics="true" pageId="32" pageNumber="433">O</emphasis>
.)
<emphasis id="B97257018D52FF8BF998FA97A75BFA91" box="[934,1084,1339,1365]" italics="true" pageId="32" pageNumber="433">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D52FF8BFE7DFA97A7E4FA91" bold="true" box="[1091,1155,1339,1365]" pageId="32" pageNumber="433">
<taxonomicNameLabel id="A241EA7A8D52FF8BFE7DFA97A7E4FA91" box="[1091,1155,1339,1365]" pageId="32" pageNumber="433" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has no such hypodermal markings; its hypodermal coloration appears just before moult to subimago, being the same as in the winged stages (
<figureCitation id="133D97968D52FF8BFA9EFA2FA265FA59" box="[160,258,1411,1437]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="32" pageNumber="433">Fig. 109</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D52FF8BFAF9FA0AA172FA05" box="[199,533,1446,1473]" pageId="32" pageNumber="433" type="distribution">
<paragraph id="8BB98B138D52FF8BFAF9FA0AA172FA05" blockId="32.[151,1437,150,1473]" box="[199,533,1446,1473]" pageId="32" pageNumber="433">
<emphasis id="B97257018D52FF8BFAF9FA0AA206FA04" bold="true" box="[199,353,1446,1472]" pageId="32" pageNumber="433">Distribution.</emphasis>
Southern
<collectingCountry id="F311CB838D52FF8BFBEAFA0BA176FA05" box="[468,529,1447,1473]" name="India" pageId="32" pageNumber="433">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>