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<mods:title id="006FC8AF6B992669B6554D1DA7CBC27A">Confirmed polyphyly, generic recircumscription and typification of Dysoxylum (Meliaceae), with revised disposition of currently accepted species</mods:title>
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<mods:namePart id="BB53329380518D8D55B503B5AEAD56F3">Holzmeyer, Laura</mods:namePart>
<mods:affiliation id="EA775ED447EDE4387DA90A34012179E4">Frank Hauenschild, &amp; David J. Mabberley &amp; Department of Molecular Evolution and Plant Systematics &amp; Herbarium (LZ), Institute of Biology, Leipzig University, Leipzig, 04103, Germany &amp; intergenic spacer support this hypothesis. Six strongly supported clades within Dysoxylum s. l. were recovered, in large part supporting the classification of Harms. Based on the largest molecular phylogenetic tree hitherto assembled, and morphological considerations, a revised disposition of 94 currently recognized Dysoxylum species, is presented, reinstating the genera Didymocheton, Epicharis, Goniocheton, Prasoxylon and Pseudocarapa. The first valid publication of the generic name Dysoxylum is identified, and it is typified with D. macrocarpum. Forty-three new combinations at species level and two at infraspecific are proposed; Didymocheton nutans, Dysoxylum alatum, D. pachyphyllum, Epicharis, Trichilia spectabilis and T. alliacea (second step) are lectotypified.</mods:affiliation>
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<mods:namePart id="C45F7998D433C18E669AD5BEBAFD4C0E">Hauenschild, Frank</mods:namePart>
<mods:affiliation id="531117707010F40B0CB9CA5C8D5ECEE7">Frank Hauenschild, &amp; David J. Mabberley &amp; Department of Molecular Evolution and Plant Systematics &amp; Herbarium (LZ), Institute of Biology, Leipzig University, Leipzig, 04103, Germany &amp; intergenic spacer support this hypothesis. Six strongly supported clades within Dysoxylum s. l. were recovered, in large part supporting the classification of Harms. Based on the largest molecular phylogenetic tree hitherto assembled, and morphological considerations, a revised disposition of 94 currently recognized Dysoxylum species, is presented, reinstating the genera Didymocheton, Epicharis, Goniocheton, Prasoxylon and Pseudocarapa. The first valid publication of the generic name Dysoxylum is identified, and it is typified with D. macrocarpum. Forty-three new combinations at species level and two at infraspecific are proposed; Didymocheton nutans, Dysoxylum alatum, D. pachyphyllum, Epicharis, Trichilia spectabilis and T. alliacea (second step) are lectotypified.</mods:affiliation>
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<mods:affiliation id="E4EFBEEA9F7CC5962284B70ADFAD1A5B">&amp; Alexandra N. Muellner-Riehl &amp; Wadham College, University of Oxford, United Kingdom; Department of Biological Sciences, Macquarie University, and Australian Institute of Botanical Science (National Herbarium of New South Wales), Sydney, Australia</mods:affiliation>
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<mods:affiliation id="89D62E4DB9C9DD1E1FEB6CA59ADB831F">Frank Hauenschild, &amp; David J. Mabberley &amp; Department of Molecular Evolution and Plant Systematics &amp; Herbarium (LZ), Institute of Biology, Leipzig University, Leipzig, 04103, Germany &amp; intergenic spacer support this hypothesis. Six strongly supported clades within Dysoxylum s. l. were recovered, in large part supporting the classification of Harms. Based on the largest molecular phylogenetic tree hitherto assembled, and morphological considerations, a revised disposition of 94 currently recognized Dysoxylum species, is presented, reinstating the genera Didymocheton, Epicharis, Goniocheton, Prasoxylon and Pseudocarapa. The first valid publication of the generic name Dysoxylum is identified, and it is typified with D. macrocarpum. Forty-three new combinations at species level and two at infraspecific are proposed; Didymocheton nutans, Dysoxylum alatum, D. pachyphyllum, Epicharis, Trichilia spectabilis and T. alliacea (second step) are lectotypified. &amp; German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, 04103, Germany</mods:affiliation>
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29.
<taxonomicName id="4C06DDD89D23FF9A46BFBDB20867FA4D" ID-CoL="8SKY5" authority="(G. Forst.) Mabb. &amp; Holzmeyer" authorityName="Mabb. &amp; Holzmeyer" authorityYear="2021" baseAuthorityName="Mabb. &amp; Holzmeyer" baseAuthorityYear="2021" box="[192,795,1435,1458]" class="Magnoliopsida" family="Meliaceae" genus="Didymocheton" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="1261" phylum="Tracheophyta" rank="species" species="spectabilis" status="comb. nov.">
<emphasis id="B9727A499D23FF9A46BFBDB20AA8FA4D" bold="true" box="[192,468,1435,1458]" italics="true" pageId="13" pageNumber="1261">Didymocheton spectabilis</emphasis>
(G.Forst.) Mabb. &amp; Holzmeyer
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,
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<emphasis id="B9727A499D23FF9A46BBBD920A46FA2D" bold="true" box="[196,314,1467,1490]" pageId="13" pageNumber="1261">comb. nov.</emphasis>
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<emphasis id="B9727A499D23FF9A473EBD940A2CFA2E" box="[321,336,1469,1489]" italics="true" pageId="13" pageNumber="1261"></emphasis>
<taxonomicName id="4C06DDD89D23FF9A4728BD920860FA2D" authority="G. Forst., Fl. Ins. Austr." authorityName="G. Forst., Fl. Ins. Austr." box="[343,796,1467,1490]" class="Magnoliopsida" family="Meliaceae" genus="Trichilia" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="1261" phylum="Tracheophyta" rank="species" species="spectabilis">
<emphasis id="B9727A499D23FF9A4728BD92095EFA2D" box="[343,546,1467,1490]" italics="true" pageId="13" pageNumber="1261">Trichilia spectabilis</emphasis>
G.Forst., Fl. Ins. Austr.
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:
</paragraph>
<paragraph id="8BB9A65B9D23FF9A46BBBDF20863FA0D" blockId="13.[151,799,1435,1938]" box="[196,799,1499,1522]" pageId="13" pageNumber="1261">
33. 1786
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<taxonomicName id="4C06DDD89D23FF9A474CBDF20863FA0D" authority="(G. Forst.) Hook. f., Handb." authorityName="Hook. f., Handb." baseAuthorityName="G. Forst." box="[307,799,1499,1522]" class="Magnoliopsida" family="Meliaceae" genus="Dysoxylum" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="1261" phylum="Tracheophyta" rank="species" species="spectabile">
<emphasis id="B9727A499D23FF9A474CBDF20973FA0D" box="[307,527,1499,1522]" italics="true" pageId="13" pageNumber="1261">Dysoxylum spectabile</emphasis>
(G.Forst.) Hook.f., Handb.
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<paragraph id="8BB9A65B9D23FF9A46BBBDD20A22F94D" blockId="13.[151,799,1435,1938]" pageId="13" pageNumber="1261">
N. Zeal. Fl.: 41. 1864
<emphasis id="B9727A499D23FF9A47B3BDD20862F9ED" bold="true" box="[460,798,1531,1554]" pageId="13" pageNumber="1261">
<typeStatus id="54BD18F99D23FF9A47B3BDD20946F9ED" box="[460,570,1531,1554]" pageId="13" pageNumber="1261" type="lectotype">Lectotype</typeStatus>
(designated here by
</emphasis>
<emphasis id="B9727A499D23FF9A46BBBE320AFCF9CD" bold="true" box="[196,384,1563,1586]" pageId="13" pageNumber="1261">D.J. Mabberley):</emphasis>
“132”, “
<taxonomicName id="4C06DDD89D23FF9A47A4BE3209D9F9CD" box="[475,677,1563,1586]" class="Magnoliopsida" family="Meliaceae" genus="Trichilia" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="1261" phylum="Tracheophyta" rank="species" species="spectabilis">Trichilia spectabilis</taxonomicName>
partially coloured graphite sketch (NHM-UK-L-5623-189-M-1), with analysis, drawn at Tōtaranui (Queen Charlotte Sound),
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,
<collectingCountry id="F311E6CB9D23FF9A472CBE520A9EF96D" box="[339,482,1659,1682]" name="New Zealand" pageId="13" pageNumber="1261">New Zealand</collectingCountry>
,
<date id="FFB8809B9D23FF9A4794BE5209EBF96D" box="[491,663,1659,1682]" pageId="13" pageNumber="1261" value="1773-11">
<collectingDate id="EFFC79739D23FF9A4794BE5209EBF96D" box="[491,663,1659,1682]" pageId="13" pageNumber="1261" value="1773-11">November 1773</collectingDate>
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, by Georg[e] Forster (BM!).
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<emphasis id="B9727A499D23FF9A46BBBE920A1CF92E" box="[196,352,1723,1746]" italics="true" pageId="13" pageNumber="1261">Distribution. </emphasis>
<collectingCountry id="F311E6CB9D23FF9A4717BE920A89F92D" box="[360,501,1723,1746]" name="New Zealand" pageId="13" pageNumber="1261">New Zealand</collectingCountry>
(see
<bibRefCitation id="EF97DBAA9D23FF9A4452BE920873F92D" author="Braggins, J. E. &amp; Large, M. F. &amp; Mabberley, D. J." box="[557,783,1723,1746]" pageId="13" pageNumber="1261" pagination="315 - 324" refId="ref21717" refString="Braggins, J. E., Large, M. F. &amp; Mabberley, D. J. 1999. Sexual arrangements in kohe-kohe (Dysoxylum spectabile, Meliaceae). Telopea 8: 315 - 324. https: // doi. org / 10.7751 / telopea 19995420" type="journal article" year="1999">Braggins &amp; al., 1999</bibRefCitation>
).
</paragraph>
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<emphasis id="B9727A499D23FF9A46BBBEF20A67F90E" box="[196,283,1755,1778]" italics="true" pageId="13" pageNumber="1261">Notes. </emphasis>
Although
<bibRefCitation id="EF97DBAA9D23FF9A47F2BEF20936F90D" author="Allan, H. H." box="[397,586,1755,1778]" pageId="13" pageNumber="1261" refId="ref21534" refString="Allan, H. H. 1961. Flora of New Zealand, vol. 1. Wellington: Government Printer." type="book" year="1961">Allan (1961: 427)</bibRefCitation>
wrote, “Type locality: Queen Charlotte Sound. Type: P, Forst. f.”, no such herbarium (or other) material has ever been found at P (see
<bibRefCitation id="EF97DBAA9D23FF9A46E8BF120A9DF8AD" author="Nicolson, D. H. &amp; Fosberg, F. R." box="[151,481,1851,1874]" pageId="13" pageNumber="1261" refId="ref24911" refString="Nicolson, D. H. &amp; Fosberg, F. R. 2004. The Forsters and the botany of the second Cook expedition (1772 - 1775). Regnum Vegetabile 139. Ruggell: Gantner." type="journal volume" year="2004">Nicolson &amp; Fosberg, 2004: 500</bibRefCitation>
) and, as no specimen has been found anywhere, the only surviving original material is artwork, of which the excellent field-sketch made by Georg[e] Forster is here selected as
<typeStatus id="54BD18F99D23FF9A4213B8920FA9FF2D" box="[1132,1237,187,210]" pageId="13" pageNumber="1261" type="lectotype">lectotype</typeStatus>
. See also:
<bibRefCitation id="EF97DBAA9D23FF9A4333B8920F54FF0D" author="Nicolson, D. H. &amp; Fosberg, F. R." pageId="13" pageNumber="1261" refId="ref24911" refString="Nicolson, D. H. &amp; Fosberg, F. R. 2004. The Forsters and the botany of the second Cook expedition (1772 - 1775). Regnum Vegetabile 139. Ruggell: Gantner." type="journal volume" year="2004">Nicolson &amp; Fosberg (2004: 500)</bibRefCitation>
;
<bibRefCitation id="EF97DBAA9D23FF9A424CB8F20E45FF0D" author="Mabberley, D. J." box="[1075,1337,219,242]" pageId="13" pageNumber="1261" refId="ref24099" refString="Mabberley, D. J. 2017 b. [Commentary on] Dysoxylon spectabile. P. 116 (&amp; pl. 50) in: Gooding, M., Mabberley, D. &amp; Studholme, J. (eds.), Banks' Florilegium: Botanical treasures from Cooks First Voyage. Londong: Thames &amp; Hudson." type="book" year="2017">Mabberley (2017b: 116</bibRefCitation>
, t. 50).
</paragraph>
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