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<document id="27D9972E5B7790CC1E761365FEC1DFBB" ID-DOI="10.1649/0010-065X-72.4.873" ID-ISSN="1938-4394" ID-Zenodo-Dep="5387219" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630304217244" checkinUser="diego" docAuthor="Sánchez-Hernández, Gibrán &amp; GóMez, Benigno" docDate="2018" docId="03AF171CFF9DF9788AF2FF33E29FFC18" docLanguage="en" docName="0010-065X-72.4.873.pdf" docOrigin="The Coleopterists Bulletin 72 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-72.4.873" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Onthophagus anewtoni Howden and Genier 2004" docType="treatment" docVersion="3" lastPageNumber="875" masterDocId="FF966F64FF9EF97B8A15FFF0E31CFFF2" masterDocTitle="First Precise Locality Data for Onthophagus atriglabrus Howden and Gill and New State Record for Onthophagus anewtoni Howden and Génier (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico" masterLastPageNumber="876" masterPageNumber="873" pageNumber="875" updateTime="1699221485953" updateUser="plazi">
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<mods:title id="668EC0822320A3329630C204ED4EFA2F">First Precise Locality Data for Onthophagus atriglabrus Howden and Gill and New State Record for Onthophagus anewtoni Howden and Génier (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico</mods:title>
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<mods:namePart id="AFFAA0F84CCCCE80FFDA4BCA4AD1689C">Sánchez-Hernández, Gibrán</mods:namePart>
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<mods:title id="CE310672ECDB9031AD6E2E9302E63A5D">The Coleopterists Bulletin</mods:title>
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<mods:date id="2FACEA46F5F8E48B5F5284AC1AE8D07D">2018</mods:date>
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<paragraph id="8BB9A60AFF9DF9788AF2FF33E2FCFF01" blockId="3.[231,600,195,269]" pageId="3" pageNumber="875">
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<taxonomicName id="4C06DD89FF9DF9788AF2FF33E2FCFF01" ID-CoL="7XMBR" authority="Howden and Genier, 2004" authorityName="Howden and Genier" authorityYear="2004" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="875" phylum="Arthropoda" rank="species" species="anewtoni">
<emphasis id="B9727A18FF9DF9788AF2FF33E2FCFF01" bold="true" pageId="3" pageNumber="875">
<emphasis id="B9727A18FF9DF9788AF2FF33E2D2FF2A" bold="true" box="[231,462,195,216]" italics="true" pageId="3" pageNumber="875">Onthophagus anewtoni</emphasis>
Howden and Génier, 2004
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31CF581FF9DF9788B71FF08E3F0FE50" pageId="3" pageNumber="875" type="description">
<paragraph id="8BB9A60AFF9DF9788B71FF08E2C7FEFF" blockId="3.[231,600,195,269]" box="[356,475,248,269]" pageId="3" pageNumber="875">
(
<figureCitation id="133DBA8FFF9DF9788B7EFF08E2A0FEFF" box="[363,444,248,269]" captionStart="Fig" captionStartId="1.[197,228,1711,1730]" captionTargetBox="[177,1187,993,1692]" captionTargetId="figure-274@1.[175,1190,991,1694]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of a) Onthophagus atriglabrus and b) Onthophagus anewtoni." figureDoi="http://doi.org/10.5281/zenodo.5387223" httpUri="https://zenodo.org/record/5387223/files/figure.png" pageId="3" pageNumber="875">Figs. 1b</figureCitation>
,
<figureCitation id="133DBA8FFF9DF9788BDDFF08E2C9FEFF" box="[456,469,248,269]" captionStart="Fig" captionStartId="2.[175,206,1663,1682]" captionTargetBox="[253,1071,1025,1645]" captionTargetId="figure-463@2.[251,1073,1023,1647]" captionTargetPageId="2" captionText="Fig. 2. Currently known distributions of Onthophagus atriglabrus and Onthophagus anewtoni. Previously known locality records for O. atriglabrus are from Howden and Gill (1987) and Kohlmann and Sol´ıs (2001); those for O. anewtoni are from Howden and Génier (2004)." figureDoi="http://doi.org/10.5281/zenodo.5387225" httpUri="https://zenodo.org/record/5387225/files/figure.png" pageId="3" pageNumber="875">2</figureCitation>
)
</paragraph>
<paragraph id="8BB9A60AFF9DF9788AD0FED3E3F0FE50" blockId="3.[173,660,291,1002]" pageId="3" pageNumber="875">
<emphasis id="B9727A18FF9DF9788AD0FED3E282FECA" bold="true" box="[197,414,291,312]" pageId="3" pageNumber="875">Known Distribution.</emphasis>
<materialsCitation id="3B6EAC57FF9DF9788BB8FED3E3F4FE50" country="Mexico" location="Guerrero" pageId="3" pageNumber="875" specimenCount="1" stateProvince="Guerrero">
Before our record reported herein, this species was only known from its
<typeStatus id="54BD18A8FF9DF9788AB8FEA8E3CBFE9F" box="[173,215,344,365]" pageId="3" pageNumber="875">type</typeStatus>
series localities in the states of
<collectingRegion id="49C268E8FF9DF9788823FEA8E18CFE9F" box="[566,656,344,365]" country="Mexico" name="Guerrero" pageId="3" pageNumber="875">Guerrero</collectingRegion>
,
<collectingRegion id="49C268E8FF9DF9788AB8FE82E3EDFE75" box="[173,241,370,391]" country="Mexico" name="Jalisco" pageId="3" pageNumber="875">Jalisco</collectingRegion>
, and
<collectingRegion id="49C268E8FF9DF9788B31FE82E273FE75" box="[292,367,370,391]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="875">Oaxaca</collectingRegion>
,
<collectingCountry id="F311E69AFF9DF9788B6CFE82E2DFFE75" box="[377,451,370,391]" name="Mexico" pageId="3" pageNumber="875">Mexico</collectingCountry>
(Howden and Génier 2004)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CF581FF9DF9788AD0FE57E15EFD63" pageId="3" pageNumber="875" type="materials_examined">
<paragraph id="8BB9A60AFF9DF9788AD0FE57E15EFD63" blockId="3.[173,660,291,1002]" pageId="3" pageNumber="875">
<emphasis id="B9727A18FF9DF9788AD0FE57E28FFE4E" bold="true" box="[197,403,423,444]" pageId="3" pageNumber="875">Examined Material.</emphasis>
<materialsCitation id="3B6EAC57FF9DF9788B8BFE57E122FD63" collectingDate="2017-06-30" collectingDateMax="2017-07-28" collectingDateMin="2017-06-30" collectionCode="R" collectorName="G. Sanchez-Hernandez &amp; Davila &amp; A. Gomez &amp; Hernandez &amp; C. Valdez" country="Mexico" elevation="875" latitude="16.728937" location="Chiapa de Corzo" longLatPrecision="1" longitude="-92.93247" municipality="Forty-two" pageId="3" pageNumber="875" specimenCount="59" specimenCount-female="35" specimenCount-male="24" stateProvince="Chiapas">
<collectingMunicipality id="6BDD3C70FF9DF9788B8BFE57E2E1FE4E" box="[414,509,423,444]" pageId="3" pageNumber="875">Forty-two</collectingMunicipality>
specimens (
<specimenCount id="9D006D83FF9DF978886FFE57E3F8FE25" pageId="3" pageNumber="875" type="male">24 males</specimenCount>
,
<specimenCount id="9D006D83FF9DF9788AF9FE32E24DFE25" box="[236,337,450,471]" pageId="3" pageNumber="875" type="female">18 females</specimenCount>
).
<emphasis id="B9727A18FF9DF9788B4AFE32E2D4FE25" bold="true" box="[351,456,450,471]" pageId="3" pageNumber="875">
<collectingCountry id="F311E69AFF9DF9788B4AFE32E2D8FE25" box="[351,452,450,471]" name="Mexico" pageId="3" pageNumber="875">MEXICO</collectingCountry>
:
</emphasis>
<collectingRegion id="49C268E8FF9DF9788BD8FE32E12DFE25" box="[461,561,450,471]" country="Mexico" name="Chiapas" pageId="3" pageNumber="875">CHIAPAS</collectingRegion>
:
<location id="8ED9F0D1FF9DF978882DFE32E3F9FE00" LSID="urn:lsid:plazi:treatment:03AF171CFF9DF9788AF2FF33E29FFC18:8ED9F0D1FF9DF978882DFE32E3F9FE00" country="Mexico" latitude="16.728937" longLatPrecision="1" longitude="-92.93247" municipality="Forty-two" name="Chiapa de Corzo" pageId="3" pageNumber="875" stateProvince="Chiapas">Chiapa de Corzo</location>
municipality,
<locationDeviation id="9A0BBF28FF9DF9788B7CFE2DE193FE00" box="[361,655,477,498]" location="Chiapa de Corzo" pageId="3" pageNumber="875">
<quantity id="4CFE0BEFFF9DF9788B7CFE2DE2B6FE00" box="[361,426,477,498]" metricMagnitude="3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="875" unit="km" value="8.2">8.2 km</quantity>
NE of Chiapa de Corzo
</locationDeviation>
, deciduous forest,
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16°43
<emphasis id="B9727A18FF9DF9788B9EFE07E28CFDFF" bold="true" box="[395,400,503,525]" pageId="3" pageNumber="875">ʹ</emphasis>
44.17
<emphasis id="B9727A18FF9DF9788BD1FE07E2D2FDFF" bold="true" box="[452,462,503,525]" pageId="3" pageNumber="875">ʹʹ</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE32C0CDFF9DF9788BE3FE07E18CFDFE" box="[502,656,503,525]" degrees="92" direction="west" minutes="55" orientation="longitude" pageId="3" pageNumber="875" precision="1" seconds="56.91" value="-92.93247">
92°55
<emphasis id="B9727A18FF9DF978883BFE07E12FFDFF" bold="true" box="[558,563,503,525]" pageId="3" pageNumber="875">ʹ</emphasis>
56.91
<emphasis id="B9727A18FF9DF9788873FE07E16CFDFF" bold="true" box="[614,624,503,525]" pageId="3" pageNumber="875">ʹʹ</emphasis>
W
</geoCoordinate>
,
<quantity id="4CFE0BEFFF9DF9788AB8FDE2E3F4FDD5" box="[173,232,530,551]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="3" pageNumber="875" unit="m" value="875.0">
<elevation id="002B4139FF9DF9788AB8FDE2E3F4FDD5" box="[173,232,530,551]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="3" pageNumber="875" unit="m" value="875.0">875 m</elevation>
</quantity>
,
<date id="FFB880CAFF9DF9788AE4FDE2E247FDD5" box="[241,347,530,551]" pageId="3" pageNumber="875" value="2017-06-30">
<collectingDate id="EFFC7922FF9DF9788AE4FDE2E247FDD5" box="[241,347,530,551]" pageId="3" pageNumber="875" value="2017-06-30">30.VI.2017</collectingDate>
</date>
, squid (216,
<specimenCount id="9D006D83FF9DF9788BCDFDE2E2E0FDD5" box="[472,508,529,551]" pageId="3" pageNumber="875" type="female">16♀</specimenCount>
), pig dung (16),
<collectorName id="26F3C3DCFF9DF9788AB8FDDCE291FDB3" box="[173,397,556,577]" pageId="3" pageNumber="875">G. Sánchez-Hernández</collectorName>
,
<collectionCode id="ED173ECFFF9DF9788B8FFDDCE2B1FDB3" box="[410,429,556,577]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="875">R</collectionCode>
.
<collectorName id="26F3C3DCFF9DF9788BACFDDCE2E4FDB3" box="[441,504,556,577]" pageId="3" pageNumber="875">Dávila</collectorName>
and
<collectorName id="26F3C3DCFF9DF978883BFDDCE18FFDB3" box="[558,659,556,577]" pageId="3" pageNumber="875">A. Gómez</collectorName>
cols.; idem except
<date id="FFB880CAFF9DF9788B64FDB7E2F4FDAE" box="[369,488,583,604]" pageId="3" pageNumber="875" value="2017-07-28">
<collectingDate id="EFFC7922FF9DF9788B64FDB7E2F4FDAE" box="[369,488,583,604]" pageId="3" pageNumber="875" value="2017-07-28">28.VII.2017</collectingDate>
</date>
, squid (26), pig dung (16), fermented fungus (
<specimenCount id="9D006D83FF9DF9788BF2FD91E2E3FD85" box="[487,511,609,631]" pageId="3" pageNumber="875" type="female">1♀</specimenCount>
), G. Sánchez-
<collectorName id="26F3C3DCFF9DF9788AB8FD8CE20AFD63" box="[173,278,636,657]" pageId="3" pageNumber="875">Hernández</collectorName>
,
<collectionCode id="ED173ECFFF9DF9788B34FD8CE228FD63" box="[289,308,636,657]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="875">R</collectionCode>
.
<collectorName id="26F3C3DCFF9DF9788B2AFD8CE262FD63" box="[319,382,636,657]" pageId="3" pageNumber="875">Dávila</collectorName>
and
<collectorName id="26F3C3DCFF9DF9788BBAFD8CE112FD63" box="[431,526,636,657]" pageId="3" pageNumber="875">C. Valdez</collectorName>
cols
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CF581FF9DF9788AD0FD67E29FFC18" pageId="3" pageNumber="875" type="discussion">
<paragraph id="8BB9A60AFF9DF9788AD0FD67E29FFC18" blockId="3.[173,660,291,1002]" pageId="3" pageNumber="875">
<emphasis id="B9727A18FF9DF9788AD0FD67E22BFD5E" bold="true" box="[197,311,663,684]" pageId="3" pageNumber="875">Comments.</emphasis>
The presence of this species in the central region of
<collectingRegion id="49C268E8FF9DF9788B78FD41E2A7FD34" box="[365,443,689,710]" country="Mexico" name="Chiapas" pageId="3" pageNumber="875">Chiapas</collectingRegion>
matches with other species of the subfamily, such as
<taxonomicName id="4C06DD89FF9DF9788BF8FD3BE251FD09" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="875" phylum="Arthropoda" rank="species" species="demon">
<emphasis id="B9727A18FF9DF9788BF8FD3BE18EFD13" box="[493,658,715,737]" italics="true" pageId="3" pageNumber="875">Phanaeus demon</emphasis>
Castelnau, 1840
</taxonomicName>
and
<taxonomicName id="4C06DD89FF9DF9788B97FD15E3FCFCE4" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="875" phylum="Arthropoda" rank="species" species="laetum">
<emphasis id="B9727A18FF9DF9788B97FD15E15DFD09" box="[386,577,741,763]" italics="true" pageId="3" pageNumber="875">Canthidium laetum</emphasis>
Harold, 1867
</taxonomicName>
, that are distributed along the Mexican Pacific coast and use the slopes of the Sierra Madre de
<collectingRegion id="49C268E8FF9DF9788AB8FCC6E3E7FCB9" box="[173,251,822,843]" country="Mexico" name="Chiapas" pageId="3" pageNumber="875">Chiapas</collectingRegion>
(Reserva de la Biósfera La Sepultura) to expand their distribution towards the Central Depression of
<collectingRegion id="49C268E8FF9DF9788B45FC9BE282FC72" box="[336,414,875,896]" country="Mexico" name="Chiapas" pageId="3" pageNumber="875">Chiapas</collectingRegion>
(Kohlmann and Sol´ıs 2006a;
<bibRefCitation id="EF97DBFBFF9DF9788AEBFC76E2C8FC69" author="Lizardo, V., F." box="[254,468,901,923]" pageId="3" pageNumber="875" pagination="271 - 294" refId="ref2152" refString="Lizardo, V., F. Escobar, and O. Rojas-Soto. 2017. Diversity and distribution of Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico. Zootaxa 4358 (2): 271 - 294." type="journal article" year="2017">
Lizardo
<emphasis id="B9727A18FF9DF9788B42FC76E28DFC69" box="[343,401,901,923]" italics="true" pageId="3" pageNumber="875">et al.</emphasis>
2017
</bibRefCitation>
). Our locality record extends the distribution range approximately
<quantity id="4CFE0BEFFF9DF9788AB8FC4BE21DFC22" box="[173,257,955,976]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="875" unit="km" value="300.0">300 km</quantity>
east of the closest known locality, Tequesistlán,
<collectingRegion id="49C268E8FF9DF9788B26FC25E263FC18" box="[307,383,981,1002]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="875">Oaxaca</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
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