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<document id="3809E26308B7BD36E299292BD9BC089E" ID-CLB-Dataset="4209" ID-DOI="10.5852/ejt.2021.767.1485" ID-GBIF-Dataset="66e2d72d-ad54-494d-80e6-56b6020ef450" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514853" ID-ZooBank="9B651F61-7F2B-4D46-9390-4F54F8AEB0AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927370404" checkinUser="felipe" docAuthor="Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2021" docId="03AEE5789D443E22770CD4EE7786F938" docLanguage="en" docName="ejt-767_40-54.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1485" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Orphnus bicolor" docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="FF979D009D4C3E2A7541D5337576FF8D" masterDocTitle="Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819" masterLastPageNumber="54" masterPageNumber="40" pageNumber="48" updateTime="1699449499239" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6FF87D9B6572EB8286E9D988AB670C69">Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819</mods:title>
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<mods:name id="063A10CB0BA4D7523497A8F4335E730F" type="personal">
<mods:role id="3E450A02159F1C9887FA39541922D30D">
<mods:roleTerm id="5A544D16CB0BF4E72205824C9B29404E">Author</mods:roleTerm>
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<mods:namePart id="4DE1C8A07C67E72FC31F61D002ED5949">Frolov, Andrey V.</mods:namePart>
<mods:nameIdentifier id="6FB11F7809524F36D4C5EEEF934CB7D1" type="LSID">5DEE612B-4252-4DB0-AD5E-BAA789A3C355</mods:nameIdentifier>
<mods:affiliation id="7E95E8F59EFAD9396CFB7F14B2451EB7">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="FFAAB4A2083940E43C7C79F86ED83C0C" type="email">frolov@scarabaeoidea.com</mods:nameIdentifier>
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<mods:name id="53163727C9479E0352B7A5BB27E24709" type="personal">
<mods:role id="BA85279EE900AC94250D89DF0542D7F2">
<mods:roleTerm id="F2EE5D96BA17C932D0E7E51F28A41768">Author</mods:roleTerm>
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<mods:namePart id="F156B1692EC7CDF6228BF75BF7AFCA12">Akhmetova, Lilia A.</mods:namePart>
<mods:nameIdentifier id="C89AF9C541DF973684CD8E1AA585CC88" type="LSID">BA35DDDD-6E6A-47A4-AB73-965EB4A92CC3</mods:nameIdentifier>
<mods:affiliation id="BC87426CC821BADBA00B57502DCA4AE5">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="0CB8019F8D66E8281900FB6947975E74" type="email">akhmetova@scarabaeoidea.com</mods:nameIdentifier>
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<mods:title id="50D39FB16A6AA812A317A47803D1A014">European Journal of Taxonomy</mods:title>
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<mods:part id="2966C1CD80BA30AB4D7838B5E8D37118">
<mods:date id="B9BA537F5B4C31EB3648AC5C60662E62">2021</mods:date>
<mods:detail id="690FCD633ACE7568C3B0F391670A5BC6" type="pubDate">
<mods:number id="8557FAEA264FD367CD727C9E8B0F2E2E">2021-09-06</mods:number>
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<mods:detail id="5C508265585D31427A7C49A0BE85F71F" type="volume">
<mods:number id="2D3ADDFDB6DBAE251873A3F15B4A767D">767</mods:number>
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<mods:number id="E85C6B812375CC4697AE663114CBC820">1</mods:number>
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<mods:classification id="34787D934937977C7324363487C1E6BE">journal article</mods:classification>
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<mods:identifier id="11B355B13D9AAB642537A171697D2A13" type="DOI">10.5852/ejt.2021.767.1485</mods:identifier>
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<treatment id="03AEE5789D443E22770CD4EE7786F938" ID-DOI="http://doi.org/10.5281/zenodo.5514749" ID-GBIF-Taxon="188498250" ID-Zenodo-Dep="5514749" LSID="urn:lsid:plazi:treatment:03AEE5789D443E22770CD4EE7786F938" httpUri="http://treatment.plazi.org/id/03AEE5789D443E22770CD4EE7786F938" lastPageNumber="48" pageId="8" pageNumber="48">
<subSubSection id="C31D07E59D443E22770CD4EE7691FE75" box="[589,999,477,504]" pageId="8" pageNumber="48" type="nomenclature">
<paragraph id="8BB8546E9D443E22770CD4EE7691FE75" blockId="8.[589,999,477,539]" box="[589,999,477,504]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E22770CD4EE7691FE75" box="[589,999,477,504]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="48" reason="2">
<taxonomicName id="4C072FED9D443E22770CD4EE7691FE75" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[589,999,477,504]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E22770CD4EE7663FE75" bold="true" box="[589,789,477,504]" italics="true" pageId="8" pageNumber="48">Orphnus bicolor</emphasis>
(
<bibRefCitation id="EF96299F9D443E227665D4EE76A8FE75" author="Fabricius J. C." box="[804,990,477,504]" pageId="8" pageNumber="48" refId="ref6461" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Impensis Bibliopolii Academici novi. Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31D07E59D443E2277B6D732764AFD96" box="[759,828,513,539]" pageId="8" pageNumber="48" type="description">
<paragraph id="8BB8546E9D443E2277B6D732764AFD96" blockId="8.[589,999,477,539]" box="[759,828,513,539]" pageId="8" pageNumber="48">
<figureCitation id="133C48EB9D443E2277B6D732764AFD96" box="[759,828,513,539]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[189,1396,343,1826]" captionTargetId="figure-16@9.[189,1398,306,1844]" captionTargetPageId="9" captionText="Fig. 4. Orphnus bicolor (Fabricius, 1801). A–F. Lectotype, ♂ (ZMUKG). G–H. ♀ (MNHN). A–B, G–H. Habitus in dorsal view. C. Labels. D. Endophallus. E. Aedeagus in lateral view. F. Parameres in dorsal view. I. Distributional record map. Scale bars: A–B, G–H = 1.0 mm; D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514861" httpUri="https://zenodo.org/record/5514861/files/figure.png" pageId="8" pageNumber="48">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31D07E59D443E2275FCD7717652FD68" pageId="8" pageNumber="48" type="reference_group">
<paragraph id="8BB8546E9D443E2275FCD77177F0FDD1" blockId="8.[189,646,578,604]" box="[189,646,578,604]" pageId="8" pageNumber="48">
<treatmentCitationGroup id="AB1773409D443E2275FCD77177F0FDD1" box="[189,646,578,604]" pageId="8" pageNumber="48">
<treatmentCitation id="0AA6727F9D443E2275FCD77177F6FDD1" author="Fabricius J. C." box="[189,640,578,604]" page="9" pageId="8" pageNumber="48" year="1801">
<taxonomicName id="4C072FED9D443E2275FCD77177F6FDD1" ID-CoL="6XVSS" authority="Fabricius, 1801: 9" authorityName="Fabricius" authorityPageNumber="9" authorityYear="1801" box="[189,640,578,604]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E2275FCD77174D4FDD1" box="[189,418,578,604]" italics="true" pageId="8" pageNumber="48">Scarabaeus bicolor</emphasis>
<bibRefCitation id="EF96299F9D443E2274E8D77177F6FDD1" author="Fabricius J. C." box="[425,640,578,604]" pageId="8" pageNumber="48" refId="ref6461" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Impensis Bibliopolii Academici novi. Kiel." type="book" year="1801">Fabricius, 1801: 9</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD7B77652FD68" blockId="8.[189,1399,643,741]" pageId="8" pageNumber="48">
<treatmentCitationGroup id="AB1773409D443E2275FCD7B77652FD68" pageId="8" pageNumber="48">
<taxonomicName id="4C072FED9D443E2275FCD7B774F6FD13" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[189,384,644,670]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E2275FCD7B774F6FD13" box="[189,384,644,670]" italics="true" pageId="8" pageNumber="48">Orphnus bicolor</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AA6727F9D443E2274DCD7B777FDFD13" author="MacLeay W. S." box="[413,651,644,670]" page="119" pageId="8" pageNumber="48" year="1819">
<bibRefCitation id="EF96299F9D443E2274DCD7B777FDFD13" author="MacLeay W. S." box="[413,651,644,670]" pageId="8" pageNumber="48" refId="ref6823" refString="MacLeay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Vol. 1 Part 1. S. Bagster, London. https: // doi. org / 10.5962 / bhl. title. 48636" type="book" year="1819">MacLeay 1819: 119</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AA6727F9D443E2277FAD7B776C1FD13" author="Westwood J. O." box="[699,951,643,670]" page="176" pageId="8" pageNumber="48" year="1845">
<bibRefCitation id="EF96299F9D443E2277FAD7B776C1FD13" author="Westwood J. O." box="[699,951,643,670]" pageId="8" pageNumber="48" pagination="155 - 180" refId="ref7114" refString="Westwood J. O. 1845. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10 - jointed antennae. Transactions of the Entomological Society of London 4: 155 - 180. https: // doi. org / 10.1111 / j. 1365 - 2311.1846. tb 01350. x" type="journal article" year="1845">Westwood 1845: 176</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AA6727F9D443E2276A6D7B77192FD13" author="Lacordaire T." box="[999,1252,643,670]" page="130" pageId="8" pageNumber="48" year="1856">
<bibRefCitation id="EF96299F9D443E2276A6D7B77192FD13" author="Lacordaire T." box="[999,1252,643,670]" pageId="8" pageNumber="48" refId="ref6766" refString="Lacordaire T. 1856. Histoire naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troiseme contenant les familles des pectinicorne et lamellicornes. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8864" type="book" year="1856">Lacordaire 1856: 130</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0AA6727F9D443E22744FD79477EDFD4C" author="Gemminger M. &amp; Harold E." box="[270,667,679,705]" page="1072" pageId="8" pageNumber="48" year="1869">
<bibRefCitation id="EF96299F9D443E22744FD79477EDFD4C" author="Gemminger M. &amp; Harold E." box="[270,667,679,705]" pageId="8" pageNumber="48" refId="ref6655" refString="Gemminger M. &amp; Harold E. 1869. Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Vol. 4 Scarabaeidae: 979 - 1346. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Gemminger &amp; Harold 1869: 1072</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0AA6727F9D443E227614D7947179FD4C" author="Arrow G. J." box="[853,1039,679,705]" page="29" pageId="8" pageNumber="48" year="1912">
<bibRefCitation id="EF96299F9D443E227614D7947179FD4C" author="Arrow G. J." box="[853,1039,679,705]" pageId="8" pageNumber="48" pagination="1 - 66" refId="ref6406" refString="Arrow G. J. 1912. Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus 43: 1 - 66. W. Junk, Berlin." type="journal article" year="1912">Arrow 1912: 29</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0AA6727F9D443E22718AD7947471FD69" author="Schmidt A." page="79" pageId="8" pageNumber="48" year="1913">
<bibRefCitation id="EF96299F9D443E22718AD7947471FD69" author="Schmidt A." pageId="8" pageNumber="48" refId="ref6967" refString="Schmidt A. 1913. Coleoptera. Lamellicornia. Fam. Scarabaeidae. Subfam. Aegialinae, Chironinae, Dynamopinae, Hybosorinae, Idiostominae, Ochodaeinae, Orphninae. Genera Insectorum 150. V. Verteneuil &amp; L. Desmet, Bruxelles." type="book" year="1913">Schmidt 1913: 79</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0AA6727F9D443E227487D7F977E7FD69" author="Frolov A. V." box="[454,657,714,740]" page="793" pageId="8" pageNumber="48" year="2012">
<bibRefCitation id="EF96299F9D443E227487D7F977E7FD69" author="Frolov A. V." box="[454,657,714,740]" pageId="8" pageNumber="48" pagination="782 - 797" refId="ref6496" refString="Frolov A. V. 2012. Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Entomological Review 92: 782 - 797. https: // doi. org / 10.1134 / S 0013873812070056" type="journal article" year="2012">Frolov 2012: 793</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D07E59D443E2275FCD63877A4FB88" pageId="8" pageNumber="48" type="diagnosis">
<paragraph id="8BB8546E9D443E2275FCD63874BFFCA8" blockId="8.[189,457,779,805]" box="[189,457,779,805]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E2275FCD63874BFFCA8" bold="true" box="[189,457,779,805]" fontSize="11" level="3" pageId="8" pageNumber="48" reason="3">
<emphasis id="B973887C9D443E2275FCD63874BFFCA8" bold="true" box="[189,457,779,805]" pageId="8" pageNumber="48">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD60977A4FB88" blockId="8.[189,1400,826,1029]" pageId="8" pageNumber="48">
<taxonomicName id="4C072FED9D443E2275FCD60974F7FCD9" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[189,385,826,852]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E2275FCD60974F7FCD9" box="[189,385,826,852]" italics="true" pageId="8" pageNumber="48">Orphnus bicolor</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C072FED9D443E22775DD60977E6FCD9" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[540,656,826,852]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B973887C9D443E22775DD60977E6FCD9" box="[540,656,826,852]" italics="true" pageId="8" pageNumber="48">O. parvus</emphasis>
</taxonomicName>
in having the pronotum without tubercle on base medially and endophallus with a few groups of spinules. It differs from the latter in having the parameres in dorsal view with small but distinct notches basad of the lateral teeth (
<figureCitation id="133C48EB9D443E2276F4D6B37166FC16" box="[949,1040,896,923]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[189,1396,343,1826]" captionTargetId="figure-16@9.[189,1398,306,1844]" captionTargetPageId="9" captionText="Fig. 4. Orphnus bicolor (Fabricius, 1801). A–F. Lectotype, ♂ (ZMUKG). G–H. ♀ (MNHN). A–B, G–H. Habitus in dorsal view. C. Labels. D. Endophallus. E. Aedeagus in lateral view. F. Parameres in dorsal view. I. Distributional record map. Scale bars: A–B, G–H = 1.0 mm; D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514861" httpUri="https://zenodo.org/record/5514861/files/figure.png" pageId="8" pageNumber="48">Fig. 4F</figureCitation>
), longer parameres (about 0.7 length of phallobase [
<figureCitation id="133C48EB9D443E227489D697775EFC33" box="[456,552,931,958]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[189,1396,343,1826]" captionTargetId="figure-16@9.[189,1398,306,1844]" captionTargetPageId="9" captionText="Fig. 4. Orphnus bicolor (Fabricius, 1801). A–F. Lectotype, ♂ (ZMUKG). G–H. ♀ (MNHN). A–B, G–H. Habitus in dorsal view. C. Labels. D. Endophallus. E. Aedeagus in lateral view. F. Parameres in dorsal view. I. Distributional record map. Scale bars: A–B, G–H = 1.0 mm; D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514861" httpUri="https://zenodo.org/record/5514861/files/figure.png" pageId="8" pageNumber="48">Fig. 4E</figureCitation>
] as opposed to 0.5 length of phallobase in
<taxonomicName id="4C072FED9D443E22717FD69771C0FC30" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[1086,1206,932,958]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B973887C9D443E22717FD69771C0FC30" box="[1086,1206,932,958]" italics="true" pageId="8" pageNumber="48">O. parvus</emphasis>
</taxonomicName>
[
<figureCitation id="133C48EB9D443E227188D697705CFC33" box="[1225,1322,931,958]" captionStart="Fig" captionStartId="10.[189,232,1886,1912]" captionTargetBox="[213,1383,283,1814]" captionTargetId="figure-11@10.[203,1385,265,1847]" captionTargetPageId="10" captionText="Fig. 5. Orphnus parvus (Wiedemann, 1823). A–C. Lectotype, ♂ (ZMUKK). D–H. Lectotype of Orphnus nanus Westwood, 1845, ♂ (ZMUKK). I. ♀ (MNHN). A–B, H–I. Habitus in dorsal view. C, E. Labels. D. Endophallus. F. Aedeagus in lateral view. G. Parameres in dorsal view. J. Distributional record map. Scale bars: A–B, H–I = 1.0 mm; D, F–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514863" httpUri="https://zenodo.org/record/5514863/files/figure.png" pageId="8" pageNumber="48">Fig. 5E</figureCitation>
]), and abdominal sternite 8 medially longer than sternites 4–7 combined (as opposed to sternite 8 medially as long as sternites 4–7 combined in
<taxonomicName id="4C072FED9D443E22770FD6D977B4FB89" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[590,706,1002,1028]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B973887C9D443E22770FD6D977B4FB89" box="[590,706,1002,1028]" italics="true" pageId="8" pageNumber="48">O. parvus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D07E59D443E2275FCD11F702EFA3E" pageId="8" pageNumber="48" type="materials_examined">
<paragraph id="8BB8546E9D443E2275FCD11F741AFBC8" blockId="8.[189,364,1067,1094]" box="[189,364,1067,1094]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E2275FCD11F741AFBC8" bold="true" box="[189,364,1067,1094]" fontSize="11" level="3" pageId="8" pageNumber="48" reason="3">
<emphasis id="B973887C9D443E2275FCD11F741AFBC8" bold="true" box="[189,364,1067,1094]" pageId="8" pageNumber="48">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8546E9D443E2275ABD16977D1FB36" blockId="8.[189,1400,1114,1211]" pageId="8" pageNumber="48">
<materialsCitation id="3B6F5E339D443E2275ABD16977D4FB36" ID-GBIF-Occurrence="3357770321" collectionCode="ZMUKG" collectorName="Daldorff Mus. T. Lund" country="India" location="INDIA" pageId="8" pageNumber="48" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B973887C9D443E2275ABD1697410FBF9" bold="true" box="[234,358,1114,1140]" pageId="8" pageNumber="48">
<typeStatus id="54BCEACC9D443E2275ABD1697410FBF9" box="[234,358,1114,1140]" pageId="8" pageNumber="48" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated,
<figureCitation id="133C48EB9D443E22777CD16977CEFBF9" box="[573,696,1114,1140]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[189,1396,343,1826]" captionTargetId="figure-16@9.[189,1398,306,1844]" captionTargetPageId="9" captionText="Fig. 4. Orphnus bicolor (Fabricius, 1801). A–F. Lectotype, ♂ (ZMUKG). G–H. ♀ (MNHN). A–B, G–H. Habitus in dorsal view. C. Labels. D. Endophallus. E. Aedeagus in lateral view. F. Parameres in dorsal view. I. Distributional record map. Scale bars: A–B, G–H = 1.0 mm; D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514861" httpUri="https://zenodo.org/record/5514861/files/figure.png" pageId="8" pageNumber="48">
Fig. 4
<collectionCode id="ED16CCAB9D443E2277C0D16977E1FBF9" box="[641,663,1114,1140]" pageId="8" pageNumber="48">A</collectionCode>
–
<collectionCode id="ED16CCAB9D443E2277E6D16977CEFBF9" box="[679,696,1114,1140]" pageId="8" pageNumber="48">F</collectionCode>
</figureCitation>
)
<collectingCountry id="F31014FE9D443E2275FCD14E7478FB1A" box="[189,270,1149,1175]" name="India" pageId="8" pageNumber="48">INDIA</collectingCountry>
•
<specimenCount id="9D019FE79D443E22746DD14D7433FB15" box="[300,325,1150,1176]" pageId="8" pageNumber="48" type="male">♂</specimenCount>
; “
<typeStatus id="54BCEACC9D443E22741CD14E74DFFB1A" box="[349,425,1149,1175]" pageId="8" pageNumber="48">TYPE</typeStatus>
/ Ind. orient.
<collectorName id="26F231B89D443E227710D14E7614FB1A" box="[593,866,1149,1176]" pageId="8" pageNumber="48">Daldorff Mus. T. Lund</collectorName>
<taxonomicName id="4C072FED9D443E22762DD14D7132FB1A" box="[876,1092,1149,1176]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E22762DD14D7132FB1A" box="[876,1092,1149,1176]" italics="true" pageId="8" pageNumber="48">Geotrupes bicolor</emphasis>
</taxonomicName>
F. /
<typeStatus id="54BCEACC9D443E22713FD14D7185FB15" box="[1150,1267,1150,1176]" pageId="8" pageNumber="48" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C072FED9D443E2271BFD14D7459FB36" authority="F." authorityName="F." class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E2271BFD14D7464FB37" italics="true" pageId="8" pageNumber="48">Geotrupes bicolor</emphasis>
F.
</taxonomicName>
A.Frolov des. 2016”;
<collectionCode id="ED16CCAB9D443E227772D19277D4FB36" box="[563,674,1185,1211]" pageId="8" pageNumber="48">ZMUKG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB8546E9D443E2275ABD1D17194FA92" blockId="8.[189,1250,1250,1312]" pageId="8" pageNumber="48">
<materialsCitation id="3B6F5E339D443E2275ABD1D171A8FA92" ID-GBIF-Occurrence="3357770398" collectionCode="ZMUKG" country="India" location="INDIA" pageId="8" pageNumber="48" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<emphasis id="B973887C9D443E2275ABD1D174E1FB71" bold="true" box="[234,407,1250,1276]" pageId="8" pageNumber="48">
<typeStatus id="54BCEACC9D443E2275ABD1D174E1FB71" box="[234,407,1250,1276]" pageId="8" pageNumber="48" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
<collectingCountry id="F31014FE9D443E2275FCD0367478FA92" box="[189,270,1285,1311]" name="India" pageId="8" pageNumber="48">INDIA</collectingCountry>
•
<specimenCount id="9D019FE79D443E227466D0367420FAAD" box="[295,342,1285,1312]" pageId="8" pageNumber="48" type="male">1 ♂</specimenCount>
; “
<typeStatus id="54BCEACC9D443E22742CD03674CFFA92" box="[365,441,1285,1311]" pageId="8" pageNumber="48">TYPE</typeStatus>
/
<typeStatus id="54BCEACC9D443E227491D0367706FA92" box="[464,624,1285,1311]" pageId="8" pageNumber="48" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C072FED9D443E227736D036761CFAAD" authority="F." authorityName="F." box="[631,874,1285,1312]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B973887C9D443E227736D0367638FA92" box="[631,846,1285,1311]" italics="true" pageId="8" pageNumber="48">Geotrupes bicolor</emphasis>
F.
</taxonomicName>
A.Frolov des. 2016”;
<collectionCode id="ED16CCAB9D443E22712ED03671A8FA92" box="[1135,1246,1285,1311]" pageId="8" pageNumber="48">ZMUKG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD0757742FAED" blockId="8.[189,564,1350,1376]" box="[189,564,1350,1376]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E2275FCD0757742FAED" bold="true" box="[189,564,1350,1376]" fontSize="11" level="3" pageId="8" pageNumber="48" reason="3">
<emphasis id="B973887C9D443E2275FCD0757742FAED" bold="true" box="[189,564,1350,1376]" pageId="8" pageNumber="48">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD046702EFA3E" blockId="8.[189,1398,1395,1459]" pageId="8" pageNumber="48">
<materialsCitation id="3B6F5E339D443E2275FCD046746FFA3E" ID-GBIF-Occurrence="3357770342" collectingDate="1953-09-13" collectionCode="NHMB" collectorName="F. Keiser" country="Sri Lanka" latitude="7.3166666" location="Weragamtota" longLatPrecision="1303" longitude="80.98333" pageId="8" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Central">
<collectingCountry id="F31014FE9D443E2275FCD0467421FA02" box="[189,343,1397,1423]" name="Sri Lanka" pageId="8" pageNumber="48">SRI LANKA</collectingCountry>
–
<emphasis id="B973887C9D443E227434D046773CFA02" bold="true" box="[373,586,1397,1423]" pageId="8" pageNumber="48">
<collectingRegion id="49C39A8C9D443E227434D046773CFA02" box="[373,586,1397,1423]" country="Sri Lanka" name="Central" pageId="8" pageNumber="48">Central Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D019FE79D443E227722D04677E4FA1D" box="[611,658,1397,1424]" pageId="8" pageNumber="48" type="male">1 ♂</specimenCount>
;
<location id="8ED802B59D443E2277DFD0457648FA1D" LSID="urn:lsid:plazi:treatment:03AEE5789D443E22770CD4EE7786F938:8ED802B59D443E2277DFD0457648FA1D" box="[670,830,1398,1424]" country="Sri Lanka" latitude="7.3166666" longLatPrecision="1303" longitude="80.98333" name="Weragamtota" pageId="8" pageNumber="48" stateProvince="Central">Weragamtota</location>
;
<geoCoordinate id="EE3332A99D443E22760BD04076D1FA1D" box="[842,935,1395,1424]" degrees="7" direction="north" minutes="19" orientation="latitude" pageId="8" pageNumber="48" precision="925" value="7.3166666">7°19′ N</geoCoordinate>
,
<geoCoordinate id="EE3332A99D443E2276F2D040716AFA1D" box="[947,1052,1395,1424]" degrees="80" direction="east" minutes="59" orientation="longitude" pageId="8" pageNumber="48" precision="925" value="80.98333">80°59′ E</geoCoordinate>
;
<date id="FFB972AE9D443E227168D04671B0FA1D" box="[1065,1222,1397,1424]" pageId="8" pageNumber="48" value="1953-09-13">
<collectingDate id="EFFD8B469D443E227168D04671B0FA1D" box="[1065,1222,1397,1424]" pageId="8" pageNumber="48" value="1953-09-13">13 Sep. 1953</collectingDate>
</date>
;
<collectorName id="26F231B89D443E227193D045704AFA02" box="[1234,1340,1397,1424]" pageId="8" pageNumber="48">F. Keiser</collectorName>
leg.;
<collectionCode id="ED16CCAB9D443E2275FCD0AA746FFA3E" box="[189,281,1433,1459]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="48">NHMB</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B6F5E339D443E22747BD0AB7023FA3E" ID-GBIF-Occurrence="3357770330" box="[314,1365,1431,1459]" collectingDate="1983-10-30" collectionCode="MNHN" country="Sri Lanka" latitude="6.2833333" location="Tissamaharana" longLatPrecision="1304" longitude="81.28333" pageId="8" pageNumber="48" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Southern">
<emphasis id="B973887C9D443E22747BD0AB7755FA3F" bold="true" box="[314,547,1432,1458]" pageId="8" pageNumber="48">
<collectingRegion id="49C39A8C9D443E22747BD0AB7755FA3F" box="[314,547,1432,1458]" country="Sri Lanka" name="Southern" pageId="8" pageNumber="48">Southern Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D019FE79D443E22777DD0AB771CFA3E" box="[572,618,1432,1459]" pageId="8" pageNumber="48" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D019FE79D443E227736D0AB77D2FA3E" box="[631,676,1432,1459]" pageId="8" pageNumber="48" type="female">1 ♀</specimenCount>
;
<location id="8ED802B59D443E2277F3D0AB7613FA3F" LSID="urn:lsid:plazi:treatment:03AEE5789D443E22770CD4EE7786F938:8ED802B59D443E2277F3D0AB7613FA3F" box="[690,869,1432,1458]" country="Sri Lanka" latitude="6.2833333" longLatPrecision="1304" longitude="81.28333" name="Tissamaharana" pageId="8" pageNumber="48" stateProvince="Southern">Tissamaharana</location>
;
<geoCoordinate id="EE3332A99D443E227631D0A476BBFA3E" box="[880,973,1431,1459]" degrees="6" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="48" precision="925" value="6.2833333">6°17′ N</geoCoordinate>
,
<geoCoordinate id="EE3332A99D443E227698D0A47134FA3E" box="[985,1090,1431,1459]" degrees="81" direction="east" minutes="17" orientation="longitude" pageId="8" pageNumber="48" precision="925" value="81.28333">81°17′ E</geoCoordinate>
;
<date id="FFB972AE9D443E22710ED0AB719DFA3F" box="[1103,1259,1432,1458]" pageId="8" pageNumber="48" value="1983-10-30">
<collectingDate id="EFFD8B469D443E22710ED0AB719DFA3F" box="[1103,1259,1432,1458]" pageId="8" pageNumber="48" value="1983-10-30">30 Oct. 1983</collectingDate>
</date>
;
<collectionCode id="ED16CCAB9D443E2271B6D0AA7023FA3E" box="[1271,1365,1433,1459]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="48">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D07E59D443E2275FCD0EA7046F9AE" box="[189,1328,1497,1571]" pageId="8" pageNumber="48" type="description">
<paragraph id="8BB8546E9D443E2275FCD0EA7445FA7E" blockId="8.[189,307,1497,1523]" box="[189,307,1497,1523]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E2275FCD0EA7445FA7E" bold="true" box="[189,307,1497,1523]" fontSize="11" level="3" pageId="8" pageNumber="48" reason="3">
<emphasis id="B973887C9D443E2275FCD0EA7445FA7E" bold="true" box="[189,307,1497,1523]" pageId="8" pageNumber="48">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD33B7046F9AE" blockId="8.[189,1328,1544,1571]" box="[189,1328,1544,1571]" pageId="8" pageNumber="48">
Body length of the examined specimens varies from 7.5 to 9.0 mm (males) and
<quantity id="4CFFF98B9D443E227125D33B71C8F9AE" box="[1124,1214,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="48" unit="mm" value="8.6">8.6 mm</quantity>
(female).
</paragraph>
</subSubSection>
<subSubSection id="C31D07E59D443E2275FCD37A7786F938" pageId="8" pageNumber="48" type="distribution">
<paragraph id="8BB8546E9D443E2275FCD37A7420F9EE" blockId="8.[189,342,1609,1635]" box="[189,342,1609,1635]" pageId="8" pageNumber="48">
<heading id="D0F0E3029D443E2275FCD37A7420F9EE" bold="true" box="[189,342,1609,1635]" fontSize="11" level="3" pageId="8" pageNumber="48" reason="3">
<emphasis id="B973887C9D443E2275FCD37A7420F9EE" bold="true" box="[189,342,1609,1635]" pageId="8" pageNumber="48">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BB8546E9D443E2275FCD34B7786F938" blockId="8.[189,1399,1656,1718]" pageId="8" pageNumber="48">
The species was described form “East
<collectingCountry id="F31014FE9D443E22773CD34B77C9F91F" box="[637,703,1656,1682]" name="India" pageId="8" pageNumber="48">India</collectingCountry>
”. In
<collectingCountry id="F31014FE9D443E2277B2D34B761EF91F" box="[755,872,1656,1682]" name="Sri Lanka" pageId="8" pageNumber="48">Sri Lanka</collectingCountry>
, it is known from two rather distant localities in the centre and on the southern coast (
<figureCitation id="133C48EB9D443E2277D1D3A87794F938" box="[656,738,1691,1717]" captionStart="Fig" captionStartId="9.[189,232,1924,1950]" captionTargetBox="[189,1396,343,1826]" captionTargetId="figure-16@9.[189,1398,306,1844]" captionTargetPageId="9" captionText="Fig. 4. Orphnus bicolor (Fabricius, 1801). A–F. Lectotype, ♂ (ZMUKG). G–H. ♀ (MNHN). A–B, G–H. Habitus in dorsal view. C. Labels. D. Endophallus. E. Aedeagus in lateral view. F. Parameres in dorsal view. I. Distributional record map. Scale bars: A–B, G–H = 1.0 mm; D–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514861" httpUri="https://zenodo.org/record/5514861/files/figure.png" pageId="8" pageNumber="48">Fig. 4I</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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