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112 lines
14 KiB
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<document ID-DOI="10.11646/zootaxa.5244.6.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7673035" ID-ZooBank="8182831B-25FC-4B92-B319-281659660565" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1677225701698" checkinUser="plazi" docAuthor="Kluge, Nikita" docDate="2023" docId="03AE0E6FFFDE161E02DC93AB64AEFDD1" docLanguage="en" docName="zootaxa.5244.6.2.pdf" docOrigin="Zootaxa 5244 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.6.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Teloganodes Eaton 1882" docType="treatment" docVersion="2" lastPageNumber="530" masterDocId="FF977617FFDC161D024B957B6076FFC8" masterDocTitle="Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae)" masterLastPageNumber="552" masterPageNumber="527" pageNumber="529" updateTime="1677226200810" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Reclassification of Teloganodes Eaton 1882 (including subgenera Dudgeodes Sartori 2008 and Derlethina Sartori 2008) with a new species from Sulawesi (Ephemeroptera, Teloganodidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kluge, Nikita</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5244</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>527</mods:start>
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<mods:end>552</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.6.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5244.6.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7673035</mods:identifier>
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<mods:identifier type="ZooBank">8182831B-25FC-4B92-B319-281659660565</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7673037" ID-Zenodo-Dep="7673037" LSID="urn:lsid:plazi:treatment:03AE0E6FFFDE161E02DC93AB64AEFDD1" httpUri="http://treatment.plazi.org/id/03AE0E6FFFDE161E02DC93AB64AEFDD1" lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529">
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<subSubSection box="[151,533,1743,1770]" pageId="2" pageNumber="529" type="nomenclature">
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<paragraph blockId="2.[151,1437,1743,2023]" box="[151,533,1743,1770]" pageId="2" pageNumber="529">
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<heading bold="true" box="[151,533,1743,1770]" fontSize="11" level="1" pageId="2" pageNumber="529" reason="1">
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<emphasis bold="true" box="[151,533,1743,1770]" pageId="2" pageNumber="529">
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Autapomorphies of
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<taxonomicName ID-CoL="7TKG" ID-ENA="1661366" authorityName="Eaton" authorityYear="1882" box="[389,533,1743,1770]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[389,533,1743,1770]" italics="true" pageId="2" pageNumber="529">Teloganodes</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529" type="description">
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<paragraph blockId="2.[151,1437,1743,2023]" pageId="2" pageNumber="529">
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<emphasis bold="true" box="[151,183,1780,1806]" pageId="2" pageNumber="529">(1)</emphasis>
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On tergalii beginning from the pair III (either pairs III–V, or pairs III–IV, or pair III only), dorsal lobe is deeply cleft (
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<figureCitation box="[343,413,1816,1842]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1428]" captionTargetBox="[151,1436,455,1358]" captionTargetId="figure-141@1.[151,1436,450,1378]" captionTargetPageId="1" captionText="FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004)." figureDoi="http://doi.org/10.5281/zenodo.7673039" httpUri="https://zenodo.org/record/7673039/files/figure.png" pageId="2" pageNumber="529">Figs 1</figureCitation>
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,
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<figureCitation box="[426,581,1816,1843]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="2" pageNumber="529">51–53, 55–56</figureCitation>
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) (unique autapomorphy). Other, non-unique features of tergalii structure are: tergalius I is absent; tergalius II is operculate, i.e. with dorsal lobe convex, larger than others and never cleft; tergalii II and next ones (either II–V, or II–IV, or II–III) retain the two-branched ventral lobe bearing processes on both its branches (that is initial for
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<taxonomicName box="[626,818,1924,1950]" pageId="2" pageNumber="529" rank="superFamily" superFamily="Ephemerelloidea">Ephemerelloidea</taxonomicName>
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) (
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<figureCitation box="[840,910,1924,1950]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1428]" captionTargetBox="[151,1436,455,1358]" captionTargetId="figure-141@1.[151,1436,450,1378]" captionTargetPageId="1" captionText="FIGURE 1. Tergalii figured by Kluge (2004: figs 96E–H) and two classifications of Teloganodes. E, tergalii of T. (Teloganodes) kodai; F, tergalii of T. (Dudgeodes) romani (= Teloganodes/fg1 sp. T4: Kluge 2004); G, tergalii of T. (Dudgeodes) jacobusi (= Teloganodes/fg1 sp. T2: Kluge 2004); H, tergalii of T. (Derlethina) eloisae (= Teloganodes/fg1 sp. T3: Kluge 2004)." figureDoi="http://doi.org/10.5281/zenodo.7673039" httpUri="https://zenodo.org/record/7673039/files/figure.png" pageId="2" pageNumber="529">Figs 1</figureCitation>
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,
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<figureCitation box="[924,999,1924,1951]" captionStart="FIGURES 50–60" captionStartId="16.[151,264,1932,1956]" captionTargetBox="[151,1436,290,1909]" captionTargetId="figure-61@16.[151,1436,273,1909]" captionTargetPageId="16" captionText="FIGURES 50–60. Tergalii. 50–53, Teloganodes (Dudgeodes) celebensis, tergalii II–V; 54–57, Teloganodes (Dudgeodes) rasus sp. n., tergalii II–V; 58–60, tergalius V of two individuals (59–60, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7673061" httpUri="https://zenodo.org/record/7673061/files/figure.png" pageId="2" pageNumber="529">50–57</figureCitation>
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).
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</paragraph>
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<paragraph blockId="2.[151,1437,1743,2023]" pageId="2" pageNumber="529">
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<emphasis bold="true" box="[151,184,1960,1986]" pageId="2" pageNumber="529">(2)</emphasis>
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Larval, subimaginal and imaginal paracercus is vestigial, 1-segmented. The same in some other mayflies; among Ephemerella/fg1, the same in
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<taxonomicName authority="Demoulin 1954" authorityName="Demoulin" authorityYear="1954" box="[616,959,1996,2023]" class="Insecta" family="Dicercomyzidae" genus="Dicercomyzon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
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<emphasis box="[616,777,1997,2022]" italics="true" pageId="2" pageNumber="529">Dicercomyzon</emphasis>
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<bibRefCitation author="Demoulin, G." box="[784,959,1996,2023]" pageId="2" pageNumber="529" pagination="1 - 4" refId="ref10533" refString="Demoulin, G. (1954) Description preliminaire d'un type larvaire nouveau d'Ephemeropteres Tricorythidae du Congo Belge. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 3 (6), 1 - 4." type="journal article" year="1954">Demoulin 1954</bibRefCitation>
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</taxonomicName>
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only.
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</paragraph>
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<paragraph blockId="3.[151,1436,150,537]" pageId="3" pageNumber="530">
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<emphasis bold="true" box="[151,184,150,176]" pageId="3" pageNumber="530">(3)</emphasis>
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In subimago, paired lateral pigmented area of mesonotum (termed here
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<emphasis bold="true" box="[1009,1427,150,177]" pageId="3" pageNumber="530">latero-posterior scutal chromozone</emphasis>
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) occupies the whole sublateroscutum, the whole submedioscutum and a latero-posterior part of medioscutum, leaving a non-pigmented median area (termed here
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<emphasis bold="true" box="[820,980,223,249]" pageId="3" pageNumber="530">achromozone</emphasis>
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) in a form of unpaired median stripe narrowing posteriorly; the latero-posterior scutal chromozone is separated from the achromozone by a distinct, evenly arched line termed here
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<emphasis bold="true" box="[559,994,294,321]" pageId="3" pageNumber="530">latero-posterior chromozonal suture</emphasis>
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(
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<figureCitation box="[1010,1143,295,321]" captionStart="FIGURES 27–30" captionStartId="12.[151,264,1697,1721]" captionTargetBox="[151,1428,182,1673]" captionTargetId="figure-23@12.[151,1436,181,1673]" captionTargetPageId="12" captionText="FIGURES 27–30. Teloganodes (Dudgeodes) celebensis. 27–28, left half of subimaginal exuviae of mesonotum; 29–30, hind wing in falling and transmitted light. Abbreviations: acz, achromozone; cz.a, anterior chromozone; cz.l-p, latero-posterior chromozone; cz.s, chromozonal suture; LPs, lateroparapsidal suture; MPs, medioparapsidal suture; MS, medioscutum; SMS, submedioscutum." figureDoi="http://doi.org/10.5281/zenodo.7673051" httpUri="https://zenodo.org/record/7673051/files/figure.png" pageId="3" pageNumber="530">Figs 27–28</figureCitation>
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). The area anteriad of the achromozone is termed here
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<emphasis bold="true" box="[521,848,330,357]" pageId="3" pageNumber="530">anterior scutal chromozone</emphasis>
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. Color of the anterior scutal chromozone, the lateroposterior scutal chromozones and the achromozone vary among species, but shape of their borders is constant for
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<taxonomicName authorityName="Eaton" authorityYear="1882" box="[240,381,403,429]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis box="[240,381,403,429]" italics="true" pageId="3" pageNumber="530">Teloganodes</emphasis>
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</taxonomicName>
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. Microtrichia cover most part of scutum, being absent on lateroscutum and posterior scutal protuberances (
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<figureCitation box="[371,456,439,465]" captionStart="FIGURES 95–101" captionStartId="23.[151,264,1926,1950]" captionTargetBox="[151,1436,273,1903]" captionTargetId="figure-56@23.[151,1436,273,1903]" captionTargetPageId="23" captionText="FIGURES 95–101. Teloganodes (Dudgeodes) rasus sp. n., male (holotype). 95–96, penis, dorsal view; 97, genitalia, ventral view; 98–99, halves of subimaginal exuviae of mesonotum, in Canadian balsam and in dry condition; 100–101, hind wing, in falling and transmitted light." figureDoi="http://doi.org/10.5281/zenodo.7673083" httpUri="https://zenodo.org/record/7673083/files/figure.png" pageId="3" pageNumber="530">Fig. 99</figureCitation>
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). In contrast to
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<taxonomicName authorityName="Klapalek" authorityYear="1909" box="[632,808,439,465]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
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and the most primitive representatives of Pantricorythi, the lateroparapsidal sutures of subimago and imago of
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<taxonomicName authorityName="Eaton" authorityYear="1882" box="[814,955,475,501]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis box="[814,955,475,501]" italics="true" pageId="3" pageNumber="530">Teloganodes</emphasis>
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</taxonomicName>
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are not curved laterally, but converge with the medioparapsidal sutures (
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<bibRefCitation author="Kluge, N. J." box="[525,662,511,537]" pageId="3" pageNumber="530" refId="ref10967" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
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: p. 294, character 10 of Ephemerella/fg1) (
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<figureCitation box="[1143,1228,511,537]" captionStart="FIGURES 27–30" captionStartId="12.[151,264,1697,1721]" captionTargetBox="[151,1428,182,1673]" captionTargetId="figure-23@12.[151,1436,181,1673]" captionTargetPageId="12" captionText="FIGURES 27–30. Teloganodes (Dudgeodes) celebensis. 27–28, left half of subimaginal exuviae of mesonotum; 29–30, hind wing in falling and transmitted light. Abbreviations: acz, achromozone; cz.a, anterior chromozone; cz.l-p, latero-posterior chromozone; cz.s, chromozonal suture; LPs, lateroparapsidal suture; MPs, medioparapsidal suture; MS, medioscutum; SMS, submedioscutum." figureDoi="http://doi.org/10.5281/zenodo.7673051" httpUri="https://zenodo.org/record/7673051/files/figure.png" pageId="3" pageNumber="530">Fig. 28</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |