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<document id="BE80149889341F9D66E874DD187C7960" ID-DOI="10.24199/j.mmv.2005.62.2" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665561" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1708004547937" checkinUser="felipe" docAuthor="Fitzgerald, Erich M. G." docDate="2005" docId="03ACB35ADC7AFF91FF7AFC82FBA0FA95" docLanguage="en" docName="MemMusVictoria.62.1.67-89.pdf" docOrigin="Memoirs of Museum Victoria 62 (1)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-62-issue-1-2005/pages-67-89/" docStyle="DocumentStyle:99484BB52C5EA01CDE4EA5EECB5782C2.2:MemMusVictoria.2003-.journal_article.type2" docStyleId="99484BB52C5EA01CDE4EA5EECB5782C2" docStyleName="MemMusVictoria.2003-.journal_article.type2" docStyleVersion="2" docTitle="Delphinoidea Gray 1821" docType="treatment" docVersion="3" lastPageNumber="75" masterDocId="FF95CB22DC72FF99FFF0FFF6FF94FF8B" masterDocTitle="Pliocene marine mammals from the Whalers Bluff Formation of Portland, Victoria, Australia" masterLastPageNumber="89" masterPageNumber="67" pageNumber="75" updateTime="1711396190357" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="424E08624201A32A41F151AD6B1AFBB1">Pliocene marine mammals from the Whalers Bluff Formation of Portland, Victoria, Australia</mods:title>
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<mods:namePart id="9081C61DDA849D1B8EAE26EC46136DCE">Fitzgerald, Erich M. G.</mods:namePart>
<mods:affiliation id="2C26C333773A4F88D97E7AD62ED9E8E2">School of Geosciences, Monash University, Vic. 3800, Australia and Museum Victoria, G. P. O. Box 666, Melbourne, Vic. 3001, Australia</mods:affiliation>
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<mods:title id="1ED2030C6ADAE55458F4760F96ED1C53">Memoirs of Museum Victoria</mods:title>
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<mods:date id="60AC0D4500A2ED3E81D9DD9B5D1424E4">2005</mods:date>
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<treatment id="03ACB35ADC7AFF91FF7AFC82FBA0FA95" LSID="urn:lsid:plazi:treatment:03ACB35ADC7AFF91FF7AFC82FBA0FA95" httpUri="http://treatment.plazi.org/id/03ACB35ADC7AFF91FF7AFC82FBA0FA95" lastPageNumber="75" pageId="8" pageNumber="75">
<subSubSection id="C31F51C7DC7AFF91FF7AFC82FE89FC3D" pageId="8" pageNumber="75" type="nomenclature">
<paragraph id="8BBA024CDC7AFF91FF7AFC82FD84FC01" blockId="8.[138,528,883,906]" box="[138,528,883,906]" pageId="8" pageNumber="75">
<heading id="D0F2B520DC7AFF91FF7AFC82FD84FC01" box="[138,528,883,906]" fontSize="9" level="2" pageId="8" pageNumber="75" reason="3">
Superfamily
<taxonomicName id="4C0579CFDC7AFF91FEFCFC85FD84FC01" authority="Gray, 1821" authorityName="Gray" authorityYear="1821" box="[268,528,883,906]" kingdom="Animalia" pageId="8" pageNumber="75" rank="superFamily" superFamily="Delphinoidea">
<emphasis id="B971DE5EDC7AFF91FEFCFC85FE0EFC02" bold="true" box="[268,410,883,905]" pageId="8" pageNumber="75">Delphinoidea</emphasis>
Gray, 1821
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</paragraph>
<paragraph id="8BBA024CDC7AFF91FF7AFC57FE89FC3D" blockId="8.[138,285,928,950]" box="[138,285,928,950]" pageId="8" pageNumber="75">
<heading id="D0F2B520DC7AFF91FF7AFC57FE89FC3D" bold="true" box="[138,285,928,950]" fontSize="9" level="2" pageId="8" pageNumber="75" reason="2">
<emphasis id="B971DE5EDC7AFF91FF7AFC57FE89FC3D" bold="true" box="[138,285,928,950]" pageId="8" pageNumber="75">Incertae sedis</emphasis>
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</paragraph>
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<subSubSection id="C31F51C7DC7AFF91FF7AFC3CFBA0FA95" pageId="8" pageNumber="75" type="materials_examined">
<paragraph id="8BBA024CDC7AFF91FF7AFC3CFEC4FC73" blockId="8.[138,769,969,1016]" pageId="8" pageNumber="75">
<emphasis id="B971DE5EDC7AFF91FF7AFC3CFED4FC56" box="[138,320,970,989]" italics="true" pageId="8" pageNumber="75">Referred specimens.</emphasis>
<collectionCode id="ED149A89DC7AFF91FEB4FC3CFEECFC56" box="[324,376,970,989]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="8" pageNumber="75" type="Museum">NMV</collectionCode>
P218283, P218284 and P218286, all isolated teeth (not figured).
</paragraph>
<paragraph id="8BBA024CDC7AFF91FF7AFBF8FD68FB5A" blockId="8.[138,770,1037,1551]" pageId="8" pageNumber="75">
<emphasis id="B971DE5EDC7AFF91FF7AFBF8FE94FBAF" box="[138,256,1038,1060]" italics="true" pageId="8" pageNumber="75">Description</emphasis>
. P21283, P218284 and P218286 all represent small odontocete teeth possessing conical enamel-covered crowns that bear fine wrinkling ornamentation, and curve lingually towards the crown apex. As in kentriodontids, there is a lingual bulge at the base of the crown but this feature is not as prominent in the teeth from Portland. None possesses an open pulp cavity suggesting that all were derived from adult individuals.
</paragraph>
<paragraph id="8BBA024CDC7AFF91FF44FB2EFE6EFACE" blockId="8.[138,770,1037,1551]" pageId="8" pageNumber="75">
P218283 is an incomplete tooth, its preserved maximum length and maximum width of the crown being
<quantity id="4CFDAFA9DC7AFF91FD87FB03FD54FA80" box="[631,704,1269,1291]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="75" unit="mm" value="16.0">16 mm</quantity>
and
<quantity id="4CFDAFA9DC7AFF91FD04FB03FF25FAA3" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="75" unit="mm" value="6.0">6 mm</quantity>
respectively. Due to the incomplete nature of this tooth it does not warrant further description.
</paragraph>
<paragraph id="8BBA024CDC7AFF91FF5FFABAFD14FA33" blockId="8.[138,770,1037,1551]" pageId="8" pageNumber="75">P218284 is the most highly polished and the most complete. It differs from the others in having a mediolaterally compressed root with a more prominent mesial-distal bulge at its midpoint. The preserved apex of the root curves posteriorly.</paragraph>
<paragraph id="8BBA024CDC7AFF91FF5AFA49FDF1F984" blockId="8.[138,770,1037,1551]" pageId="8" pageNumber="75">The most notable feature of P218286 distinguishing it from the other teeth is its bulbous root, which contrasts with the transversely flattened morphology of P218284.</paragraph>
<paragraph id="8BBA024CDC7AFF91FF7AF9D3FACCFB21" blockId="8.[138,769,1572,1941]" lastBlockId="8.[806,1438,883,1310]" pageId="8" pageNumber="75">
<emphasis id="B971DE5EDC7AFF91FF7AF9D3FF6CF9B0" box="[138,248,1573,1595]" italics="true" pageId="8" pageNumber="75">Discussion</emphasis>
. Only one delphinoid odontocete has previously been described from the Tertiary of
<collectingCountry id="F31242DCDC7AFF91FDDBF9B7FD1EF9DC" box="[555,650,1601,1623]" name="Australia" pageId="8" pageNumber="75">Australia</collectingCountry>
, the latest Miocene-earliest Pliocene “
<taxonomicName id="4C0579CFDC7AFF91FE55F9A9FE48F9FE" authorityName="Gray" authorityYear="1846" box="[421,476,1631,1653]" class="Mammalia" family="Delphinidae" genus="Steno" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="B971DE5EDC7AFF91FE55F9A9FE48F9FE" box="[421,476,1631,1653]" italics="true" pageId="8" pageNumber="75">Steno</emphasis>
</taxonomicName>
<emphasis id="B971DE5EDC7AFF91FE00F9A9FDD9F9FE" box="[496,589,1631,1653]" italics="true" pageId="8" pageNumber="75">cudmorei</emphasis>
<bibRefCitation id="EF947FBDDC7AFF91FDA6F9A8FC95F9FF" author="Chapman, F." box="[598,769,1630,1652]" pageId="8" pageNumber="75" pagination="32 - 43" refId="ref13758" refString="Chapman, F. 1917. New or little-known Victorian fossils in the National Museum. Part XXI. Some Tertiary cetacean remains. Proceedings of the Royal Society of Victoria 30: 32 - 43." type="journal article" year="1917">Chapman (1917)</bibRefCitation>
from the Black Rock Sandstone of Beaumaris,
<collectingRegion id="49C1CCAEDC7AFF91FD5EF98DFC95F91A" box="[686,769,1659,1681]" country="Australia" name="Victoria" pageId="8" pageNumber="75">Victoria</collectingRegion>
(
<bibRefCitation id="EF947FBDDC7AFF91FF61F96EFEDCF925" author="Fitzgerald, E. M. G." box="[145,328,1688,1710]" pageId="8" pageNumber="75" pagination="183 - 208" refId="ref14100" refString="Fitzgerald, E. M. G. 2004 b. A review of the Tertiary fossil Cetacea (Mammalia) localities in Australia. Memoirs of Museum Victoria 61: 183 - 208." type="journal article" year="2004">Fitzgerald, 2004b</bibRefCitation>
).
<bibRefCitation id="EF947FBDDC7AFF91FEAAF96EFE62F925" author="Fordyce, R. E." box="[346,502,1688,1710]" pageId="8" pageNumber="75" pagination="43 - 58" refId="ref14135" refString="Fordyce, R. E. 1982. A review of Australian fossil Cetacea. Memoirs of the National Museum of Victoria 43: 43 - 58." type="journal article" year="1982">Fordyce (1982)</bibRefCitation>
questioned the taxonomic identity of “
<emphasis id="B971DE5EDC7AFF91FEF5F943FE85F940" box="[261,273,1717,1739]" italics="true" pageId="8" pageNumber="75">S</emphasis>
.”
<emphasis id="B971DE5EDC7AFF91FEDBF943FE1CF940" box="[299,392,1717,1739]" italics="true" pageId="8" pageNumber="75">cudmorei</emphasis>
(the
<typeStatus id="54BEBCEEDC7AFF91FE4EF943FD8CF940" box="[446,536,1717,1739]" pageId="8" pageNumber="75" type="holotype">holotype</typeStatus>
, P13033, being an isolated tooth) and
<bibRefCitation id="EF947FBDDC7AFF91FEC6F924FE60F963" author="Fitzgerald, E. M. G." box="[310,500,1746,1768]" pageId="8" pageNumber="75" pagination="183 - 208" refId="ref14100" refString="Fitzgerald, E. M. G. 2004 b. A review of the Tertiary fossil Cetacea (Mammalia) localities in Australia. Memoirs of Museum Victoria 61: 183 - 208." type="journal article" year="2004">Fitzgerald (2004b)</bibRefCitation>
referred “
<emphasis id="B971DE5EDC7AFF91FD90F924FDF8F963" box="[608,620,1746,1768]" italics="true" pageId="8" pageNumber="75">S</emphasis>
.”
<emphasis id="B971DE5EDC7AFF91FD77F924FD70F963" box="[647,740,1746,1768]" italics="true" pageId="8" pageNumber="75">cudmorei</emphasis>
to
<taxonomicName id="4C0579CFDC7AFF91FF7AF919FE9FF88E" authorityName="Gray" authorityYear="1821" box="[138,267,1775,1797]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
, genus and species indeterminate.
<bibRefCitation id="EF947FBDDC7AFF91FD6FF919FF59F8A9" author="Chapman, F." pageId="8" pageNumber="75" pagination="32 - 43" refId="ref13758" refString="Chapman, F. 1917. New or little-known Victorian fossils in the National Museum. Part XXI. Some Tertiary cetacean remains. Proceedings of the Royal Society of Victoria 30: 32 - 43." type="journal article" year="1917">Chapman (1917)</bibRefCitation>
assigned P13033 to
<taxonomicName id="4C0579CFDC7AFF91FE5BF8FAFE76F8A9" authorityName="Gray" authorityYear="1846" box="[427,482,1804,1826]" class="Mammalia" family="Delphinidae" genus="Steno" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="B971DE5EDC7AFF91FE5BF8FAFE76F8A9" box="[427,482,1804,1826]" italics="true" pageId="8" pageNumber="75">Steno</emphasis>
</taxonomicName>
on the basis of the resemblance of its crown ornamentation to that seen in the teeth of the extant
<taxonomicName id="4C0579CFDC7AFF91FF0EF8B0FD9FF8D7" authority="Cuvier" authorityName="Cuvier" box="[254,523,1862,1884]" class="Mammalia" family="Delphinidae" genus="Steno" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="bredanensis">
<emphasis id="B971DE5EDC7AFF91FF0EF8B0FE2EF8D7" box="[254,442,1862,1884]" italics="true" pageId="8" pageNumber="75">Steno bredanensis</emphasis>
Cuvier
</taxonomicName>
in Lesson, 1828 (e.g.,
<bibRefCitation id="EF947FBDDC7AFF91FF7AF895FE09F8F2" author="Miyazaki, N. &amp; Perrin, W. F." box="[138,413,1891,1913]" pageId="8" pageNumber="75" pagination="1 - 21" refId="ref16126" refString="Miyazaki, N., and Perrin, W. F. 1994. Rough-toothed dolphin Steno bredanensis (Lesson, 1828). Pp. 1 - 21 in: Ridgway, S. H., and Harrison, R. (eds), Handbook of Marine Mammals, Volume 5: The First Book of Dolphins. Academic Press: London." type="book chapter" year="1994">Miyazaki and Perrin, 1994</bibRefCitation>
). Given that
<taxonomicName id="4C0579CFDC7AFF91FDD2F895FDCDF8F2" authorityName="Gray" authorityYear="1846" box="[546,601,1891,1913]" class="Mammalia" family="Delphinidae" genus="Steno" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="B971DE5EDC7AFF91FDD2F895FDCDF8F2" box="[546,601,1891,1913]" italics="true" pageId="8" pageNumber="75">Steno</emphasis>
</taxonomicName>
is probably in a basal position in the phylogeny of
<taxonomicName id="4C0579CFDC7AFF91FE1DF889FDF3F81E" authorityName="Gray" authorityYear="1821" box="[493,615,1919,1941]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
(
<bibRefCitation id="EF947FBDDC7AFF91FD85F889FC3DFC02" author="Miyazaki, N. &amp; Perrin, W. F." box="[629,937,883,1941]" pageId="8" pageNumber="75" pagination="1 - 21" refId="ref16126" refString="Miyazaki, N., and Perrin, W. F. 1994. Rough-toothed dolphin Steno bredanensis (Lesson, 1828). Pp. 1 - 21 in: Ridgway, S. H., and Harrison, R. (eds), Handbook of Marine Mammals, Volume 5: The First Book of Dolphins. Academic Press: London." type="book chapter" year="1994">Miyazaki and Perrin, 1994</bibRefCitation>
;
<bibRefCitation id="EF947FBDDC7AFF91FC47FC85FBEAFC02" author="LeDuc, R. G. &amp; Perrin, W. F. &amp; Dizon, A. E." box="[951,1150,883,905]" pageId="8" pageNumber="75" pagination="619 - 648" refId="ref15772" refString="LeDuc, R. G., Perrin, W. F., and Dizon, A. E. 1999. Phylogenetic relationships among the delphinoid cetaceans based on full cytochrome b sequences. Marine Mammal Science 15: 619 - 648." type="journal article" year="1999">LeDuc et al., 1999</bibRefCitation>
) and that some of the presumed ancestors of
<taxonomicName id="4C0579CFDC7AFF91FC01FC66FBFAFC2D" authorityName="Gray" authorityYear="1821" box="[1009,1134,912,934]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
, the
<taxonomicName id="4C0579CFDC7AFF91FB6FFC66FAD5FC2D" box="[1183,1345,912,934]" class="Mammalia" family="Kentriodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="family">Kentriodontidae</taxonomicName>
(
<bibRefCitation id="EF947FBDDC7AFF91FAA0FC66FCCEFC48" author="Barnes, L. G." pageId="8" pageNumber="75" pagination="1 - 35" refId="ref12890" refString="Barnes, L. G. 1978. A review of Lophocetus and Liolithax and their relationships to the delphinoid family Kentriodontidae (Cetacea: Odontoceti). Bulletin of the Natural History Museum of Los Angeles County 28: 1 - 35." type="journal article" year="1978">Barnes, 1978</bibRefCitation>
,
<bibRefCitation id="EF947FBDDC7AFF91FC98FC5BFC09FC48" author="Barnes, L. G." box="[872,925,941,963]" pageId="8" pageNumber="75" pagination="314 - 316" refId="ref13093" refString="Barnes, L. G. 2002. Delphinoids, evolution of the modern families. Pp. 314 - 316 in: Perrin, W. F., Wursig, B., and Thewissen, J. G. M. (eds), Encyclopedia of Marine Mammals. Academic Press: San Diego." type="book chapter" year="2002">2002</bibRefCitation>
;
<bibRefCitation id="EF947FBDDC7AFF91FC5AFC5BFBA0FC48" author="LeDuc, R. G." box="[938,1076,941,963]" pageId="8" pageNumber="75" pagination="310 - 314" refId="ref15817" refString="LeDuc, R. G. 2002. Delphinids, overview. Pp. 310 - 314 in: Perrin, W. F., Wursig, B., and Thewissen, J. G. M. (eds), Encyclopedia of Marine Mammals. Academic Press: San Diego." type="book chapter" year="2002">LeDuc, 2002</bibRefCitation>
), possessed
<emphasis id="B971DE5EDC7AFF91FB44FC5BFA8CFC48" box="[1204,1304,941,963]" italics="true" pageId="8" pageNumber="75">Steno-like</emphasis>
crown ornamentation (e.g.,
<bibRefCitation id="EF947FBDDC7AFF91FC3AFC3CFBC8FC6B" author="Kellogg, R." box="[970,1116,970,992]" pageId="8" pageNumber="75" pagination="99 - 101" refId="ref15692" refString="Kellogg, R. 1966. Fossil marine mammals from the Miocene Calvert Formation of Maryland and Virginia. Part 4. A new odontocete from the Calvert Miocene of Maryland. United States National Museum Bulletin 247: 99 - 101." type="journal article" year="1966">Kellogg, 1966</bibRefCitation>
), the anastomosing striae on the crown of P13033 (and P218283, P218284, P218286) are of dubious use in assessing the phylogenetic affinities of isolated teeth. Furthermore, non-delphinid small odontocetes such as
<taxonomicName id="4C0579CFDC7AFF91FCD6FBC8FBC7FBDF" authority="Miller, 1918" authorityName="Miller" authorityYear="1918" box="[806,1107,1086,1108]" class="Mammalia" family="Lipotidae" genus="Lipotes" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="vexillifer">
<emphasis id="B971DE5EDC7AFF91FCD6FBC8FC5BFBDF" box="[806,975,1086,1108]" italics="true" pageId="8" pageNumber="75">Lipotes vexillifer</emphasis>
<bibRefCitation id="EF947FBDDC7AFF91FC26FBC8FBC7FBDF" author="Miller, G. S., Jr." box="[982,1107,1086,1108]" pageId="8" pageNumber="75" pagination="1 - 12" refId="ref16053" refString="Miller, G. S., Jr. 1918. A new river-dolphin from China. Smithsonian Miscellaneous Collections 68 (2486): 1 - 12." type="journal article" year="1918">Miller, 1918</bibRefCitation>
</taxonomicName>
possess anastomosing wrinkling on tooth crown enamel (
<bibRefCitation id="EF947FBDDC7AFF91FBDAFBADFB3BFBFA" author="Miller, G. S., Jr." box="[1066,1199,1115,1137]" pageId="8" pageNumber="75" pagination="1 - 12" refId="ref16053" refString="Miller, G. S., Jr. 1918. A new river-dolphin from China. Smithsonian Miscellaneous Collections 68 (2486): 1 - 12." type="journal article" year="1918">Miller, 1918</bibRefCitation>
;
<bibRefCitation id="EF947FBDDC7AFF91FB4DFBADFCCFFB05" author="Brownell, R. L., Jr. &amp; Herald, E. S." pageId="8" pageNumber="75" pagination="1 - 4" refId="ref13728" refString="Brownell, R. L., Jr., and Herald, E. S. 1972. Lipotes vexillifer. Mammalian Species 10: 1 - 4." type="journal article" year="1972">Brownell and Herald, 1972</bibRefCitation>
;
<bibRefCitation id="EF947FBDDC7AFF91FC9EFB8EFC6FFB05" author="Barnes, L. G." box="[878,1019,1144,1166]" pageId="8" pageNumber="75" pagination="1 - 34" refId="ref12966" refString="Barnes, L. G. 1985. Fossil pontoporiid dolphins (Mammalia: Cetacea) from the Pacific coast of North America. Contributions in Science, Natural History Museum of Los Angeles County 363: 1 - 34." type="journal article" year="1985">Barnes, 1985</bibRefCitation>
) which casts doubt on any perceived taxonomic or phylogenetic signal present in these teeth.
</paragraph>
<paragraph id="8BBA024CDC7AFF91FCB6FB47FBA0FA95" blockId="8.[806,1438,883,1310]" pageId="8" pageNumber="75">
The isolated teeth from Portland are assigned to
<taxonomicName id="4C0579CFDC7AFF91FCD6FB38FC39FB6F" authorityName="Gray" authorityYear="1821" box="[806,941,1230,1252]" pageId="8" pageNumber="75" rank="superFamily" superFamily="Delphinoidea">Delphinoidea</taxonomicName>
incertae sedis. None of the Portland Pliocene teeth share demonstrably close affinities with the
<typeStatus id="54BEBCEEDC7AFF91FAB6FB1DFA09FA8A" box="[1350,1437,1259,1281]" pageId="8" pageNumber="75" type="holotype">holotype</typeStatus>
tooth of “
<taxonomicName id="4C0579CFDC7AFF91FC77FAFEFC2AFA95" authorityName="Gray" authorityYear="1846" box="[903,958,1288,1310]" class="Mammalia" family="Delphinidae" genus="Steno" kingdom="Animalia" order="Cetacea" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="B971DE5EDC7AFF91FC77FAFEFC2AFA95" box="[903,958,1288,1310]" italics="true" pageId="8" pageNumber="75">Steno</emphasis>
</taxonomicName>
<emphasis id="B971DE5EDC7AFF91FC21FAFEFBBAFA95" box="[977,1070,1288,1310]" italics="true" pageId="8" pageNumber="75">cudmorei</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>