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<document id="E5642AB3DC766D62D27C7C6F95906218" ID-CLB-Dataset="32146" ID-DOI="10.15553/c2017v721a12" ID-GBIF-Dataset="b2bae7e5-7a3f-4d18-b2b1-c445afbdb5e2" ID-ISSN="2235-3658" ID-Zenodo-Dep="888344" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505072655702" checkinUser="plazi" docAuthor="Eva Maier, Michelle J. Price &amp; Terry A. Hedderson" docDate="2017" docId="03AC87ED6D4FFF86FCBFE4B722C2FD7C" docLanguage="en" docName="Candollea.72.1.199-230.pdf" docOrigin="Candollea 72 (1)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Grimmia pulvinata (Hedw.) Sm." docType="treatment" docVersion="13" lastPageNumber="223" masterDocId="FF95FF956D5AFF9EFF9EE6642008FFBA" masterDocTitle="A revision of Grimmia (Grimmiaceae) from South Africa and Lesotho" masterLastPageNumber="230" masterPageNumber="199" pageNumber="220" updateTime="1698468194828" updateUser="ExternalLinkService">
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<mods:title id="2831CFE1A182A98A8A581CCEF84D23C9">A revision of Grimmia (Grimmiaceae) from South Africa and Lesotho</mods:title>
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<mods:namePart id="FD9B8EEC8816F6EE6DB6B2BEC0A671CC">Eva Maier</mods:namePart>
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<mods:namePart id="B96FFD1EF3D55433B6005A571BD632F1">Michelle J. Price</mods:namePart>
<mods:affiliation id="4DFEFF27EC0CFF3AA3C46DC9247DB87F">Conservatoire et Jardin botaniques de la Ville de Genève, C. P. 60, 1292 Chambésy, Switzerland.</mods:affiliation>
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<mods:namePart id="467106B124715A19FA3BBC4011C6E845">Terry A. Hedderson</mods:namePart>
<mods:affiliation id="73ABCCEBA47C1CB73C6572BC958B1350">Bolus Herbarium, University of Cape Town, Private Bag, Rondebosch 7701, South Africa</mods:affiliation>
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<mods:title id="A23C1261A4DA02ABA3F0C7A278603BA0">Candollea</mods:title>
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<treatment id="03AC87ED6D4FFF86FCBFE4B722C2FD7C" ID-DOI="http://doi.org/10.5281/zenodo.5722027" ID-GBIF-Taxon="134016925" ID-Zenodo-Dep="5722027" LSID="urn:lsid:plazi:treatment:03AC87ED6D4FFF86FCBFE4B722C2FD7C" httpUri="http://treatment.plazi.org/id/03AC87ED6D4FFF86FCBFE4B722C2FD7C" lastPageId="24" lastPageNumber="223" pageId="21" pageNumber="220">
<paragraph id="8BBA36FB6D4FFF8BFCBFE4B723B7FCB1" blockId="21.[801,1453,722,1273]" pageId="21" pageNumber="220">
<subSubSection id="C31F65706D4FFF8BFCBFE4B724A5FD51" box="[801,1197,722,747]" pageId="21" pageNumber="220" type="nomenclature">
10.
<taxonomicName id="4C054D786D4FFF8BFCD7E4B724A5FD51" ID-CoL="6L4QJ" authority="(Hedw.) Sm." authorityName="(Hedw.) Sm." box="[841,1197,722,747]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="21" pageNumber="220" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4FFF8BFCD7E4B72414FD50" bold="true" box="[841,1052,722,747]" italics="true" pageId="21" pageNumber="220">Grimmia pulvinata</emphasis>
(Hedw.) Sm.
</taxonomicName>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFB2AE4B7235EFCB1" pageId="21" pageNumber="220" type="reference_group">
<treatmentCitation id="0AA410EA6D4FFF8BFB2AE4B7235EFCB1" author="Sm." page="1728" pageId="21" pageNumber="220" year="1807">in Engl. Bot. 24: 1728. 1807</treatmentCitation>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFCC3E49723B7FCB1" box="[861,959,755,779]" pageId="21" pageNumber="220" type="description">
(
<figureCitation id="133E2A7E6D4FFF8BFCFBE49723BAFCB1" box="[869,946,755,779]" captionStart="Fig. 12" captionStartId="22.[136,163,1876,1895]" captionTargetBox="[136,1388,293,1828]" captionTargetId="figure@22.[136,1399,276,1832]" captionTargetPageId="22" captionText="Fig. 12. Grimmia pulvinata (Hedw.) Sm. A. Transverse section of stem; B, C. Leaves; D, E. Outlines of transverse sections of leaves; F, G. Cells in leaf base; H, I. Transverse sections of leaves. [A, D, H: Hedderson 13601, BOL; C, F: Hedderson 13148, BOL; B, E, I: Hedderson 13851, BOL; G: Hedderson 13677, BOL]" httpUri="https://zenodo.org/record/888354/files/figure.png" pageId="21" pageNumber="220">Fig. 12</figureCitation>
).
</subSubSection>
</paragraph>
<subSubSection id="C31F65706D4FFF8BFCD9E57B25A5FC8C" box="[839,1453,798,822]" pageId="21" pageNumber="220" type="reference_group">
<paragraph id="8BBA36FB6D4FFF8BFCD9E57B25A5FC8C" blockId="21.[801,1453,722,1273]" box="[839,1453,798,822]" pageId="21" pageNumber="220">
[
<treatmentCitation id="0AA410EA6D4FFF8BFCF3E57A25A5FC8C" author="Hedw." box="[877,1453,798,822]" page="158" pageId="21" pageNumber="220" year="1801">
<taxonomicName id="4C054D786D4FFF8BFCF3E57A2533FC8C" authority="Hedw., Sp. Musc. Frond." authorityName="Hedw., Sp. Musc. Frond." box="[877,1339,798,822]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="21" pageNumber="220" phylum="Bryophyta" rank="species" species="pulvinatus">
<emphasis id="B971EAE96D4FFF8BFCF3E57A243DFC8C" box="[877,1077,798,822]" italics="true" pageId="21" pageNumber="220">Fissidens pulvinatus</emphasis>
Hedw., Sp. Musc. Frond.
</taxonomicName>
158. 1801.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFCD9E52D24BBFC1B" pageId="21" pageNumber="220" type="materials_examined">
<paragraph id="8BBA36FB6D4FFF8BFCD9E52D24BBFC1B" blockId="21.[801,1453,722,1273]" pageId="21" pageNumber="220">
<materialsCitation id="3B6D3CA66D4FFF8BFCD9E52D24BBFC1B" ID-GBIF-Occurrence="3467920347" collectionCode="G" location="&quot; In tectis et muris vulgaris, vere maturans &quot;" pageId="21" pageNumber="220" specimenCode="G00040350" specimenCount="1" typeStatus="lectotype">
<emphasis id="B971EAE96D4FFF8BFCD9E52D23C3FCDB" bold="true" box="[839,971,841,865]" pageId="21" pageNumber="220">
<typeStatus id="54BE88596D4FFF8BFCD9E52D23C3FCDB" box="[839,971,841,865]" pageId="21" pageNumber="220" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated by
<bibRefCitation id="EF944B0A6D4FFF8BFB1FE52D2590FCDB" author="Cao, T. &amp; D. H. Vitt" box="[1153,1432,841,865]" firstAuthor="Cao" journalOrPublisher="J. Hattori Bot. Lab" page="191" pageId="21" pageNumber="220" pagination="123 - 247" part="61" refId="ref21091" refString="Cao, T. &amp; D. H. Vitt (1986). A taxonomic revision and phylogenetic analysis of Grimmia and Schistidium (Bryopsida, Grimmiaceae) in China. J. Hattori Bot. Lab. 61: 123 - 247." title="A taxonomic revision and phylogenetic analysis of Grimmia and Schistidium (Bryopsida, Grimmiaceae) in China" type="journal article" year="1986">Cao &amp; Vitt, 1986: 191</bibRefCitation>
):
<emphasis id="B971EAE96D4FFF8BFCD9E50D23CDFC3B" bold="true" box="[839,965,873,897]" pageId="21" pageNumber="220">Sine loC.:</emphasis>
<location id="8EDA60206D4FFF8BFC53E50D25AFFC3B" LSID="urn:lsid:plazi:treatment:03AC87ED6D4FFF86FCBFE4B722C2FD7C:8EDA60206D4FFF8BFC53E50D25AFFC3B" box="[973,1447,873,897]" name="&quot; In tectis et muris vulgaris, vere maturans &quot;" pageId="21" pageNumber="220">“In tectis et muris vulgaris, vere maturans”</location>
, s.d.,
<emphasis id="B971EAE96D4FFF8BFCE6E5EE23D6FC1B" box="[888,990,906,929]" italics="true" pageId="21" pageNumber="220">Anon. s.n.</emphasis>
(
<collectionCode id="ED14AE3E6D4FFF8BFC72E5ED240AFC1B" box="[1004,1026,905,929]" pageId="21" pageNumber="220">G</collectionCode>
[
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]!).
</materialsCitation>
</paragraph>
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<paragraph id="8BBA36FB6D4FFF8BFCD9E5D12420FC56" blockId="21.[801,1453,722,1273]" pageId="21" pageNumber="220">
5
<treatmentCitation id="0AA410EA6D4FFF8BFCF3E5D02420FC56" author="Hedw." page="159" pageId="21" pageNumber="220" year="1801">
<taxonomicName id="4C054D786D4FFF8BFCF3E5D023BCFC56" authority="Hedw., Sp. Musc. Frond." authorityName="Hedw., Sp. Musc. Frond." class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="21" pageNumber="220" phylum="Bryophyta" rank="variety" species="pulvinatus" variety="africanus">
<emphasis id="B971EAE96D4FFF8BFCF3E5D0244EFC76" box="[877,1094,948,972]" italics="true" pageId="21" pageNumber="220">Fissidens pulvinatus</emphasis>
var.
<emphasis id="B971EAE96D4FFF8BFBE2E5D024D6FC76" box="[1148,1246,948,972]" italics="true" pageId="21" pageNumber="220">africanus</emphasis>
Hedw., Sp. Musc. Frond.
</taxonomicName>
159. 1801.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFBB0E5B025ACFBF7" pageId="21" pageNumber="220" type="materials_examined">
<paragraph id="8BBA36FB6D4FFF8BFBB0E5B025ACFBF7" blockId="21.[801,1453,722,1273]" pageId="21" pageNumber="220">
<materialsCitation id="3B6D3CA66D4FFF8BFBB0E5B025ACFBF7" ID-GBIF-Occurrence="3467920345" collectionCode="G" country="South Africa" location="&quot; Ad Cap. bonae spei lecta, specimina misit O. Swartz &quot;" pageId="21" pageNumber="220" specimenCode="G0004035" specimenCount="1" typeStatus="lectotype">
<emphasis id="B971EAE96D4FFF8BFBB0E5B024A4FC56" bold="true" box="[1070,1196,980,1004]" pageId="21" pageNumber="220">
<typeStatus id="54BE88596D4FFF8BFBB0E5B024A4FC56" box="[1070,1196,980,1004]" pageId="21" pageNumber="220" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated by
<bibRefCitation id="EF944B0A6D4FFF8BFACCE5B123EBFBB6" author="Munoz, J." firstAuthor="Munoz" journalOrPublisher="Ann. Missouri Bot. Gard" page="168" pageId="21" pageNumber="220" pagination="118 - 191" part="86" refId="ref21654" refString="Munoz, J. (1999). A revision of Grimmia (Musci, Grimmiaceae) in the Americas. 1: Latin America. Ann. Missouri Bot. Gard. 86: 118 - 191." title="A revision of Grimmia (Musci, Grimmiaceae) in the Americas. 1: Latin America" type="journal article" year="1999">Muñoz, 1999: 168</bibRefCitation>
):
<emphasis id="B971EAE96D4FFF8BFC61E59024B4FBB7" bold="true" box="[1023,1212,1012,1037]" pageId="21" pageNumber="220">
<collectingCountry id="F312766B6D4FFF8BFC61E59024B8FBB6" box="[1023,1200,1012,1036]" pageId="21" pageNumber="220">South AfriCA</collectingCountry>
:
</emphasis>
<location id="8EDA60206D4FFF8BFB5AE59024D9FB97" LSID="urn:lsid:plazi:treatment:03AC87ED6D4FFF86FCBFE4B722C2FD7C:8EDA60206D4FFF8BFB5AE59024D9FB97" country="South Africa" name="&quot; Ad Cap. bonae spei lecta, specimina misit O. Swartz &quot;" pageId="21" pageNumber="220">“Ad Cap. bonae spei lecta, specimina misit O. Swartz”</location>
, s.d.,
<emphasis id="B971EAE96D4FFF8BFA91E271258EFB96" box="[1295,1414,1045,1068]" italics="true" pageId="21" pageNumber="220">Swartz s.n.</emphasis>
(
<collectionCode id="ED14AE3E6D4FFF8BFA0BE27125A3FB97" box="[1429,1451,1045,1069]" pageId="21" pageNumber="220">G</collectionCode>
[
<specimenCode id="DBA39E806D4FFF8BFCEDE25123F9FBF7" box="[883,1009,1077,1101]" pageId="21" pageNumber="220">G0004035</specimenCode>
]!), synonymized by
<bibRefCitation id="EF944B0A6D4FFF8BFB5FE25125A9FBF7" author="Magill, R. E." box="[1217,1441,1077,1101]" editor="Leistner, O. A." firstAuthor="Magill" journalOrPublisher="Southern Africa" page="275" pageId="21" pageNumber="220" pagination="1" refId="ref21423" refString="Magill, R. E. (1981). Bryophyta. I (1). In: Leistner, O. A. (ed.), Fl. Southern Africa." title="Bryophyta. I (1)." type="book chapter" volumeTitle="Fl. Southern Africa." year="1981">Magill (1981: 275)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFCD9E20425A4FBC2" box="[839,1452,1120,1144]" pageId="21" pageNumber="220" type="reference_group">
<paragraph id="8BBA36FB6D4FFF8BFCD9E20425A4FBC2" blockId="21.[801,1453,722,1273]" box="[839,1452,1120,1144]" pageId="21" pageNumber="220">
5
<treatmentCitation id="0AA410EA6D4FFF8BFCF3E20425A4FBC2" author="Spreng." box="[877,1452,1120,1144]" page="321" pageId="21" pageNumber="220" year="1827">
<taxonomicName id="4C054D786D4FFF8BFCF3E2042503FBC2" ID-CoL="9S6FK" authority="Spreng., Syst. Veg." authorityName="Spreng., Syst. Veg." box="[877,1291,1120,1144]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="21" pageNumber="220" phylum="Bryophyta" rank="species" species="ecklonii">
<emphasis id="B971EAE96D4FFF8BFCF3E2042424FBC2" box="[877,1068,1120,1144]" italics="true" pageId="21" pageNumber="220">Grimmia ecklonii</emphasis>
Spreng., Syst. Veg.
</taxonomicName>
4: 321. 1827.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4FFF8BFCF3E2E4256AFB43" pageId="21" pageNumber="220" type="materials_examined">
<paragraph id="8BBA36FB6D4FFF8BFCF3E2E4256AFB43" blockId="21.[801,1453,722,1273]" pageId="21" pageNumber="220">
<materialsCitation id="3B6D3CA66D4FFF8BFCF3E2E4243CFB62" ID-GBIF-Occurrence="3467920350" collectionCode="BM" country="South Africa" location="Swellendam" municipality="Swellendam" pageId="21" pageNumber="220" specimenCode="BM000575940" specimenCount="1" typeStatus="lectotype">
<emphasis id="B971EAE96D4FFF8BFCF3E2E423E5FB22" bold="true" box="[877,1005,1152,1176]" pageId="21" pageNumber="220">
<typeStatus id="54BE88596D4FFF8BFCF3E2E423E5FB22" box="[877,1005,1152,1176]" pageId="21" pageNumber="220" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated by
<bibRefCitation id="EF944B0A6D4FFF8BFB06E2E42380FB02" author="Munoz, J. &amp; F. Pando" firstAuthor="Munoz" journalOrPublisher="Monogr. Syst. Bot. Missouri Bot. Gard" page="67" pageId="21" pageNumber="220" pagination="83" part="83" refId="ref21693" refString="Munoz, J. &amp; F. Pando (2000). A world synopsis of the genus Grimmia (Musci, Grimmiaceae). Monogr. Syst. Bot. Missouri Bot. Gard. 83." title="A world synopsis of the genus Grimmia (Musci, Grimmiaceae)" type="book" year="2000">Muñoz &amp; Pando, 2000: 67</bibRefCitation>
):
<emphasis id="B971EAE96D4FFF8BFC00E2C42444FB02" bold="true" box="[926,1100,1184,1208]" pageId="21" pageNumber="220">
<collectingCountry id="F312766B6D4FFF8BFC00E2C4244BFB02" box="[926,1091,1184,1208]" pageId="21" pageNumber="220">South AfriCA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BDEAC816D4FFF8BFBCFE2C424D8FB02" box="[1105,1232,1184,1208]" pageId="21" pageNumber="220">Swellendam</collectingMunicipality>
, s.d.,
<emphasis id="B971EAE96D4FFF8BFA96E2C4257AFB02" box="[1288,1394,1184,1208]" italics="true" pageId="21" pageNumber="220">Ecklon s.n.</emphasis>
(
<collectionCode id="ED14AE3E6D4FFF8BFAE1E2C425A3FB02" box="[1407,1451,1184,1208]" pageId="21" pageNumber="220">BM</collectionCode>
[
<specimenCode id="DBA39E806D4FFF8BFCEDE2A42428FB62" box="[883,1056,1216,1240]" pageId="21" pageNumber="220">BM000575940</specimenCode>
]!;
</materialsCitation>
<materialsCitation id="3B6D3CA66D4FFF8BFBA5E2A4256AFB43" ID-GBIF-Occurrence="3467920356" collectionCode="BM" country="South Africa" location="Swellendam" municipality="Swellendam" pageId="21" pageNumber="220" specimenCode="BM000670219" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54BE88596D4FFF8BFBA5E2A4249FFB62" box="[1083,1175,1216,1240]" pageId="21" pageNumber="220" type="isolectotype">isolecto-</typeStatus>
:
<collectionCode id="ED14AE3E6D4FFF8BFB38E2A524DAFB62" box="[1190,1234,1217,1240]" pageId="21" pageNumber="220">BM</collectionCode>
[
<specimenCode id="DBA39E806D4FFF8BFB41E2A42582FB62" box="[1247,1418,1216,1240]" pageId="21" pageNumber="220">BM000670219</specimenCode>
]!), synonymized by
<bibRefCitation id="EF944B0A6D4FFF8BFBBEE2852555FB43" author="Munoz, J. &amp; F. Pando" box="[1056,1373,1249,1273]" firstAuthor="Munoz" journalOrPublisher="Monogr. Syst. Bot. Missouri Bot. Gard" page="67" pageId="21" pageNumber="220" pagination="83" part="83" refId="ref21693" refString="Munoz, J. &amp; F. Pando (2000). A world synopsis of the genus Grimmia (Musci, Grimmiaceae). Monogr. Syst. Bot. Missouri Bot. Gard. 83." title="A world synopsis of the genus Grimmia (Musci, Grimmiaceae)" type="book" year="2000">Muñoz &amp; Pando (2000: 67)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4FFF89FCD9E34520D7F9CF" lastPageId="23" lastPageNumber="222" pageId="21" pageNumber="220" type="description">
<paragraph id="8BBA36FB6D4FFF89FCD9E34522AFFB9F" blockId="21.[801,1452,1313,1949]" lastBlockId="23.[113,764,282,1653]" lastPageId="23" lastPageNumber="222" pageId="21" pageNumber="220">
Gametophyte.
<emphasis id="B971EAE96D4FFF8BFC73E34624CAFA83" box="[1005,1218,1313,1337]" italics="true" pageId="21" pageNumber="220">Monoicous. Female:</emphasis>
innermost perichaetial leaf 2.2-2.5 mm long, sheathing up to apical part, tubulose, hyaline in lower part of leaf, costa excurrent to long, bluntly denticulate hair-point;
<emphasis id="B971EAE96D4FFF8BFB58E3E6250CFA23" box="[1222,1284,1410,1434]" italics="true" pageId="21" pageNumber="220">male:</emphasis>
perigonium as bud of 4 or 5 leaves, on short stalk in leaf axil on stem of fruiting plant, close to perichaetium, innermost perigonial leaf 1 mm long, sheathing up to apex, concave, ovate, apex acute or mucronate, costa vanishing below apex, hyaline except at apex, paraphyses few.
<emphasis id="B971EAE96D4FFF8BFA8BE04725AAF981" box="[1301,1442,1571,1595]" italics="true" pageId="21" pageNumber="220">Growth form</emphasis>
: cushion dense, lax, young shoots originating from detritus, leaflets with hair-point, plants radiculose at base, erect, dichotomously branched, stems up to 15 mm high, central strand developed.
<emphasis id="B971EAE96D4FFF8BFC70E0C0243FF901" box="[1006,1079,1700,1723]" italics="true" pageId="21" pageNumber="220">Leaves</emphasis>
in lower part of stem small, with hair-point, becoming gradually longer, up to 2.5 mm long, crowded, loosely disposed on stem, upper part of leaf slightly flexuose when dry, especially the comal ones, apical part quickly bending backwards when moistened, erecto-patent when wet, broadest part of leaf in lower third, broad-lanceolate or lanceolate from a short, ovate leaf base, apex obtuse, hair-point denticulate;
<emphasis id="B971EAE96D4FFF89FAD6E1E120B1FE88" italics="true" lastPageId="23" lastPageNumber="222" pageId="21" pageNumber="220">leaf form in situ</emphasis>
, leaf base widely concave or concave, laminal part widely keeled, margin recurved on one side from insertion or leaf base up to above mid-leaf, rarely on the other side and then, at most, in the middle of the leaf; in leaf base some paracostal cell rows of slightly elongate-rectangular cells, walls smooth, towards margin cells short-rectangular, at margin one or two rows of rectangular cells which may appear hyaline, transverse walls thickened, all cell walls smooth, in transitional part cells rectangular, walls sinuose, in laminal part mostly isodiametric, walls somewhat sinuose, thickened, at apex cells isodiametric, lumina rounded; lamina, seen in transverse section, unistratose, margin at insertion and leaf base unistratose, in laminal part, at least in the very apical region, several marginal cell rows are bistratose on both sides or on one side only, even completely unistratose margins may be seen.
<emphasis id="B971EAE96D4DFF89FDE1E56C22B4FCA5" box="[639,700,776,799]" italics="true" pageId="23" pageNumber="222">Costa</emphasis>
, seen on dorsal side, of nearly uniform width, faintly smaller in leaf base, percurrent, seen in transverse section, on dorsal side rounded, on ventral side at insertion and leaf base slightly concave, in laminal part above widest part of leaf widely channelled, at insertion and leaf base 4 guide cells, in laminal part reduced to two, at insertion and leaf base a small band of substereids or stereids and a median group of hydroids, gradually vanishing up to apical part.
</paragraph>
<caption id="DF7A66736D4CFF88FF16E13024B4F820" httpUri="https://zenodo.org/record/888354/files/figure.png" pageId="22" pageNumber="221" targetBox="[136,1388,293,1828]" targetPageId="22">
<paragraph id="8BBA36FB6D4CFF88FF16E1302284F83A" blockId="22.[136,1300,1876,1947]" pageId="22" pageNumber="221">
<emphasis id="B971EAE96D4CFF88FF16E13020DAF8D2" bold="true" box="[136,210,1876,1896]" pageId="22" pageNumber="221">Fig. 12. </emphasis>
<taxonomicName id="4C054D786D4CFF88FF49E131226BF8D2" authority="(Hedw.) Sm. A. Transverse" authorityName="(Hedw.) Sm. A. Transverse" box="[215,611,1876,1896]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="22" pageNumber="221" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4CFF88FF49E131217FF8DD" box="[215,375,1876,1896]" italics="true" pageId="22" pageNumber="221">Grimmia pulvinata</emphasis>
(Hedw.) Sm.
<emphasis id="B971EAE96D4CFF88FE74E13021F4F8DD" bold="true" box="[490,508,1876,1895]" pageId="22" pageNumber="221">A.</emphasis>
Transverse
</taxonomicName>
section of stem;
<emphasis id="B971EAE96D4CFF88FD60E1312320F8DD" bold="true" box="[766,808,1876,1896]" pageId="22" pageNumber="221">B, C.</emphasis>
Leaves;
<emphasis id="B971EAE96D4CFF88FCE9E13123A9F8D2" bold="true" box="[887,929,1877,1896]" pageId="22" pageNumber="221">D, E.</emphasis>
Outlines of transverse sections of leaves;
<emphasis id="B971EAE96D4CFF88FF16E10A20B9F83A" bold="true" box="[136,177,1901,1921]" pageId="22" pageNumber="221">F, G.</emphasis>
Cells in leaf base;
<emphasis id="B971EAE96D4CFF88FEC9E10A216DF83B" bold="true" box="[343,357,1902,1921]" pageId="22" pageNumber="221">H</emphasis>
,
<emphasis id="B971EAE96D4CFF88FEF1E10A2172F83B" bold="true" box="[367,378,1902,1921]" pageId="22" pageNumber="221">I.</emphasis>
Transverse sections of leaves.
</paragraph>
<paragraph id="8BBA36FB6D4CFF88FF16E1EC24B4F820" blockId="22.[136,1300,1876,1947]" box="[136,1212,1927,1947]" pageId="22" pageNumber="221">
[
<emphasis id="B971EAE96D4CFF88FF13E1E320C4F821" bold="true" box="[141,204,1927,1947]" pageId="22" pageNumber="221">A, D, H:</emphasis>
<emphasis id="B971EAE96D4CFF88FF4FE1E32179F820" box="[209,369,1927,1946]" italics="true" pageId="22" pageNumber="221">Hedderson 13601</emphasis>
, BOL;
<emphasis id="B971EAE96D4CFF88FE35E1E321DDF821" bold="true" box="[427,469,1927,1947]" pageId="22" pageNumber="221">C, F:</emphasis>
<emphasis id="B971EAE96D4CFF88FE44E1E32277F820" box="[474,639,1927,1946]" italics="true" pageId="22" pageNumber="221">Hedderson 13148,</emphasis>
BOL;
<emphasis id="B971EAE96D4CFF88FD2BE1EC22CAF821" bold="true" box="[693,706,1928,1947]" pageId="22" pageNumber="221">B</emphasis>
,
<emphasis id="B971EAE96D4CFF88FD55E1EC22DFF821" bold="true" box="[715,727,1928,1947]" pageId="22" pageNumber="221">E</emphasis>
,
<emphasis id="B971EAE96D4CFF88FD7FE1EC22E6F821" bold="true" box="[737,750,1928,1947]" pageId="22" pageNumber="221">I:</emphasis>
<emphasis id="B971EAE96D4CFF88FD6DE1E32390F820" box="[755,920,1927,1946]" italics="true" pageId="22" pageNumber="221">Hedderson 13851,</emphasis>
BOL;
<emphasis id="B971EAE96D4CFF88FC53E1E323EAF821" bold="true" box="[973,994,1927,1947]" pageId="22" pageNumber="221">G:</emphasis>
<emphasis id="B971EAE96D4CFF88FC76E1E32485F820" box="[1000,1165,1927,1946]" italics="true" pageId="22" pageNumber="221">Hedderson 13677,</emphasis>
BOL]
</paragraph>
</caption>
<paragraph id="8BBA36FB6D4DFF89FF09E24B20D7F9CF" blockId="23.[113,764,282,1653]" pageId="23" pageNumber="222">
Sporophyte.
<emphasis id="B971EAE96D4DFF89FEBEE24B2143FBFC" box="[288,331,1071,1094]" italics="true" pageId="23" pageNumber="222">Seta</emphasis>
up to 4.5 mm long, erect, twisted when old and dry, in immature and mature state arcuate or curved when wet, vaginula 0.8 mm long, cylindrical.
<emphasis id="B971EAE96D4DFF89FDF3E21422B7FB32" box="[621,703,1136,1160]" italics="true" pageId="23" pageNumber="222">Capsule</emphasis>
erect when old, in immature and mature state pendent and hidden in cushion, obloid, in mature state with eight to ten ribs, exothecial cells elongated, walls curvilinear, thin, stomata numerous in short apophysis, annulus of three or four rows of cell, detaching in a spiral, cells of capsule at orifice with crenulate outer walls.
<taxonomicName id="4C054D786D4DFF89FEBCE3522175FAF4" box="[290,381,1334,1358]" class="Insecta" family="Noctuidae" genus="Calyptra" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAE96D4DFF89FEBCE3522175FAF4" box="[290,381,1334,1358]" italics="true" pageId="23" pageNumber="222">Calyptra</emphasis>
</taxonomicName>
mitrate, lobed, covering upper part of capsule.
<emphasis id="B971EAE96D4DFF89FF72E3322155FAD4" box="[236,349,1366,1390]" italics="true" pageId="23" pageNumber="222">Operculum</emphasis>
with straight beak of variable length, margin crenulate, a marginal row of oval cells, cells in conical part mostly elongated of different shape.
<emphasis id="B971EAE96D4DFF89FD84E3FD2271FA0A" box="[538,633,1433,1456]" italics="true" pageId="23" pageNumber="222">Peristome</emphasis>
, teeth erectspreading when dry, broad at base, elongate-lanceolate, in upper half split into two or three divisions, separated down to insertion, dorsal side smooth near insertion, the upper part of dorsal side and ventral side densely covered with fine papillae, trabeculae thin, neared, slightly protruding.
<emphasis id="B971EAE96D4DFF89FDCEE0592298F9EE" box="[592,656,1597,1620]" italics="true" pageId="23" pageNumber="222">Spores</emphasis>
8-11 µm, granulose.
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4DFF89FF09E0C4210BF827" pageId="23" pageNumber="222" type="diagnosis">
<paragraph id="8BBA36FB6D4DFF89FF09E0C4210BF827" blockId="23.[113,764,1695,1949]" pageId="23" pageNumber="222">
<emphasis id="B971EAE96D4DFF89FF09E0C42181F90D" box="[151,393,1695,1719]" italics="true" pageId="23" pageNumber="222">Diagnostic characters. </emphasis>
Gametophyte. Leaves broadest in the lower third; margin recurved on one side from insertion or leaf base up to above mid-leaf, rarely so on the other side, and if so only in the middle of the leaf; leaf with one or two basal paracostal cell rows rectangular, towards margin cells short-rectangular, at margin one or two rows rectangular with thickened transverse walls which may appear hyaline, all cell walls smooth.
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4DFF89FCD9E77E2566FC0D" pageId="23" pageNumber="222" type="distribution">
<paragraph id="8BBA36FB6D4DFF89FCD9E77E24CEFD6F" blockId="23.[801,1452,282,951]" pageId="23" pageNumber="222">
<emphasis id="B971EAE96D4DFF89FCD9E77E2589FE88" box="[839,1409,282,306]" italics="true" pageId="23" pageNumber="222">
Distribution, habitat and ecology.
<taxonomicName id="4C054D786D4DFF89FB29E77F2589FE88" box="[1207,1409,282,306]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">Grimmia pulvinata</taxonomicName>
</emphasis>
is a very widespread species that occurs on every continent except Antarctica. It is often common in temperate areas of the northern hemisphere, with some extensions northward into boreal zones, and (rarely) southward to subtropical areas. In the southern hemisphere this species is recorded from Australia, New Zealand, southern South America (Chile and Uruguay), and from southern Africa. Although frequently considered cosmopolitan, it is actually largely absent from the tropics and subtropics. In sub-Saharan Africa, for example, it appears to be largely restricted to the study area and adjacent Namibia, and is not recorded for any other country on the continent except Ethiopia. It is thus probably better considered a temperate species with some poleward extensions.
</paragraph>
<paragraph id="8BBA36FB6D4DFF89FCD9E4BA2566FC0D" blockId="23.[801,1452,282,951]" pageId="23" pageNumber="222">
In South Africa and Lesotho (
<figureCitation id="133E2A7E6D4DFF89FB1AE4B924C9FD4F" box="[1156,1217,733,757]" captionStart="Fig. 2" captionStartId="3.[113,140,1889,1908]" captionTargetPageId="3" captionText="Fig. 2. Geographic distribution of Grimmia Hedw. species in South Africa and Lesotho. A. Grimmia longirostris Hook.; B. Grimmia montana Bruch &amp; Schimp.; C. Grimmia orbicularis Wilson; D. Grimmia pulvinata (Hedw.) Sm.; E. Grimmia pygmaea Müll. Hal.; F. Grimmia sessitana De Not. (circles) and G. tortuosa Hook. f. &amp; Wilson (star)." httpUri="https://zenodo.org/record/888401/files/figure.png" pageId="23" pageNumber="222">Fig. 2</figureCitation>
D),
<taxonomicName id="4C054D786D4DFF89FB77E4BA2561FD4F" box="[1257,1385,733,757]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FB77E4BA2561FD4F" box="[1257,1385,733,757]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
</taxonomicName>
is one of the two most common species of
<taxonomicName id="4C054D786D4DFF89FB3DE49A250AFCAF" box="[1187,1282,766,789]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="genus">
<emphasis id="B971EAE96D4DFF89FB3DE49A250AFCAF" box="[1187,1282,766,789]" italics="true" pageId="23" pageNumber="222">Grimmia</emphasis>
</taxonomicName>
encountered. It occurs over a broad altitudinal range (50 to 1,910 m), and in most of the major biomes, except for the various savannah and tropical thicket and forest types. Most collections are from quartzitic sandstones, but it also occurs on most other rock types (e.g. shales, granites, basalt) present in the region.
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D4DFF86FCD9E584231DFE0F" lastPageId="24" lastPageNumber="223" pageId="23" pageNumber="222" type="discussion">
<paragraph id="8BBA36FB6D4DFF89FCD9E58423EAFB02" blockId="23.[801,1455,991,1949]" pageId="23" pageNumber="222">
<emphasis id="B971EAE96D4DFF89FCD9E5842392FC4D" box="[839,922,992,1015]" italics="true" pageId="23" pageNumber="222">Notes. </emphasis>
A total of 167 specimens were seen of which 67 had sporophytes, although only 6 had capsules that were in a suitable state for examination whilst 9 were immature, and the remaining 52 were decomposed. The leaf shape in
<taxonomicName id="4C054D786D4DFF89FAD4E2242385FBC2" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FAD4E2242385FBC2" italics="true" pageId="23" pageNumber="222">Grimmia pulvinata</emphasis>
</taxonomicName>
is variable. Leaves can vary from short ovatelanceolate to elongated ovate-lanceolate, and small lanceolate forms also appear.
</paragraph>
<paragraph id="8BBA36FB6D4DFF89FCD9E2A423D9FA20" blockId="23.[801,1455,991,1949]" pageId="23" pageNumber="222">
The hair-points can be longer than the lamina, whilst at the other extreme some plants have leaves with only a few hyaline cells at the apex or even lack the hair-point altogether. A specimen from the summit of the Zuurberge Range (
<emphasis id="B971EAE96D4DFF89FAEDE345238CFAE3" italics="true" pageId="23" pageNumber="222">Alexander s.n.</emphasis>
, BOL) where all leaves are muticous was annotated by Dixon as “
<taxonomicName id="4C054D786D4DFF89FC26E30624D7FAC0" box="[952,1247,1378,1402]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="variety" species="pulvinata" variety="submutica">
<emphasis id="B971EAE96D4DFF89FC26E3062449FAC0" box="[952,1089,1378,1402]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
var.
<emphasis id="B971EAE96D4DFF89FBE8E30624D7FAC0" box="[1142,1247,1378,1402]" italics="true" pageId="23" pageNumber="222">submutica</emphasis>
</taxonomicName>
”, a name that was never published.
</paragraph>
<paragraph id="8BBA36FB6D4DFF89FCD9E3C625A7F946" blockId="23.[801,1455,991,1949]" pageId="23" pageNumber="222">
The stratosity of the leaf margins is variable. In different leaves, even on the same plant, unistratose or bistratose margins may be observed. For the correct identification of
<taxonomicName id="4C054D786D4DFF89FAB8E38725ADFA40" box="[1318,1445,1506,1530]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FAB8E38725ADFA40" box="[1318,1445,1506,1530]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
</taxonomicName>
, and for its differentiation from
<taxonomicName id="4C054D786D4DFF89FBEEE06724F0F9A1" box="[1136,1272,1539,1563]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="orbicularis">
<emphasis id="B971EAE96D4DFF89FBEEE06724F0F9A1" box="[1136,1272,1539,1563]" italics="true" pageId="23" pageNumber="222">G. orbicularis</emphasis>
</taxonomicName>
, the stratosity of the leaf margins is thus not very useful. A stable distinguishing character is provided by the paracostal cell rows in the leaf base. In
<taxonomicName id="4C054D786D4DFF89FCBFE0002395F9C1" box="[801,925,1635,1659]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FCBFE0002395F9C1" box="[801,925,1635,1659]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
</taxonomicName>
these (
<figureCitation id="133E2A7E6D4DFF89FC7AE0072421F9C1" box="[996,1065,1635,1659]" captionStart="Fig. 12" captionStartId="22.[136,163,1876,1895]" captionTargetBox="[136,1388,293,1828]" captionTargetId="figure@22.[136,1399,276,1832]" captionTargetPageId="22" captionText="Fig. 12. Grimmia pulvinata (Hedw.) Sm. A. Transverse section of stem; B, C. Leaves; D, E. Outlines of transverse sections of leaves; F, G. Cells in leaf base; H, I. Transverse sections of leaves. [A, D, H: Hedderson 13601, BOL; C, F: Hedderson 13148, BOL; B, E, I: Hedderson 13851, BOL; G: Hedderson 13677, BOL]" httpUri="https://zenodo.org/record/888354/files/figure.png" pageId="23" pageNumber="222">Fig. 12</figureCitation>
F, G) are slightly elongate-rectangular, becoming short-rectangular towards the margins and all cell walls are smooth. In
<taxonomicName id="4C054D786D4DFF89FC28E0C0243AF906" box="[950,1074,1700,1724]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="orbicularis">
<emphasis id="B971EAE96D4DFF89FC28E0C0243AF906" box="[950,1074,1700,1724]" italics="true" pageId="23" pageNumber="222">G. orbicularis</emphasis>
</taxonomicName>
on the other hand (
<figureCitation id="133E2A7E6D4DFF89FB68E0C02533F906" box="[1270,1339,1700,1724]" captionStart="Fig" captionStartId="20.[136,163,1876,1895]" captionTargetBox="[220,1393,862,1852]" captionTargetId="figure@20.[208,1403,867,1836]" captionTargetPageId="20" captionText="Fig. 11. Grimmia orbicularis Wilson. A. Transverse section of stem; B, C. Leaves; D. Outlines of transverse section of leaf; E, G. Cells in leaf base; F. Cells in transitional part; H. Transverse section of leaf. [A: Hedderson 15285, BOL; B, E: MacOwen s.n., BM; C, F-H: Hedderson 15264, BOL; D, Hedderson 13047, BOL]" httpUri="https://zenodo.org/record/888352/files/figure.png" pageId="23" pageNumber="222">Fig. 11</figureCitation>
E, F), these are elongate-rectangular, with nodulose walls towards the costa, but rectangular to quadrate with smooth walls at the margins.
</paragraph>
<paragraph id="8BBA36FB6D4DFF89FCD9E16023A3F827" blockId="23.[801,1455,991,1949]" pageId="23" pageNumber="222">
Other species have been subject to frequent misidentification as
<taxonomicName id="4C054D786D4DFF89FCECE14123F0F887" box="[882,1016,1829,1853]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FCECE14123F0F887" box="[882,1016,1829,1853]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
</taxonomicName>
, and amongst the material at BOL and PRE specimens of
<taxonomicName id="4C054D786D4DFF89FC76E1212466F8E7" box="[1000,1134,1861,1885]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="consobrina">
<emphasis id="B971EAE96D4DFF89FC76E1212466F8E7" box="[1000,1134,1861,1885]" italics="true" pageId="23" pageNumber="222">G. consobrina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D786D4DFF89FBE7E12124FDF8E7" box="[1145,1269,1861,1885]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="fuscolutea">
<emphasis id="B971EAE96D4DFF89FBE7E12124FDF8E7" box="[1145,1269,1861,1885]" italics="true" pageId="23" pageNumber="222">G. fuscolutea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D786D4DFF89FA9EE1212571F8E7" box="[1280,1401,1861,1885]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="laevigata">
<emphasis id="B971EAE96D4DFF89FA9EE1212571F8E7" box="[1280,1401,1861,1885]" italics="true" pageId="23" pageNumber="222">G. laevigata</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C054D786D4DFF89FCBFE101239FF8C7" box="[801,919,1893,1917]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pygmaea">
<emphasis id="B971EAE96D4DFF89FCBFE101239FF8C7" box="[801,919,1893,1917]" italics="true" pageId="23" pageNumber="222">G. pygmaea</emphasis>
</taxonomicName>
were found to have been erroneously attributed to
<taxonomicName id="4C054D786D4DFF89FCBFE1E223ACF827" box="[801,932,1925,1949]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="23" pageNumber="222" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D4DFF89FCBFE1E223ACF827" box="[801,932,1925,1949]" italics="true" pageId="23" pageNumber="222">G. pulvinata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBA36FB6D42FF86FF30E77F231DFE0F" blockId="24.[136,789,282,437]" pageId="24" pageNumber="223">
<taxonomicName id="4C054D786D42FF86FF30E77F217EFE88" box="[174,374,282,306]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="24" pageNumber="223" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D42FF86FF30E77F217EFE88" box="[174,374,282,306]" italics="true" pageId="24" pageNumber="223">Grimmia pulvinata</emphasis>
</taxonomicName>
grows from 250 to 2,000 m in Europe and North America (records from above 2,000 m should be checked), from 1,000 to 1,800 m in Asia, and from 50 to 1,910 m in South Africa.
<taxonomicName id="4C054D786D42FF86FE09E7182255FE2E" box="[407,605,380,404]" class="Bryopsida" family="Grimmiaceae" genus="Grimmia" kingdom="Plantae" order="Grimmiales" pageId="24" pageNumber="223" phylum="Bryophyta" rank="species" species="pulvinata">
<emphasis id="B971EAE96D42FF86FE09E7182255FE2E" box="[407,605,380,404]" italics="true" pageId="24" pageNumber="223">Grimmia pulvinata</emphasis>
</taxonomicName>
thus seems to be restricted to altitudes from sea level to 2,000 m across its range.
</paragraph>
</subSubSection>
<subSubSection id="C31F65706D42FF86FF30E7BE22C2FD7C" pageId="24" pageNumber="223" type="materials_examined">
<paragraph id="8BBA36FB6D42FF86FF30E7BE22C2FD7C" blockId="24.[136,787,474,711]" pageId="24" pageNumber="223">
<emphasis id="B971EAE96D42FF86FF30E7BE2192FE57" box="[174,410,474,493]" italics="true" pageId="24" pageNumber="223">Selected specimens examined. </emphasis>
<emphasis id="B971EAE96D42FF86FE01E7BE22D2FE57" bold="true" box="[415,730,474,494]" pageId="24" pageNumber="223">South AfriCA. Prov. Western Cape</emphasis>
: Cape Peninsula, Silvermine NR, Silvermine Valley, just above “Sunbird Environmental Recreation Centre”, 34°0628”S 18°2437”E, c. 50 m, 6.IV.2000,
<emphasis id="B971EAE96D42FF86FD20E47520B3FD85" italics="true" pageId="24" pageNumber="223">Hedderson 13148</emphasis>
(BOL); Ceres District, Oliphants River Mountains. Beaverlac area, Ratels River, c. 400 m, 32°5427”S 19°0401”E, 8.I.2001,
<emphasis id="B971EAE96D42FF86FDA6E42322C1FDE0" box="[568,713,583,602]" italics="true" pageId="24" pageNumber="223">Hedderson 13601</emphasis>
(BOL); Citrusdal Region, Cederberg, Cederberg State Forest, Welbedacht Kloof, 900- 1300 m, 32°2430”S 19°1024”E, 17.II.2001,
<emphasis id="B971EAE96D42FF86FE62E4192280FD2A" box="[508,648,637,656]" italics="true" pageId="24" pageNumber="223">Hedderson 13677</emphasis>
(BOL); c. 4 km along R355 from intersection with R46 between Ceres and Touwsrivier Steep, c. 850 m, 33°1110”S 19°4705”E, 4.VI.2001,
<emphasis id="B971EAE96D42FF86FE64E4D7228DFD7C" box="[506,645,691,710]" italics="true" pageId="24" pageNumber="223">Hedderson 13851</emphasis>
(BOL).
</paragraph>
</subSubSection>
</treatment>
</document>