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635 lines
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<document id="EB21A6CC0E831358C7763E6F2C28FB33" ID-CLB-Dataset="4210" ID-DOI="10.5852/ejt.2021.767.1495" ID-GBIF-Dataset="67bc6a73-4137-4de3-9dc4-ce87972f742d" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514839" ID-ZooBank="9F019E02-95E7-4B6A-B5BC-01F8E26F3466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927261920" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn" docDate="2021" docId="03AC87E7C329FFC49F70F9BF90D8BF13" docLanguage="en" docName="ejt-767_142-166.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1495" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria circinata Nahok & Tumpeesuwan & Tumpeesuwan 2021, sp. nov." docType="treatment" docUuid="B434E62B-9FA6-43FB-B27F-A4B1B0413225" docUuidSource="ZooBank" docVersion="11" lastPageNumber="153" masterDocId="FF95FF9FC32EFFCF9D1FFF9B931BBE42" masterDocTitle="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)" masterLastPageNumber="166" masterPageNumber="142" pageNumber="149" updateTime="1699449512379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="400197AE60E7AA7B7B99863B88B2571B">Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)</mods:title>
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<mods:namePart id="6472EB30C873A2C464B416C168ECB14A">Nahok, Benchawan</mods:namePart>
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<mods:affiliation id="FEC18B523F5307A990FCAF801546C029">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Educational Research Development and Demonstration Institute, Srinakharinwirot University, Ongkharak District, Nakhon Nayok, 26120 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="85751EDF75DB1F1E2C6AA59EF579EDB9" type="email">nahok.b@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AE14683781B9E22C75E6F54A9B2E2D25">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="B7F8AE7B29A61DC4CF6FBC9FA1AE285B">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="CF04415E5BBB26955D478E4658D67C24" type="email">stumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:affiliation id="63424C8EC78678BA7079482261DC90E4">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="618A7F75999ED301BE137AB4FEE540C8" type="email">ctumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:title id="C04117075DA2C97BF6D17DCBC98BAADD">European Journal of Taxonomy</mods:title>
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<heading id="D0F2819DC329FFC89F70F9BF90DEB87D" box="[623,965,1572,1599]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="149" reason="2">
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<taxonomicName id="4C054D72C329FFC89F70F9BF9078B87C" ID-CoL="8QGBF" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[623,867,1572,1599]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
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<emphasis id="B971EAE3C329FFC89F70F9BF9078B87C" bold="true" box="[623,867,1572,1599]" italics="true" pageId="7" pageNumber="149">Landouria circinata</emphasis>
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<taxonomicNameLabel id="A2425798C329FFC89E75F9BE90DEB87D" box="[874,965,1573,1599]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BBA36F1C329FFC89F58F9F090F7B8C7" blockId="7.[369,1219,1572,1670]" box="[583,1004,1643,1670]" pageId="7" pageNumber="149">
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<figureCitation id="133E2A74C329FFC89F58F9F091BDB8C7" box="[583,678,1643,1669]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="7" pageNumber="149">Figs 3A</figureCitation>
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,
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<figureCitation id="133E2A74C329FFC89FACF9F091C1B8C7" box="[691,730,1643,1669]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="7" pageNumber="149">4A</figureCitation>
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,
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<figureCitation id="133E2A74C329FFC89FF9F9F09017B8C4" box="[742,780,1643,1670]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="7" pageNumber="149">5A</figureCitation>
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,
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<figureCitation id="133E2A74C329FFC89E07F9F09079B8C7" box="[792,866,1643,1669]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="7" pageNumber="149">6A–C</figureCitation>
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;
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<tableCitation id="C687034AC329FFC89E72F9F090F7B8C7" box="[877,1004,1643,1669]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="2.[185,255,857,885]" captionStartId-1="4.[186,256,600,628]" captionStartId-2="6.[185,255,695,723]" captionStartId-3="10.[185,255,544,572]" captionTargetBox-0="[195,1921,987,1444]" captionTargetBox-1="[197,1907,827,1685]" captionTargetBox-2="[212,1909,868,1619]" captionTargetBox-3="[196,1908,758,1739]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="10" captionText-0="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." captionText-1="Table 2. Locality data and identification of specimens of Landouria Godwin-Austen, 1918 used for mtDNA sequencing, including collection catalogue numbers and GenBank accession numbers. Abbreviations: NHMSU = Natural History Museum, Mahasarakham University; HNHM = Hungarian Natural History Museum; AM = Australian Museum. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Hirano et al. (2014); 3 = Köhler et al. (2018)." captionText-2="Table 3. Pairwise p-distance values of Landouria Godwin-Austen, 1918 based on mitochondrial COI sequences." captionText-3="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri-0="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" httpUri-1="http://table.plazi.org/id/DF7A6679C32AFFCB9DA5FDC39249BC9F" httpUri-2="http://table.plazi.org/id/DF7A6679C328FFC99DA6FD2C96FFBC91" httpUri-3="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="7" pageNumber="149" tableUuid-0="DF7A6679C32CFFCD9DA6FCC295B0BD37" tableUuid-1="DF7A6679C32AFFCB9DA5FDC39249BC9F" tableUuid-2="DF7A6679C328FFC99DA6FD2C96FFBC91" tableUuid-3="DF7A6679C324FFC59DA6FDBB969CBCE7">Tables 1–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F657AC329FFC89DA2F937927CB916" pageId="7" pageNumber="149" type="reference_group">
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<paragraph id="8BBA36F1C329FFC89DA2F937923EB8A8" blockId="7.[189,1398,1708,1876]" pageId="7" pageNumber="149">
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<treatmentCitationGroup id="AB1511DFC329FFC89DA2F937966DB884" box="[189,1398,1708,1735]" pageId="7" pageNumber="149">
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<taxonomicName id="4C054D72C329FFC89DA2F9379278B885" box="[189,355,1708,1735]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B971EAE3C329FFC89DA2F9379222B884" box="[189,313,1708,1734]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
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sp.
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</taxonomicName>
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–
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<treatmentCitation id="0AA410E0C329FFC89C9CF93791CEB885" author="Tanmuangpak K. & Tumpeesuwan C. & Tumpeesuwan S." box="[387,725,1708,1735]" page="7" pageId="7" pageNumber="149" year="2012">
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<bibRefCitation id="EF944B00C329FFC89C9CF93791CEB885" author="Tanmuangpak K. & Tumpeesuwan C. & Tumpeesuwan S." box="[387,725,1708,1735]" pageId="7" pageNumber="149" pagination="15 - 26" refId="ref11296" refString="Tanmuangpak K., Tumpeesuwan C. & Tumpeesuwan S. 2012. Diversity of land snails in Phu Pha Lom Forest Park at Muang District, Loei Province. Journal of Science and Technology Mahasarakham University Special Issue 8: 15 - 26. [In Thai with English abstract.]" type="journal article" year="2012">
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Tanmuangpak
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<emphasis id="B971EAE3C329FFC89F2CF936916BB884" box="[563,624,1708,1734]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
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2012: 7
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</bibRefCitation>
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</treatmentCitation>
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||
, 12, fig. 5y. —
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<treatmentCitation id="0AA410E0C329FFC89E8EF93797EFB885" author="Tanmuangpak K." box="[913,1268,1708,1735]" page="125" pageId="7" pageNumber="149" year="2016">
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<bibRefCitation id="EF944B00C329FFC89E8EF93797EFB885" author="Tanmuangpak K." box="[913,1268,1708,1735]" pageId="7" pageNumber="149" refId="ref11254" refString="Tanmuangpak K. 2016. Species Diversity, Abundance, Distribution and Habitat Relationship of Land Snails in Phu Pha Lom Limestone Area, Loei Province. Unpublished PhD thesis, Department of Biology, Faculty of Science, Mahasarakham University." type="book" year="2016">Tanmuangpak 2016: 125–126</bibRefCitation>
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</treatmentCitation>
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, figs 4.74–
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||
</treatmentCitationGroup>
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4.75.
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</paragraph>
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<paragraph id="8BBA36F1C329FFC89DA2F96890C7B94F" blockId="7.[189,1398,1708,1876]" box="[189,988,1778,1805]" pageId="7" pageNumber="149">
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<taxonomicName id="4C054D72C329FFC89DA2F968926BB94F" box="[189,368,1779,1805]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="undefined-4">
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<emphasis id="B971EAE3C329FFC89DA2F9689222B94F" box="[189,313,1779,1805]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
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||
sp.4
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EF944B00C329FFC89C91F96891A1B94F" author="Ounchareon J." box="[398,698,1778,1805]" pageId="7" pageNumber="149" refId="ref10673" refString="Ounchareon J. 2015. Taxonomy of Land Snail Genus Landouria Godwin-Austen, 1918 in Northeastern Thailand. Unpublished Master thesis. Department Faculty of Science, Mahasakham University, Maha Sarakham. [In Thai with English abstract.]" type="book" year="2015">Ounchareon 2015: 40–42</bibRefCitation>
|
||
, figs 4.12, 4.13, 5.1, 5.7.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C329FFC89DA2F88D927CB916" blockId="7.[189,1398,1708,1876]" pageId="7" pageNumber="149">
|
||
<taxonomicName id="4C054D72C329FFC89DA2F88D926AB972" box="[189,369,1814,1840]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="undefined-2">
|
||
<emphasis id="B971EAE3C329FFC89DA2F88D9222B972" box="[189,313,1814,1840]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
|
||
sp.2
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EF944B00C329FFC89C92F88D91C4B972" author="Yingkhamhang A." box="[397,735,1814,1840]" pageId="7" pageNumber="149" refId="ref11408" refString="Yingkhamhang A. 2016. Shell Morphology and Reproductive Anatomy of Land Snails Genus Landouria Godwin-Austen, 1918 in Northeastern Thailand. Unpublished Bachelor's Degree Science Project. Department Faculty of Science, Mahasakham University, Maha Sarakham. [In Thai with English abstract.]" type="book" year="2016">Yingkhamhang 2016: 21–22</bibRefCitation>
|
||
, 39–41, 43–45, 47–48, figs 4.3–4.4, 4.21b, 4.22c, 4.23b, 5.1b, 5.2b.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC326FFC79DA2FE93924EBFD5" pageId="8" pageNumber="150" type="diagnosis">
|
||
<paragraph id="8BBA36F1C326FFC79DA2FE93922DBF60" blockId="8.[189,310,264,290]" box="[189,310,264,290]" pageId="8" pageNumber="150">
|
||
<heading id="D0F2819DC326FFC79DA2FE93922DBF60" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="8" pageNumber="150" reason="3">
|
||
<emphasis id="B971EAE3C326FFC79DA2FE93922DBF60" bold="true" box="[189,310,264,290]" pageId="8" pageNumber="150">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2FEAC924EBFD5" blockId="8.[189,1399,311,407]" pageId="8" pageNumber="150">Shell brownish-corneous, periphery angular, sharply keeled. Flagellum circinate; penis cylindrical and longer than vagina; base of gametolytic sac thick and stout. Radula with unicuspid short tongue-shaped central teeth.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC326FFC79DA2FE249274BC69" pageId="8" pageNumber="150" type="etymology">
|
||
<paragraph id="8BBA36F1C326FFC79DA2FE24925FBF9B" blockId="8.[189,324,447,473]" box="[189,324,447,473]" pageId="8" pageNumber="150">
|
||
<heading id="D0F2819DC326FFC79DA2FE24925FBF9B" bold="true" box="[189,324,447,473]" fontSize="11" level="3" pageId="8" pageNumber="150" reason="3">
|
||
<emphasis id="B971EAE3C326FFC79DA2FE24925FBF9B" bold="true" box="[189,324,447,473]" pageId="8" pageNumber="150">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2FE769274BC69" blockId="8.[189,1399,493,555]" pageId="8" pageNumber="150">
|
||
The Latin word ‘
|
||
<taxonomicName id="4C054D72C326FFC79C61FE7592FCBC45" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[382,487,494,519]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="circinata">
|
||
<emphasis id="B971EAE3C326FFC79C61FE7592FCBC45" box="[382,487,494,519]" italics="true" pageId="8" pageNumber="150">circinata</emphasis>
|
||
</taxonomicName>
|
||
’ refers to the coiled or curled flagellum which resembles the circinate vernation of a fern shoot.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC326FFC79DA2FDC9915CBDCE" pageId="8" pageNumber="150" type="materials_examined">
|
||
<paragraph id="8BBA36F1C326FFC79DA2FDC99277BC2E" blockId="8.[189,364,594,620]" box="[189,364,594,620]" pageId="8" pageNumber="150">
|
||
<heading id="D0F2819DC326FFC79DA2FDC99277BC2E" bold="true" box="[189,364,594,620]" fontSize="11" level="3" pageId="8" pageNumber="150" reason="3">
|
||
<emphasis id="B971EAE3C326FFC79DA2FDC99277BC2E" bold="true" box="[189,364,594,620]" pageId="8" pageNumber="150">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DF5FD1A970DBD46" blockId="8.[189,1399,641,773]" pageId="8" pageNumber="150">
|
||
<materialsCitation id="3B6D3CACC326FFC79DF5FD1A970DBD46" ID-GBIF-Occurrence="3357769310" collectingDate="2017-06-07" collectionCode="UW" collectorName="B. Nahok & U. Chanlabut" country="Thailand" county="Mueang District" elevation="415" latitude="17.555008" location="Phu Pha Lom Forest Park" longLatPrecision="1" longitude="101.86853" municipality="Phu Pha Lom Forest Park" pageId="8" pageNumber="150" specimenCode="NHMSU-00023" specimenCount="1" stateProvince="Loei Province" typeStatus="holotype">
|
||
<emphasis id="B971EAE3C326FFC79DF5FD1A9247BCD9" bold="true" box="[234,348,641,667]" pageId="8" pageNumber="150">
|
||
<typeStatus id="54BE8853C326FFC79DF5FD1A9247BCD9" box="[234,348,641,667]" pageId="8" pageNumber="150" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C326FFC79DA2FD3F924EBCFC" box="[189,341,676,702]" name="Thailand" pageId="8" pageNumber="150">THAILAND</collectingCountry>
|
||
• shell (SH =
|
||
<quantity id="4CFD9B14C326FFC79CE7FD3F914EBCFC" box="[504,597,676,702]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="150" unit="mm" value="8.2">8.2 mm</quantity>
|
||
, SW =
|
||
<quantity id="4CFD9B14C326FFC79FB5FD3F900EBCFC" box="[682,789,676,702]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="8" pageNumber="150" unit="mm" value="13.8">13.8 mm</quantity>
|
||
, AH =
|
||
<quantity id="4CFD9B14C326FFC79E78FD3F90D8BCFC" box="[871,963,676,703]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="150" unit="mm" value="5.1">5.1 mm</quantity>
|
||
, AW =
|
||
<quantity id="4CFD9B14C326FFC79906FD3F976EBCFC" box="[1049,1141,676,703]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="150" unit="mm" value="5.1">5.1 mm</quantity>
|
||
, UW =
|
||
<quantity id="4CFD9B14C326FFC799D0FD3F9636BCFC" box="[1231,1325,676,702]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="150" unit="mm" value="3.8">3.8 mm</quantity>
|
||
);
|
||
<collectingRegion id="49C1F813C326FFC7985EFD3F9233BCA3" country="Thailand" name="Loei" pageId="8" pageNumber="150">Loei Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62DB4E7DC326FFC79C28FD5392E6BCA3" box="[311,509,711,738]" pageId="8" pageNumber="150">Mueang District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BDEAC8BC326FFC79F13FD5C9050BCA3" box="[524,843,711,738]" pageId="8" pageNumber="150">Phu Pha Lom Forest Park</collectingMunicipality>
|
||
;
|
||
<geoCoordinate id="EE315036C326FFC79E45FD5E9703BCA0" box="[858,1048,709,738]" degrees="17" direction="north" minutes="33" orientation="latitude" pageId="8" pageNumber="150" precision="1" seconds="18.03" value="17.555008">17°33′18.03″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE315036C326FFC79938FD5E97F9BCA0" box="[1063,1250,709,738]" degrees="101" direction="east" minutes="52" orientation="longitude" pageId="8" pageNumber="150" precision="1" seconds="6.71" value="101.86853">101°52′6.71″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CFD9B14C326FFC7983EFD5C966ABCA0" box="[1313,1393,711,738]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="8" pageNumber="150" unit="m" value="415.0">
|
||
<elevation id="0028D1C2C326FFC7983EFD5C966ABCA0" box="[1313,1393,711,738]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="8" pageNumber="150" unit="m" value="415.0">415 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFBB1031C326FFC79DA2FD709253BD46" box="[189,328,746,773]" pageId="8" pageNumber="150" value="2017-06-07">
|
||
<collectingDate id="EFFFE9D9C326FFC79DA2FD709253BD46" box="[189,328,746,773]" pageId="8" pageNumber="150" value="2017-06-07">7 Jun. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F05327C326FFC79C4AFD7092DEBD46" box="[341,453,746,773]" pageId="8" pageNumber="150">B. Nahok</collectorName>
|
||
and
|
||
<collectorName id="26F05327C326FFC79CE1FD709180BD46" box="[510,667,746,773]" pageId="8" pageNumber="150">U. Chanlabut</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBA39E8AC326FFC79FC3FD719086BD46" box="[732,925,746,772]" pageId="8" pageNumber="150">NHMSU-00023</specimenCode>
|
||
. (
|
||
<figureCitation id="133E2A74C326FFC79EAEFD719716BD46" box="[945,1037,746,772]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="8" pageNumber="150">Fig. 3A</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DF5FCB7915CBDCE" blockId="8.[189,1398,812,909]" pageId="8" pageNumber="150">
|
||
<materialsCitation id="3B6D3CACC326FFC79DF5FCB79158BDCE" ID-GBIF-Occurrence="3357769303" accessionNumber="MN449401 and MZ435746" collectingDate="2017-06-07" collectionCode="MN, MZ" collectorName="GenBank No." country="Thailand" county="Mueang District" elevation="415" latitude="17.555008" location="THAILAND" longLatPrecision="1" longitude="101.86853" municipality="Phu Pha Lom Forest Park" pageId="8" pageNumber="150" specimenCode="NHMSU-00024" specimenCount="54" stateProvince="Loei" typeStatus="paratype">
|
||
<emphasis id="B971EAE3C326FFC79DF5FCB7927CBD04" bold="true" box="[234,359,812,838]" pageId="8" pageNumber="150">
|
||
<typeStatus id="54BE8853C326FFC79DF5FCB7927CBD04" box="[234,359,812,838]" pageId="8" pageNumber="150" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C326FFC79DA2FCD4924EBD2B" box="[189,341,847,873]" name="Thailand" pageId="8" pageNumber="150">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C326FFC79C74FCD492C8BD2B" box="[363,467,847,873]" pageId="8" pageNumber="150">11 shells</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD78C326FFC79CC1FCD4915EBD2B" box="[478,581,847,873]" pageId="8" pageNumber="150">43 living</specimenCount>
|
||
specs; same collection data as for holotype; GenBank No.
|
||
<accessionNumber id="9456AB12C326FFC799F0FCD4966DBD2B" box="[1263,1398,847,873]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN449401" pageId="8" pageNumber="150">MN449401</accessionNumber>
|
||
and
|
||
<accessionNumber id="9456AB12C326FFC79DF0FCE9926CBDCF" box="[239,375,882,909]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MZ435746" pageId="8" pageNumber="150" type="GenBank">MZ435746</accessionNumber>
|
||
;
|
||
<specimenCode id="DBA39E8AC326FFC79C9DFCE99158BDCE" box="[386,579,882,908]" pageId="8" pageNumber="150">NHMSU-00024</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC326FFC79DA2FC289728B82F" pageId="8" pageNumber="150" type="description">
|
||
<paragraph id="8BBA36F1C326FFC79DA2FC289255BD8F" blockId="8.[189,334,947,973]" box="[189,334,947,973]" pageId="8" pageNumber="150">
|
||
<heading id="D0F2819DC326FFC79DA2FC289255BD8F" bold="true" box="[189,334,947,973]" fontSize="11" level="3" pageId="8" pageNumber="150" reason="3">
|
||
<emphasis id="B971EAE3C326FFC79DA2FC289255BD8F" bold="true" box="[189,334,947,973]" pageId="8" pageNumber="150">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2FC799070BA27" blockId="8.[189,1401,994,1126]" pageId="8" pageNumber="150">
|
||
SHELL (
|
||
<figureCitation id="133E2A74C326FFC79C09FC79926EBDBE" box="[278,373,994,1021]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="8" pageNumber="150">Fig. 3A</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC326FFC79C9BFC7992C7BDBE" box="[388,476,994,1020]" captionStart="Table 1" captionStartId="2.[185,255,857,885]" captionTargetBox="[195,1921,987,1444]" captionTargetPageId="2" captionText="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." httpUri="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" pageId="8" pageNumber="150" tableUuid="DF7A6679C32CFFCD9DA6FCC295B0BD37">Table 1</tableCitation>
|
||
). Dextral, medium sized, conical-lenticular, with six slightly convex whorls. Suture shallow. Apex obtuse, depressed and low-conical. Light brown at 4 early whorls, dark brown at 2 last whorls. Sharply keeled, with pale brown band on the peripheral keel. Umbilicus very deep and wide. Aperture lip somewhat thickened, rounded rhombic.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2FB169750BB83" blockId="8.[187,1401,1165,1473]" pageId="8" pageNumber="150">
|
||
GENITAL SYSTEM (n = 3) (
|
||
<figureCitation id="133E2A74C326FFC79CF5FB169152BAEA" box="[490,585,1165,1192]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="8" pageNumber="150">Fig. 5A</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC326FFC79F48FB1691A8BAE5" box="[599,691,1165,1191]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="8" pageNumber="150" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
). Penis very long, cylindrical, and curled, divided into two portions of equal length, internally with parallel, transverse folds, containing a short, rather rounded verge. Epiphallus short cylindrical, thinner than the penis. Flagellum slightly crenated and curled up, resembling the circinate vernation of a fern shoot, with thin grooves internally and tubercles on its surface at the end, internally with transverse robust folds. Vas deferens is a slender, cylindrical tube, thicker at the end where it connects to free oviduct; entering flagellum basally. Vagina less than half as long as penis, internally with four thin longitudinal pilasters. Free oviduct shorter than vagina. Gametolytic sac thickened at base, stout and gradually tapering distally to form a slender distal tube ending in a balloonshaped, medium-sized sac. Prostate gland very long, uterus long and swollen.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2FA729728B82F" blockId="8.[188,1399,1512,1645]" pageId="8" pageNumber="150">
|
||
RADULA (n = 3). Comprises 95 transverse rows with 57–61 teeth per row; radular formula: (20)-(8–10)- 1-(8–10)-(20). Central tooth symmetric, unicuspid, tongue-shaped; lateral teeth longer than central tooth and oblique, unicuspid, tongue-shaped (
|
||
<figureCitation id="133E2A74C326FFC79F95F9B591F3B80B" box="[650,744,1582,1609]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="8" pageNumber="150">Fig. 6A</figureCitation>
|
||
). Marginal teeth tricuspid with medium-sized endocone; mesocone rather large, with rounded cusp; ectocone triangular (
|
||
<figureCitation id="133E2A74C326FFC79EB8F9C9973DB82F" box="[935,1062,1618,1645]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="8" pageNumber="150">Fig. 6B–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC326FFC79DA2F9089606B9CE" pageId="8" pageNumber="150" type="discussion">
|
||
<paragraph id="8BBA36F1C326FFC79DA2F9089234B8EF" blockId="8.[189,303,1683,1709]" box="[189,303,1683,1709]" pageId="8" pageNumber="150">
|
||
<heading id="D0F2819DC326FFC79DA2F9089234B8EF" bold="true" box="[189,303,1683,1709]" fontSize="11" level="3" pageId="8" pageNumber="150" reason="3">
|
||
<emphasis id="B971EAE3C326FFC79DA2F9089234B8EF" bold="true" box="[189,303,1683,1709]" pageId="8" pageNumber="150">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C326FFC79DA2F9599606B9CE" blockId="8.[189,1399,1730,1932]" pageId="8" pageNumber="150">
|
||
<taxonomicName id="4C054D72C326FFC79DA2F95992B3B899" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,424,1730,1756]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C326FFC79DA2F95992B3B899" box="[189,424,1730,1756]" italics="true" pageId="8" pageNumber="150">Landouria circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C326FFC79CB1F9599113B89E" box="[430,520,1730,1756]" pageId="8" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
resembles
|
||
<taxonomicName id="4C054D72C326FFC79F97F9599034B89E" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" box="[648,815,1730,1756]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C326FFC79F97F9599034B89E" box="[648,815,1730,1756]" italics="true" pageId="8" pageNumber="150">L. strobiloides</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF944B00C326FFC79E75F959963DB89E" author="Tumpeesuwan C. & Tumpeesuwan S." box="[874,1318,1730,1756]" pageId="8" pageNumber="150" pagination="298 - 305" refId="ref11349" refString="Tumpeesuwan C. & Tumpeesuwan S. 2019. First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species. Raffles Bulletin of Zoology 67: 298 - 305. https: // doi. org / 10.26107 / RBZ- 2019 - 0021" type="journal article" year="2019">Tumpeesuwan & Tumpeesuwan 2019</bibRefCitation>
|
||
: fig. 3) in shell morphology,
|
||
<taxonomicName id="4C054D72C326FFC79CA6F97D915AB8BD" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[441,577,1766,1791]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C326FFC79CA6F97D915AB8BD" box="[441,577,1766,1791]" italics="true" pageId="8" pageNumber="150">L. circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C326FFC79F57F97E91BFB8BD" box="[584,676,1765,1791]" pageId="8" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a smaller shell, a shallower suture, and darker peripheral zone than
|
||
<taxonomicName id="4C054D72C326FFC79C2EF89292C3B960" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" box="[305,472,1800,1826]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C326FFC79C2EF89292C3B960" box="[305,472,1800,1826]" italics="true" pageId="8" pageNumber="150">L. strobiloides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C687034AC326FFC79CF8F893915AB960" box="[487,577,1800,1826]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="8" pageNumber="150" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
,
|
||
<figureCitation id="133E2A74C326FFC79F53F89391BCB960" box="[588,679,1800,1826]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="8" pageNumber="150">Fig. 3A</figureCitation>
|
||
). Its central and lateral radular teeth are unicuspid and tongueshaped, whereas they are lanceolate in
|
||
<taxonomicName id="4C054D72C326FFC79F81F8B79051B907" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" box="[670,842,1835,1861]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C326FFC79F81F8B79051B907" box="[670,842,1835,1861]" italics="true" pageId="8" pageNumber="150">L. strobiloides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C687034AC326FFC79E41F8B090A6B907" box="[862,957,1835,1861]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="8" pageNumber="150" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
,
|
||
<figureCitation id="133E2A74C326FFC79ED1F8B0972BB904" box="[974,1072,1835,1862]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="8" pageNumber="150">Fig. 6A</figureCitation>
|
||
). The unique characters of
|
||
<taxonomicName id="4C054D72C326FFC79DA2F8D4925FB92A" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,324,1871,1896]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C326FFC79DA2F8D4925FB92A" box="[189,324,1871,1896]" italics="true" pageId="8" pageNumber="150">L. circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C326FFC79C56F8D492B8B92B" box="[329,419,1871,1897]" pageId="8" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are its circinate flagellum and its penis that is much longer than the vagina (
|
||
<figureCitation id="133E2A74C326FFC79813F8D59672B92B" box="[1292,1385,1870,1897]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="8" pageNumber="150">Fig. 5A</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4C054D72C326FFC79C3BF8E992D7B9CE" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" box="[292,460,1906,1932]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="150" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C326FFC79C3BF8E992D7B9CE" box="[292,460,1906,1932]" italics="true" pageId="8" pageNumber="150">L. strobiloides</emphasis>
|
||
</taxonomicName>
|
||
has a strobilus-like flagellum and a penis that is shorter than the vagina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7A6679C327FFC69DA2F8A2904DB9A2" ID-DOI="http://doi.org/10.5281/zenodo.5514845" ID-Zenodo-Dep="5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="9" pageNumber="151" startId="9.[189,232,1849,1875]" targetBox="[220,1368,265,1811]" targetPageId="9">
|
||
<paragraph id="8BBA36F1C327FFC69DA2F8A2904DB9A2" blockId="9.[189,1399,1849,2016]" pageId="9" pageNumber="151">
|
||
<emphasis id="B971EAE3C327FFC69DA2F8A29208B911" bold="true" box="[189,275,1849,1875]" pageId="9" pageNumber="151">Fig. 3.</emphasis>
|
||
External shell morphology of
|
||
<taxonomicName id="4C054D72C327FFC69FBFF8A29040B916" box="[672,859,1849,1876]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C327FFC69FBFF8A29007B911" box="[672,796,1849,1875]" italics="true" pageId="9" pageNumber="151">Landouria</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
from northeastern Thailand.
|
||
<emphasis id="B971EAE3C327FFC699D0F8A297FFB911" bold="true" box="[1231,1252,1849,1875]" pageId="9" pageNumber="151">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC699E4F8A2923DB934" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C327FFC699E4F8A2923DB934" italics="true" pageId="9" pageNumber="151">Landouria circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69C30F8C69295B935" box="[303,398,1885,1911]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMSU-00023).
|
||
<emphasis id="B971EAE3C327FFC69FF0F8C7901AB934" bold="true" box="[751,769,1884,1910]" pageId="9" pageNumber="151">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC69E0EF8C690A3B934" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[785,952,1884,1910]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C327FFC69E0EF8C690A3B934" box="[785,952,1884,1910]" italics="true" pageId="9" pageNumber="151">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69EDDF8C6973AB935" box="[962,1057,1885,1911]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMSU-00025).
|
||
<emphasis id="B971EAE3C327FFC69DA0F8E493C9B9DB" bold="true" box="[191,210,1919,1945]" pageId="9" pageNumber="151">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC69DF9F81B92FBB9DB" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[230,480,1919,1945]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C327FFC69DF9F81B92FBB9DB" box="[230,480,1919,1945]" italics="true" pageId="9" pageNumber="151">L. trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69CF3F81B9154B9D8" box="[492,591,1920,1946]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMSU-00027).
|
||
<emphasis id="B971EAE3C327FFC69EA8F81B90D7B9D8" bold="true" box="[951,972,1920,1946]" pageId="9" pageNumber="151">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC69EFFF81B978BB9DB" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[992,1168,1919,1945]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C327FFC69EFFF81B978BB9DB" box="[992,1168,1919,1945]" italics="true" pageId="9" pageNumber="151">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69982F81B97E4B9D8" box="[1181,1279,1920,1946]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMSU-00029).
|
||
<emphasis id="B971EAE3C327FFC69C83F83892ABB9FF" bold="true" box="[412,432,1955,1981]" pageId="9" pageNumber="151">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC69CA1F838912CB9FE" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[446,567,1954,1980]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B971EAE3C327FFC69CA1F838912CB9FE" box="[446,567,1954,1980]" italics="true" pageId="9" pageNumber="151">L. elegans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69F22F8389181B9FF" box="[573,666,1955,1981]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (NHMSU-00032).
|
||
<emphasis id="B971EAE3C327FFC69EEDF838971FB9FF" bold="true" box="[1010,1028,1955,1981]" pageId="9" pageNumber="151">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C327FFC6990CF838921AB9A2" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="151" phylum="Mollusca" rank="species" species="diplogramma" status="comb. nov.">
|
||
<emphasis id="B971EAE3C327FFC6990CF83897D4B9FE" box="[1043,1231,1954,1980]" italics="true" pageId="9" pageNumber="151">L. diplogramma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF944B00C327FFC699C1F83993E2B9A2" author="Mollendorff O. F." pageId="9" pageNumber="151" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C327FFC69C17F85D9291B9A2" box="[264,394,1990,2016]" pageId="9" pageNumber="151" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(NHMSU-00034). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A6679C324FFC59DA6FDBB969CBCE7" ID-Table-UUID="DF7A6679C324FFC59DA6FDBB969CBCE7" httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="10" pageNumber="152" startId="10.[185,255,544,572]" targetBox="[196,1908,758,1739]" targetIsTable="true" targetPageId="10">
|
||
<paragraph id="8BBA36F1C324FFC59DA6FDBB969CBCE7" blockId="10.[185,1939,544,677]" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59DA6FDBB9239BC7E" bold="true" box="[185,290,544,572]" pageId="10" pageNumber="152">Table 4.</emphasis>
|
||
Comparison of shells, radulae, and genital systems of
|
||
<taxonomicName id="4C054D72C324FFC59ECCFDBB9790BC7E" box="[979,1163,544,572]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C324FFC59ECCFDBB9754BC7E" box="[979,1103,544,572]" italics="true" pageId="10" pageNumber="152">Landouria</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
from northeastern Thailand. References: 1 =
|
||
<bibRefCitation id="EF944B00C324FFC59BD9FDBB92A7BC1D" author="Tumpeesuwan C. & Tumpeesuwan S." pageId="10" pageNumber="152" pagination="298 - 305" refId="ref11349" refString="Tumpeesuwan C. & Tumpeesuwan S. 2019. First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species. Raffles Bulletin of Zoology 67: 298 - 305. https: // doi. org / 10.26107 / RBZ- 2019 - 0021" type="journal article" year="2019">Tumpeesuwan & Tumpeesuwan (2019)</bibRefCitation>
|
||
; 2 =
|
||
<bibRefCitation id="EF944B00C324FFC59CEDFDD89183BC1D" author="Nahok B." box="[498,664,579,607]" pageId="10" pageNumber="152" refId="ref10522" refString="Nahok B. 2020. Systematics and Taxonomy of Land Snails Genera Aegista, Thaitropis, Landouria, and Pseudobuliminus in Thailand. Unpublished PhD thesis, Department of Biology, Faculty of Science, Mahasarakham University." type="book" year="2020">Nahok (2020)</bibRefCitation>
|
||
; 3 = this study.
|
||
<superScript id="7C709BB9C324FFC59E56FDDA9049BC10" attach="right" box="[841,850,577,594]" fontSize="6" pageId="10" pageNumber="152">*</superScript>
|
||
In
|
||
<bibRefCitation id="EF944B00C324FFC59E6FFDD89620BC1D" author="Tumpeesuwan C. & Tumpeesuwan S." box="[880,1339,579,607]" pageId="10" pageNumber="152" pagination="298 - 305" refId="ref11349" refString="Tumpeesuwan C. & Tumpeesuwan S. 2019. First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species. Raffles Bulletin of Zoology 67: 298 - 305. https: // doi. org / 10.26107 / RBZ- 2019 - 0021" type="journal article" year="2019">Tumpeesuwan & Tumpeesuwan (2019)</bibRefCitation>
|
||
indicated the number of teeth per row in
|
||
<taxonomicName id="4C054D72C324FFC59A09FDD89259BCC0" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C324FFC59A09FDD89259BCC0" italics="true" pageId="10" pageNumber="152">Landouria strobiloides</emphasis>
|
||
</taxonomicName>
|
||
varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBA36F1C324FFC59DDBFD6D9413B889" pageId="10" pageNumber="152">
|
||
<table id="F905C451C32400309DDBFD6D946FB889" box="[196,1908,758,1739]" gridcols="8" gridrows="20" pageId="10" pageNumber="152">
|
||
<tr id="353534B3C32400309DDBFD6D946FBD4B" box="[196,1908,758,777]" gridrow="0" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFD6D92A7BD4B" box="[196,444,758,777]" gridcol="0" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59DDBFD6D923FBD4B" bold="true" box="[196,292,758,777]" pageId="10" pageNumber="152">Characters</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309CC6FD6D9168BD4B" box="[473,627,758,777]" gridcol="1" gridrow="0" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59CC6FD6D9156BD4B" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2019" box="[473,589,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="strobiloides">
|
||
<emphasis id="B971EAE3C324FFC59CC6FD6D9156BD4B" bold="true" box="[473,589,758,777]" italics="true" pageId="10" pageNumber="152">L. strobiloides</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76E45DCFC32400309F8BFD6D9022BD4B" box="[660,825,758,777]" gridcol="2" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59F8BFD6D9022BD4B" bold="true" box="[660,825,758,777]" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59F8BFD6D91EFBD4B" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[660,756,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C324FFC59F8BFD6D91EFBD4B" bold="true" box="[660,756,758,777]" italics="true" pageId="10" pageNumber="152">L. circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C324FFC59FE6FD6D9022BD4B" box="[761,825,758,777]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309E49FD6D970BBD4B" box="[854,1040,758,777]" gridcol="3" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59E49FD6D970BBD4B" bold="true" box="[854,1040,758,777]" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59E49FD6D90D1BD4B" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[854,970,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C324FFC59E49FD6D90D1BD4B" bold="true" box="[854,970,758,777]" italics="true" pageId="10" pageNumber="152">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C324FFC59ED0FD6D970BBD4B" box="[975,1040,758,777]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309937FD6D9601BD4B" box="[1064,1306,758,777]" gridcol="4" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59937FD6D9601BD4B" bold="true" box="[1064,1306,758,777]" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59937FD6D97CFBD4B" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[1064,1236,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C324FFC59937FD6D97CFBD4B" bold="true" box="[1064,1236,758,777]" italics="true" pageId="10" pageNumber="152">L. trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C324FFC599C6FD6D9601BD4B" box="[1241,1306,758,777]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309853FD6D9513BD4B" box="[1356,1544,758,777]" gridcol="5" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59853FD6D9513BD4B" bold="true" box="[1356,1544,758,777]" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59853FD6D96D8BD4B" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[1356,1475,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C324FFC59853FD6D96D8BD4B" bold="true" box="[1356,1475,758,777]" italics="true" pageId="10" pageNumber="152">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C324FFC598D7FD6D9513BD4B" box="[1480,1544,758,777]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309B30FD6D95D3BD4B" box="[1583,1736,758,777]" gridcol="6" gridrow="0" pageId="10" pageNumber="152">
|
||
<emphasis id="B971EAE3C324FFC59B30FD6D95D3BD4B" bold="true" box="[1583,1736,758,777]" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59B30FD6D9598BD4B" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[1583,1667,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B971EAE3C324FFC59B30FD6D9598BD4B" bold="true" box="[1583,1667,758,777]" italics="true" pageId="10" pageNumber="152">L. elegans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C324FFC59B97FD6D95D3BD4B" box="[1672,1736,758,777]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="76E45DCFC32400309BF5FD6D946FBD4B" box="[1770,1908,758,777]" gridcol="7" gridrow="0" pageId="10" pageNumber="152">
|
||
<taxonomicName id="4C054D72C324FFC59BF5FD6D9475BD4B" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[1770,1902,758,777]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="152" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C324FFC59BF5FD6D9475BD4B" bold="true" box="[1770,1902,758,777]" italics="true" pageId="10" pageNumber="152">L. diplogramma</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFCB2946FBD7E" box="[196,1908,809,828]" gridrow="1" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFCB2946FBD7E" box="[196,1908,809,828]" colspan="8" colspanRight="7" gridcol="0" gridrow="1" pageId="10" pageNumber="152">Shell:</th>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFCC0946FBD2C" box="[196,1908,859,878]" gridrow="2" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFCC092A7BD2C" box="[196,444,859,878]" gridcol="0" gridrow="2" pageId="10" pageNumber="152">Shell height (mm)</th>
|
||
<td id="76E45DCFC32400309CC6FCC09168BD2C" box="[473,627,859,878]" gridcol="1" gridrow="2" pageId="10" pageNumber="152">7.10–9.60</td>
|
||
<td id="76E45DCFC32400309F8BFCC09022BD2C" box="[660,825,859,878]" gridcol="2" gridrow="2" pageId="10" pageNumber="152">5.60–8.10</td>
|
||
<td id="76E45DCFC32400309E49FCC0970BBD2C" box="[854,1040,859,878]" gridcol="3" gridrow="2" pageId="10" pageNumber="152">7.00–9.60</td>
|
||
<td id="76E45DCFC32400309937FCC09601BD2C" box="[1064,1306,859,878]" gridcol="4" gridrow="2" pageId="10" pageNumber="152">7.60–11.30</td>
|
||
<td id="76E45DCFC32400309853FCC09513BD2C" box="[1356,1544,859,878]" gridcol="5" gridrow="2" pageId="10" pageNumber="152">6.10–12.00</td>
|
||
<td id="76E45DCFC32400309B30FCC095D3BD2C" box="[1583,1736,859,878]" gridcol="6" gridrow="2" pageId="10" pageNumber="152">6.00–10.00</td>
|
||
<td id="76E45DCFC32400309BF5FCC0946FBD2C" box="[1770,1908,859,878]" gridcol="7" gridrow="2" pageId="10" pageNumber="152">6.00–8.70</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFC15946FBDE3" box="[196,1908,910,929]" gridrow="3" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFC1592A7BDE3" box="[196,444,910,929]" gridcol="0" gridrow="3" pageId="10" pageNumber="152">Shell colour</th>
|
||
<td id="76E45DCFC32400309CC6FC159168BDE3" box="[473,627,910,929]" gridcol="1" gridrow="3" pageId="10" pageNumber="152">pale brown</td>
|
||
<td id="76E45DCFC32400309F8BFC159022BDE3" box="[660,825,910,929]" gridcol="2" gridrow="3" pageId="10" pageNumber="152">brownish-corneous</td>
|
||
<td id="76E45DCFC32400309E49FC15970BBDE3" box="[854,1040,910,929]" gridcol="3" gridrow="3" pageId="10" pageNumber="152">red brown</td>
|
||
<td id="76E45DCFC32400309937FC159601BDE3" box="[1064,1306,910,929]" gridcol="4" gridrow="3" pageId="10" pageNumber="152">brownish-corneous</td>
|
||
<td id="76E45DCFC32400309853FC159513BDE3" box="[1356,1544,910,929]" gridcol="5" gridrow="3" pageId="10" pageNumber="152">red brown</td>
|
||
<td id="76E45DCFC32400309B30FC1595D3BDE3" box="[1583,1736,910,929]" gridcol="6" gridrow="3" pageId="10" pageNumber="152">brownish-corneous</td>
|
||
<td id="76E45DCFC32400309BF5FC15946FBDE3" box="[1770,1908,910,929]" gridcol="7" gridrow="3" pageId="10" pageNumber="152">whitish-corneous</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFC5A946FBD96" box="[196,1908,961,980]" gridrow="4" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFC5A92A7BD96" box="[196,444,961,980]" gridcol="0" gridrow="4" pageId="10" pageNumber="152">Suture</th>
|
||
<td id="76E45DCFC32400309CC6FC5A9168BD96" box="[473,627,961,980]" gridcol="1" gridrow="4" pageId="10" pageNumber="152">deep</td>
|
||
<td id="76E45DCFC32400309F8BFC5A9022BD96" box="[660,825,961,980]" gridcol="2" gridrow="4" pageId="10" pageNumber="152">shallow</td>
|
||
<td id="76E45DCFC32400309E49FC5A970BBD96" box="[854,1040,961,980]" gridcol="3" gridrow="4" pageId="10" pageNumber="152">slightly deep</td>
|
||
<td id="76E45DCFC32400309937FC5A9601BD96" box="[1064,1306,961,980]" gridcol="4" gridrow="4" pageId="10" pageNumber="152">shallow</td>
|
||
<td id="76E45DCFC32400309853FC5A9513BD96" box="[1356,1544,961,980]" gridcol="5" gridrow="4" pageId="10" pageNumber="152">deep</td>
|
||
<td id="76E45DCFC32400309B30FC5A95D3BD96" box="[1583,1736,961,980]" gridcol="6" gridrow="4" pageId="10" pageNumber="152">shallow</td>
|
||
<td id="76E45DCFC32400309BF5FC5A946FBD96" box="[1770,1908,961,980]" gridcol="7" gridrow="4" pageId="10" pageNumber="152">shallow</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFC68946FBA44" box="[196,1908,1011,1030]" gridrow="5" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFC6892A7BA44" box="[196,444,1011,1030]" gridcol="0" gridrow="5" pageId="10" pageNumber="152">Peripheral band</th>
|
||
<td id="76E45DCFC32400309CC6FC689168BA44" box="[473,627,1011,1030]" gridcol="1" gridrow="5" pageId="10" pageNumber="152">dark brown</td>
|
||
<td id="76E45DCFC32400309F8BFC689022BA44" box="[660,825,1011,1030]" gridcol="2" gridrow="5" pageId="10" pageNumber="152">pale brown</td>
|
||
<td id="76E45DCFC32400309E49FC68970BBA44" box="[854,1040,1011,1030]" gridcol="3" gridrow="5" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309937FC689601BA44" box="[1064,1306,1011,1030]" gridcol="4" gridrow="5" pageId="10" pageNumber="152">brown</td>
|
||
<td id="76E45DCFC32400309853FC689513BA44" box="[1356,1544,1011,1030]" gridcol="5" gridrow="5" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309B30FC6895D3BA44" box="[1583,1736,1011,1030]" gridcol="6" gridrow="5" pageId="10" pageNumber="152">dark brown</td>
|
||
<td id="76E45DCFC32400309BF5FC68946FBA44" box="[1770,1908,1011,1030]" gridcol="7" gridrow="5" pageId="10" pageNumber="152">pale brown</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFBBD946FBA7B" box="[196,1908,1062,1081]" gridrow="6" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFBBD92A7BA7B" box="[196,444,1062,1081]" gridcol="0" gridrow="6" pageId="10" pageNumber="152">Peripheral keel</th>
|
||
<td id="76E45DCFC32400309CC6FBBD9168BA7B" box="[473,627,1062,1081]" gridcol="1" gridrow="6" pageId="10" pageNumber="152">sharp</td>
|
||
<td id="76E45DCFC32400309F8BFBBD9022BA7B" box="[660,825,1062,1081]" gridcol="2" gridrow="6" pageId="10" pageNumber="152">sharp</td>
|
||
<td id="76E45DCFC32400309E49FBBD970BBA7B" box="[854,1040,1062,1081]" gridcol="3" gridrow="6" pageId="10" pageNumber="152">moderate</td>
|
||
<td id="76E45DCFC32400309937FBBD9601BA7B" box="[1064,1306,1062,1081]" gridcol="4" gridrow="6" pageId="10" pageNumber="152">sharp</td>
|
||
<td id="76E45DCFC32400309853FBBD9513BA7B" box="[1356,1544,1062,1081]" gridcol="5" gridrow="6" pageId="10" pageNumber="152">weak</td>
|
||
<td id="76E45DCFC32400309B30FBBD95D3BA7B" box="[1583,1736,1062,1081]" gridcol="6" gridrow="6" pageId="10" pageNumber="152">sharp</td>
|
||
<td id="76E45DCFC32400309BF5FBBD946FBA7B" box="[1770,1908,1062,1081]" gridcol="7" gridrow="6" pageId="10" pageNumber="152">very weak</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFBC2946FBA2E" box="[196,1908,1113,1132]" gridrow="7" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFBC292A7BA2E" box="[196,444,1113,1132]" gridcol="0" gridrow="7" pageId="10" pageNumber="152">Sculpture</th>
|
||
<td id="76E45DCFC32400309CC6FBC29168BA2E" box="[473,627,1113,1132]" gridcol="1" gridrow="7" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309F8BFBC29022BA2E" box="[660,825,1113,1132]" gridcol="2" gridrow="7" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309E49FBC2970BBA2E" box="[854,1040,1113,1132]" gridcol="3" gridrow="7" pageId="10" pageNumber="152">tubercles</td>
|
||
<td id="76E45DCFC32400309937FBC29601BA2E" box="[1064,1306,1113,1132]" gridcol="4" gridrow="7" pageId="10" pageNumber="152">scaly</td>
|
||
<td id="76E45DCFC32400309853FBC29513BA2E" box="[1356,1544,1113,1132]" gridcol="5" gridrow="7" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309B30FBC295D3BA2E" box="[1583,1736,1113,1132]" gridcol="6" gridrow="7" pageId="10" pageNumber="152">absent</td>
|
||
<td id="76E45DCFC32400309BF5FBC2946FBA2E" box="[1770,1908,1113,1132]" gridcol="7" gridrow="7" pageId="10" pageNumber="152">absent</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFB12946FBADC" box="[196,1908,1161,1182]" gridrow="8" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFB1292A7BADC" box="[196,444,1161,1182]" gridcol="0" gridrow="8" pageId="10" pageNumber="152">Number of whorls</th>
|
||
<td id="76E45DCFC32400309CC6FB129168BADC" box="[473,627,1161,1182]" gridcol="1" gridrow="8" pageId="10" pageNumber="152">5½</td>
|
||
<td id="76E45DCFC32400309F8BFB129022BADC" box="[660,825,1161,1182]" gridcol="2" gridrow="8" pageId="10" pageNumber="152">6</td>
|
||
<td id="76E45DCFC32400309E49FB12970BBADC" box="[854,1040,1161,1182]" gridcol="3" gridrow="8" pageId="10" pageNumber="152">6½</td>
|
||
<td id="76E45DCFC32400309937FB129601BADC" box="[1064,1306,1161,1182]" gridcol="4" gridrow="8" pageId="10" pageNumber="152">6</td>
|
||
<td id="76E45DCFC32400309853FB129513BADC" box="[1356,1544,1161,1182]" gridcol="5" gridrow="8" pageId="10" pageNumber="152">6½</td>
|
||
<td id="76E45DCFC32400309B30FB1295D3BADC" box="[1583,1736,1161,1182]" gridcol="6" gridrow="8" pageId="10" pageNumber="152">6</td>
|
||
<td id="76E45DCFC32400309BF5FB12946FBADC" box="[1770,1908,1161,1182]" gridcol="7" gridrow="8" pageId="10" pageNumber="152">5½</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFB25946FBA93" box="[196,1908,1214,1233]" gridrow="9" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFB25946FBA93" box="[196,1908,1214,1233]" colspan="8" colspanRight="7" gridcol="0" gridrow="9" pageId="10" pageNumber="152">Radula:</th>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFB6A946FBB46" box="[196,1908,1265,1284]" gridrow="10" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFB6A92A7BB46" box="[196,444,1265,1284]" gridcol="0" gridrow="10" pageId="10" pageNumber="152">Number of rows</th>
|
||
<td id="76E45DCFC32400309CC6FB6A9168BB46" box="[473,627,1265,1284]" gridcol="1" gridrow="10" pageId="10" pageNumber="152">100</td>
|
||
<td id="76E45DCFC32400309F8BFB6A9022BB46" box="[660,825,1265,1284]" gridcol="2" gridrow="10" pageId="10" pageNumber="152">93–95</td>
|
||
<td id="76E45DCFC32400309E49FB6A970BBB46" box="[854,1040,1265,1284]" gridcol="3" gridrow="10" pageId="10" pageNumber="152">126–130</td>
|
||
<td id="76E45DCFC32400309937FB6A9601BB46" box="[1064,1306,1265,1284]" gridcol="4" gridrow="10" pageId="10" pageNumber="152">107–111</td>
|
||
<td id="76E45DCFC32400309853FB6A9513BB46" box="[1356,1544,1265,1284]" gridcol="5" gridrow="10" pageId="10" pageNumber="152">125–129</td>
|
||
<td id="76E45DCFC32400309B30FB6A95D3BB46" box="[1583,1736,1265,1284]" gridcol="6" gridrow="10" pageId="10" pageNumber="152">110–114</td>
|
||
<td id="76E45DCFC32400309BF5FB6A946FBB46" box="[1770,1908,1265,1284]" gridcol="7" gridrow="10" pageId="10" pageNumber="152">121–125</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFAB9946FBB74" box="[196,1908,1314,1334]" gridrow="11" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFAB992A7BB74" box="[196,444,1314,1334]" gridcol="0" gridrow="11" pageId="10" pageNumber="152">Number of teeth in each row</th>
|
||
<td id="76E45DCFC32400309CC6FAB99168BB74" box="[473,627,1314,1334]" gridcol="1" gridrow="11" pageId="10" pageNumber="152">3*–53</td>
|
||
<td id="76E45DCFC32400309F8BFAB99022BB74" box="[660,825,1314,1334]" gridcol="2" gridrow="11" pageId="10" pageNumber="152">55–63</td>
|
||
<td id="76E45DCFC32400309E49FAB9970BBB74" box="[854,1040,1314,1334]" gridcol="3" gridrow="11" pageId="10" pageNumber="152">79–87</td>
|
||
<td id="76E45DCFC32400309937FAB99601BB74" box="[1064,1306,1314,1334]" gridcol="4" gridrow="11" pageId="10" pageNumber="152">65–73</td>
|
||
<td id="76E45DCFC32400309853FAB99513BB74" box="[1356,1544,1314,1334]" gridcol="5" gridrow="11" pageId="10" pageNumber="152">75–83</td>
|
||
<td id="76E45DCFC32400309B30FAB995D3BB74" box="[1583,1736,1314,1334]" gridcol="6" gridrow="11" pageId="10" pageNumber="152">69–77</td>
|
||
<td id="76E45DCFC32400309BF5FAB9946FBB74" box="[1770,1908,1314,1334]" gridcol="7" gridrow="11" pageId="10" pageNumber="152">61–69</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFACD946FBB2B" box="[196,1908,1366,1385]" gridrow="12" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFACD92A7BB2B" box="[196,444,1366,1385]" gridcol="0" gridrow="12" pageId="10" pageNumber="152">- Lateral teeth</th>
|
||
<td id="76E45DCFC32400309CC6FACD9168BB2B" box="[473,627,1366,1385]" gridcol="1" gridrow="12" pageId="10" pageNumber="152">8–10</td>
|
||
<td id="76E45DCFC32400309F8BFACD9022BB2B" box="[660,825,1366,1385]" gridcol="2" gridrow="12" pageId="10" pageNumber="152">8–10</td>
|
||
<td id="76E45DCFC32400309E49FACD970BBB2B" box="[854,1040,1366,1385]" gridcol="3" gridrow="12" pageId="10" pageNumber="152">11–13</td>
|
||
<td id="76E45DCFC32400309937FACD9601BB2B" box="[1064,1306,1366,1385]" gridcol="4" gridrow="12" pageId="10" pageNumber="152">8–10</td>
|
||
<td id="76E45DCFC32400309853FACD9513BB2B" box="[1356,1544,1366,1385]" gridcol="5" gridrow="12" pageId="10" pageNumber="152">11–13</td>
|
||
<td id="76E45DCFC32400309B30FACD95D3BB2B" box="[1583,1736,1366,1385]" gridcol="6" gridrow="12" pageId="10" pageNumber="152">12–14</td>
|
||
<td id="76E45DCFC32400309BF5FACD946FBB2B" box="[1770,1908,1366,1385]" gridcol="7" gridrow="12" pageId="10" pageNumber="152">7–9</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFA12946FBBDE" box="[196,1908,1417,1436]" gridrow="13" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFA1292A7BBDE" box="[196,444,1417,1436]" gridcol="0" gridrow="13" pageId="10" pageNumber="152">- Marginal teeth</th>
|
||
<td id="76E45DCFC32400309CC6FA129168BBDE" box="[473,627,1417,1436]" gridcol="1" gridrow="13" pageId="10" pageNumber="152">15–17</td>
|
||
<td id="76E45DCFC32400309F8BFA129022BBDE" box="[660,825,1417,1436]" gridcol="2" gridrow="13" pageId="10" pageNumber="152">19–21</td>
|
||
<td id="76E45DCFC32400309E49FA12970BBBDE" box="[854,1040,1417,1436]" gridcol="3" gridrow="13" pageId="10" pageNumber="152">28–30</td>
|
||
<td id="76E45DCFC32400309937FA129601BBDE" box="[1064,1306,1417,1436]" gridcol="4" gridrow="13" pageId="10" pageNumber="152">24–26</td>
|
||
<td id="76E45DCFC32400309853FA129513BBDE" box="[1356,1544,1417,1436]" gridcol="5" gridrow="13" pageId="10" pageNumber="152">26–28</td>
|
||
<td id="76E45DCFC32400309B30FA1295D3BBDE" box="[1583,1736,1417,1436]" gridcol="6" gridrow="13" pageId="10" pageNumber="152">22–24</td>
|
||
<td id="76E45DCFC32400309BF5FA12946FBBDE" box="[1770,1908,1417,1436]" gridcol="7" gridrow="13" pageId="10" pageNumber="152">23–25</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFA20946FBB8C" box="[196,1908,1467,1486]" gridrow="14" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFA2092A7BB8C" box="[196,444,1467,1486]" gridcol="0" gridrow="14" pageId="10" pageNumber="152">Central tooth</th>
|
||
<td id="76E45DCFC32400309CC6FA209168BB8C" box="[473,627,1467,1486]" gridcol="1" gridrow="14" pageId="10" pageNumber="152">elongate lanceolate</td>
|
||
<td id="76E45DCFC32400309F8BFA209022BB8C" box="[660,825,1467,1486]" gridcol="2" gridrow="14" pageId="10" pageNumber="152">tongue-shaped</td>
|
||
<td id="76E45DCFC32400309E49FA20970BBB8C" box="[854,1040,1467,1486]" gridcol="3" gridrow="14" pageId="10" pageNumber="152">slender lanceolate</td>
|
||
<td id="76E45DCFC32400309937FA209601BB8C" box="[1064,1306,1467,1486]" gridcol="4" gridrow="14" pageId="10" pageNumber="152">short lanceolate</td>
|
||
<td id="76E45DCFC32400309853FA209513BB8C" box="[1356,1544,1467,1486]" gridcol="5" gridrow="14" pageId="10" pageNumber="152">lanceolate</td>
|
||
<td id="76E45DCFC32400309B30FA2095D3BB8C" box="[1583,1736,1467,1486]" gridcol="6" gridrow="14" pageId="10" pageNumber="152">tongue-shaped</td>
|
||
<td id="76E45DCFC32400309BF5FA20946FBB8C" box="[1770,1908,1467,1486]" gridcol="7" gridrow="14" pageId="10" pageNumber="152">triangular</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBFA75946FB843" box="[196,1908,1518,1537]" gridrow="15" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBFA75946FB843" box="[196,1908,1518,1537]" colspan="8" colspanRight="7" gridcol="0" gridrow="15" pageId="10" pageNumber="152">Genital system:</th>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBF9BA946FB876" box="[196,1908,1569,1588]" gridrow="16" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBF9BA92A7B876" box="[196,444,1569,1588]" gridcol="0" gridrow="16" pageId="10" pageNumber="152">Penis</th>
|
||
<td id="76E45DCFC32400309CC6F9BA9168B876" box="[473,627,1569,1588]" gridcol="1" gridrow="16" pageId="10" pageNumber="152">short, slightly stout</td>
|
||
<td id="76E45DCFC32400309F8BF9BA9022B876" box="[660,825,1569,1588]" gridcol="2" gridrow="16" pageId="10" pageNumber="152">long cylindrical</td>
|
||
<td id="76E45DCFC32400309E49F9BA970BB876" box="[854,1040,1569,1588]" gridcol="3" gridrow="16" pageId="10" pageNumber="152">swollen</td>
|
||
<td id="76E45DCFC32400309937F9BA9601B876" box="[1064,1306,1569,1588]" gridcol="4" gridrow="16" pageId="10" pageNumber="152">long slender</td>
|
||
<td id="76E45DCFC32400309853F9BA9513B876" box="[1356,1544,1569,1588]" gridcol="5" gridrow="16" pageId="10" pageNumber="152">very large and stout</td>
|
||
<td id="76E45DCFC32400309B30F9BA95D3B876" box="[1583,1736,1569,1588]" gridcol="6" gridrow="16" pageId="10" pageNumber="152">long cylindrical</td>
|
||
<td id="76E45DCFC32400309BF5F9BA946FB876" box="[1770,1908,1569,1588]" gridcol="7" gridrow="16" pageId="10" pageNumber="152">swollen</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBF9C8946FB824" box="[196,1908,1619,1638]" gridrow="17" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBF9C892A7B824" box="[196,444,1619,1638]" gridcol="0" gridrow="17" pageId="10" pageNumber="152">Flagellum</th>
|
||
<td id="76E45DCFC32400309CC6F9C89168B824" box="[473,627,1619,1638]" gridcol="1" gridrow="17" pageId="10" pageNumber="152">strobilus-shaped</td>
|
||
<td id="76E45DCFC32400309F8BF9C89022B824" box="[660,825,1619,1638]" gridcol="2" gridrow="17" pageId="10" pageNumber="152">circinate</td>
|
||
<td id="76E45DCFC32400309E49F9C8970BB824" box="[854,1040,1619,1638]" gridcol="3" gridrow="17" pageId="10" pageNumber="152">short protrusion</td>
|
||
<td id="76E45DCFC32400309937F9C89601B824" box="[1064,1306,1619,1638]" gridcol="4" gridrow="17" pageId="10" pageNumber="152">slender towards tip</td>
|
||
<td id="76E45DCFC32400309853F9C89513B824" box="[1356,1544,1619,1638]" gridcol="5" gridrow="17" pageId="10" pageNumber="152">narrow ovate</td>
|
||
<td id="76E45DCFC32400309B30F9C895D3B824" box="[1583,1736,1619,1638]" gridcol="6" gridrow="17" pageId="10" pageNumber="152">water drop-shaped</td>
|
||
<td id="76E45DCFC32400309BF5F9C8946FB824" box="[1770,1908,1619,1638]" gridcol="7" gridrow="17" pageId="10" pageNumber="152">short protrusion</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBF91D946FB8DB" box="[196,1908,1670,1689]" gridrow="18" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBF91D92A7B8DB" box="[196,444,1670,1689]" gridcol="0" gridrow="18" pageId="10" pageNumber="152">Basal part of gametolytic sac</th>
|
||
<td id="76E45DCFC32400309CC6F91D9168B8DB" box="[473,627,1670,1689]" gridcol="1" gridrow="18" pageId="10" pageNumber="152">swollen</td>
|
||
<td id="76E45DCFC32400309F8BF91D9022B8DB" box="[660,825,1670,1689]" gridcol="2" gridrow="18" pageId="10" pageNumber="152">thick and stout</td>
|
||
<td id="76E45DCFC32400309E49F91D970BB8DB" box="[854,1040,1670,1689]" gridcol="3" gridrow="18" pageId="10" pageNumber="152">rhin</td>
|
||
<td id="76E45DCFC32400309937F91D9601B8DB" box="[1064,1306,1670,1689]" gridcol="4" gridrow="18" pageId="10" pageNumber="152">thin</td>
|
||
<td id="76E45DCFC32400309853F91D9513B8DB" box="[1356,1544,1670,1689]" gridcol="5" gridrow="18" pageId="10" pageNumber="152">thick</td>
|
||
<td id="76E45DCFC32400309B30F91D95D3B8DB" box="[1583,1736,1670,1689]" gridcol="6" gridrow="18" pageId="10" pageNumber="152">stout</td>
|
||
<td id="76E45DCFC32400309BF5F91D946FB8DB" box="[1770,1908,1670,1689]" gridcol="7" gridrow="18" pageId="10" pageNumber="152">thick</td>
|
||
</tr>
|
||
<tr id="353534B3C32400309DDBF923946FB889" box="[196,1908,1720,1739]" gridrow="19" pageId="10" pageNumber="152">
|
||
<th id="76E45DCFC32400309DDBF92392A7B889" box="[196,444,1720,1739]" gridcol="0" gridrow="19" pageId="10" pageNumber="152">References</th>
|
||
<td id="76E45DCFC32400309CC6F9239168B889" box="[473,627,1720,1739]" gridcol="1" gridrow="19" pageId="10" pageNumber="152">1, 2</td>
|
||
<td id="76E45DCFC32400309F8BF9239022B889" box="[660,825,1720,1739]" gridcol="2" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
<td id="76E45DCFC32400309E49F923970BB889" box="[854,1040,1720,1739]" gridcol="3" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
<td id="76E45DCFC32400309937F9239601B889" box="[1064,1306,1720,1739]" gridcol="4" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
<td id="76E45DCFC32400309853F9239513B889" box="[1356,1544,1720,1739]" gridcol="5" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
<td id="76E45DCFC32400309B30F92395D3B889" box="[1583,1736,1720,1739]" gridcol="6" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
<td id="76E45DCFC32400309BF5F923946FB889" box="[1770,1908,1720,1739]" gridcol="7" gridrow="19" pageId="10" pageNumber="152">2, 3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C31F657AC325FFC49DA2FE9390D8BF13" box="[189,963,264,338]" pageId="11" pageNumber="153" type="distribution">
|
||
<paragraph id="8BBA36F1C325FFC49DA2FE93924DBF60" blockId="11.[189,342,264,290]" box="[189,342,264,290]" pageId="11" pageNumber="153">
|
||
<heading id="D0F2819DC325FFC49DA2FE93924DBF60" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="11" pageNumber="153" reason="3">
|
||
<emphasis id="B971EAE3C325FFC49DA2FE93924DBF60" bold="true" box="[189,342,264,290]" pageId="11" pageNumber="153">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C325FFC49DA2FEAC90D8BF13" blockId="11.[189,963,311,338]" box="[189,963,311,338]" pageId="11" pageNumber="153">
|
||
<taxonomicName id="4C054D72C325FFC49DA2FEAC92B2BF13" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,425,311,337]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="153" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C325FFC49DA2FEAC92B2BF13" box="[189,425,311,337]" italics="true" pageId="11" pageNumber="153">Landouria circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C325FFC49CAFFEA39110BF10" box="[432,523,312,338]" pageId="11" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="54BE8853C325FFC49E34FEA39045BF10" box="[811,862,312,338]" pageId="11" pageNumber="153">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |