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<document ID-DOI="http://doi.org/10.5281/zenodo.3788330" ID-GBIF-Dataset="1900f828-2e5e-41fa-9917-752c41a5315e" ID-GBIF-Taxon="163749774" ID-ISSN="13132970" ID-Zenodo-Dep="3788330" approvalRequired="95" approvalRequired_for_textStreams="95" checkinTime="1587977121657" checkinUser="plazi" docAuthor="Tauber, Catherine" docDate="2010" docId="03AC87E3FFAB586064F3E3A4C3C2FAAB" docLanguage="en" docName="ZK_article_2195.pdf" docOrigin="ZooKeys 44 (44)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@79747166" docStyleName="zookeys.2008.journal_article" docTitle="Chrysopodes (Neosuarius) oswaldi Penny. The 2002" docType="treatment" docVersion="4" lastPageId="56" lastPageNumber="57" masterDocId="FF95FF9BFF985858647FE74DC004FFE0" masterDocTitle="Revision of Neosuarius, a subgenus of Chrysopodes (Neuroptera, Chrysopidae)" masterLastPageNumber="104" masterPageNumber="1" pageId="51" pageNumber="52" updateTime="1643563224695" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Neosuarius, a subgenus of Chrysopodes (Neuroptera, Chrysopidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tauber, Catherine</mods:namePart>
<mods:affiliation>Cornell University, Davis ,, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-04-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>44</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>104</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.44.387</mods:identifier>
<mods:identifier type="GBIF-Dataset">1900f828-2e5e-41fa-9917-752c41a5315e</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576660</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788330" ID-GBIF-Taxon="163749774" ID-Zenodo-Dep="3788330" LSID="urn:lsid:plazi:treatment:03AC87E3FFAB586064F3E3A4C3C2FAAB" httpUri="http://treatment.plazi.org/id/03AC87E3FFAB586064F3E3A4C3C2FAAB" lastPageId="56" lastPageNumber="57" pageId="51" pageNumber="52">
<subSubSection pageId="51" pageNumber="52" type="nomenclature">
<paragraph blockId="51.[140,699,1257,1319]" box="[140,699,1257,1284]" pageId="51" pageNumber="52">
<heading box="[140,699,1257,1284]" fontSize="11" level="2" pageId="51" pageNumber="52" reason="3">
<taxonomicName authority="Penny, 2002" authorityName="Penny. The" authorityYear="2002" box="[140,699,1257,1284]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis bold="true" box="[140,699,1257,1284]" pageId="51" pageNumber="52">
<emphasis bold="true" box="[140,545,1257,1284]" italics="true" pageId="51" pageNumber="52">Chrysopodes (Neosuarius) oswaldi</emphasis>
<bibRefCitation author="Penny ND" box="[552,699,1257,1284]" pageId="51" pageNumber="52" pagination="161 - 457" refId="ref59468" refString="Penny ND (2002) Family Chrysopidae. Pp. 187 - 227 (text), 301 - 373 (figures) in Penny ND (Ed) A Guide to the Lacewings (Neuroptera) of Costa Rica. Proceedings of the California Academy of Sciences (4) 53: 161 - 457." type="journal article" year="2002">Penny, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="51.[140,699,1257,1319]" box="[140,384,1293,1319]" pageId="51" pageNumber="52">Figs 11d, 12d, 3639</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="reference_group">
<paragraph blockId="51.[140,1108,1363,1460]" pageId="51" pageNumber="52">
<treatmentCitationGroup pageId="51" pageNumber="52">
<treatmentCitation author="Penny ND" box="[140,707,1363,1390]" page="224" pageId="51" pageNumber="52" year="2002">
<taxonomicName authority="Penny, 2002: 224" authorityName="Penny. The" authorityPageNumber="224" authorityYear="2002" box="[140,707,1363,1390]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[140,271,1363,1389]" italics="true" pageId="51" pageNumber="52">Chrysopodes</emphasis>
(
<emphasis box="[287,406,1364,1389]" italics="true" pageId="51" pageNumber="52">Neosuarius</emphasis>
)
<emphasis box="[421,502,1363,1389]" italics="true" pageId="51" pageNumber="52">oswaldi</emphasis>
<bibRefCitation author="Penny ND" box="[509,707,1364,1390]" pageId="51" pageNumber="52" pagination="161 - 457" refId="ref59468" refString="Penny ND (2002) Family Chrysopidae. Pp. 187 - 227 (text), 301 - 373 (figures) in Penny ND (Ed) A Guide to the Lacewings (Neuroptera) of Costa Rica. Proceedings of the California Academy of Sciences (4) 53: 161 - 457." type="journal article" year="2002">Penny, 2002: 224</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[CAS,
<typeStatus box="[791,898,1363,1389]" pageId="51" pageNumber="52">Holotype</typeStatus>
(by original designation): “
<quantity box="[293,368,1399,1425]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="51" pageNumber="52" unit="mi" value="10.0">10 mi.</quantity>
S.
<collectingRegion box="[403,486,1399,1425]" country="Puerto Rico" name="Rincon" pageId="51" pageNumber="52">Rincon</collectingRegion>
de Osa /
<collectingRegion box="[594,718,1399,1425]" country="Costa Rica" name="Puntarenas" pageId="51" pageNumber="52">Puntarenas</collectingRegion>
Proince [Sic!], Costa / Rica, 412. III.69 / J. K. Sheldon Field #/.”].
<bibRefCitation author="Oswald JD" box="[564,717,1434,1460]" pageId="51" pageNumber="52" refId="ref59168" refString="Oswald JD (2007) Neuropterida Species of the World. Version 2.0. http: // lacewing. tamu. edu / species-catalogue /. [accessed April 1, 2008 - January 12, 2010]." type="url" year="2007">Oswald 2007</bibRefCitation>
[catalog listing].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="diagnosis">
<paragraph blockId="51.[140,1108,1504,1672]" lastBlockId="52.[140,1108,765,1672]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis bold="true" box="[140,267,1504,1530]" pageId="51" pageNumber="52">
<emphasis box="[140,262,1504,1530]" inLineHeading="true" pageId="51" pageNumber="52" reason="1">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName authorityName="Penny. The" authorityYear="2002" box="[275,641,1504,1531]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[275,406,1504,1530]" italics="true" pageId="51" pageNumber="52">Chrysopodes</emphasis>
(
<emphasis box="[424,543,1505,1530]" italics="true" pageId="51" pageNumber="52">Neosuarius</emphasis>
)
<emphasis box="[560,641,1504,1530]" italics="true" pageId="51" pageNumber="52">oswaldi</emphasis>
</taxonomicName>
adults most closely resemble those of
<taxonomicName authorityName="Schneider" authorityYear="1851" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="51" pageNumber="24" phylum="Arthropoda" rank="species" species="collaris" subGenus="Neosuarius">
<emphasis box="[1081,1100,1505,1531]" italics="true" pageId="51" pageNumber="52">C</emphasis>
. (
<emphasis box="[149,171,1540,1565]" italics="true" pageId="51" pageNumber="52">N</emphasis>
.)
<emphasis box="[195,270,1539,1565]" italics="true" pageId="51" pageNumber="52">collaris</emphasis>
</taxonomicName>
. They can be recognized by the distinctive head markings (Figs 11d, 12d). There is a deep red, Y-shaped mark on the vertex above the antennae, and on the face is a pair of red, concave marks below the toruli. The dorsal surface of the scapes has a broad, red, longitudinal stripe; the frons is white; and the entire gena is red. The wings are broader and more rounded distally than those of
<taxonomicName authorityName="Schneider" authorityYear="1851" box="[724,886,765,792]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="52" pageNumber="24" phylum="Arthropoda" rank="species" species="collaris" subGenus="Neosuarius">
<emphasis box="[724,743,766,792]" italics="true" pageId="52" pageNumber="53">C</emphasis>
. (
<emphasis box="[766,788,766,791]" italics="true" pageId="52" pageNumber="53">N</emphasis>
.)
<emphasis box="[811,886,765,791]" italics="true" pageId="52" pageNumber="53">collaris</emphasis>
</taxonomicName>
; the forewing venation is mostly green, but the costal veinlets, the bases of radial crossveins #18, and the icu crossveins are dark brown.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768132" ID-Zenodo-Dep="3768132" box="[140,855,691,713]" httpUri="https://zenodo.org/record/3768132/files/figure.png" pageId="52" pageNumber="53" startId="52.[140,212,691,713]" targetBox="[287,960,165,650]" targetPageId="52">
<paragraph blockId="52.[140,855,691,713]" box="[140,855,691,713]" pageId="52" pageNumber="53">
<emphasis bold="true" box="[140,251,691,713]" pageId="52" pageNumber="53">Figure 36.</emphasis>
<taxonomicName authorityName="Penny. The" authorityYear="2002" box="[257,554,691,713]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[257,364,691,712]" italics="true" pageId="52" pageNumber="53">Chrysopodes</emphasis>
(
<emphasis box="[377,474,692,713]" italics="true" pageId="52" pageNumber="53">Neosuarius</emphasis>
)
<emphasis box="[488,554,691,712]" italics="true" pageId="52" pageNumber="53">oswaldi</emphasis>
</taxonomicName>
: Forewing, hindwing (Panama).
</paragraph>
</caption>
<paragraph blockId="52.[140,1108,765,1672]" pageId="52" pageNumber="53">
In the male, the arms and bridge of the gonarcus extend widely. The triangular arcessus is supported basally by a pair of dorsal rods that arise from the front of the gonarcal bridge, not from beneath it. The arcessus terminates in a setose, hook-shaped apical beak (
<figureCitation box="[285,371,976,1002]" captionStart="Figure 38" captionStartId="54.[140,212,669,691]" captionTargetBox="[249,1003,184,635]" captionTargetId="figure@54.[237,1010,165,650]" captionTargetPageId="54" captionText="Figure 38. Chrysopodes (Neosuarius) oswaldi: Male genitalia. (a) gonarcus, dorsal (b) gonarcus, lateral (hypandrium internum, not shown). g.b. gonarcal bridge rod internal rod of mediuncus." figureDoi="http://doi.org/10.5281/zenodo.3768136" httpUri="https://zenodo.org/record/3768136/files/figure.png" pageId="52" pageNumber="53">Fig. 38</figureCitation>
). The female terminalia are almost identical to those of
<taxonomicName authorityName="Schneider" authorityYear="1851" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="52" pageNumber="24" phylum="Arthropoda" rank="species" species="collaris" subGenus="Neosuarius">
<emphasis box="[1024,1043,977,1003]" italics="true" pageId="52" pageNumber="53">C</emphasis>
. (
<emphasis box="[1069,1091,977,1002]" italics="true" pageId="52" pageNumber="53">N</emphasis>
.)
<emphasis box="[140,216,1011,1037]" italics="true" pageId="52" pageNumber="53">collaris</emphasis>
</taxonomicName>
; i.e., they are typified by a relatively wide, funnel-shaped spermatheca with a broad ventral invagination and an elongate, curved velum (
<figureCitation box="[808,892,1047,1073]" captionStart="Figure 39" captionStartId="55.[140,212,669,691]" captionTargetBox="[226,1021,176,640]" captionTargetId="figure@55.[219,1028,165,650]" captionTargetPageId="55" captionText="Figure 39. Chrysopodes (Neosuarius) oswaldi: Female genitalia, ventral. b.c. bursa copulatrix b.d. bursal duct b.g. bursal gland inv invagination sg subgenitale sp spermatheca sp.d. spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.3768138" httpUri="https://zenodo.org/record/3768138/files/figure.png" pageId="52" pageNumber="53">Fig. 39</figureCitation>
). The bursa is large and leathery; the spermathecal duct is long and coiled; and the bursal duct is very long and narrow.
</paragraph>
</subSubSection>
<subSubSection lastPageId="56" lastPageNumber="57" pageId="52" pageNumber="53" type="description">
<paragraph blockId="52.[140,1108,765,1672]" pageId="52" pageNumber="53">
<emphasis bold="true" box="[185,335,1152,1178]" pageId="52" pageNumber="53">
<emphasis box="[185,330,1152,1178]" inLineHeading="true" pageId="52" pageNumber="53" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[342,401,1152,1178]" italics="true" pageId="52" pageNumber="53">Head</emphasis>
(Figs 11d, 12d): Width (frontal, including eyes)
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; ratio, head: eye width = 2.22.5:1; distance (straight-line) between tentorial pits
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.5" metricValueMin="4.9" pageId="52" pageNumber="53" unit="mm" value="0.52" valueMax="0.55" valueMin="0.49">0.49 0.55 mm</quantity>
. Distance between antennae ̴
<quantity box="[588,766,1223,1249]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="52" pageNumber="53" unit="mm" value="0.125" valueMax="0.14" valueMin="0.11">0.110.14 mm</quantity>
; length of frons (mid-antenna midway between tentorial pits) ̴
<quantity box="[529,706,1258,1284]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.9" metricValueMin="4.0" pageId="52" pageNumber="53" unit="mm" value="0.445" valueMax="0.49" valueMin="0.4">0.400.49 mm</quantity>
; clypeus ̴
<quantity box="[818,993,1258,1284]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="52" pageNumber="53" unit="mm" value="0.265" valueMax="0.29" valueMin="0.24">0.240.29 mm</quantity>
long. Antenna ̴0.61× length of forewing (
<quantity box="[537,633,1293,1320]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="52" pageNumber="53" unit="mm" value="8.5">8.5 mm</quantity>
; n=1). Vertex slightly raised, flat, round- ed throughout, with very small, upward fold posteriorly; surface of vertex smooth, without setae. Frons relatively flat, unsculptured; surface smooth, shiny throughout. Clypeus unsculptured; surface smooth, mostly flat, very slightly raised in middle. Labrum flat; surface smooth; distal margin straight.
</paragraph>
<paragraph blockId="52.[140,1108,765,1672]" pageId="52" pageNumber="53">
<emphasis box="[185,360,1469,1495]" italics="true" pageId="52" pageNumber="53">Head coloration</emphasis>
: Head cream with deep red marks; vertex with single, V-shaped mark extending along posteromesal margin of toruli, part way around lateral edge of vertex; torulus cream colored, unmarked; posterolateral region unmarked. Genae red throughout; frons creamy white, with pair of deep red, scalloped marks below toruli; clypeus creamy white to amber distally. Scapes creamy white, with broad, longitudinal, red, mesal stripe on dorsum, unmarked frontally; pedicel cream, with brown
</paragraph>
<paragraph blockId="53.[478,504,176,198]" box="[478,504,176,198]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[478,504,176,198]" pageId="53" pageNumber="54">T7</emphasis>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768134" ID-Zenodo-Dep="3768134" httpUri="https://zenodo.org/record/3768134/files/figure.png" pageId="53" pageNumber="54" startId="53.[140,212,809,831]" targetBox="[297,942,190,784]" targetPageId="53">
<paragraph blockId="53.[140,1109,809,927]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[140,250,809,831]" pageId="53" pageNumber="54">Figure 37.</emphasis>
<taxonomicName authorityName="Penny. The" authorityYear="2002" box="[254,549,809,830]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[254,361,809,830]" italics="true" pageId="53" pageNumber="54">Chrysopodes</emphasis>
(
<emphasis box="[374,471,809,830]" italics="true" pageId="53" pageNumber="54">Neosuarius</emphasis>
)
<emphasis box="[483,549,809,830]" italics="true" pageId="53" pageNumber="54">oswaldi</emphasis>
</taxonomicName>
: Male terminus, lateral (membrane at tip of S8+9 expanded).
<emphasis bold="true" box="[140,179,841,863]" pageId="53" pageNumber="54">c.a.</emphasis>
caudal arm of apodeme on T9+ectoproct
<emphasis bold="true" box="[574,613,841,863]" pageId="53" pageNumber="54">c.c.</emphasis>
callus cerci
<emphasis bold="true" box="[727,786,841,863]" pageId="53" pageNumber="54">ev.m.</emphasis>
eversible membrane
<emphasis bold="true" box="[984,1022,841,863]" pageId="53" pageNumber="54">g.c.</emphasis>
gonarcal complex
<emphasis bold="true" box="[225,261,873,895]" pageId="53" pageNumber="54">s.p.</emphasis>
setose subrectal plate
<emphasis bold="true" box="[468,524,873,895]" pageId="53" pageNumber="54">S8+9</emphasis>
fused eighth and ninth sternites
<emphasis bold="true" box="[833,863,873,895]" pageId="53" pageNumber="54">T7</emphasis>
seventh tergite
<emphasis bold="true" box="[1012,1042,873,895]" pageId="53" pageNumber="54">T8</emphasis>
eighth tergite
<emphasis bold="true" box="[204,284,905,927]" pageId="53" pageNumber="54">T9+ect</emphasis>
fused ninth tergite and ectoproct
<emphasis bold="true" box="[600,635,905,927]" pageId="53" pageNumber="54">v.a.</emphasis>
ventral arm of apodeme on T9+ectoproct.
</paragraph>
</caption>
<paragraph blockId="53.[140,1108,976,1672]" pageId="53" pageNumber="54">with small red horizontal stipe laterally; flagellum creamy white throughout. Maxillary palp: segments 35 amber, with brown dorsal mark; basal two segments cream. Labial palp with terminal segment amber to brown, basal segments cream. Venter creamcolored, unmarked.</paragraph>
<paragraph blockId="53.[140,1108,976,1672]" pageId="53" pageNumber="54">
<emphasis box="[185,261,1117,1143]" italics="true" pageId="53" pageNumber="54">Thorax</emphasis>
(Fig. 11d): Pronotum wider than long: ̴
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long; ̴
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.2799999999999998" metricValueMax="1.4" metricValueMin="1.16" pageId="53" pageNumber="54" unit="mm" value="1.2799999999999998" valueMax="1.4" valueMin="1.16">1.161.40 mm</quantity>
wide; cream (probably green in life), unmarked; numerous thin, pale setae (longest ̴
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long). Meso-, metanota cream (probably green in life), probably with middorsal yellow stripe, unmarked; setae thin, pale. Pleural areas cream colored, without markings. Legs cream without markings, setae golden. Tarsal claws amber, long, narrow, with broad cleft, small base with dark mark.
</paragraph>
<paragraph blockId="53.[140,1108,976,1672]" lastBlockId="54.[140,1108,765,1672]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<emphasis box="[185,253,1329,1354]" italics="true" pageId="53" pageNumber="54">Wings</emphasis>
(
<figureCitation box="[271,359,1328,1355]" captionStart="Figure 36" captionStartId="52.[140,212,691,713]" captionTargetBox="[287,960,165,650]" captionTargetId="figure@52.[287,961,165,672]" captionTargetPageId="52" captionText="Figure 36. Chrysopodes (Neosuarius) oswaldi: Forewing, hindwing (Panama)." figureDoi="http://doi.org/10.5281/zenodo.3768132" httpUri="https://zenodo.org/record/3768132/files/figure.png" pageId="53" pageNumber="54">Fig. 36</figureCitation>
): Forewing
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long,
<quantity box="[753,898,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.8" metricValueMin="4.1" pageId="53" pageNumber="54" unit="mm" value="4.449999999999999" valueMax="4.8" valueMin="4.1">4.14.8 mm</quantity>
wide; L:W (midpoint) ratio, 3.03.1; width greatest near midpoint, tapering at basal 1/4th and after distal 3/4th of wing; costal margin fairly straight, sloping gradually at base; apex broad, rounded. Media (before and after furcation of M1 and M2), ma, base of Cu1 (above icu1) crassate. Costal area slightly enlarged; height of tallest costal cell (#57) = 0.180.19× width of wing, 4.56.4× length of first costal vein. Subcosta, radius sinuate; most subcostal veinlets, radial crossveins straight. Twelve closed radial cells (between R and Rs), height of tallest radial cell 1.52.1× width; other than radial crossveins, only gradate veins in contact with PsM. Four b cells, four b cells; First intramedian cell ovate, 0.60.9× width of third medial cell; inner row of five to seven gradate veins; eight to ten outer gradate veins; both rows regularly stepped, parallel to each other distally. Second cell beneath Rs with i.g. at base =
<quantity box="[851,995,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="54" pageNumber="55" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
tall, 2.6 2.9× width; third cell =
<quantity box="[407,549,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="54" pageNumber="55" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
tall, 2.62.9× width. Second gradate cell
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="54" pageNumber="55" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
tall, 3.13.2× width; third gradate cell
<quantity box="[635,777,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="54" pageNumber="55" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
tall, 3.03.2× width. Length of second cell beneath Rs with i.g. at base = 1.1× length of second gradate cell. Three intracubital cells; distal one open, icu1, icu2 each shorter than icu3; icu1 slightly shorter than icu2. Vein 1A forked. Hindwing narrow, with tip angulate;
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.26" metricValueMax="1.31" metricValueMin="1.21" pageId="54" pageNumber="55" unit="mm" value="12.6" valueMax="13.1" valueMin="12.1">12.113.1 mm</quantity>
long, 3.74.0 mm wide. Eleven to twelve radial crossveins; four to five inner gradates; seven to eight outer gradate veins; three b cells, plus small t cell; four 4 b cells; two intracubital cells, distal one open.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768136" ID-Zenodo-Dep="3768136" httpUri="https://zenodo.org/record/3768136/files/figure.png" pageId="54" pageNumber="55" startId="54.[140,212,669,691]" targetBox="[249,1003,184,635]" targetPageId="54">
<paragraph blockId="54.[140,1107,669,723]" pageId="54" pageNumber="55">
<emphasis bold="true" box="[140,252,669,691]" pageId="54" pageNumber="55">Figure 38.</emphasis>
<taxonomicName authorityName="Penny. The" authorityYear="2002" box="[260,560,669,691]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[260,367,669,690]" italics="true" pageId="54" pageNumber="55">Chrysopodes</emphasis>
(
<emphasis box="[382,479,670,691]" italics="true" pageId="54" pageNumber="55">Neosuarius</emphasis>
)
<emphasis box="[494,560,669,690]" italics="true" pageId="54" pageNumber="55">oswaldi</emphasis>
</taxonomicName>
: Male genitalia.
<emphasis bold="true" box="[719,750,669,691]" pageId="54" pageNumber="55">(a)</emphasis>
gonarcus, dorsal
<emphasis bold="true" box="[916,948,669,691]" pageId="54" pageNumber="55">(b)</emphasis>
gonarcus, lateral (hypandrium internum, not shown).
<emphasis bold="true" box="[485,524,701,723]" pageId="54" pageNumber="55">g.b.</emphasis>
gonarcal bridge
<emphasis bold="true" box="[679,717,701,723]" pageId="54" pageNumber="55">rod</emphasis>
internal rod of mediuncus.
</paragraph>
</caption>
<paragraph blockId="54.[140,1108,765,1672]" pageId="54" pageNumber="55">
<emphasis box="[185,357,1117,1143]" italics="true" pageId="54" pageNumber="55">Wing coloration</emphasis>
: Hyaline; stigma tinged with brown. Longitudinal veins green, unmarked; costal veinlets #510 brownish; tip of im1 lower vein, rs-m crossvein brown or partially brown; gradates with tinge of brown; base of
<taxonomicName box="[790,919,1187,1214]" pageId="54" pageNumber="55" rank="subSpecies" subSpecies="crossveins">r crossveins</taxonomicName>
#18, icu crossvein #1 dark brown.
</paragraph>
<paragraph blockId="54.[140,1108,765,1672]" pageId="54" pageNumber="55">
<emphasis box="[185,287,1258,1284]" italics="true" pageId="54" pageNumber="55">Abdomen</emphasis>
(
<figureCitation box="[304,390,1258,1284]" captionStart="Figure 37" captionStartId="53.[140,212,809,831]" captionTargetBox="[297,942,190,784]" captionTargetId="figure@53.[296,952,165,789]" captionTargetPageId="53" captionText="Figure 37. Chrysopodes (Neosuarius) oswaldi: Male terminus, lateral (membrane at tip of S8+9 expanded). c.a. caudal arm of apodeme on T9+ectoproct c.c. callus cerci ev.m. eversible membrane g.c. gonarcal complex s.p. setose subrectal plate S8+9 fused eighth and ninth sternites T7 seventh tergite T8 eighth tergite T9+ect fused ninth tergite and ectoproct v.a. ventral arm of apodeme on T9+ectoproct." figureDoi="http://doi.org/10.5281/zenodo.3768134" httpUri="https://zenodo.org/record/3768134/files/figure.png" pageId="54" pageNumber="55">Fig. 37</figureCitation>
): Tergites 6, 7: roughly quadrate, with convex ventral margins, straight basal margins, rounded distal margins; length 2.42.7 times greater than height (lateral view); with sparse, long, thin setae. Pleural region P7 with very few, medium-length, thin, setae. Sternite 6 quadrate, with dorsal margin relatively straight; length 0.900.95× height; with long, slender setae. Spiracles oval, not enlarged (̴
<quantity metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="54" pageNumber="55" unit="mm" value="0.07">0. 07 mm</quantity>
diameter); atria not enlarged.
</paragraph>
<paragraph blockId="54.[140,1108,765,1672]" lastBlockId="55.[140,1108,765,1672]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis box="[185,242,1469,1495]" italics="true" pageId="54" pageNumber="55">Male</emphasis>
(
<figureCitation box="[261,353,1469,1495]" captionStart="Figure 37" captionStartId="53.[140,212,809,831]" captionTargetBox="[297,942,190,784]" captionTargetId="figure@53.[296,952,165,789]" captionTargetPageId="53" captionText="Figure 37. Chrysopodes (Neosuarius) oswaldi: Male terminus, lateral (membrane at tip of S8+9 expanded). c.a. caudal arm of apodeme on T9+ectoproct c.c. callus cerci ev.m. eversible membrane g.c. gonarcal complex s.p. setose subrectal plate S8+9 fused eighth and ninth sternites T7 seventh tergite T8 eighth tergite T9+ect fused ninth tergite and ectoproct v.a. ventral arm of apodeme on T9+ectoproct." figureDoi="http://doi.org/10.5281/zenodo.3768134" httpUri="https://zenodo.org/record/3768134/files/figure.png" pageId="54" pageNumber="55">Figs 37</figureCitation>
,
<figureCitation box="[368,402,1469,1495]" captionStart="Figure 38" captionStartId="54.[140,212,669,691]" captionTargetBox="[249,1003,184,635]" captionTargetId="figure@54.[237,1010,165,650]" captionTargetPageId="54" captionText="Figure 38. Chrysopodes (Neosuarius) oswaldi: Male genitalia. (a) gonarcus, dorsal (b) gonarcus, lateral (hypandrium internum, not shown). g.b. gonarcal bridge rod internal rod of mediuncus." figureDoi="http://doi.org/10.5281/zenodo.3768136" httpUri="https://zenodo.org/record/3768136/files/figure.png" pageId="54" pageNumber="55">38</figureCitation>
): T8 rounded anteriorly, posteriorly (lateral view), with setose ventrolateral extension reaching below spiracle. Left, right T9+ectoproct fused dorsally; terminal edge straight vertically, bearing long, thin setae; with distinct apodeme along ventral margin, apodeme straight basally, with knob and ventral arm subapically, caudal arm bent downward, with arrow-like apical process; area around callus cerci weakly sclerotized. Callus cerci oblong (
<quantity box="[747,925,1645,1672]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="54" pageNumber="55" unit="mm" value="0.25" valueMax="0.28" valueMin="0.22">0.220.28 mm</quantity>
tall,
<quantity lastPageId="55" lastPageNumber="56" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="54" pageNumber="55" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.150.17 mm</quantity>
wide), with ̴30 short, thin trichobothria. S8+9 length ̴1.25× proximal height; dorsal margin sloping throughout; ventral margin slightly bent upward, without knob, invagination, or apodeme; terminus short, blunt, with distal margin bearing long, heavy setae on somewhat enlarged setal bases, heavy membrane covered with dense field of small gonocristae. Subanal plate distinct, with dense, robust setae. Gonarcus broadly arcuate, tightly attached to T9+ectoproct dorsally via short, folded subanal membrane; broadly V-shaped in frontal view, straight in lateral view; with apodemes attached distally on bridge; bridge smooth, rod-like; span of gonarcus near arch ̴
<quantity box="[263,376,1047,1073]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="56" unit="mm" value="0.6">0.60 mm</quantity>
, distance between gonarcal apodemes distally ̴1.0 mm. Gonarcal apodemes round, smooth. Mediuncus membranous basally, with pair of robust, sclerotized rods internally, extending from below distal margin of gonarcal bridge, curving upward, approaching each other, fading distally; apical section of mediuncus rounded, dome-like dorsally; terminus bent perpendicularly into blunt, ventrally projecting beak; dorsal surface with dense covering of microsetae; membrane below mediuncus rounded, gathered upward, apparently with large, dense field of robust gonocristae. Gonosaccus beneath mediuncus delicate, thin, without gonosetae or much structure. Hypandrium internum expanded in dorsal view, flat in lateral view, with arms rounded, meeting at apex in smooth, round surface, with narrow, hooklike combs, not extending beyond arms.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768138" ID-Zenodo-Dep="3768138" httpUri="https://zenodo.org/record/3768138/files/figure.png" pageId="55" pageNumber="56" startId="55.[140,212,669,691]" targetBox="[226,1021,176,640]" targetPageId="55">
<paragraph blockId="55.[140,1108,669,723]" pageId="55" pageNumber="56">
<emphasis bold="true" box="[140,251,669,691]" pageId="55" pageNumber="56">Figure 39.</emphasis>
<taxonomicName authorityName="Penny. The" authorityYear="2002" box="[257,555,669,691]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis box="[257,364,669,690]" italics="true" pageId="55" pageNumber="56">Chrysopodes</emphasis>
(
<emphasis box="[378,475,670,691]" italics="true" pageId="55" pageNumber="56">Neosuarius</emphasis>
)
<emphasis box="[489,555,669,690]" italics="true" pageId="55" pageNumber="56">oswaldi</emphasis>
</taxonomicName>
: Female genitalia, ventral.
<emphasis bold="true" box="[804,843,669,691]" pageId="55" pageNumber="56">b.c.</emphasis>
bursa copulatrix
<emphasis bold="true" box="[1005,1046,669,691]" pageId="55" pageNumber="56">b.d.</emphasis>
bursal duct
<emphasis bold="true" box="[187,227,701,723]" pageId="55" pageNumber="56">b.g.</emphasis>
bursal gland
<emphasis bold="true" box="[352,384,701,723]" pageId="55" pageNumber="56">inv</emphasis>
invagination
<emphasis bold="true" box="[512,535,701,723]" pageId="55" pageNumber="56">sg</emphasis>
subgenitale
<emphasis bold="true" box="[650,674,701,723]" pageId="55" pageNumber="56">sp</emphasis>
spermatheca
<emphasis bold="true" box="[801,852,701,723]" pageId="55" pageNumber="56">sp.d.</emphasis>
spermathecal duct.
</paragraph>
</caption>
<paragraph blockId="55.[140,1108,765,1672]" lastBlockId="56.[140,1108,167,1355]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
<emphasis box="[185,264,1434,1460]" italics="true" pageId="55" pageNumber="56">Female</emphasis>
(
<figureCitation box="[282,371,1434,1460]" captionStart="Figure 39" captionStartId="55.[140,212,669,691]" captionTargetBox="[226,1021,176,640]" captionTargetId="figure@55.[219,1028,165,650]" captionTargetPageId="55" captionText="Figure 39. Chrysopodes (Neosuarius) oswaldi: Female genitalia, ventral. b.c. bursa copulatrix b.d. bursal duct b.g. bursal gland inv invagination sg subgenitale sp spermatheca sp.d. spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.3768138" httpUri="https://zenodo.org/record/3768138/files/figure.png" pageId="55" pageNumber="56">Fig. 39</figureCitation>
): Very similar to
<taxonomicName authorityName="Schneider" authorityYear="1851" box="[577,750,1434,1460]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="55" pageNumber="24" phylum="Arthropoda" rank="species" species="collaris" subGenus="Neosuarius">
<emphasis box="[577,596,1434,1460]" italics="true" pageId="55" pageNumber="56">C</emphasis>
. (
<emphasis box="[622,644,1435,1460]" italics="true" pageId="55" pageNumber="56">N</emphasis>
.)
<emphasis box="[671,750,1434,1460]" italics="true" pageId="55" pageNumber="56">collaris</emphasis>
</taxonomicName>
(see Figs 18, 19). Callus cerci very slightly taller than broad (
<quantity box="[507,617,1469,1496]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="55" pageNumber="56" unit="mm" value="0.13">0.13 mm</quantity>
tall,
<quantity box="[680,789,1469,1496]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="55" pageNumber="56" unit="mm" value="0.12">0.12 mm</quantity>
wide), with approximately 24 trichobothria. Gonapophyses laterales fairly tall, approximately 2× taller than wide, rounded dorsally, ventrally. Subgenitale broad, thick, fleshy, with short, broad neck, very small ventrally projecting process; base with thick, textured, folded membrane, attached to S7 via leathery, textured, invaginated membrane. Pair of large, bulbous bursal glands, with elongate ducts opening on dorsolateral margin of bursa, without accessory ducts. Bursa fairly large, extending laterally full width of segment, posteriorly beyond spermatheca; membrane leathery basally, with numerous, small, transverse folds, bursal section becoming thinner distally, folds becoming longitudinal. Bursal duct elongate, much longer than length of abdomen); section arising from bursa flat, fluted, becoming flat, ribbon-like, coiled, surface with small transverse folds (bursal section somewhat longer than 1/2 length of bursal duct); section arising from spermatheca tubular, with smooth surface, initially straight, becoming coiled, bent (spermathecal section slightly less than 1/2 length of bursal duct). Spermatheca funnel-shaped, with large mesal, invaginated velum that makes two full bends, tapers into bursal duct; ̴
<quantity box="[748,860,483,510]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="56" pageNumber="57" unit="mm" value="0.58">0.58 mm</quantity>
diameter, ̴
<quantity box="[996,1108,483,510]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="56" pageNumber="57" unit="mm" value="0.53">0.53 mm</quantity>
in length to first bend; invagination elongate, tapered, extending through first two bends, perhaps into bursal duct, as internal duct (total length ̴
<quantity box="[876,953,554,580]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="mm" value="2.0">2 mm</quantity>
). Spermathecal duct ̴
<quantity box="[251,323,589,616]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="56" pageNumber="57" unit="mm" value="4.0">4 mm</quantity>
long, narrow, arising from posterior surface of spermatheca slightly on left side, with full coil within subgenitale; basal 1/3rd sclerotized, brownish; distal 2/3rd lightly sclerotized, brushy, very pale (only visible under compound microscope), becoming enlarged distally. Colleterial gland relatively short, bulbous, probably not extending into seventh segment, with slightly grainy texture; no reservoir or accessory glands found. Transverse sclerotization well-formed, curved, ellipsoid plate, located ventrally within gonapophyses laterales, with longitudinal (to the body) rows of setae.
</paragraph>
<paragraph blockId="56.[140,1108,167,1355]" box="[185,397,871,897]" pageId="56" pageNumber="57">
<emphasis box="[185,260,871,896]" italics="true" pageId="56" pageNumber="57">Larvae</emphasis>
. Unknown.
</paragraph>
<paragraph blockId="56.[140,1108,167,1355]" box="[185,368,906,932]" pageId="56" pageNumber="57">
<emphasis box="[185,231,907,932]" italics="true" pageId="56" pageNumber="57">Eggs</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="56" pageNumber="57" type="biology_ecology">
<paragraph blockId="56.[140,1108,167,1355]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[185,283,941,967]" pageId="56" pageNumber="57">
<emphasis box="[185,278,941,967]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Biology</emphasis>
.
</emphasis>
Unknown. The only known adults were collected in January, March, August and December.
</paragraph>
</subSubSection>
<subSubSection pageId="56" pageNumber="57" type="materials_examined">
<paragraph blockId="56.[140,1108,167,1355]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[185,350,1011,1037]" pageId="56" pageNumber="57">
<emphasis box="[185,243,1011,1037]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Type</emphasis>
material
</emphasis>
.
<taxonomicName authority="Penny. The" authorityName="Penny. The" authorityYear="2002" box="[366,912,1011,1038]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="oswaldi" subGenus="Neosuarius">
<emphasis bold="true" box="[366,861,1011,1038]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[366,773,1011,1038]" italics="true" pageId="56" pageNumber="57">Chrysopodes (Neosuarius) oswaldi</emphasis>
Penny.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608859722" box="[869,912,1011,1037]" collectionCode="CAS" collectorName="J. K. Sheldon Field" determinerName="Penny" location="Rincon" pageId="56" pageNumber="57" specimenCount="2" stateProvince="Rincon" typeStatus="holotype">The</materialsCitation>
</taxonomicName>
unique type (examined), the
<typeStatus box="[298,405,1047,1073]" pageId="56" pageNumber="57">Holotype</typeStatus>
(by original designation), is in the
<collectionCode box="[827,883,1047,1073]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="56" pageNumber="57" type="Museum">CAS</collectionCode>
; its labels read: (1) “
<quantity box="[151,219,1082,1108]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="56" pageNumber="57" unit="mi" value="10.0">10mi.</quantity>
S.
<collectingRegion box="[263,346,1082,1108]" country="Puerto Rico" name="Rincon" pageId="56" pageNumber="57">Rincon</collectingRegion>
de Osa /
<collectingRegion box="[471,707,1082,1108]" country="Costa Rica" name="Puntarenas" pageId="56" pageNumber="57">Puntarenas Province</collectingRegion>
, Costa / Rica, 412.III.69 /
<collectorName pageId="56" pageNumber="57">J. K. Sheldon Field</collectorName>
#”; (2) “
<typeStatus box="[395,552,1117,1143]" pageId="56" pageNumber="57">HOLOTYPE</typeStatus>
/
<taxonomicName authorityName="Navas" authorityYear="1913" box="[576,724,1117,1143]" class="Insecta" family="Chrysopidae" genus="Chrysopodes" kingdom="Animalia" order="Neuroptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">Chrysopodes</taxonomicName>
/ oswaldi / det.
<determinerName box="[908,979,1118,1144]" pageId="56" pageNumber="57">Penny</determinerName>
, 02” [red]. One pair of wings is attached to a tab below the labels.
</paragraph>
<paragraph blockId="56.[140,1108,167,1355]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[185,435,1187,1213]" pageId="56" pageNumber="57">
<emphasis box="[185,311,1187,1213]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Specimens</emphasis>
examined
</emphasis>
(in addition to the
<typeStatus box="[670,718,1188,1214]" pageId="56" pageNumber="57">type</typeStatus>
specimen above).
<emphasis bold="true" box="[934,1041,1187,1213]" pageId="56" pageNumber="57">
<collectingCountry box="[934,1036,1187,1213]" name="Belize" pageId="56" pageNumber="57">BELIZE</collectingCountry>
.
</emphasis>
Twin Cays, I/5/89, red mangrove (1M,
<collectionCode box="[517,608,1223,1249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="56" pageNumber="57" type="Museum">USNM</collectionCode>
).
<emphasis bold="true" box="[626,948,1222,1249]" pageId="56" pageNumber="57">
<collectingCountry box="[626,796,1222,1248]" name="Costa Rica" pageId="56" pageNumber="57">COSTA RICA</collectingCountry>
.
<collectingRegion box="[807,943,1223,1249]" country="Costa Rica" name="Puntarenas" pageId="56" pageNumber="57">Puntarenas</collectingRegion>
:
</emphasis>
Rio Barranca,
<quantity box="[140,217,1258,1284]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="56" pageNumber="57" unit="km" value="11.0">11 km</quantity>
. E. Puntarenas,
<date box="[396,540,1258,1284]" pageId="56" pageNumber="57" value="1964-08-07">VIII/7/1964</date>
, G. C. Eickwort (1M,
<collectionCode box="[796,842,1258,1284]" country="USA" name="Biodiversity Institute, University of Kansas" pageId="56" pageNumber="57" type="University or college">KU</collectionCode>
).
<emphasis bold="true" box="[861,987,1257,1283]" pageId="56" pageNumber="57">
<collectingCountry box="[861,982,1257,1283]" name="Panama" pageId="56" pageNumber="57">PANAMA</collectingCountry>
.
</emphasis>
<date box="[993,1103,1258,1285]" bridgedPair="/" pageId="56" pageNumber="57" value="1935-12">XII/1935</date>
, No. 14,735 (1F,
<collectionCode box="[329,420,1293,1319]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="56" pageNumber="57" type="Museum">USNM</collectionCode>
).
</paragraph>
<paragraph blockId="56.[140,1108,167,1355]" box="[185,966,1328,1355]" pageId="56" pageNumber="57">
<emphasis bold="true" box="[185,424,1328,1354]" pageId="56" pageNumber="57">
<emphasis box="[185,273,1328,1354]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Known</emphasis>
distribution
</emphasis>
. Central America (
<collectingCountry box="[640,712,1328,1354]" name="Belize" pageId="56" pageNumber="57">Belize</collectingCountry>
,
<collectingCountry box="[723,848,1328,1354]" name="Costa Rica" pageId="56" pageNumber="57">Costa Rica</collectingCountry>
,
<collectingCountry box="[860,955,1329,1355]" name="Panama" pageId="56" pageNumber="57">Panama</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>