362 lines
42 KiB
XML
362 lines
42 KiB
XML
<document id="26F3DEFE65846A883B82DDED5D816D00" ID-DOI="10.11646/zootaxa.3635.5.3" ID-GBIF-Dataset="012c682c-6953-4e8e-a3fe-8af53403d142" ID-ISSN="1175-5326" ID-Zenodo-Dep="220591" ID-ZooBank="40372BBF-2235-4770-AEB8-41E342DFF770" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460587444626" checkinUser="plazi" docAuthor="Mikhaljova, Elena V., Burkitbaeva, Ulzhan D., Tuf, Ivan H. & Ulykpan, Kaman" docDate="2013" docId="03AC87CDFFC7DC73FF52FB0305B206A3" docLanguage="en" docName="zt03635p544.pdf" docOrigin="Zootaxa 3635 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tarbagataya zaisanica Mikhaljova, sp. nov." docType="treatment" docVersion="8" lastPageNumber="542" masterDocId="FF95FFB5FFC2DC7AFFC5FF9C046F024B" masterDocTitle="The millipede order Chordeumatida (Diplopoda) in Kazakhstan, with descriptions of three new species" masterLastPageNumber="544" masterPageNumber="533" pageNumber="538" updateTime="1698325424133" updateUser="plazi">
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<mods:titleInfo id="2E38C2735A43CA39038CA499A8FBF726">
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<mods:title id="A5F2314D0898A07C15389F631BCA3DA6">The millipede order Chordeumatida (Diplopoda) in Kazakhstan, with descriptions of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="39DABA044E5AF7686304A2B9AEA47347" type="personal">
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<mods:namePart id="D8D089D7D170BEEB26DB0A54DFAE6A45">Mikhaljova, Elena V.</mods:namePart>
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<mods:roleTerm id="1AC102928FD5BF99FC02FF99F71AD34F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AC1BE140148BEE6EC6C21F9DB2A6859C">Burkitbaeva, Ulzhan D.</mods:namePart>
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</mods:role>
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<mods:namePart id="A1F10482F7EC4E5AA9D6D418E7166348">Tuf, Ivan H.</mods:namePart>
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</mods:name>
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<mods:name id="245BC957C500A8B2412F0B7D419706CA" type="personal">
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<mods:roleTerm id="ED7B0BA94C01E4EE7A2DDEC110F2B471">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0A815F6646F9C6B39B7EB96F6E10F9D6">Ulykpan, Kaman</mods:namePart>
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<mods:typeOfResource id="A6758BCABE15C02C10702E82FB1266C1">text</mods:typeOfResource>
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<mods:title id="D1A3E31BCAA04E0D4AD77BD6481FE4BE">Zootaxa</mods:title>
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<mods:part id="E607F79B57C8BC8166AC6EF06FCDB525">
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<mods:date id="07451FB6ABE9B40BCCEE71E9954F2E72">2013</mods:date>
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<mods:number id="21759DC1FE85C86F3CF020B923140F6B">3635</mods:number>
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<mods:detail id="A68C31458C919003DD0F1A9B7EC70014" type="issue">
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<mods:classification id="63925679E063370C5FB5FB8BBED7A1C9">journal article</mods:classification>
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<mods:identifier id="A82A537B1E67593DBE9845922B50BBCD" type="DOI">10.11646/zootaxa.3635.5.3</mods:identifier>
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<treatment id="03AC87CDFFC7DC73FF52FB0305B206A3" ID-DOI="http://doi.org/10.5281/zenodo.6145106" ID-GBIF-Taxon="119565218" ID-Zenodo-Dep="6145106" LSID="urn:lsid:plazi:treatment:03AC87CDFFC7DC73FF52FB0305B206A3" httpUri="http://treatment.plazi.org/id/03AC87CDFFC7DC73FF52FB0305B206A3" lastPageId="9" lastPageNumber="542" pageId="5" pageNumber="538">
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<subSubSection id="C31F6550FFC7DC7FFF52FB03057B0691" pageId="5" pageNumber="538" type="nomenclature">
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<paragraph id="8BBA36DBFFC7DC7FFF52FB0306CE06F2" blockId="5.[151,673,1183,1242]" box="[151,673,1183,1209]" pageId="5" pageNumber="538">
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<heading id="D0F281B7FFC7DC7FFF52FB0306CE06F2" bold="true" box="[151,673,1183,1209]" fontSize="11" level="1" pageId="5" pageNumber="538" reason="1">
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<emphasis id="B971EAC9FFC7DC7FFF52FB0306CE06F2" bold="true" box="[151,673,1183,1209]" pageId="5" pageNumber="538">
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<taxonomicName id="4C054D58FFC7DC7FFF52FB03065306F2" authority="Mikhaljova" authorityName="Mikhaljova" box="[151,572,1183,1209]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
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<emphasis id="B971EAC9FFC7DC7FFF52FB0305CB06F2" bold="true" box="[151,420,1183,1209]" italics="true" pageId="5" pageNumber="538">Tarbagataya zaisanica</emphasis>
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Mikhaljova
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</taxonomicName>
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<taxonomicNameLabel id="A24257B2FFC7DC7FFD81FB0306CE06F2" box="[580,673,1183,1209]" pageId="5" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36DBFFC7DC7FFF52FB5E057B0691" blockId="5.[151,673,1183,1242]" box="[151,276,1218,1242]" pageId="5" pageNumber="538">
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<figureCitation id="133E2A5EFFC7DC7FFF52FB5E057B0691" box="[151,276,1218,1242]" captionStart-0="FIGURES 10 – 14" captionStart-1="FIGURES 15 – 16" captionStart-2="FIGURES 17 – 23" captionStartId-0="6.[151,264,1892,1915]" captionStartId-1="7.[151,264,1954,1977]" captionStartId-2="8.[151,264,1896,1919]" captionTargetBox-0="[165,1424,409,1859]" captionTargetBox-1="[301,1275,212,1913]" captionTargetBox-2="[199,1382,213,1869]" captionTargetId-0="figure@6.[165,1425,409,1876]" captionTargetId-1="figure@7.[280,1286,193,1932]" captionTargetId-2="figure@8.[187,1387,193,1872]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 10 – 14. Tarbagataya zaisanica sp. nov., male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod." captionText-1="FIGURES 15 – 16. Tarbagataya zaisanica sp. nov., female paratype. 15, vulva, ventral view; 16, vulva, lateral view." captionText-2="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri-0="https://zenodo.org/record/220595/files/figure.png" httpUri-1="https://zenodo.org/record/220596/files/figure.png" httpUri-2="https://zenodo.org/record/220597/files/figure.png" pageId="5" pageNumber="538">Figs 10–23</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F6550FFC7DC7FFF52FA96062104E5" pageId="5" pageNumber="538" type="materials_examined">
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<emphasis id="B971EAC9FFC7DC7FFF52FA9605EB0768" bold="true" box="[151,388,1290,1315]" pageId="5" pageNumber="538">Material examined.</emphasis>
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<emphasis id="B971EAC9FFC7DC7FFE4AFA97059A0768" box="[399,501,1291,1315]" italics="true" pageId="5" pageNumber="538">Holotype</emphasis>
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</typeStatus>
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:
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(IBSS),
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<collectingCountry id="F312764BFFC7DC7FFD07FA9607280769" box="[706,839,1290,1314]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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a.s.l.,
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<date id="FFBB101BFFC7DC7FFE80FAB2058C070D" box="[325,483,1326,1350]" pageId="5" pageNumber="538" value="2011-07-12">12 July 2011</date>
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, leg. K. Ulykpan and U.D. Burkitbaeva.
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<emphasis id="B971EAC9FFC7DC7FFC14FAAC002C070C" box="[977,1091,1328,1351]" italics="true" pageId="5" pageNumber="538">Paratypes</emphasis>
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:
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,
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(IBSS),
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<collectingCountry id="F312764BFFC7DC7FFF52FACE05730721" box="[151,284,1362,1386]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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<quantity id="4CFD9B3EFFC7DC7FFCB5FACE07A80721" box="[880,967,1362,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.755" pageId="5" pageNumber="538" unit="m" value="1755.0">1755 m</quantity>
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a.s.l.,
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;
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,
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(IBSS),
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<collectingCountry id="F312764BFFC7DC7FFEE7FAEA05C807C5" box="[290,423,1398,1422]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
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, Vostochno-Kazakhstanskaya Area, Zaisan District, forest,
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<date id="FFBB101BFFC7DC7FFB85FAEA00AB07C5" box="[1088,1220,1398,1422]" pageId="5" pageNumber="538" value="2011-07-08">8 July 2011</date>
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;
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,
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(IBSS),
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<collectingCountry id="F312764BFFC7DC7FFEB4FA06059807F9" box="[369,503,1434,1458]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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<quantity id="4CFD9B3EFFC7DC7FFB95FA0600C807F9" box="[1104,1191,1434,1458]" metricMagnitude="3" metricUnit="m" metricValue="1.848" pageId="5" pageNumber="538" unit="m" value="1848.0">1848 m</quantity>
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a.s.l.,
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;
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(ZMUM),
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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a.s.l.,
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;
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,
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(IBSS),
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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a.s.l.,
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<date id="FFBB101BFFC7DC7FFF52F99A05450455" box="[151,298,1542,1566]" pageId="5" pageNumber="538" value="2011-07-11">11 July 2011</date>
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;
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,
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(IBSS), together with
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,
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;
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,
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,
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(PU),
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<collectingCountry id="F312764BFFC7DC7FFF52F9B605730409" box="[151,284,1578,1602]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
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<quantity id="4CFD9B3EFFC7DC7FFC46F9B607B20409" box="[899,989,1578,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="5" pageNumber="538" unit="m" value="1680.0">1680 m</quantity>
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a.s.l.,
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;
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(IBSS),
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, Vostochno-Kazakhstanskaya Area, Zaisan District,
|
||
<quantity id="4CFD9B3EFFC7DC7FFCAEF9D207AE042D" box="[875,961,1614,1638]" metricMagnitude="3" metricUnit="m" metricValue="1.6619999999999997" pageId="5" pageNumber="538" unit="m" value="1662.0">1662 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFBB101BFFC7DC7FFBCEF9D200F0042D" box="[1035,1183,1614,1638]" pageId="5" pageNumber="538" value="2011-07-13">13 July 2011</date>
|
||
;
|
||
<specimenCount id="9D03FD52FFC7DC7FFB6EF9D20096042D" box="[1195,1273,1614,1638]" pageId="5" pageNumber="538" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD52FFC7DC7FFAC3F9D201ED042D" box="[1286,1410,1614,1638]" pageId="5" pageNumber="538" type="female">11 females</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD52FFC7DC7FFA4BF9D2049304C1" pageId="5" pageNumber="538" type="juvenile">4 juveniles</specimenCount>
|
||
(IBSS),
|
||
<collectingCountry id="F312764BFFC7DC7FFEA7F9EE058804C1" box="[354,487,1650,1674]" name="Kazakhstan" pageId="5" pageNumber="538">Kazakhstan</collectingCountry>
|
||
, Vostochno-Kazakhstanskaya Area, Zaisan District,
|
||
<quantity id="4CFD9B3EFFC7DC7FFBFFF9EE00FE04C1" box="[1082,1169,1650,1674]" metricMagnitude="3" metricUnit="m" metricValue="1.735" pageId="5" pageNumber="538" unit="m" value="1735.0">1735 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFBB101BFFC7DC7FFB18F9EE011C04C1" box="[1245,1395,1650,1674]" pageId="5" pageNumber="538" value="2011-07-17">17 July 2011</date>
|
||
; all leg. K. Ulykpan and U.D. Burkitbaeva.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F6550FFC7DC7FFF0CF92606480575" pageId="5" pageNumber="538" type="diagnosis">
|
||
<paragraph id="8BBA36DBFFC7DC7FFF0CF92606480575" blockId="5.[151,1436,1290,2034]" pageId="5" pageNumber="538">
|
||
<emphasis id="B971EAC9FFC7DC7FFF0CF926052C0498" bold="true" box="[201,323,1722,1747]" pageId="5" pageNumber="538">Diagnosis.</emphasis>
|
||
Differs from its only congener
|
||
<taxonomicName id="4C054D58FFC7DC7FFD6AF92700990499" authority="Golovatch & Wytwer, 2003" authorityName="Golovatch & Wytwer" authorityYear="2003" box="[687,1270,1722,1747]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="B971EAC9FFC7DC7FFD6AF92707DD0498" box="[687,946,1723,1747]" italics="true" pageId="5" pageNumber="538">Tarbagataya splendida</emphasis>
|
||
Golovatch & Wytwer, 2003
|
||
</taxonomicName>
|
||
mainly by the structure of the posterior gonopods, with the coxal process supplied with three teeth, the structure of leg pair 7 with tongue-shaped, long, apically bilobate sternal protuberance, the structure of the anterior gonopods as well as the presence of tarsal papillae in males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F6550FFC7DC73FF0CF8D607730644" lastPageId="9" lastPageNumber="542" pageId="5" pageNumber="538" type="description">
|
||
<paragraph id="8BBA36DBFFC7DC7FFF0CF8D6073F05B9" blockId="5.[151,1436,1290,2034]" pageId="5" pageNumber="538">
|
||
<emphasis id="B971EAC9FFC7DC7FFF0CF8D605340528" bold="true" box="[201,347,1866,1891]" pageId="5" pageNumber="538">Description.</emphasis>
|
||
Male. Length
|
||
<quantity id="4CFD9B3EFFC7DC7FFDC1F8D606EE0529" box="[516,641,1866,1890]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="5" pageNumber="538" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
|
||
, width of midbody metazona 0.7–0.8 mm. Coloration in alcohol light brown or brown with marbled, annulated pattern due to pale stricture between pro- and metazona as well as with light marbled spots on sides; this spots gradually decrease towards the telson. Ventral side pale. Clypeolabral region of head with pattern of small light spots of different shapes. Antennae brown, marbled brown distally. Legs pale with marbled light brown distal portions. Ocellaria black.
|
||
</paragraph>
|
||
<paragraph id="8BBA36DBFFC4DC7CFF0CFF0B0740032F" blockId="6.[151,1436,151,356]" pageId="6" pageNumber="539">
|
||
Body with 28 segments. Head covered with short and relatively long setae. Clypeolabral region slightly convex. Eye patches triangular, each composed of at least 18 ocelli. Antennae
|
||
<emphasis id="B971EAC9FFC4DC7CFBCBFF21003B029F" box="[1038,1108,189,212]" italics="true" pageId="6" pageNumber="539">in situ</emphasis>
|
||
reaching to posterior margin of body segment 4. Collum semi-circular. Body juloid, subcylindrical, somewhat moniliform. Paraterga missing. Metazona of anterior body segments with low bulges laterally. Metazonital macrochaetae in a transverse row on somites 25–27, like an elongate (to different degrees) triangle on preceding somites. Numerous macrochaetae broken off, remaining ones middle length, pointed apically.
|
||
</paragraph>
|
||
<caption id="DF7A6653FFC4DC7CFF52F8F807DB05F3" httpUri="https://zenodo.org/record/220595/files/figure.png" pageId="6" pageNumber="539" targetBox="[165,1424,409,1859]" targetPageId="6">
|
||
<paragraph id="8BBA36DBFFC4DC7CFF52F8F807DB05F3" blockId="6.[151,1436,1892,1976]" pageId="6" pageNumber="539">
|
||
<emphasis id="B971EAC9FFC4DC7CFF52F8F805470531" bold="true" box="[151,296,1892,1915]" pageId="6" pageNumber="539">FIGURES 10</emphasis>
|
||
–
|
||
<emphasis id="B971EAC9FFC4DC7CFEF0F8F9053B0531" bold="true" box="[309,340,1893,1914]" pageId="6" pageNumber="539">14.</emphasis>
|
||
<taxonomicName id="4C054D58FFC4DC7CFE99F8F9062B0530" box="[348,580,1893,1915]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="539" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
|
||
<emphasis id="B971EAC9FFC4DC7CFE99F8F9062B0530" box="[348,580,1893,1915]" italics="true" pageId="6" pageNumber="539">Tarbagataya zaisanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24257B2FFC4DC7CFD89F8F906F10531" box="[588,670,1893,1914]" pageId="6" pageNumber="539" rank="species">
|
||
<emphasis id="B971EAC9FFC4DC7CFD89F8F906F10531" bold="true" box="[588,670,1893,1914]" pageId="6" pageNumber="539">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view;
|
||
<emphasis id="B971EAC9FFC4DC7CFB00F818008705D2" bold="true" box="[1221,1256,1924,1945]" pageId="6" pageNumber="539">mb</emphasis>
|
||
, mesal branch of coxite;
|
||
<emphasis id="B971EAC9FFC4DC7CFF27F83E049805FC" bold="true" box="[226,247,1954,1975]" pageId="6" pageNumber="539">lb</emphasis>
|
||
, lateral branch of coxite;
|
||
<emphasis id="B971EAC9FFC4DC7CFE3EF83E066A05FC" bold="true" box="[507,517,1954,1975]" pageId="6" pageNumber="539">s</emphasis>
|
||
, saculiform structure of anterior gonopod.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A6653FFC5DC7DFF52F83E015E05F3" box="[151,1329,1954,1977]" httpUri="https://zenodo.org/record/220596/files/figure.png" pageId="7" pageNumber="540" targetBox="[301,1275,212,1913]" targetPageId="7">
|
||
<paragraph id="8BBA36DBFFC5DC7DFF52F83E015E05F3" blockId="7.[151,1329,1954,1977]" box="[151,1329,1954,1977]" pageId="7" pageNumber="540">
|
||
<emphasis id="B971EAC9FFC5DC7DFF52F83E054805F3" bold="true" box="[151,295,1954,1977]" pageId="7" pageNumber="540">FIGURES 15</emphasis>
|
||
–
|
||
<emphasis id="B971EAC9FFC5DC7DFEF1F83F053C05F3" bold="true" box="[308,339,1955,1976]" pageId="7" pageNumber="540">16.</emphasis>
|
||
<taxonomicName id="4C054D58FFC5DC7DFE9FF83F062F05F2" box="[346,576,1955,1977]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="540" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
|
||
<emphasis id="B971EAC9FFC5DC7DFE9FF83F062F05F2" box="[346,576,1955,1977]" italics="true" pageId="7" pageNumber="540">Tarbagataya zaisanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24257B2FFC5DC7DFD83F83F06F805F3" box="[582,663,1955,1976]" pageId="7" pageNumber="540" rank="species">
|
||
<emphasis id="B971EAC9FFC5DC7DFD83F83F06F805F3" bold="true" box="[582,663,1955,1976]" pageId="7" pageNumber="540">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female paratype. 15, vulva, ventral view; 16, vulva, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A6653FFCADC72FF52F8F407F90590" httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="8" pageNumber="541" targetBox="[199,1382,213,1869]" targetPageId="8">
|
||
<paragraph id="8BBA36DBFFCADC72FF52F8F407F90590" blockId="8.[151,1436,1896,2011]" pageId="8" pageNumber="541">
|
||
<emphasis id="B971EAC9FFCADC72FF52F8F405460535" bold="true" box="[151,297,1896,1919]" pageId="8" pageNumber="541">FIGURES 17</emphasis>
|
||
–
|
||
<emphasis id="B971EAC9FFCADC72FEF3F8F505390535" bold="true" box="[310,342,1897,1918]" pageId="8" pageNumber="541">23.</emphasis>
|
||
<taxonomicName id="4C054D58FFCADC72FE9AF8F506260534" box="[351,585,1897,1919]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
|
||
<emphasis id="B971EAC9FFCADC72FE9AF8F506260534" box="[351,585,1897,1919]" italics="true" pageId="8" pageNumber="541">Tarbagataya zaisanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24257B2FFCADC72FD94F8F506CA0535" box="[593,677,1897,1918]" pageId="8" pageNumber="541" rank="species">
|
||
<emphasis id="B971EAC9FFCADC72FD94F8F506CA0535" bold="true" box="[593,677,1897,1918]" pageId="8" pageNumber="541">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male paratype (17–21), female paratype (22–23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view;
|
||
<emphasis id="B971EAC9FFCADC72FBC1F83B004805F7" bold="true" box="[1028,1063,1959,1980]" pageId="8" pageNumber="541">mb</emphasis>
|
||
, mesal branch of coxite;
|
||
<emphasis id="B971EAC9FFCADC72FAFDF83B012205F7" bold="true" box="[1336,1357,1959,1980]" pageId="8" pageNumber="541">lb</emphasis>
|
||
, lateral branch of coxite;
|
||
<emphasis id="B971EAC9FFCADC72FE8CF859053C0591" bold="true" box="[329,339,1989,2010]" pageId="8" pageNumber="541">s</emphasis>
|
||
, saculiform structure of anterior gonopod. Scales in mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBA36DBFFCBDC73FF0CFF04072703E7" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">Leg pairs 3–6 slightly enlarged, with funnel-shaped tarsal papillae throughout ventrally; claw at base with two small additional claws dorsally and short setoid filament ventrally; coxae micropapillate ventrally. Postgonopodal legs (including leg pairs 10 and 11) with funnel-shaped tarsal papillae gradually missing toward end of body (tarsal papillae of leg pairs 10 and 11 occupying 2/3 length of tarsus i.e. space near claw free from papillae; tarsal papillae of midbody legs reaching to middle of tarsus i.e. distal part of tarsus free from papillae; hindmost legs without tarsal papillae); claw at base with two small additional claws dorsally and short setoid filament ventrally. In some individuals small dorsal additional claws on claws of hindmost legs missing. Claw of leg pairs 1 and 2 at base with two small additional claws dorsally and short setoid filament.</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF0CFE24059F005C" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">
|
||
Leg 7 strongly modified (
|
||
<figureCitation id="133E2A5EFFCBDC73FE28FE2B06000384" box="[493,623,439,463]" captionStart="FIGURES 10 – 14" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[165,1424,409,1859]" captionTargetId="figure@6.[165,1425,409,1876]" captionTargetPageId="6" captionText="FIGURES 10 – 14. Tarbagataya zaisanica sp. nov., male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod." httpUri="https://zenodo.org/record/220595/files/figure.png" pageId="9" pageNumber="542">Figs 10–11</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A5EFFCBDC73FDB9FE2B06F40384" box="[636,667,439,463]" captionStart="FIGURES 17 – 23" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[199,1382,213,1869]" captionTargetId="figure@8.[187,1387,193,1872]" captionTargetPageId="8" captionText="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="9" pageNumber="542">17</figureCitation>
|
||
); each coxite ventrally elongated in tongue-shaped process covered with setae medially and papillae laterally; telopodites 2-segmented, setose ventrally; sternite with long protuberance bilobate apically.
|
||
</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF0CFDB905C60014" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">Legs 10 with coxal glands but without other modifications. Legs 11 without coxal glands; coxae micropapillate ventrally.</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF0CFDF00621015F" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">
|
||
Anterior gonopods (
|
||
<figureCitation id="133E2A5EFFCBDC73FE6FFDF0064400CF" box="[426,555,620,644]" captionStart="FIGURES 10 – 14" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[165,1424,409,1859]" captionTargetId="figure@6.[165,1425,409,1876]" captionTargetPageId="6" captionText="FIGURES 10 – 14. Tarbagataya zaisanica sp. nov., male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod." httpUri="https://zenodo.org/record/220595/files/figure.png" pageId="9" pageNumber="542">Figs 12–13</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A5EFFCBDC73FDF2FDF0061100CF" box="[567,638,620,644]" captionStart="FIGURES 17 – 23" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[199,1382,213,1869]" captionTargetId="figure@8.[187,1387,193,1872]" captionTargetPageId="8" captionText="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="9" pageNumber="542">19–21</figureCitation>
|
||
) with subtriangular, ventrally slightly bilobed sternite. Coxite with two branches. Mesal branch (
|
||
<emphasis id="B971EAC9FFCBDC73FE71FD0C05B500E3" bold="true" box="[436,474,656,680]" pageId="9" pageNumber="542">mb</emphasis>
|
||
) as numerous long filaments one or two of which longer than others. Lateral branch (
|
||
<emphasis id="B971EAC9FFCBDC73FF65FD2804D80087" bold="true" box="[160,183,692,716]" pageId="9" pageNumber="542">lb</emphasis>
|
||
) curved caudad, its mesal part thin fold covering
|
||
<emphasis id="B971EAC9FFCBDC73FD3CFD28074F0087" bold="true" box="[761,800,692,716]" pageId="9" pageNumber="542">mb</emphasis>
|
||
frontally and enveloping
|
||
<emphasis id="B971EAC9FFCBDC73FB8FFD28001E0087" bold="true" box="[1098,1137,692,716]" pageId="9" pageNumber="542">mb</emphasis>
|
||
mesodistally. Solenomere barely traceable on lateral branch. In posterior view (
|
||
<figureCitation id="133E2A5EFFCBDC73FCCFFD4B070A00A4" box="[778,869,727,751]" captionStart="FIGURES 10 – 14" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[165,1424,409,1859]" captionTargetId="figure@6.[165,1425,409,1876]" captionTargetPageId="6" captionText="FIGURES 10 – 14. Tarbagataya zaisanica sp. nov., male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod." httpUri="https://zenodo.org/record/220595/files/figure.png" pageId="9" pageNumber="542">Figs 13</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A5EFFCBDC73FCB0FD4B07FC00A4" box="[885,915,727,751]" captionStart="FIGURES 17 – 23" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[199,1382,213,1869]" captionTargetId="figure@8.[187,1387,193,1872]" captionTargetPageId="8" captionText="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="9" pageNumber="542">20</figureCitation>
|
||
) anterior gonopods with pair of sacculiform structures (
|
||
<emphasis id="B971EAC9FFCBDC73FED1FD600570015F" bold="true" box="[276,287,764,788]" pageId="9" pageNumber="542">s</emphasis>
|
||
) (modified coxal glands?).
|
||
</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF02FC83001A0117" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">
|
||
Posterior gonopods (
|
||
<figureCitation id="133E2A5EFFCBDC73FE6BFC83066A017C" box="[430,517,799,823]" captionStart="FIGURES 10 – 14" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[165,1424,409,1859]" captionTargetId="figure@6.[165,1425,409,1876]" captionTargetPageId="6" captionText="FIGURES 10 – 14. Tarbagataya zaisanica sp. nov., male paratype. 10, leg pair 7, caudal view; 11, leg pair 7, front view; 12, anterior gonopods, front view; 13, anterior gonopods, caudal view; 14, posterior gonopods, front view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod." httpUri="https://zenodo.org/record/220595/files/figure.png" pageId="9" pageNumber="542">Figs 14</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A5EFFCBDC73FDD4FC830642017C" box="[529,557,799,823]" captionStart="FIGURES 17 – 23" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[199,1382,213,1869]" captionTargetId="figure@8.[187,1387,193,1872]" captionTargetPageId="8" captionText="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="9" pageNumber="542">18</figureCitation>
|
||
) reduced to a cupped coxosternum with normal tracheal apodemes. Coxal part with process supplied with three teeth. Sternal part with a high median process frontally.
|
||
</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF02FCFB049901A7" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">
|
||
Female. Body with 28 segments. Length
|
||
<quantity id="4CFD9B3EFFCBDC73FD5FFCFB077A0134" box="[666,789,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="9" pageNumber="542" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
|
||
, width of midbody metazona 0.9–1.0 mm. Leg 2 typical, not reduced. Vulvae (
|
||
<figureCitation id="133E2A5EFFCBDC73FE54FC10067901EF" box="[401,534,908,932]" captionStart="FIGURES 15 – 16" captionStartId="7.[151,264,1954,1977]" captionTargetBox="[301,1275,212,1913]" captionTargetId="figure@7.[280,1286,193,1932]" captionTargetPageId="7" captionText="FIGURES 15 – 16. Tarbagataya zaisanica sp. nov., female paratype. 15, vulva, ventral view; 16, vulva, lateral view." httpUri="https://zenodo.org/record/220596/files/figure.png" pageId="9" pageNumber="542">Figs 15–16</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A5EFFCBDC73FDE0FC10060301EF" box="[549,620,908,932]" captionStart="FIGURES 17 – 23" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[199,1382,213,1869]" captionTargetId="figure@8.[187,1387,193,1872]" captionTargetPageId="8" captionText="FIGURES 17 – 23. Tarbagataya zaisanica sp. nov., male paratype (17 – 21), female paratype (22 – 23). 17, leg pair 7, caudal view; 18, posterior gonopods, front view; 19, left anterior gonopod, lateral view; 20, anterior gonopods, caudal view; 21, anterior gonopods, front view; 22, left vulva, mesal view; 23, vulva, ventral view; mb, mesal branch of coxite; lb, lateral branch of coxite; s, saculiform structure of anterior gonopod. Scales in mm." httpUri="https://zenodo.org/record/220597/files/figure.png" pageId="9" pageNumber="542">22–23</figureCitation>
|
||
) oblong with long setae ventrally. Operculum bilobate. Receptaculum seminis central in position. In some individuals small dorsal additional claws at base of claws of hindmost legs missing.
|
||
</paragraph>
|
||
<paragraph id="8BBA36DBFFCBDC73FF02FC6407730644" blockId="9.[151,1437,151,1256]" box="[199,796,1015,1040]" pageId="9" pageNumber="542">
|
||
<emphasis id="B971EAC9FFCBDC73FF02FC64057B065B" bold="true" box="[199,276,1016,1040]" pageId="9" pageNumber="542">Name.</emphasis>
|
||
The specific epithet refers to the
|
||
<typeStatus id="54BE8879FFCBDC73FD4EFC6406D4065B" box="[651,699,1016,1040]" pageId="9" pageNumber="542">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F6550FFCBDC73FF02FB8005B206A3" pageId="9" pageNumber="542" type="discussion">
|
||
<paragraph id="8BBA36DBFFCBDC73FF02FB8005B206A3" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="542">
|
||
<emphasis id="B971EAC9FFCBDC73FF02FB800554067F" bold="true" box="[199,315,1052,1076]" pageId="9" pageNumber="542">Remarks.</emphasis>
|
||
Up to now
|
||
<taxonomicName id="4C054D58FFCBDC73FE14FB800630067F" box="[465,607,1052,1076]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971EAC9FFCBDC73FE14FB800630067F" box="[465,607,1052,1076]" italics="true" pageId="9" pageNumber="542">Tarbagataya</emphasis>
|
||
</taxonomicName>
|
||
included only one species -
|
||
<taxonomicName id="4C054D58FFCBDC73FC7CFB8101F8067F" authority="Golovatch & Wytwer, 2003" authorityName="Golovatch & Wytwer" authorityYear="2003" box="[953,1431,1052,1077]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="B971EAC9FFCBDC73FC7CFB810029067F" box="[953,1094,1052,1076]" italics="true" pageId="9" pageNumber="542">T. splendida</emphasis>
|
||
Golovatch & Wytwer, 2003
|
||
</taxonomicName>
|
||
, described from East
|
||
<collectingCountry id="F312764BFFCBDC73FE41FBA30668061C" box="[388,519,1087,1111]" name="Kazakhstan" pageId="9" pageNumber="542">Kazakhstan</collectingCountry>
|
||
(Golovatch & Wytwer 2003).
|
||
<taxonomicName id="4C054D58FFCBDC73FCA2FBDD07800613" box="[871,1007,1089,1112]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
|
||
<emphasis id="B971EAC9FFCBDC73FCA2FBDD07800613" box="[871,1007,1089,1112]" italics="true" pageId="9" pageNumber="542">T. zaisanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24257B2FFCBDC73FC32FBDC003C0613" box="[1015,1107,1088,1112]" pageId="9" pageNumber="542" rank="species">
|
||
<emphasis id="B971EAC9FFCBDC73FC32FBDC003C0613" bold="true" box="[1015,1107,1088,1112]" pageId="9" pageNumber="542">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the second species of this genus. One of the distinguishing characters of
|
||
<taxonomicName id="4C054D58FFCBDC73FD60FBF8075C0637" box="[677,819,1124,1148]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971EAC9FFCBDC73FD60FBF8075C0637" box="[677,819,1124,1148]" italics="true" pageId="9" pageNumber="542">Tarbagataya</emphasis>
|
||
</taxonomicName>
|
||
is the absence of tarsal papillae in males because they are absent in the
|
||
<typeStatus id="54BE8879FFCBDC73FE91FB1505EB06EA" box="[340,388,1161,1185]" pageId="9" pageNumber="542">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C054D58FFCBDC73FE26FB15060406EB" box="[483,619,1160,1184]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="splendida">
|
||
<emphasis id="B971EAC9FFCBDC73FE26FB15060406EB" box="[483,619,1160,1184]" italics="true" pageId="9" pageNumber="542">T. splendida</emphasis>
|
||
</taxonomicName>
|
||
. However because tarsal papillae are present in
|
||
<taxonomicName id="4C054D58FFCBDC73FB43FB15016306EB" box="[1158,1292,1161,1184]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="zaisanica" status="sp. nov.">
|
||
<emphasis id="B971EAC9FFCBDC73FB43FB15016306EB" box="[1158,1292,1161,1184]" italics="true" pageId="9" pageNumber="542">T. zaisanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24257B2FFCBDC73FAD7FB14010306EB" box="[1298,1388,1160,1184]" pageId="9" pageNumber="542" rank="species">
|
||
<emphasis id="B971EAC9FFCBDC73FAD7FB14010306EB" bold="true" box="[1298,1388,1160,1184]" pageId="9" pageNumber="542">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the original diagnosis of the genus must be changed accordingly. Thus, tarsal papillae in the males of
|
||
<taxonomicName id="4C054D58FFCBDC73FACBFB3001F3068F" box="[1294,1436,1196,1220]" class="Diplopoda" family="Altajellidae" genus="Tarbagataya" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971EAC9FFCBDC73FACBFB3001F3068F" box="[1294,1436,1196,1220]" italics="true" pageId="9" pageNumber="542">Tarbagataya</emphasis>
|
||
</taxonomicName>
|
||
species are absent or present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |