170 lines
21 KiB
XML
170 lines
21 KiB
XML
<document id="259AA961D2CA784E105A37DA23735BCC" ID-DOI="10.1649/0010-065X-67.2.61" ID-Zenodo-Dep="5368429" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630260625427" checkinUser="carolina" docAuthor="Anderson, Robert S. & Marvaldi, Adriana E." docDate="2013" docId="03AC87C0093EFF883982ED2E063043E5" docLanguage="en" docName="0010-065X-67.2.61.pdf" docOrigin="The Coleopterists Bulletin 67 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-67.2.61" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Archicorynini Anderson and Marvaldi 2013, new tribe" docType="treatment" docVersion="4" lastPageNumber="71" masterDocId="FF95FFB80938FF8E3922EE640511465D" masterDocTitle="Finding Unexpected Beetles in Odd Places: Archicorynus kuscheliAnderson and Marvaldi, a New Genus and Species Representing a New Tribe, Archicorynini, of Oxycoryninae (Coleoptera: Belidae) from Nicaragua" masterLastPageNumber="71" masterPageNumber="61" pageNumber="61" updateTime="1699201200614" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F1B1EAA302AEF715F17E0CA62E0C30F0">Finding Unexpected Beetles in Odd Places: Archicorynus kuscheliAnderson and Marvaldi, a New Genus and Species Representing a New Tribe, Archicorynini, of Oxycoryninae (Coleoptera: Belidae) from Nicaragua</mods:title>
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<mods:namePart id="1082E96BFA221FF4DF02622C374F7A11">Anderson, Robert S.</mods:namePart>
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<mods:namePart id="90D90F08C13E48B6300A3BC8249E8E8A">Marvaldi, Adriana E.</mods:namePart>
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<mods:title id="B52C1A628493FA6503CE6937AE26FAA4">The Coleopterists Bulletin</mods:title>
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<mods:part id="492AD64FCCA2597256E59450B7AAD002">
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<mods:date id="FA26062DB32BFBB6BA9387A3E2130ADB">2013</mods:date>
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<mods:number id="A15CEF4377C74520989F6314CD5AA50F">2013-06-30</mods:number>
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<mods:number id="6DACB391B78FC7E227678D2115D2D077">67</mods:number>
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<mods:url id="D45D06013AA2CB12E6B00600973508B7">http://dx.doi.org/10.1649/0010-065x-67.2.61</mods:url>
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<treatment id="03AC87C0093EFF883982ED2E063043E5" ID-DOI="http://doi.org/10.5281/zenodo.5460745" ID-Zenodo-Dep="5460745" LSID="urn:lsid:plazi:treatment:03AC87C0093EFF883982ED2E063043E5" httpUri="http://treatment.plazi.org/id/03AC87C0093EFF883982ED2E063043E5" pageId="6">
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<subSubSection id="C31F655D093EFF883982ED2E04AA4527" pageId="6" type="nomenclature">
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<paragraph id="8BBA36D6093EFF883982ED2E04AA4527" blockId="6.[160,629,842,890]" pageId="6">
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<heading id="D0F281BA093EFF883982ED2E04AA4527" centered="true" fontSize="10" level="2" pageId="6" reason="2">
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<emphasis id="B971EAC4093EFF883982ED2E04AA4527" bold="true" pageId="6">
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Tribe
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<taxonomicName id="4C054D55093EFF8839FFED2E07614502" authority="Anderson and Marvaldi" authorityName="Anderson and Marvaldi" authorityYear="2013" box="[221,624,842,863]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Archicorynini">Archicorynini Anderson and Marvaldi</taxonomicName>
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,
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<taxonomicNameLabel id="A24257BF093EFF883878ED0104AA4527" box="[346,443,869,890]" pageId="6" rank="tribe">new tribe</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F655D093EFF88398DEDEB05E845E2" pageId="6" type="type_taxon">
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<paragraph id="8BBA36D6093EFF88398DEDEB05E845E2" blockId="6.[151,638,911,1729]" pageId="6">
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<emphasis id="B971EAC4093EFF88398DEDEB042745F9" bold="true" box="[175,310,911,932]" pageId="6">
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<typeStatus id="54BE8874093EFF88398DEDEB05F145F9" box="[175,224,911,932]" pageId="6">Type</typeStatus>
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Genus.
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</emphasis>
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<taxonomicName id="4C054D55093EFF883866EDF405E845E2" authority="Anderson and Marvaldi." authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883866EDF404C545F8" box="[324,468,912,933]" italics="true" pageId="6">Archicorynus</emphasis>
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Anderson and Marvaldi.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F655D093EFF88398DEDA0063043E5" pageId="6" type="diagnosis">
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<paragraph id="8BBA36D6093EFF88398DEDA004654224" blockId="6.[151,638,911,1729]" pageId="6">
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<emphasis id="B971EAC4093EFF88398DEDA004094584" bold="true" box="[175,280,964,985]" pageId="6">Diagnosis.</emphasis>
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<taxonomicName id="4C054D55093EFF883804EDA004A44584" box="[294,437,964,985]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
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characterized by a more ventral rostral insertion (than species in other tribes) not forming a rounded ventral side of head, anterior face of procoxae with suture visible and without protruding apex of trochanter, and by compact antennal clubs comprising tightly joined segments 9–11.
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</paragraph>
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<paragraph id="8BBA36D6093EFF88398DEA1B068044DD" blockId="6.[151,638,911,1729]" lastBlockId="6.[685,1172,194,1464]" pageId="6">
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<emphasis id="B971EAC4093EFF88398DEA1B048D42C9" bold="true" box="[175,412,1151,1172]" pageId="6">Phylogenetic Position.</emphasis>
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The monophyly of
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<taxonomicName id="4C054D55093EFF8839B5EAFD040842F3" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[151,281,1177,1198]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF8839B5EAFD040842F3" box="[151,281,1177,1198]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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and the remaining oxycorynines is supported by at least 13 unambiguous synapomorphies (see
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<figureCitation id="133E2A53093EFF883806EAAA047A42BE" box="[292,363,1230,1251]" captionStart="Fig" captionStartId="9.[197,229,950,969]" captionTargetBox="[190,1170,215,927]" captionTargetId="figure-400@9.[173,1193,183,948]" captionTargetPageId="9" captionText="Fig. 14. Most parsimonious tree for Belidae obtained from analysis of the matrix in Table 1. Only unambiguous character changes are mapped on the cladogram, unique and homoplastic changes are in black and white, respectively." figureDoi="http://doi.org/10.5281/zenodo.5368439" httpUri="https://zenodo.org/record/5368439/files/figure.png" pageId="6">Fig. 14</figureCitation>
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), the clearest being: funicles with second segment subequal in length to first; lateral pronotal margins crenulate; meso- and metatibiae with carinate dorsal margin; tarsi with basal segment very short, shorter than second and almost concealed; hind margin of wings devoid of setae on anal lobe. As the sole representative of
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<taxonomicName id="4C054D55093EFF8839B5EBEC040E43C0" box="[151,287,1416,1437]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Archicorynini">Archicorynini</taxonomicName>
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,
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<taxonomicName id="4C054D55093EFF88380FEBEC04BD43C0" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[301,428,1416,1437]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF88380FEBEC04BD43C0" box="[301,428,1416,1437]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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appears to represent the sister-group of all other known
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<taxonomicName id="4C054D55093EFF8838DBEBC7076F43E5" box="[505,638,1443,1464]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
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(
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<figureCitation id="133E2A53093EFF8839BCEBD905F8438F" box="[158,233,1469,1490]" captionStart="Fig" captionStartId="9.[197,229,950,969]" captionTargetBox="[190,1170,215,927]" captionTargetId="figure-400@9.[173,1193,183,948]" captionTargetPageId="9" captionText="Fig. 14. Most parsimonious tree for Belidae obtained from analysis of the matrix in Table 1. Only unambiguous character changes are mapped on the cladogram, unique and homoplastic changes are in black and white, respectively." figureDoi="http://doi.org/10.5281/zenodo.5368439" httpUri="https://zenodo.org/record/5368439/files/figure.png" pageId="6">Fig. 14</figureCitation>
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). This earliest divergent position is supported by plesiomorphic retention of the rostral insertion on the ventral side of head in lateral view (36-0) and by the anterior face of the procoxae with a visible suture but without the protruding apex of the trochanter (70-0). These characters exclude the genus from a monophyletic clade consisting of all other
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<taxonomicName id="4C054D55093EFF8838D6E813076F40D1" box="[500,638,1655,1676]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
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which share the apomorphic conditions of a convex, usually rounded, ventral side of head (36-1-2) and the anterior face of the procoxae with suture obliterated and the trochanteral apex protruding out of the pit (70-1). In addition, the underside of the elytra of
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<taxonomicName id="4C054D55093EFF883AE5EF76015A477A" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[967,1099,274,295]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883AE5EF76015A477A" box="[967,1099,274,295]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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in the region opposite to the metacoxae has an odd ‘pocket-like’ ridge (
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<figureCitation id="133E2A53093EFF883AA7EF2306D64701" box="[901,967,327,348]" captionStart="Figs" captionStartId="8.[175,215,1313,1332]" captionTargetBox="[157,1166,190,1291]" captionTargetId="figure-167@8.[151,1171,183,1307]" captionTargetPageId="8" captionText="Figs. 8–13. Archicorynus kuscheli. 8) Male, ventral view; 9) Ventral surface of elytra, male (PLR = pocket-like cavity); 10) Female genitalia (BC = bursa copulatrix; HLE = hook-like extensions of T VIII; HS = gonocoxites; S = spermatheca; SD = spermathecal duct; SG = spermathecal gland; S VIII = sternite VIII; T VIII = tergite VIII); 11) Aedeagus, lateral view (PN = penis; PE = pedon; TC = tectum; TG = tegmen); 12) Penis (a) and tegmen (b), separated; 13) Hindwing, male." figureDoi="http://doi.org/10.5281/zenodo.5368437" httpUri="https://zenodo.org/record/5368437/files/figure.png" pageId="6">Fig. 9</figureCitation>
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) not seen in other
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<taxonomicName id="4C054D55093EFF883B8FEF050628472B" box="[685,825,353,374]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
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. A similar structure was observed in the inner surface of elytra of several
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<taxonomicName id="4C054D55093EFF883D68EF18018547CC" box="[1098,1172,380,401]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Belinae">Belinae</taxonomicName>
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(i.e.
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<taxonomicName id="4C054D55093EFF883BC3EFF3066747F1" box="[737,886,407,428]" class="Insecta" family="Belidae" genus="Agnesiotis" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B971EAC4093EFF883BC3EFF3065B47F1" box="[737,842,407,428]" italics="true" pageId="6">Agnesiotis</emphasis>
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sp.
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</taxonomicName>
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,
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<taxonomicName id="4C054D55093EFF883AA4EFF3010547F1" box="[902,1044,407,428]" class="Insecta" family="Belidae" genus="Rhinotia" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B971EAC4093EFF883AA4EFF306CC47F1" box="[902,989,407,428]" italics="true" pageId="6">Rhinotia</emphasis>
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spp.
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</taxonomicName>
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) suggesting that this feature might be another plesiomorphic retention of
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<taxonomicName id="4C054D55093EFF883A1BEFA806D547BC" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[825,964,460,481]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883A1BEFA806D547BC" box="[825,964,460,481]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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. Besides the morphological evidence presented here, the DNA sequences (from 18S and 28S rRNA and COI markers) already obtained for
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<taxonomicName id="4C054D55093EFF883AF1EC78015C446C" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[979,1101,540,561]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883AF1EC78015C446C" box="[979,1101,540,561]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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(A. E. Marvaldi and M. S. Ferrer, unpublished data) also suggest a close relationship of
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<taxonomicName id="4C054D55093EFF883AC6EC350171443B" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[996,1120,593,614]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883AC6EC350171443B" box="[996,1120,593,614]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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with the other oxycorynines.
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</paragraph>
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<paragraph id="8BBA36D6093EFF883BE7ECE2069B42F3" blockId="6.[685,1172,194,1464]" pageId="6">
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Alternative relationships that might be easier to justify by vicariance biogeography are that
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<taxonomicName id="4C054D55093EFF883B8FECDF063A448D" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[685,811,699,720]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883B8FECDF063A448D" box="[685,811,699,720]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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is the sister taxon to just the New World
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<taxonomicName id="4C054D55093EFF883BD6ECB1011D44B7" authority="(Oxycorynini)" baseAuthorityName="Oxycorynini" box="[756,1036,725,746]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae (Oxycorynini)</taxonomicName>
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or to a subgroup of these. This hypothesis requires assumption of a reversal in the structure of the trochanter as well as a reversal in the positioning of the rostrum. It is also less consistent with the biological attributes of the taxa as all
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<taxonomicName id="4C054D55093EFF883D3AED3E01824532" authorityName="Schoenherr" authorityYear="1840" box="[1048,1171,858,879]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Oxycorynini">Oxycorynini</taxonomicName>
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are associated with the reproductive structures of various plants, whereas
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<taxonomicName id="4C054D55093EFF883A9DEDF4015145F8" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[959,1088,912,933]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971EAC4093EFF883A9DEDF4015145F8" box="[959,1088,912,933]" italics="true" pageId="6">Archicorynus</emphasis>
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</taxonomicName>
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appears to be associated with dead vegetative plant parts, the presumed ancestral habit of the subfamily (
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<bibRefCitation id="EF944B27093EFF883B96EDBB069745A9" author="Marvaldi" box="[692,902,991,1012]" pageId="6" pagination="447 - 476" refId="ref8161" refString="Marvaldi, A. E., C. H. C. Lyal, R. G. Oberprieler, T. Bradbury, and R. S. Anderson. 2006. Phylogeny of the Oxycoryninae s. l. (Coleoptera: Belidae) and evolution of host-plant associations. Invertebrate Systematics 20: 447 - 476." type="journal article" year="2006">
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Marvaldi
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<emphasis id="B971EAC4093EFF883A34EDBB065B45A9" box="[790,842,991,1012]" italics="true" pageId="6">et al.</emphasis>
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2006
|
||
</bibRefCitation>
|
||
). Thus, while
|
||
<taxonomicName id="4C054D55093EFF883D37EDBB018245A9" baseAuthorityName="Voss" baseAuthorityYear="1953" box="[1045,1171,991,1012]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Metrioxenini">Metrioxenini</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C054D55093EFF883BF9ED9E064A4252" baseAuthorityName="Wollaston" baseAuthorityYear="1864" box="[731,859,1018,1039]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Aglycyderini">Aglycyderini</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C054D55093EFF883AB9ED9E01344252" box="[923,1061,1018,1039]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Archicorynini">Archicorynini</taxonomicName>
|
||
) appear to have retained an association with dead vegetative plant parts,
|
||
<taxonomicName id="4C054D55093EFF883A77EA4B06DE4219" authorityName="Schoenherr" authorityYear="1840" box="[853,975,1071,1092]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="tribe" tribe="Oxycorynini">Oxycorynini</taxonomicName>
|
||
by contrast shifted their development to reproductive organs, initially retaining an association with conifers but later colonizing cycads and two groups of rootparasitic angiosperms.
|
||
</paragraph>
|
||
<paragraph id="8BBA36D6093EFF883BE7EAD0063043E5" blockId="6.[685,1172,194,1464]" pageId="6">
|
||
<taxonomicName id="4C054D55093EFF883BE7EAD0065A4294" box="[709,843,1204,1225]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
|
||
are clearly an ancient lineage and the presence of the taxon sister to the remaining taxa in
|
||
<collectingCountry id="F3127646093EFF883BDEEA8D064F42A3" box="[764,862,1257,1278]" name="Nicaragua" pageId="6">Nicaragua</collectingCountry>
|
||
may be unexpected but is not implausible. This is a relictual taxon in which numerous extinctions and range contractions would be expected simply because of their antiquity, widespread distribution at the subfamily level but highly localized distribution at the generic level, and their generalized non-specialized ancestral biology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |