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<document id="F5C64C1A89D9C70278AAEAB364529460" ID-CLB-Dataset="112023" ID-DOI="10.11646/zootaxa.5103.1.1" ID-GBIF-Dataset="1bf32ba1-8e0d-4435-ba04-6d7dbd0d7238" ID-ISSN="1175-5326" ID-Zenodo-Dep="6245709" ID-ZooBank="9E1A72E7-3862-44F7-B69F-ECE64B239FF9" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1645601651449" checkinUser="plazi" docAuthor="Kirejtshuk, Alexander G. &amp; Kovalev, Alexey V." docDate="2022" docId="03AC73267602D60175E0FB54FC05FE12" docLanguage="en" docName="zootaxa.5103.1.1.pdf" docOrigin="Zootaxa 5103 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Oniphenes bicoloratus Kirejtshuk &amp; Kovalev 2022, sp. nov." docType="treatment" docUuid="B0F295BE-30CF-473C-AE2E-EF87C9847573" docUuidSource="ZooBank" docVersion="9" lastPageNumber="122" masterDocId="FF950B5E7674D6787577FFF3FFF4FFE7" masterDocTitle="Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily" masterLastPageNumber="133" masterPageNumber="1" pageNumber="119" updateTime="1699299454408" updateUser="plazi">
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<mods:title id="7342AA2828A496C6B6D9FD22FC5CA490">Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily</mods:title>
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<mods:namePart id="BBC5212362033187F59D9469207861ED">Kirejtshuk, Alexander G.</mods:namePart>
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<mods:namePart id="5B4B9380593DC41E9BEAC6B51EAF1305">Kovalev, Alexey V.</mods:namePart>
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<mods:nameIdentifier id="0AAC0B8C5294D0B9CF80900C86A3F633" type="email">agk@zin.ru</mods:nameIdentifier>
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<mods:title id="D4305ED107415DEDCAE630FAD8B3D3B9">Zootaxa</mods:title>
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<mods:date id="A839F88954AC913897143B540E7486C8">2022</mods:date>
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<treatment id="03AC73267602D60175E0FB54FC05FE12" ID-DOI="http://doi.org/10.5281/zenodo.6823500" ID-GBIF-Taxon="198574623" ID-Zenodo-Dep="6823500" LSID="urn:lsid:zoobank.org:act:B0F295BE-30CF-473C-AE2E-EF87C9847573" httpUri="http://treatment.plazi.org/id/03AC73267602D60175E0FB54FC05FE12" lastPageId="121" lastPageNumber="122" pageId="118" pageNumber="119">
<subSubSection id="C31F91BB7602D60E75E0FB54FCAFFB26" box="[151,859,1191,1218]" pageId="118" pageNumber="119" type="nomenclature">
<paragraph id="8BBAC2307602D60E75E0FB54FCAFFB26" blockId="118.[151,859,1191,1253]" box="[151,859,1191,1218]" pageId="118" pageNumber="119">
<emphasis id="B9711E227602D60E75E0FB54FCAFFB26" bold="true" box="[151,859,1191,1218]" pageId="118" pageNumber="119">
50.
<taxonomicName id="4C05B9B37602D60E75B4FB54FD90FB26" authority="Kirejtshuk" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[195,612,1191,1218]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="bicoloratus">
<emphasis id="B9711E227602D60E75B4FB54FE20FB25" bold="true" box="[195,468,1191,1218]" italics="true" pageId="118" pageNumber="119">Oniphenes bicoloratus</emphasis>
Kirejtshuk
</taxonomicName>
et Kovalev,
<taxonomicNameLabel id="A242A3597602D60E778AFB54FCAFFB26" box="[765,859,1191,1217]" pageId="118" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7602D60E75E0FB38FC4BFAC0" pageId="118" pageNumber="119" type="description">
<paragraph id="8BBAC2307602D60E75E0FB38FED7FB02" blockId="118.[151,859,1191,1253]" box="[151,291,1227,1253]" pageId="118" pageNumber="119">
<figureCitation id="133EDEB57602D60E75E0FB38FF18FB02" box="[151,236,1227,1253]" captionStart="FIGURE 36" captionStartId="119.[151,250,1864,1888]" captionTargetBox="[182,1404,185,1832]" captionTargetId="figure-19@119.[177,1411,181,1840]" captionTargetPageId="119" captionText="FIGURE 36. Oniphenes bicoloratus gen. et sp. nov., orig. AH: holotype, male and I: paratype, female from QLD (“… 3 km ENE of Mt. Tozer …”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: male anal sclerite, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 0.5 mm to Fig. A; b = 0.5 mm to Figs BD; с = 0.25 mm to Figs EI." figureDoi="http://doi.org/10.5281/zenodo.6245898" httpUri="https://zenodo.org/record/6245898/files/figure.png" pageId="118" pageNumber="119">Figs 36</figureCitation>
, AI
</paragraph>
<paragraph id="8BBAC2307602D60E75E0FAE3FC4BFAC0" blockId="118.[151,959,1295,1320]" box="[151,959,1295,1320]" pageId="118" pageNumber="119">
LSID
<uri id="FF94CE327602D60E75A1FAFCFC4BFAC0" box="[214,959,1295,1319]" pageId="118" pageNumber="119">
urn:lsid:zoobank.org:act:
<uuid id="FFA3F8E57602D60E74BBFAFCFC4BFAC0" box="[460,959,1295,1319]" pageId="118" pageNumber="119">CD950A5F-B6A6-4D76-816F-CE94A169A657</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7602D60E75E0FAA5FBC3FA73" pageId="118" pageNumber="119" type="materials_examined">
<paragraph id="8BBAC2307602D60E75E0FAA5FBC3FA73" blockId="118.[151,1437,1365,2040]" pageId="118" pageNumber="119">
<emphasis id="B9711E227602D60E75E0FAA5FE23FA88" bold="true" box="[151,471,1365,1392]" pageId="118" pageNumber="119">Type specimens examined.</emphasis>
<materialsCitation id="3B6DC86D7602D60E74AAFAA6FBC3FA73" ID-GBIF-Occurrence="3878844422" collectingDate="1986-07-01" collectingDateMax="1986-07-04" collectingDateMin="1986-07-01" collectionCode="ANIC, ZIN" collectorName="T. Weir" country="Australia" latitude="-12.44" location="Mt. Tozer" longLatPrecision="776" longitude="143.14" pageId="118" pageNumber="119" specimenCount="2" typeStatus="holotype">
<emphasis id="B9711E227602D60E74AAFAA6FD6EFA88" bold="true" box="[477,666,1365,1391]" pageId="118" pageNumber="119">
<collectingCountry id="F31282A07602D60E74AAFAA6FDBBFA88" box="[477,591,1365,1391]" name="Australia" pageId="118" pageNumber="119">Australia</collectingCountry>
, QLD
</emphasis>
:
<typeStatus id="54BE7C927602D60E77DDFAA5FCE1FA97" box="[682,789,1366,1392]" pageId="118" pageNumber="119" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED145AF57602D60E761FFAA5FC44FA97" box="[872,944,1366,1392]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="118" pageNumber="119">ANIC</collectionCode>
) and
<specimenCount id="9D0309B97602D60E7686FAA5FB82FA97" box="[1009,1142,1366,1392]" pageId="118" pageNumber="119" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
, male and female (
<collectionCode id="ED145AF57602D60E7038FAA5FA6CFA97" box="[1359,1432,1366,1392]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="118" pageNumber="119">ANIC</collectionCode>
,
<collectionCode id="ED145AF57602D60E75E0FA89FF3CFA73" box="[151,200,1402,1428]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="118" pageNumber="119" type="Museum">ZIN</collectionCode>
)—“
<geoCoordinate id="EE31A4F77602D60E7580FA89FEBAFA73" box="[247,334,1402,1428]" degrees="12.44" direction="south" orientation="latitude" pageId="118" pageNumber="119" precision="555" value="-12.44">12.44 S</geoCoordinate>
<geoCoordinate id="EE31A4F77602D60E7423FA89FE4FFA73" box="[340,443,1402,1428]" degrees="143.14" direction="east" orientation="longitude" pageId="118" pageNumber="119" precision="555" value="143.14">143.14 E</geoCoordinate>
,
<quantity id="4CFD6FD57602D60E74B0FA89FDF4FA73" box="[455,512,1402,1428]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="118" pageNumber="119" unit="km" value="3.0">3 km</quantity>
ENE of
<location id="8EDA94EB7602D60E7714FA89FD26FA73" LSID="urn:lsid:plazi:treatment:03AC73267602D60175E0FB54FC05FE12:8EDA94EB7602D60E7714FA89FD26FA73" box="[611,722,1402,1428]" country="Australia" latitude="-12.44" longLatPrecision="776" longitude="143.14" name="Mt. Tozer" pageId="118" pageNumber="119">Mt. Tozer</location>
, QLD,
<date id="FFBBE4F07602D60E7652FA89FC32FA73" box="[805,966,1402,1428]" pageId="118" pageNumber="119" value="1986-07-01" valueMax="1986-07-04" valueMin="1986-07-01">
<collectingDate id="EFFF1D187602D60E7652FA89FC32FA73" box="[805,966,1402,1428]" pageId="118" pageNumber="119" value="1986-07-01" valueMax="1986-07-04" valueMin="1986-07-01">14 July 1986</collectingDate>
</date>
,
<collectorName id="26F0A7E67602D60E76A6FA89FBDDFA73" box="[977,1065,1402,1428]" pageId="118" pageNumber="119">T. Weir</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7602D60E75B0FA6EFB87F833" pageId="118" pageNumber="119" type="diagnosis">
<paragraph id="8BBAC2307602D60E75B0FA6EFDD4F953" blockId="118.[151,1437,1365,2040]" pageId="118" pageNumber="119">
<emphasis id="B9711E227602D60E75B0FA6EFECEFA50" bold="true" box="[199,314,1437,1463]" pageId="118" pageNumber="119">Diagnosis</emphasis>
. This new species has a very peculiar coloration and make it easy to distinguish it from other congeners (although in lighter
<typeStatus id="54BE7C927602D60E7485FA31FDA6FA3B" box="[498,594,1474,1500]" pageId="118" pageNumber="119" type="paratype">paratype</typeStatus>
it is not clear). Besides,
<taxonomicName id="4C05B9B37602D60E761BFA31FB9FFA3B" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[876,1131,1474,1500]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="bicoloratus" status="sp. nov.">
<emphasis id="B9711E227602D60E761BFA31FB9FFA3B" box="[876,1131,1474,1500]" italics="true" pageId="118" pageNumber="119">Oniphenes bicoloratus</emphasis>
</taxonomicName>
<emphasis id="B9711E227602D60E7105FA31FB39FA3B" bold="true" box="[1138,1229,1474,1500]" pageId="118" pageNumber="119">
<taxonomicNameLabel id="A242A3597602D60E7105FA31FB39FA3B" box="[1138,1229,1474,1500]" pageId="118" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has, in contrast to other congeners, the body more robust with larger and more convex pronotum and more arcuate elytral sides, scape strongly swollen and not more than 1.3
<emphasis id="B9711E227602D60E773CF9F8FDAAF9C3" box="[587,606,1547,1572]" italics="true" pageId="118" pageNumber="119">×</emphasis>
as long as thick, and also very characteristic outline of the apex of the male anal sclerite and structure of aedeagus and comparatively wide gonocoxites of its ovipositor. All congeners have the peculiar shape of pronotum (anterior and posterior edges, posterior angles and explanation of sides), shape of mouthparts (labrum, mandibles, mentum, terminal maxillary palpomeres), pregenal processes and antennal grooves. This new species also differs from:
</paragraph>
<paragraph id="8BBAC2307602D60E75B0F94DFBE7F8C7" blockId="118.[151,1437,1365,2040]" pageId="118" pageNumber="119">
<emphasis id="B9711E227602D60E75A9F94DFE90F93F" box="[222,356,1726,1752]" italics="true" pageId="118" pageNumber="119">
O,
<taxonomicName id="4C05B9B37602D60E7475F94DFE90F93F" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[258,356,1726,1752]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="lobanovi" status="sp. nov.">lobanovi</taxonomicName>
</emphasis>
<emphasis id="B9711E227602D60E741BF94DFE33F93F" bold="true" box="[364,455,1726,1752]" pageId="118" pageNumber="119">
<taxonomicNameLabel id="A242A3597602D60E741BF94DFE33F93F" box="[364,455,1726,1752]" pageId="118" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the much shorter elytra, temples not projecting externally behind eyes; distinct 8-9 striae on elytra (not only five); abdominal laterosternites V and VI about 4.0 × as long as wide (not at least 5.0 × as long as wide); arcuately convergent antennal grooves (not S-likely convergent);
</paragraph>
<paragraph id="8BBAC2307602D60E75B0F8D9FB87F833" blockId="118.[151,1437,1365,2040]" pageId="118" pageNumber="119">
<taxonomicName id="4C05B9B37602D60E75ACF8D9FE75F8A3" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[219,385,1834,1860]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="subunicolor" status="sp. nov.">
<emphasis id="B9711E227602D60E75ACF8D9FE75F8A3" box="[219,385,1834,1860]" italics="true" pageId="118" pageNumber="119">O. subunicolor</emphasis>
</taxonomicName>
<emphasis id="B9711E227602D60E74F1F8D9FE2AF8A3" bold="true" box="[390,478,1834,1860]" pageId="118" pageNumber="119">
<taxonomicNameLabel id="A242A3597602D60E74F1F8D9FE2AF8A3" box="[390,478,1834,1860]" pageId="118" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the narrowly explanate elytral sides; subrectiliner temples (not arcuate); subtruncate anterior edge of frons (not emarginate); gently curved outer edge of mandible (not dilated and forming an anterior angular excision); distinct 8-9 striae on elytra (not only six); sparser dorsal punctation; more medially convex prosternal process (not somewhat subflattened); abdominal laterosternites V and VI about 4.0 × as long as wide (not 4.5 × as long as wide); arcuately convergent antennal grooves (not S-likely convergent);
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB7602D60E75B0F82DFC61F81F" box="[199,917,2014,2040]" pageId="118" pageNumber="119" type="etymology">
<paragraph id="8BBAC2307602D60E75B0F82DFC61F81F" blockId="118.[151,1437,1365,2040]" box="[199,917,2014,2040]" pageId="118" pageNumber="119">
<emphasis id="B9711E227602D60E75B0F82DFEBCF81F" bold="true" box="[199,328,2014,2040]" pageId="118" pageNumber="119">Etymology</emphasis>
.The epithet of this new species means “
<emphasis id="B9711E227602D60E767CF82DFC76F81F" box="[779,898,2014,2040]" italics="true" pageId="118" pageNumber="119">bicolorous</emphasis>
”.
</paragraph>
</subSubSection>
<caption id="DF7A92B87603D60F75E0F8BBFAC3F82C" ID-DOI="http://doi.org/10.5281/zenodo.6245898" ID-Zenodo-Dep="6245898" httpUri="https://zenodo.org/record/6245898/files/figure.png" pageId="119" pageNumber="120" startId="119.[151,250,1864,1888]" targetBox="[182,1404,185,1832]" targetPageId="119">
<paragraph id="8BBAC2307603D60F75E0F8BBFAC3F82C" blockId="119.[151,1436,1864,1996]" pageId="119" pageNumber="120">
<emphasis id="B9711E227603D60F75E0F8BBFED4F887" bold="true" box="[151,288,1864,1888]" pageId="119" pageNumber="120">FIGURE 36.</emphasis>
<taxonomicName id="4C05B9B37603D60F7450F8BBFDF9F887" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[295,525,1864,1888]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="bicoloratus" status="gen. et sp. nov.">
<emphasis id="B9711E227603D60F7450F8BBFDF9F887" box="[295,525,1864,1888]" italics="true" pageId="119" pageNumber="120">Oniphenes bicoloratus</emphasis>
</taxonomicName>
<emphasis id="B9711E227603D60F7763F8BBFD46F887" bold="true" box="[532,690,1864,1888]" pageId="119" pageNumber="120">
<taxonomicNameLabel id="A242A3597603D60F7763F8BBFD46F887" box="[532,690,1864,1888]" pageId="119" pageNumber="120" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, orig. AH: holotype, male and I: paratype, female from QLD (“… 3 km ENE of Mt. Tozer …”);
<emphasis id="B9711E227603D60F74E2F89FFE53F863" bold="true" box="[405,423,1900,1924]" pageId="119" pageNumber="120">A</emphasis>
: body, dorsal;
<emphasis id="B9711E227603D60F7735F89FFDA7F863" bold="true" box="[578,595,1900,1924]" pageId="119" pageNumber="120">B</emphasis>
: head, dorsal;
<emphasis id="B9711E227603D60F779AF89FFD0BF863" bold="true" box="[749,767,1900,1924]" pageId="119" pageNumber="120">C</emphasis>
: head and thorax, ventral;
<emphasis id="B9711E227603D60F7161F89FFBDCF863" bold="true" box="[1046,1064,1900,1924]" pageId="119" pageNumber="120">D</emphasis>
: lateral side of abdominal segments 5 and 6, dorsal;
<emphasis id="B9711E227603D60F7436F863FEA6F84F" bold="true" box="[321,338,1936,1960]" pageId="119" pageNumber="120">E</emphasis>
: male anal sclerite, ventral;
<emphasis id="B9711E227603D60F770EF863FD7CF84F" bold="true" box="[633,648,1936,1960]" pageId="119" pageNumber="120">F</emphasis>
: tegmen, ventral;
<emphasis id="B9711E227603D60F7631F863FCAEF84F" bold="true" box="[838,858,1936,1960]" pageId="119" pageNumber="120">G</emphasis>
: penis trunk, dorsal;
<emphasis id="B9711E227603D60F714FF863FBB8F84F" bold="true" box="[1080,1100,1936,1960]" pageId="119" pageNumber="120">H</emphasis>
: proximal part of inner sac with armature;
<emphasis id="B9711E227603D60F7588F847FEFDF82B" bold="true" box="[255,265,1972,1996]" pageId="119" pageNumber="120">I</emphasis>
: ovipositor, ventral. Scale bars:
<emphasis id="B9711E227603D60F7727F847FDA9F82B" bold="true" box="[592,605,1972,1996]" pageId="119" pageNumber="120">a</emphasis>
= 0.5 mm to Fig. A;
<emphasis id="B9711E227603D60F7645F847FCB4F82B" bold="true" box="[818,832,1972,1996]" pageId="119" pageNumber="120">b</emphasis>
= 0.5 mm to Figs BD;
<emphasis id="B9711E227603D60F714FF847FBB7F82B" bold="true" box="[1080,1091,1972,1996]" pageId="119" pageNumber="120">с</emphasis>
= 0.25 mm to Figs EI.
</paragraph>
</caption>
<subSubSection id="C31F91BB760CD60175B0FF65FB73FE4A" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121" type="description">
<paragraph id="8BBAC230760CD60075B0FF65FB2BFE6F" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">
<emphasis id="B9711E22760CD60075B0FF65FDC2FF57" bold="true" box="[199,566,150,176]" pageId="120" pageNumber="121">
Description of
<typeStatus id="54BE7C92760CD600740DFF65FE15FF57" box="[378,481,150,176]" pageId="120" pageNumber="121" type="holotype">holotype</typeStatus>
(male)
</emphasis>
. Length of body with abdomen 3.4, length without abdomen 2.6, width 1.1, height
<quantity id="4CFD6FD5760CD6007592FF49FECBFF32" box="[229,319,186,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="120" pageNumber="121" unit="mm" value="0.4">0.4 mm</quantity>
. Body moderately elongate, subparallel-sided; moderately convex dorsally (particularly prothorax) and ventrally; chestnut brown dorsally with yellowish spot at elytral shoulder; coloration of underside becoming lighter from lateral side to middle; anterior part of head, antennal flagelli and legs reddish; body shining dorsally and ventrally; dorsum without pubescence (only very short and almost inconspicuous setae insided punctures on pygidium), sides of pronotum and elytra not ciliate, although middle of anterior and posterior edges of pronotum with dense and short microsetae; underside with moderately short and sparse fine yellowish hairs.
</paragraph>
<paragraph id="8BBAC230760CD60075B0FE61FEA8FD17" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">Head and pronotum with punctures, about 1.5 × as coarse as eye facets, separated by 24 puncture diameters, interspaces between them shining. Elytra with punctures somewhat coarser and densely arranged in longitudinal rows on bottom of weakly deepened nine striae, interstriae about as wide as 57 puncture diameters, very finely and very sparsely punctured and shining. Exposed abdominal dorsal sclerites with punctures slightly finer and denser than those on head and pronotum, shining to somewhat alutaceous. Prosternum and metaventrite with finer, shallower and sparser punctures than those on head and pronotum, separated by about 23 puncture diameters on prosternum and 45 puncture diameters on metaventrite, and interspaces between punctures finely and densely microreticulate on prosternum and smoothed on metasternum. Abdominal ventrites with somewhat finer punctation than that on thoracic underside and becoming denser and coarser apically; interspaces between punctures smoothed on ventrirtes 14.</paragraph>
<paragraph id="8BBAC230760CD60075B0FD09FF4EFB47" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">
Head subflattened, with moderately large and finely faceted eyes located anteriorly from arcuate temples, anterior edge of frons truncate. Labrum moderately short and with subtruncate anterior edge. Mandibles stout and short, externally gently curved to narrow bidentate apices (without dilatation at base). Antennal grooves distinctly outlined and C-shaped convergent and with minimal distance about 0.3
<emphasis id="B9711E22760CD60076BAFC94FC14FC67" box="[973,992,871,896]" italics="true" pageId="120" pageNumber="121">×</emphasis>
as great as width of mentum. Mentum subtrapezoid with anterior edge subbi-sinuate, moderately wide and arcuately widening anteriorly, and with anterior lateral angles rounded and slightly projecting anteriorly. Pregenal process at hypostomal sinus moderately wide and slightly curved, with oblique apex (outer angle more projecting than inner one). Terminal labial palpomere short, subcylindrical and slightly longer than wide, truncate at apex; terminal maxillary palpomere moderately long and subconical about 2.5 × as long as thick. Antennae with scape strongly swollen and not more than 1.3
<emphasis id="B9711E22760CD6007066FBE8FAD0FBD3" box="[1297,1316,1051,1076]" italics="true" pageId="120" pageNumber="121">×</emphasis>
as long as thick, pedicel about 0.4 × as long as scape, antennomere 3 about 1.7 × as long as pedicel, following antennomeres 47 about 0.3 × as long antennomeres 3, compact club 1.5 × as long thick and with comparable widest antennomere 10.
</paragraph>
<paragraph id="8BBAC230760CD60075B0FB58FD5CFA7A" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">Pronotum with anterior edge subemarginate at middle; posterior edge slightly convex and with slightly expressed posterior angles; sides gently sloping, all edges narrowly bordered. Scutellar shield about 1.5 × as wide as long and widely rounded at apex. Elytra nearly as long as combined width, somewhat arcuate along elytral sides and widely rounded at outer apical angles, leaving last three abdominal segments completely exposed, sides steeply sloping to extremely narrowly explanate at edge. Abdominal laterosternites V and VI about 4.0× as long as wide. Three last abdominal tergites medially rather convex and weakly subdepressed at sides. Male pygidium about 1.8 × as wide as long, arcuately narrowinig to subtruncate apex.</paragraph>
<paragraph id="8BBAC230760CD60075B0FA54FB8CF9B6" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">Prosternum medially convex and with process very narrow between procoxae, very strongly and arcuately widened at subtruncate apex, with maximum width comparable with that of antennal club. Pro- and metacoxae extremely narrowly separated and distance between mesocoxae as great as scape thickness. Metaventrite with a distinct median suture nearly reaching anterior third. Metepisterna very narrow, at anterior end markedly narrower than thickness of scape. Hypopygidium about 1.8 × as wide as long, subtruncate at apex.</paragraph>
<paragraph id="8BBAC230760CD60075B0F9A9FC56F8E2" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">Tibiae subtriangilar, protibia about as wide as antennal club and with preapical teeth moderately raised, meso-and metatibiae widely rounded at outer subapical angles, spurs comparatively long and rather stout; outer side of meso- and metatibiae with with raised spines in distal part along their outer edge. Femora rather wide and moderately short; profemur about 1.5 ×, meso- and metafemora nearly 2.0 × as wide as corresponding tibiae. Protarsus nearly as wide as antennal pedicel, meso- and metatarsi somewhat narrower.</paragraph>
<paragraph id="8BBAC230760CD60075B0F8FDFDE3F85F" blockId="120.[151,1437,150,2013]" pageId="120" pageNumber="121">Anal sclerite well exposed, rounded to subangular at apex. Aedeagus moderately sclerotized and more than 2.0 × as long as wide; tegmen slightly arcuate at sides basal two thirds and and with median long apical process rounded, in turn, at setose apex; penis trunk slightly shorter than tegmen and with lobes of apical orifice surpassing apex of penis trunk. Inner sac with many small and slightly sclerotized granules in most part of sac and with one pair arcuate sclerites behind them.</paragraph>
<paragraph id="8BBAC230760CD60175B0F831FCBEFEFA" blockId="120.[151,1437,150,2013]" lastBlockId="121.[151,1436,151,501]" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121">
<emphasis id="B9711E22760CD60075B0F831FEEAF83B" bold="true" box="[199,286,1986,2012]" pageId="120" pageNumber="121">Female</emphasis>
. Differs from the male only in the narrower and less convex pronotum, slightly narrower protarsi, and also in the length and shape of sclerites of the last abdominal segment (pygidium about 1.4 × as wide as long and hypopygidum about 1.3 × as long as wide and widely rounded at apex). Ovipositor moderately sclerotized; rather wide, inner and outer lobes of gonocoxites clearly isolated and outer lobe about 0.5 × as long as inner lobe; styli very short and slightly visible, located at distance from blunt apex.
</paragraph>
<paragraph id="8BBAC230760DD60175B0FED5FB73FE4A" blockId="121.[151,1436,151,501]" pageId="121" pageNumber="122">
<emphasis id="B9711E22760DD60175B0FED5FEB6FEA7" bold="true" box="[199,322,294,320]" pageId="121" pageNumber="122">Variations</emphasis>
. The body of the complete
<typeStatus id="54BE7C92760DD60177FBFED4FD18FEA6" box="[652,748,295,321]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
<quantity id="4CFD6FD5760DD6017781FED4FCA4FEA6" box="[758,848,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="121" pageNumber="122" unit="mm" value="3.4">3.4 mm</quantity>
long with coloration and sculpture as those in the
<typeStatus id="54BE7C92760DD60175E0FEB8FF09FE82" box="[151,253,331,357]" pageId="121" pageNumber="122" type="holotype">holotype</typeStatus>
. The second
<typeStatus id="54BE7C92760DD60174FCFEB8FE1FFE82" box="[395,491,331,357]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
without head and prothorax, but other parts of body and appendages are lighter (with light body coloration as that in
<taxonomicName id="4C05B9B3760DD6017481FE9CFD8EFE6E" authority="Kirejtshuk &amp; Kovalev, 2022" authorityName="Kirejtshuk &amp; Kovalev" authorityYear="2022" box="[502,634,367,393]" class="Insecta" family="Nitidulidae" genus="Oniphenes" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="lobanovi" status="sp. nov.">
<emphasis id="B9711E22760DD6017481FE9CFD8EFE6E" box="[502,634,367,393]" italics="true" pageId="121" pageNumber="122">O. lobanovi</emphasis>
</taxonomicName>
<emphasis id="B9711E22760DD60177F6FE9CFD2FFE6E" bold="true" box="[641,731,367,393]" pageId="121" pageNumber="122">
<taxonomicNameLabel id="A242A359760DD60177F6FE9CFD2FFE6E" box="[641,731,367,393]" pageId="121" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the spot at elytral shoulder is distinctly the same as that in other specimens of the type series). The expression of the elytral stria 9 can be very weak.
</paragraph>
</subSubSection>
<subSubSection id="C31F91BB760DD60175B0FE45FC05FE12" pageId="121" pageNumber="122" type="distribution">
<paragraph id="8BBAC230760DD60175B0FE45FD8EFE37" blockId="121.[151,1436,151,501]" box="[199,634,438,465]" pageId="121" pageNumber="122">
<emphasis id="B9711E22760DD60175B0FE45FEADFE37" bold="true" box="[199,345,438,464]" pageId="121" pageNumber="122">Distribution</emphasis>
.
<collectingRegion id="49C10CD2760DD6017410FE44FE1FFE36" box="[359,491,439,465]" country="Australia" name="Queensland" pageId="121" pageNumber="122">Queensland</collectingRegion>
(Mt. Tozer).
</paragraph>
<paragraph id="8BBAC230760DD60175B0FE28FC05FE12" blockId="121.[151,1436,151,501]" box="[199,1009,474,501]" pageId="121" pageNumber="122">
<emphasis id="B9711E22760DD60175B0FE28FE6AFE13" bold="true" box="[199,414,474,501]" pageId="121" pageNumber="122">Notes on bionomy</emphasis>
. This species is apparenly associated with rainforest.
</paragraph>
</subSubSection>
</treatment>
</document>