treatments-xml/data/03/AC/52/03AC526E15205F23FEE8FCFBFB91E538.xml
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<document id="F077232EC1F83454516D2DBA3D090944" ID-DOI="10.5281/zenodo.173127" ID-GBIF-Dataset="fe8a4769-447c-45b0-bc5f-0f6b77809511" ID-ISSN="1175­5326" ID-Zenodo-Dep="173127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459770052372" checkinUser="plazi" docAuthor="Ibáñez, Miguel, Siverio, Felipe, Alonso, María R. &amp; Ponte-Lira, Carmen E." docDate="2006" docId="03AC526E15205F23FEE8FCFBFB91E538" docLanguage="en" docName="zt01258p045.pdf" docOrigin="Zootaxa 1258" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Canariella giustii Ibanez &amp; Alonso, new species" docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FF952A1615245F29FFE0FFBAFFE6E703" masterDocTitle="Two Canariella species (Gastropoda: Helicodea: Hygromiidae) endemic to the Northwest Tenerife (Canary Islands)" masterLastPageNumber="45" masterPageNumber="33" pageNumber="37" updateTime="1698217154900" updateUser="plazi">
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<mods:title id="8CC7CBAF5A9A3CF95901466A95E9ABBA">Two Canariella species (Gastropoda: Helicodea: Hygromiidae) endemic to the Northwest Tenerife (Canary Islands)</mods:title>
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<mods:namePart id="83A67F75B8C03F149FA07EF4BD746FD9">Ibáñez, Miguel</mods:namePart>
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<mods:namePart id="46327D928F68C508C5457013D879454E">Siverio, Felipe</mods:namePart>
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<mods:namePart id="059C5245917772DBED1CC3A0EA89BBFB">Alonso, María R.</mods:namePart>
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<mods:namePart id="76DA4AE96CC5C3A122E46AE1A21F4A9B">Ponte-Lira, Carmen E.</mods:namePart>
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<mods:date id="DC0E918618FC8DD9418EE448214C4EB0">2006</mods:date>
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<mods:number id="F36F9CF1BEB92775D6020B449F94174A">1258</mods:number>
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<treatment id="03AC526E15205F23FEE8FCFBFB91E538" ID-DOI="http://doi.org/10.5281/zenodo.6262754" ID-GBIF-Taxon="119346943" ID-Zenodo-Dep="6262754" LSID="urn:lsid:plazi:treatment:03AC526E15205F23FEE8FCFBFB91E538" httpUri="http://treatment.plazi.org/id/03AC526E15205F23FEE8FCFBFB91E538" lastPageId="10" lastPageNumber="42" pageId="4" pageNumber="37">
<subSubSection id="C31FB0F315205F2DFEE8FCFBFCADE458" box="[264,843,833,859]" pageId="4" pageNumber="37" type="nomenclature">
<paragraph id="8BBAE37815205F2DFEE8FCFBFCADE458" blockId="4.[264,843,833,859]" box="[264,843,833,859]" pageId="4" pageNumber="37">
<heading id="D0F2541415205F2DFEE8FCFBFCADE458" bold="true" box="[264,843,833,859]" fontSize="11" level="1" pageId="4" pageNumber="37" reason="1">
<emphasis id="B9713F6A15205F2DFEE8FCFBFCADE458" bold="true" box="[264,843,833,859]" pageId="4" pageNumber="37">
<taxonomicName id="4C0598FB15205F2DFEE8FCFBFD52E458" authority="Ibanez &amp; Alonso" authorityName="Ibanez &amp; Alonso" box="[264,692,833,859]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="37" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B9713F6A15205F2DFEE8FCFBFE3DE458" bold="true" box="[264,475,833,859]" italics="true" pageId="4" pageNumber="37">Canariella giustii</emphasis>
Ibáñez &amp; Alonso
</taxonomicName>
<taxonomicNameLabel id="A242821115205F2DFD5BFCFBFCADE458" box="[699,843,833,859]" pageId="4" pageNumber="37" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31FB0F315205F2DFEE8FC3EFE6CE362" pageId="4" pageNumber="37" type="materials_examined">
<paragraph id="8BBAE37815205F2DFEE8FC3EFE97E4C0" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<emphasis id="B9713F6A15205F2DFEE8FC3EFE48E49D" bold="true" box="[264,430,900,926]" pageId="4" pageNumber="37">
<typeStatus id="54BE5DDA15205F2DFEE8FC3EFEA3E49D" box="[264,325,900,926]" pageId="4" pageNumber="37">Type</typeStatus>
locality
</emphasis>
. Barranco de Masca (Tenerife; UTM coordinates: 28
<collectionCode id="ED147BBD15205F2DFBB1FC3EFB6AE49D" box="[1105,1164,900,926]" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="4" pageNumber="37">RCS</collectionCode>
<quantity id="4CFD4E9D15205F2DFB6CFC3EFACDE49D" box="[1164,1323,900,926]" metricMagnitude="3" metricUnit="m" metricValue="1.83125" pageId="4" pageNumber="37" unit="m" value="1831.25">1831, 250 m</quantity>
altitude).
</paragraph>
<paragraph id="8BBAE37815205F2DFED8FC74FBBDE4EA" blockId="4.[264,1323,900,1902]" box="[312,1115,974,1001]" pageId="4" pageNumber="37">
<typeStatus id="54BE5DDA15205F2DFED8FC74FE4CE4EB" box="[312,426,974,1000]" pageId="4" pageNumber="37" type="holotype">
<emphasis id="B9713F6A15205F2DFED8FC74FE4CE4EB" bold="true" box="[312,426,974,1000]" pageId="4" pageNumber="37">Holotype</emphasis>
</typeStatus>
(
<figureCitation id="133EFFFD15205F2DFE5AFC75FDE7E4EA" box="[442,513,975,1001]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="4" pageNumber="37">Fig. 4</figureCitation>
B):
<collectionCode id="ED147BBD15205F2DFDCCFC75FD9AE4EA" box="[556,636,975,1001]" pageId="4" pageNumber="37">TFMC</collectionCode>
(MT 0387); leg. M. Ibáñez,
<date id="FFBBC5B815205F2DFC2CFC75FBB1E4EA" box="[972,1111,975,1001]" pageId="4" pageNumber="37" value="1983-11-14">14­11­1983</date>
.
</paragraph>
<paragraph id="8BBAE37815205F2DFED8FC4CFE6CE362" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<typeStatus id="54BE5DDA15205F2DFED8FC4CFE53E313" box="[312,437,1014,1040]" pageId="4" pageNumber="37" type="paratype">
<emphasis id="B9713F6A15205F2DFED8FC4CFE53E313" bold="true" box="[312,437,1014,1040]" pageId="4" pageNumber="37">Paratypes</emphasis>
</typeStatus>
. 256
<typeStatus id="54BE5DDA15205F2DFE1DFC4DFD8BE312" box="[509,621,1015,1041]" pageId="4" pageNumber="37" type="paratype">paratypes</typeStatus>
(49 alcohol specimens and 207 shells, leg. between
<date id="FFBBC5B815205F2DFB0EFC4DFE9AE33A" pageId="4" pageNumber="37" value="1982" valueMax="2005">1982 and 2005</date>
).
<collectionCode id="ED147BBD15205F2DFE75FBA5FE03E33A" box="[405,485,1055,1081]" pageId="4" pageNumber="37">TFMC</collectionCode>
(MT 0277/1),
<collectionCode id="ED147BBD15205F2DFD7EFBA5FD3AE33A" box="[670,732,1055,1081]" httpUri="http://grbio.org/cool/zeas-t0vi" name="National Museum of Prague" pageId="4" pageNumber="37">CGH</collectionCode>
(1
<typeStatus id="54BE5DDA15205F2DFCEBFBA5FC92E33A" box="[779,884,1055,1081]" pageId="4" pageNumber="37" type="paratype">paratype</typeStatus>
),
<collectionCode id="ED147BBD15205F2DFC6CFBA5FC26E33A" box="[908,960,1055,1081]" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="4" pageNumber="37">CFS</collectionCode>
(20
<typeStatus id="54BE5DDA15205F2DFC1EFBA5FB96E33A" box="[1022,1136,1055,1081]" pageId="4" pageNumber="37" type="paratype">paratypes</typeStatus>
) and AIT (235
<typeStatus id="54BE5DDA15205F2DFEE8FBFDFE98E362" box="[264,382,1095,1121]" pageId="4" pageNumber="37" type="paratype">paratypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C31FB0F315205F2DFED8FBD4FCF2E3DA" pageId="4" pageNumber="37" type="etymology">
<paragraph id="8BBAE37815205F2DFED8FBD4FCF2E3DA" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<emphasis id="B9713F6A15205F2DFED8FBD4FE5BE38B" bold="true" box="[312,445,1134,1160]" pageId="4" pageNumber="37">Etymology</emphasis>
. The specific name derives from the family name of Dr. Folco Giusti (University of Siena), to whom we dedicate this species for his remarkable contributions to the knowledge of land and freshwater snails.
</paragraph>
</subSubSection>
<subSubSection id="C31FB0F315205F2DFED8FB5CFCB3E27A" pageId="4" pageNumber="37" type="distribution">
<paragraph id="8BBAE37815205F2DFED8FB5CFCB3E27A" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<emphasis id="B9713F6A15205F2DFED8FB5CFD88E203" bold="true" box="[312,622,1254,1280]" pageId="4" pageNumber="37">Distribution and habitat</emphasis>
(
<figureCitation id="133EFFFD15205F2DFD60FB5DFD2EE202" box="[640,712,1255,1281]" captionStart="FIGURE 1" captionStartId="1.[264,367,1802,1826]" captionTargetBox="[275,1313,896,1765]" captionTargetId="figure@1.[264,1323,889,1778]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of Canariella pontelirae and C. giustii sp. nov.; symbols represent 1 x 1 km UTM squares." httpUri="https://zenodo.org/record/173128/files/figure.png" pageId="4" pageNumber="37">Fig. 1</figureCitation>
): The species is endemic to Tenerife, occupying a range of habitats, from dry open areas through to humid laurel forests (“laurisilva”), between
<quantity id="4CFD4E9D15205F2DFE92FA8DFDF4E252" box="[370,530,1335,1361]" metricMagnitude="2" metricUnit="m" metricValue="5.45" metricValueMax="10.5" metricValueMin="0.4" pageId="4" pageNumber="37" unit="m" value="545.0" valueMax="1050.0" valueMin="40.0">40 to 1050 m</quantity>
altitude. In the area in which it co­occurs with
<taxonomicName id="4C0598FB15205F2DFBAAFA8DFB00E253" box="[1098,1254,1335,1360]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="37" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A15205F2DFBAAFA8DFB00E253" box="[1098,1254,1335,1360]" italics="true" pageId="4" pageNumber="37">C. pontelirae</emphasis>
</taxonomicName>
, both species are also found subfossil to depths of
<specimenCount id="9D0328F115205F2DFCF2FAE5FCB3E27A" box="[786,853,1375,1401]" pageId="4" pageNumber="37" type="generic">
<quantity id="4CFD4E9D15205F2DFCF2FAE5FCB7E27A" box="[786,849,1375,1401]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="37" unit="m" value="15.0">15 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C31FB0F315205F2DFED8FA3CFE4EE142" pageId="4" pageNumber="37" type="diagnosis">
<paragraph id="8BBAE37815205F2DFED8FA3CFE4EE142" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<emphasis id="B9713F6A15205F2DFED8FA3CFE56E2A3" bold="true" box="[312,432,1414,1440]" pageId="4" pageNumber="37">Diagnosis</emphasis>
: A medium­sized
<taxonomicName id="4C0598FB15205F2DFD75FA3DFCF2E2A3" box="[661,788,1415,1440]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="37" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A15205F2DFD75FA3DFCF2E2A3" box="[661,788,1415,1440]" italics="true" pageId="4" pageNumber="37">Canariella</emphasis>
</taxonomicName>
with a depressed, dorso­ventrally flattened, discoidal, umbilicate shell, angular at the periphery; 4¾5½ whorls. Epiphallus opening laterally on the proximal side of the broadened penis. Two epiphallar folds extending in the proximal part of the penial cavity without forming the penial papilla that is present in other species.
</paragraph>
</subSubSection>
<subSubSection id="C31FB0F315205F2EFED8F9F4FC7DE538" lastPageId="7" lastPageNumber="40" pageId="4" pageNumber="37" type="description">
<paragraph id="8BBAE37815205F2DFED8F9F4FC86E06D" blockId="4.[264,1323,900,1902]" pageId="4" pageNumber="37">
<emphasis id="B9713F6A15205F2DFED8F9F4FE2FE16B" bold="true" box="[312,457,1614,1640]" pageId="4" pageNumber="37">Description</emphasis>
: Shell (
<figureCitation id="133EFFFD15205F2DFDC9F9F5FD94E16A" box="[553,626,1615,1641]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="4" pageNumber="37">Fig. 4</figureCitation>
B) dextral, depressed, dorso­ventrally flattened, angular at periphery, with a small to medium­sized diameter and a spire of 4¾5½ slightly convex whorls and a marked suture; umbilicus about 14% of shell diameter, slightly obscured by the reflected lip. Mouth rounded­ovate with a discontinuous peristome, slightly expanded as a pale lip at the basal­columellar edge. Colour uniform matt brown. Protoconch with small granulations; teleoconch densely, regularly, radially ribbed, ventral ribs slightly smoother than dorsal ones. Thin periostracal hairs present on periphery (up to 800 μm long) but only up to 140 μm long on the umbilicus.
</paragraph>
<paragraph id="8BBAE37815215F2CFEE8FE9CFCDDE63C" blockId="5.[264,827,291,319]" box="[264,827,291,319]" pageId="5" pageNumber="38">
<emphasis id="B9713F6A15215F2CFEE8FE9CFE9DE63D" bold="true" box="[264,379,294,318]" pageId="5" pageNumber="38">TABLE 1.</emphasis>
Data of known living
<taxonomicName id="4C0598FB15215F2CFD8CFE9DFD06E63D" box="[620,736,295,318]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A15215F2CFD8CFE9DFD06E63D" box="[620,736,295,318]" italics="true" pageId="5" pageNumber="38">Canariella</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BBAE37815215F2CFEF0FEE6FB9BE320" pageId="5" pageNumber="38">
<table id="F90511D81521A0D6FEE8FEE6FAC5E320" box="[264,1315,348,1059]" gridcols="3" gridrows="6" pageId="5" pageNumber="38">
<tr id="3535E13A1521A0D6FEE8FEE6FAC5E592" box="[264,1315,348,657]" gridrow="0" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FEE6FE8EE592" box="[264,360,348,657]" gridcol="0" gridrow="0" pageId="5" pageNumber="38">Island Tenerife</th>
<th id="76E488461521A0D6FE23FEE6FC59E592" box="[451,959,348,657]" gridcol="1" gridrow="0" pageId="5" pageNumber="38">
Species name and synonyms
<taxonomicName id="4C0598FB15215F2CFE23FE32FC59E69D" authority="Webb &amp; Berthelot, 1833" authorityName="Webb &amp; Berthelot" authorityYear="1833" box="[451,959,391,415]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A15215F2CFE23FE32FD46E69C" box="[451,672,392,415]" italics="true" pageId="5" pageNumber="38">Canariella hispidula</emphasis>
(Webb &amp; Berthelot, 1833)
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FE10FCD0E6C2" authority="A. Ferussac, 1835" authorityName="A. Ferussac" authorityYear="1835" box="[451,822,426,449]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="bertheloti">
<emphasis id="B9713F6A15215F2CFE23FE10FD8DE6C2" box="[451,619,426,449]" italics="true" pageId="5" pageNumber="38">Helix bertheloti</emphasis>
A. Férussac, 1835
</taxonomicName>
.
<taxonomicName id="4C0598FB15215F2CFE23FE77FCDAE6E0" authority="Shuttleworth, 1852" authorityName="Shuttleworth" authorityYear="1852" box="[451,828,460,484]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="fortunata">
<emphasis id="B9713F6A15215F2CFE23FE77FD80E6E7" box="[451,614,461,484]" italics="true" pageId="5" pageNumber="38">Helix fortunata</emphasis>
Shuttleworth, 1852
</taxonomicName>
a.
<taxonomicName id="4C0598FB15215F2CFE23FE4AFD13E505" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[451,757,495,519]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="lanosa">
<emphasis id="B9713F6A15215F2CFE23FE4AFDACE504" box="[451,586,496,519]" italics="true" pageId="5" pageNumber="38">Helix lanosa</emphasis>
Mousson, 1872
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FDA8FC46E52A" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[451,928,530,553]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="subSpecies" species="hispidula" subSpecies="subhispidula">
<emphasis id="B9713F6A15215F2CFE23FDA8FD13E52A" box="[451,757,530,553]" italics="true" pageId="5" pageNumber="38">Helix hispidula subhispidula</emphasis>
Mousson, 1872
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FD8FFD15E548" authority="Wollaston, 1878" authorityName="Wollaston" authorityYear="1878" box="[451,755,564,588]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="beata">
<emphasis id="B9713F6A15215F2CFE23FD8FFDD8E54F" box="[451,574,565,588]" italics="true" pageId="5" pageNumber="38">Helix beata</emphasis>
Wollaston, 1878
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FDE2FD04E56D" authority="Mabille, 1882" authorityName="Mabille" authorityYear="1882" box="[451,738,599,623]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="everia">
<emphasis id="B9713F6A15215F2CFE23FDE2FDA0E56C" box="[451,582,600,623]" italics="true" pageId="5" pageNumber="38">Helix everia</emphasis>
Mabille, 1882
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FDC0FCF5E592" authority="Gude, 1911" authorityName="Gude" authorityYear="1911" box="[451,787,634,657]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="salteri">
<emphasis id="B9713F6A15215F2CFE23FDC0FD77E592" box="[451,657,634,657]" italics="true" pageId="5" pageNumber="38">Helicodonta salteri</emphasis>
Gude, 1911
</taxonomicName>
</th>
<th id="76E488461521A0D6FBEBFEE6FAC5E592" box="[1035,1315,348,657]" gridcol="2" gridrow="0" pageId="5" pageNumber="38">
Main data Type species Webb &amp; Berthelot (1833), Ibáñez
<emphasis id="B9713F6A15215F2CFBB9FE4AFB6EE504" box="[1113,1160,496,519]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</th>
</tr>
<tr id="3535E13A1521A0D6FEE8FD11FAC5E5EE" box="[264,1315,683,749]" gridrow="1" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FD11FE8EE5EE" box="[264,360,683,749]" gridcol="0" gridrow="1" pageId="5" pageNumber="38">Tenerife</th>
<td id="76E488461521A0D6FE23FD11FC59E5EE" box="[451,959,683,749]" gridcol="1" gridrow="1" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FD11FD1EE5C1" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[451,760,683,706]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="planaria">
<emphasis id="B9713F6A15215F2CFE23FD11FDD8E5C1" box="[451,574,683,706]" italics="true" pageId="5" pageNumber="38">C. planaria</emphasis>
(Lamarck, 1822)
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFD11FAC5E5EE" box="[1035,1315,683,749]" gridcol="2" gridrow="1" pageId="5" pageNumber="38">
Lamarck (1822), Ibáñez
<emphasis id="B9713F6A15215F2CFAEDFD11FBC7E5EE" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FCB9FAC5E446" box="[264,1315,771,837]" gridrow="2" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FCB9FE8EE446" box="[264,360,771,837]" gridcol="0" gridrow="2" pageId="5" pageNumber="38">Tenerife</th>
<td id="76E488461521A0D6FE23FCB9FC59E446" box="[451,959,771,837]" gridcol="1" gridrow="2" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FCB9FCF0E419" authority="Shuttleworth, 1852" authorityName="Shuttleworth" authorityYear="1852" box="[451,790,771,794]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="leprosa">
<emphasis id="B9713F6A15215F2CFE23FCB9FDD7E419" box="[451,561,771,794]" italics="true" pageId="5" pageNumber="38">C. leprosa</emphasis>
(Shuttleworth, 1852)
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFCB9FAC5E446" box="[1035,1315,771,837]" gridcol="2" gridrow="2" pageId="5" pageNumber="38">
Shuttleworth (1852a), Ibáñez
<emphasis id="B9713F6A15215F2CFBB9FC94FB6EE446" box="[1113,1160,814,837]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FCE1FAC5E49E" box="[264,1315,859,925]" gridrow="3" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FCE1FE8EE49E" box="[264,360,859,925]" gridcol="0" gridrow="3" pageId="5" pageNumber="38">Tenerife</th>
<td id="76E488461521A0D6FE23FCE1FC59E49E" box="[451,959,859,925]" gridcol="1" gridrow="3" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FCE1FCEEE471" authority="J. Mabille, 1883" authorityName="J. Mabille" authorityYear="1883" box="[451,776,859,882]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="pthonera">
<emphasis id="B9713F6A15215F2CFE23FCE1FDA4E471" box="[451,578,859,882]" italics="true" pageId="5" pageNumber="38">C. pthonera</emphasis>
(J. Mabille, 1883)
</taxonomicName>
<taxonomicName id="4C0598FB15215F2CFE23FC3CFCBCE49E" authority="Ponsonby &amp; Sykes, 1894" authorityName="Ponsonby &amp; Sykes" authorityYear="1894" box="[451,858,902,925]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="parryi">
<emphasis id="B9713F6A15215F2CFE23FC3CFDA0E49E" box="[451,582,902,925]" italics="true" pageId="5" pageNumber="38">Helix parryi</emphasis>
Ponsonby &amp; Sykes, 1894
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFCE1FAC5E49E" box="[1035,1315,859,925]" gridcol="2" gridrow="3" pageId="5" pageNumber="38">
Mabille (1883), Groh
<emphasis id="B9713F6A15215F2CFB17FCE1FBC7E49E" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1994)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FC0EFAC5E4F6" box="[264,1315,948,1013]" gridrow="4" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FC0EFE8EE4F6" box="[264,360,948,1013]" gridcol="0" gridrow="4" pageId="5" pageNumber="38">Tenerife</th>
<td id="76E488461521A0D6FE23FC0EFC59E4F6" box="[451,959,948,1013]" gridcol="1" gridrow="4" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FC0EFD16E4C8" authority="Hutterer, 1994" authorityName="Hutterer" authorityYear="1994" box="[451,752,948,971]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A15215F2CFE23FC0EFDA9E4C8" box="[451,591,948,971]" italics="true" pageId="5" pageNumber="38">C. pontelirae</emphasis>
Hutterer, 1994
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFC0EFAC5E4F6" box="[1035,1315,948,1013]" gridcol="2" gridrow="4" pageId="5" pageNumber="38">Hutterer (1994), this paper</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FBB1FAC5E320" box="[264,1315,1035,1059]" gridrow="5" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FBB1FE8EE320" box="[264,360,1035,1059]" gridcol="0" gridrow="5" pageId="5" pageNumber="38">Tenerife</th>
<td id="76E488461521A0D6FE23FBB1FC59E320" box="[451,959,1035,1059]" gridcol="1" gridrow="5" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FBB6FD02E320" authority="Ibanez &amp; Alonso" authorityName="Ibanez &amp; Alonso" box="[451,740,1036,1059]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B9713F6A15215F2CFE23FBB6FDC2E320" box="[451,548,1036,1059]" italics="true" pageId="5" pageNumber="38">C. giustii</emphasis>
Ibáñez &amp; Alonso
</taxonomicName>
,
<taxonomicNameLabel id="A242821115215F2CFD0FFBB1FCA2E320" box="[751,836,1035,1059]" pageId="5" pageNumber="38" rank="species">
<emphasis id="B9713F6A15215F2CFD0FFBB1FCA2E320" bold="true" box="[751,836,1035,1059]" pageId="5" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
</td>
<td id="76E488461521A0D6FBEBFBB1FAC5E320" box="[1035,1315,1035,1059]" gridcol="2" gridrow="5" pageId="5" pageNumber="38">This paper</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBAE37815215F2CFEF0FB83FB11E353" blockId="5.[272,1271,1081,1190]" box="[272,1271,1081,1105]" pageId="5" pageNumber="38">
La Gomera
<taxonomicName id="4C0598FB15215F2CFE23FB80FCDDE353" authority="Shuttleworth, 1852" authorityName="Shuttleworth" authorityYear="1852" box="[451,827,1081,1105]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="discobolus">
<emphasis id="B9713F6A15215F2CFE23FB80FDB2E352" box="[451,596,1082,1105]" italics="true" pageId="5" pageNumber="38">C. discobolus</emphasis>
(Shuttleworth, 1852)
</taxonomicName>
,
<bibRefCitation id="EF949E8915215F2CFBECFB83FB15E353" author="Shuttleworth" box="[1036,1267,1081,1104]" pageId="5" pageNumber="45" refString="Shuttleworth, R. J. (1852 b) Diagnosen neuer Mollusken. Mittheilungen der naturforschenden Gesellschaft in Bern, 260 / 261, 289 - 304." type="journal article" year="1852" yearSuffix="b">Shuttleworth (1852b)</bibRefCitation>
,
</paragraph>
<paragraph id="8BBAE37815215F2CFE23FBDEFB3BE378" blockId="5.[272,1271,1081,1190]" box="[451,1245,1124,1147]" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FBDEFD28E378" authority="A. Ferussac" authorityName="A. Ferussac" box="[451,718,1124,1147]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="afficta">
<emphasis id="B9713F6A15215F2CFE23FBDEFDA1E378" box="[451,583,1124,1147]" italics="true" pageId="5" pageNumber="38">Helix afficta</emphasis>
A. Férussac
</taxonomicName>
,
<emphasis id="B9713F6A15215F2CFD39FBDEFD08E378" box="[729,750,1124,1147]" italics="true" pageId="5" pageNumber="38">in</emphasis>
Férussac &amp; Deshayes, Ibáñez
<emphasis id="B9713F6A15215F2CFBB9FBDEFB6EE378" box="[1113,1160,1124,1147]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</paragraph>
<paragraph id="8BBAE37815215F2CFE23FB35FE1EE3A5" blockId="5.[272,1271,1081,1190]" box="[451,504,1167,1190]" pageId="5" pageNumber="38">1832</paragraph>
<paragraph id="8BBAE37815215F2CFEF0FB07FB05E2DC" pageId="5" pageNumber="38">
<table id="F90511D81521A0D6FEE8FB07FAC5E2E3" box="[264,1315,1213,1504]" gridcols="3" gridrows="5" pageId="5" pageNumber="38">
<tr id="3535E13A1521A0D6FEE8FB07FAC5E3FC" box="[264,1315,1213,1279]" gridrow="0" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FB07FE6FE3FC" box="[264,393,1213,1279]" gridcol="0" gridrow="0" pageId="5" pageNumber="38">La Gomera</th>
<th id="76E488461521A0D6FE23FB07FC2EE3FC" box="[451,968,1213,1279]" gridcol="1" gridrow="0" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FB07FCCBE3D7" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[451,813,1213,1236]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="multigranosa">
<emphasis id="B9713F6A15215F2CFE23FB07FD96E3D7" box="[451,624,1213,1236]" italics="true" pageId="5" pageNumber="38">C. multigranosa</emphasis>
(Mousson, 1872)
</taxonomicName>
</th>
<th id="76E488461521A0D6FBEBFB07FAC5E3FC" box="[1035,1315,1213,1279]" gridcol="2" gridrow="0" pageId="5" pageNumber="38">
Mousson (1872), Groh
<emphasis id="B9713F6A15215F2CFAE6FB07FBC7E3FC" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1994)
</th>
</tr>
<tr id="3535E13A1521A0D6FEE8FAAFFAC5E254" box="[264,1315,1301,1367]" gridrow="1" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FAAFFE6FE254" box="[264,393,1301,1367]" gridcol="0" gridrow="1" pageId="5" pageNumber="38">La Gomera</th>
<td id="76E488461521A0D6FE23FAAFFC2EE254" box="[451,968,1301,1367]" gridcol="1" gridrow="1" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FAAFFCE0E22F" authority="Wollaston, 1878" authorityName="Wollaston" authorityYear="1878" box="[451,774,1301,1324]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="gomerae">
<emphasis id="B9713F6A15215F2CFE23FAAFFDD9E22F" box="[451,575,1301,1324]" italics="true" pageId="5" pageNumber="38">C. gomerae</emphasis>
(Wollaston, 1878)
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFAAFFAC5E254" box="[1035,1315,1301,1367]" gridcol="2" gridrow="1" pageId="5" pageNumber="38">
Wollaston (1878), Ibáñez
<emphasis id="B9713F6A15215F2CFBECFAFAFBDDE254" box="[1036,1083,1344,1367]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FAD7FAC5E286" box="[264,1315,1389,1413]" gridrow="2" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FAD7FE6FE286" box="[264,393,1389,1413]" gridcol="0" gridrow="2" pageId="5" pageNumber="38">La Gomera</th>
<td id="76E488461521A0D6FE23FAD7FC2EE286" box="[451,968,1389,1413]" gridcol="1" gridrow="2" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FAD4FC55E287" authority="Alonso, Ibanez &amp; Ponte-Lira, 2002" authorityName="Alonso, Ibanez &amp; Ponte-Lira" authorityYear="2002" box="[451,947,1389,1413]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="falkneri">
<emphasis id="B9713F6A15215F2CFE23FAD4FDD3E286" box="[451,565,1390,1413]" italics="true" pageId="5" pageNumber="38">C. falkneri</emphasis>
Alonso, Ibáñez &amp; Ponte­Lira, 2002
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFAD7FAC5E286" box="[1035,1315,1389,1413]" gridcol="2" gridrow="2" pageId="5" pageNumber="38">
Alonso
<emphasis id="B9713F6A15215F2CFBBFFAD4FB6BE286" box="[1119,1165,1390,1413]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2002)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FA21FAC5E2B1" box="[264,1315,1435,1458]" gridrow="3" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FA21FE6FE2B1" box="[264,393,1435,1458]" gridcol="0" gridrow="3" pageId="5" pageNumber="38">La Gomera</th>
<td id="76E488461521A0D6FE23FA21FC2EE2B1" box="[451,968,1435,1458]" gridcol="1" gridrow="3" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FA21FD10E2B1" authority="Ponte-Lira, 2002" authorityName="Ponte-Lira" authorityYear="2002" box="[451,758,1435,1458]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="ronceroi">
<emphasis id="B9713F6A15215F2CFE23FA21FDDAE2B1" box="[451,572,1435,1458]" italics="true" pageId="5" pageNumber="38">C. ronceroi</emphasis>
Ponte­Lira, 2002
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFA21FAC5E2B1" box="[1035,1315,1435,1458]" gridcol="2" gridrow="3" pageId="5" pageNumber="38">
Ibáñez
<emphasis id="B9713F6A15215F2CFBB9FA21FB6EE2B1" box="[1113,1160,1435,1458]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2002)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8FA72FAC5E2E3" box="[264,1315,1480,1504]" gridrow="4" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8FA72FE6FE2E3" box="[264,393,1480,1504]" gridcol="0" gridrow="4" pageId="5" pageNumber="38">La Gomera</th>
<td id="76E488461521A0D6FE23FA72FC2EE2E3" box="[451,968,1480,1504]" gridcol="1" gridrow="4" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23FA73FC2EE2DC" authority="Alonso, Ibanez &amp; Ponte-Lira, 2003" authorityName="Alonso, Ibanez &amp; Ponte-Lira" authorityYear="2003" box="[451,968,1480,1504]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="squamata">
<emphasis id="B9713F6A15215F2CFE23FA73FDACE2E3" box="[451,586,1481,1504]" italics="true" pageId="5" pageNumber="38">C. squamata</emphasis>
Alonso, Ibáñez &amp; Ponte­Lira, 2003
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBFA72FAC5E2E3" box="[1035,1315,1480,1504]" gridcol="2" gridrow="4" pageId="5" pageNumber="38">
Alonso
<emphasis id="B9713F6A15215F2CFBBFFA73FB6BE2E3" box="[1119,1165,1481,1504]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2003)
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBAE37815215F2FFEF0FA4CFD61E03D" blockId="5.[272,1251,1526,1549]" lastBlockId="6.[264,1324,1679,1891]" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
La Gomera
<taxonomicName id="4C0598FB15215F2FFE23FA4CFE68E1AA" authority="Alonso, Ibanez &amp; Ponte-Lira, 2003 Alonso et al. (2003) Genital" authorityName="Alonso, Ibanez &amp; Ponte-Lira, 2003 Alonso et al. (2003) Genital" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="6" lastPageNumber="39" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="tenuicostulata">
<emphasis id="B9713F6A15215F2CFE23FA4CFD9FE10E" box="[451,633,1526,1549]" italics="true" pageId="5" pageNumber="38">C. tenuicostulata</emphasis>
<bibRefCitation id="EF949E8915215F2CFD9FFA4CFC11E10E" author="Alonso" box="[639,1015,1526,1549]" pageId="5" pageNumber="44" refString="Alonso, M. R., Ibanez, M. &amp; Ponte-Lira, C. E. (2003) Canariella (Salvinia), new subgenus, and three new species of Canariella Hesse, 1918 (Gastropoda: Pulmonata: Hygromiidae). The Veliger, 46, 69 - 76." type="journal article" year="2003">Alonso, Ibáñez &amp; Ponte­Lira, 2003</bibRefCitation>
<bibRefCitation id="EF949E8915215F2CFBECFA4CFB05E10E" box="[1036,1251,1526,1549]" pageId="5" pageNumber="44" refString="Alonso, M. R., Ibanez, M. &amp; Ponte-Lira, C. E. (2003) Canariella (Salvinia), new subgenus, and three new species of Canariella Hesse, 1918 (Gastropoda: Pulmonata: Hygromiidae). The Veliger, 46, 69 - 76." type="journal article">
Alonso
<emphasis id="B9713F6A15215F2CFBBFFA4CFB6BE10E" box="[1119,1165,1526,1549]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2003)
</bibRefCitation>
Genital
</taxonomicName>
system (
<specimenCount id="9D0328F115225F2FFE19F935FD6DE1AA" box="[505,651,1679,1705]" pageId="6" pageNumber="39" type="generic">7 specimens</specimenCount>
dissected;
<figureCitation id="133EFFFD15225F2FFCEFF935FC86E1AA" box="[783,864,1679,1705]" captionStart="FIGURE 5" captionStartId="8.[264,367,1556,1580]" captionTargetBox="[360,1237,312,1549]" captionTargetId="figure@8.[350,1245,306,1558]" captionTargetPageId="8" captionText="FIGURE 5. Drawings of genital systems and details of distal portions. A. Canariella bimbachensis from Las Lajas (El Hierro; taken from Ibáñez et al. 2002, Fig. 1 B). B. C. discobolus from Barranco de La Rajita (La Gomera; taken from Ibáñez et al. 1995, Fig. 34). C. C. hispidula from Tabaiba Alta (Tenerife; taken from Ibáñez et al. 1995, Fig. 28). D. C. giustii sp. nov. from La Punta (Teno massif, Tenerife). a, general appearance of the whole genital system; at, atrium; ag, albumen gland; b, detail of the distal female duct; bc, bursa copulatrix; c, detail of the distal male duct; d, epiphallus­penis diagram (without scale); e, vaginal cross­section diagram with arrangement of vaginal glands (without scale); ef, epiphallar fold; ep, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; p, penis; pi, pilaster; pp, penial papilla; r, retractor muscle; s, sheath; t, body tegument; v, vagina; vd, vas deferens; vg, vaginal gland." httpUri="https://zenodo.org/record/173131/files/figure.png" pageId="6" pageNumber="39">Figs. 5</figureCitation>
D, 6B): Atrium short. Distal male duct between atrium and penis retractor muscle insertion almost twice as long as the epiphallar proximal portion and 56 times longer than flagellum. Flagellum and epiphallus tubular, the latter opening laterally on the proximal side of the broadened penis, whose diameter is twice than that of the epiphallus.
</paragraph>
<paragraph id="8BBAE37815215F2CFEF0F999FB69E045" pageId="5" pageNumber="38">
<table id="F90511D81521A0D6FEE8F999FAC5E045" box="[264,1315,1571,1862]" gridcols="3" gridrows="5" pageId="5" pageNumber="38">
<tr id="3535E13A1521A0D6FEE8F999FAC5E139" box="[264,1315,1571,1594]" gridrow="0" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8F999FE4EE139" box="[264,424,1571,1594]" gridcol="0" gridrow="0" pageId="5" pageNumber="38">La Palma</th>
<th id="76E488461521A0D6FE23F999FC45E139" box="[451,931,1571,1594]" gridcol="1" gridrow="0" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23F999FC45E139" authority="Alonso, Ibanez &amp; Ponte-Lira, 2003" authorityName="Alonso, Ibanez &amp; Ponte-Lira" authorityYear="2003" box="[451,931,1571,1594]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="tillieri">
<emphasis id="B9713F6A15215F2CFE23F999FDC3E139" box="[451,549,1571,1594]" italics="true" pageId="5" pageNumber="38">C. tillieri</emphasis>
Alonso, Ibáñez &amp; Ponte­Lira, 2003
</taxonomicName>
</th>
<th id="76E488461521A0D6FBEBF999FAC5E139" box="[1035,1315,1571,1594]" gridcol="2" gridrow="0" pageId="5" pageNumber="38">
Alonso
<emphasis id="B9713F6A15215F2CFBBFF999FB6BE139" box="[1119,1165,1571,1594]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2003)
</th>
</tr>
<tr id="3535E13A1521A0D6FEE8F9EBFAC5E16B" box="[264,1315,1617,1640]" gridrow="1" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8F9EBFE4EE16B" box="[264,424,1617,1640]" gridcol="0" gridrow="1" pageId="5" pageNumber="38">El Hierro</th>
<td id="76E488461521A0D6FE23F9EBFC45E16B" box="[451,931,1617,1640]" gridcol="1" gridrow="1" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23F9EBFCB4E16B" authority="Ponte-Lira &amp; Groh, 1994" authorityName="Ponte-Lira &amp; Groh" authorityYear="1994" box="[451,850,1617,1640]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="huttereri">
<emphasis id="B9713F6A15215F2CFE23F9EBFDD8E16B" box="[451,574,1617,1640]" italics="true" pageId="5" pageNumber="38">C. huttereri</emphasis>
Ponte­Lira &amp; Groh, 1994
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBF9EBFAC5E16B" box="[1035,1315,1617,1640]" gridcol="2" gridrow="1" pageId="5" pageNumber="38">
Groh
<emphasis id="B9713F6A15215F2CFBA9F9EBFB9EE16B" box="[1097,1144,1617,1640]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1994)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8F9C4FAC5E195" box="[264,1315,1662,1686]" gridrow="2" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8F9C4FE4EE195" box="[264,424,1662,1686]" gridcol="0" gridrow="2" pageId="5" pageNumber="38">El Hierro</th>
<td id="76E488461521A0D6FE23F9C4FC45E195" box="[451,931,1662,1686]" gridcol="1" gridrow="2" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23F9C5FC94E196" authority="Ibanez &amp; Alonso, 2002" authorityName="Ibanez &amp; Alonso" authorityYear="2002" box="[451,882,1662,1686]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="bimbachensis">
<emphasis id="B9713F6A15215F2CFE23F9C5FD95E195" box="[451,627,1663,1686]" italics="true" pageId="5" pageNumber="38">C. bimbachensis</emphasis>
Ibáñez &amp; Alonso, 2002
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBF9C4FAC5E195" box="[1035,1315,1662,1686]" gridcol="2" gridrow="2" pageId="5" pageNumber="38">
Ibáñez
<emphasis id="B9713F6A15215F2CFBB9F9C5FB6EE195" box="[1113,1160,1663,1686]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (2002)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8F916FAC5E1ED" box="[264,1315,1708,1774]" gridrow="3" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8F916FE4EE1ED" box="[264,424,1708,1774]" gridcol="0" gridrow="3" pageId="5" pageNumber="38">Fuerteventura</th>
<td id="76E488461521A0D6FE23F916FC45E1ED" box="[451,931,1708,1774]" gridcol="1" gridrow="3" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23F916FDDEE1C0" box="[451,568,1708,1731]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="eutropis">
<emphasis id="B9713F6A15215F2CFE23F916FDDEE1C0" box="[451,568,1708,1731]" italics="true" pageId="5" pageNumber="38">C. eutropis</emphasis>
</taxonomicName>
(Shuttleworth
<emphasis id="B9713F6A15215F2CFD38F916FD0BE1C0" box="[728,749,1708,1731]" italics="true" pageId="5" pageNumber="38">in</emphasis>
Pfeiffer, 1861)
</td>
<td id="76E488461521A0D6FBEBF916FAC5E1ED" box="[1035,1315,1708,1774]" gridcol="2" gridrow="3" pageId="5" pageNumber="38">
Pfeiffer (1861), Ibáñez
<emphasis id="B9713F6A15215F2CFAE5F916FBC7E1ED" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1995)
</td>
</tr>
<tr id="3535E13A1521A0D6FEE8F8BEFAC5E045" box="[264,1315,1796,1862]" gridrow="4" pageId="5" pageNumber="38">
<th id="76E488461521A0D6FEE8F8BEFE4EE045" box="[264,424,1796,1862]" gridcol="0" gridrow="4" pageId="5" pageNumber="38">Lanzarote and Fuerteventura</th>
<td id="76E488461521A0D6FE23F8BEFC45E045" box="[451,931,1796,1862]" gridcol="1" gridrow="4" pageId="5" pageNumber="38">
<taxonomicName id="4C0598FB15215F2CFE23F8BEFD35E018" authority="Lowe, 1861" authorityName="Lowe" authorityYear="1861" box="[451,723,1796,1819]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="38" phylum="Mollusca" rank="species" species="plutonia">
<emphasis id="B9713F6A15215F2CFE23F8BEFDDDE018" box="[451,571,1796,1819]" italics="true" pageId="5" pageNumber="38">C. plutonia</emphasis>
(Lowe, 1861)
</taxonomicName>
</td>
<td id="76E488461521A0D6FBEBF8BEFAC5E045" box="[1035,1315,1796,1862]" gridcol="2" gridrow="4" pageId="5" pageNumber="38">
Lowe (1861), Ponte­Lira
<emphasis id="B9713F6A15215F2CFBECF895FBDDE045" box="[1036,1083,1839,1862]" italics="true" pageId="5" pageNumber="38">et al</emphasis>
. (1997)
</td>
</tr>
</table>
</paragraph>
<caption id="DF7AB3F015225F2FFEE8F993FCFBE163" httpUri="https://zenodo.org/record/173129/files/figure.png" pageId="6" pageNumber="39" targetBox="[273,1312,335,1546]" targetPageId="6">
<paragraph id="8BBAE37815225F2FFEE8F993FCFBE163" blockId="6.[264,1323,1577,1632]" pageId="6" pageNumber="39">
<emphasis id="B9713F6A15225F2FFEE8F993FE65E142" bold="true" box="[264,387,1577,1601]" pageId="6" pageNumber="39">FIGURE 3</emphasis>
. Scatter plots of some shell measurements for
<emphasis id="B9713F6A15225F2FFC97F993FB29E143" box="[887,1231,1577,1600]" italics="true" pageId="6" pageNumber="39">
<taxonomicName id="4C0598FB15225F2FFC97F993FB84E143" box="[887,1122,1577,1600]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="39" phylum="Mollusca" rank="species" species="pontelirae">Canariella pontelirae</taxonomicName>
,
<taxonomicName id="4C0598FB15225F2FFB8CF993FB29E143" box="[1132,1231,1577,1600]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="39" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">C. giustii</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A242821115225F2FFB35F993FACDE142" box="[1237,1323,1577,1601]" pageId="6" pageNumber="39" rank="species">
<emphasis id="B9713F6A15225F2FFB35F993FACDE142" bold="true" box="[1237,1323,1577,1601]" pageId="6" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C0598FB15225F2FFED5F9F3FE51E163" box="[309,439,1609,1632]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="39" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A15225F2FFED5F9F3FE51E163" box="[309,439,1609,1632]" italics="true" pageId="6" pageNumber="39">C. hispidula</emphasis>
</taxonomicName>
. Abbreviations same as in Fig. 2.
</paragraph>
</caption>
<paragraph id="8BBAE37815225F2EFED8F8F3FE7EE6C8" blockId="6.[264,1324,1679,1891]" lastBlockId="7.[264,1323,284,571]" lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
Epiphallus with two internal, distally expanded, longitudinal folds which extend in the penis tilting towards the proximal penial concavity opposite the epiphallar insertion, ending separately but very close to each other, not merged into a penial papilla. Each epiphallar fold distally has a transverse constriction, forming a small, terminal papilla (indicated by arrows in the
<figureCitation id="133EFFFD15235F2EFDAFFE36FD75E6A5" box="[591,659,396,422]" captionStart="FIGURE 5" captionStartId="8.[264,367,1556,1580]" captionTargetBox="[360,1237,312,1549]" captionTargetId="figure@8.[350,1245,306,1558]" captionTargetPageId="8" captionText="FIGURE 5. Drawings of genital systems and details of distal portions. A. Canariella bimbachensis from Las Lajas (El Hierro; taken from Ibáñez et al. 2002, Fig. 1 B). B. C. discobolus from Barranco de La Rajita (La Gomera; taken from Ibáñez et al. 1995, Fig. 34). C. C. hispidula from Tabaiba Alta (Tenerife; taken from Ibáñez et al. 1995, Fig. 28). D. C. giustii sp. nov. from La Punta (Teno massif, Tenerife). a, general appearance of the whole genital system; at, atrium; ag, albumen gland; b, detail of the distal female duct; bc, bursa copulatrix; c, detail of the distal male duct; d, epiphallus­penis diagram (without scale); e, vaginal cross­section diagram with arrangement of vaginal glands (without scale); ef, epiphallar fold; ep, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; p, penis; pi, pilaster; pp, penial papilla; r, retractor muscle; s, sheath; t, body tegument; v, vagina; vd, vas deferens; vg, vaginal gland." httpUri="https://zenodo.org/record/173131/files/figure.png" pageId="7" pageNumber="40">Fig. 5</figureCitation>
Da,c). The penis has 78 longitudinal internal pilasters (
<figureCitation id="133EFFFD15235F2EFEF1FE0BFEB2E6C8" box="[273,340,433,459]" captionStart="FIGURE 5" captionStartId="8.[264,367,1556,1580]" captionTargetBox="[360,1237,312,1549]" captionTargetId="figure@8.[350,1245,306,1558]" captionTargetPageId="8" captionText="FIGURE 5. Drawings of genital systems and details of distal portions. A. Canariella bimbachensis from Las Lajas (El Hierro; taken from Ibáñez et al. 2002, Fig. 1 B). B. C. discobolus from Barranco de La Rajita (La Gomera; taken from Ibáñez et al. 1995, Fig. 34). C. C. hispidula from Tabaiba Alta (Tenerife; taken from Ibáñez et al. 1995, Fig. 28). D. C. giustii sp. nov. from La Punta (Teno massif, Tenerife). a, general appearance of the whole genital system; at, atrium; ag, albumen gland; b, detail of the distal female duct; bc, bursa copulatrix; c, detail of the distal male duct; d, epiphallus­penis diagram (without scale); e, vaginal cross­section diagram with arrangement of vaginal glands (without scale); ef, epiphallar fold; ep, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; p, penis; pi, pilaster; pp, penial papilla; r, retractor muscle; s, sheath; t, body tegument; v, vagina; vd, vas deferens; vg, vaginal gland." httpUri="https://zenodo.org/record/173131/files/figure.png" pageId="7" pageNumber="40">Fig. 5</figureCitation>
Dc).
</paragraph>
<paragraph id="8BBAE37815235F2EFED8FE6DFC7DE538" blockId="7.[264,1323,284,571]" pageId="7" pageNumber="40">
Short vagina proximally widened, with numerous irregular longitudinal folds, some of them anastomosed, and 38 crown­shaped, finger­like vaginal glands, which open near the orifice connecting the vagina to the oviduct (
<figureCitation id="133EFFFD15235F2EFCF5FD9BFCBFE538" box="[789,857,545,571]" captionStart="FIGURE 5" captionStartId="8.[264,367,1556,1580]" captionTargetBox="[360,1237,312,1549]" captionTargetId="figure@8.[350,1245,306,1558]" captionTargetPageId="8" captionText="FIGURE 5. Drawings of genital systems and details of distal portions. A. Canariella bimbachensis from Las Lajas (El Hierro; taken from Ibáñez et al. 2002, Fig. 1 B). B. C. discobolus from Barranco de La Rajita (La Gomera; taken from Ibáñez et al. 1995, Fig. 34). C. C. hispidula from Tabaiba Alta (Tenerife; taken from Ibáñez et al. 1995, Fig. 28). D. C. giustii sp. nov. from La Punta (Teno massif, Tenerife). a, general appearance of the whole genital system; at, atrium; ag, albumen gland; b, detail of the distal female duct; bc, bursa copulatrix; c, detail of the distal male duct; d, epiphallus­penis diagram (without scale); e, vaginal cross­section diagram with arrangement of vaginal glands (without scale); ef, epiphallar fold; ep, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; p, penis; pi, pilaster; pp, penial papilla; r, retractor muscle; s, sheath; t, body tegument; v, vagina; vd, vas deferens; vg, vaginal gland." httpUri="https://zenodo.org/record/173131/files/figure.png" pageId="7" pageNumber="40">Fig. 5</figureCitation>
Da).
</paragraph>
</subSubSection>
<caption id="DF7AB3F015235F2EFEE8F91BFACAE1DB" httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="7" pageNumber="40" targetBox="[264,1323,625,1676]" targetPageId="7">
<paragraph id="8BBAE37815235F2EFEE8F91BFACAE1DB" blockId="7.[264,1324,1697,1753]" pageId="7" pageNumber="40">
<emphasis id="B9713F6A15235F2EFEE8F91BFE6EE1BA" bold="true" box="[264,392,1697,1721]" pageId="7" pageNumber="40">FIGURE 4</emphasis>
. A.
<taxonomicName id="4C0598FB15235F2EFE5FF918FD4DE1BA" box="[447,683,1698,1721]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="40" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A15235F2EFE5FF918FD4DE1BA" box="[447,683,1698,1721]" italics="true" pageId="7" pageNumber="40">Canariella pontelirae</emphasis>
</taxonomicName>
living specimen and shell, from La Punta (Teno massif, Tenerife). B.
<taxonomicName id="4C0598FB15235F2EFE75F978FE11E1DA" box="[405,503,1730,1753]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="40" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B9713F6A15235F2EFE75F978FE11E1DA" box="[405,503,1730,1753]" italics="true" pageId="7" pageNumber="40">C. giustii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242821115235F2EFE1DF97BFDB4E1DA" box="[509,594,1729,1753]" pageId="7" pageNumber="40" rank="species">
<emphasis id="B9713F6A15235F2EFE1DF97BFDB4E1DA" bold="true" box="[509,594,1729,1753]" pageId="7" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype shell. C.
<taxonomicName id="4C0598FB15235F2EFCC0F978FC44E1DA" box="[800,930,1730,1753]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="40" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A15235F2EFCC0F978FC44E1DA" box="[800,930,1730,1753]" italics="true" pageId="7" pageNumber="40">C. hispidula</emphasis>
</taxonomicName>
shell, from Las Caletillas (Tenerife).
</paragraph>
</caption>
<caption id="DF7AB3F0152C5F21FEE8F9AEFD17E066" httpUri="https://zenodo.org/record/173131/files/figure.png" pageId="8" pageNumber="41" targetBox="[360,1237,312,1549]" targetPageId="8">
<paragraph id="8BBAE378152C5F21FEE8F9AEFD17E066" blockId="8.[264,1323,1556,1893]" pageId="8" pageNumber="41">
<emphasis id="B9713F6A152C5F21FEE8F9AEFE65E12F" bold="true" box="[264,387,1556,1580]" pageId="8" pageNumber="41">FIGURE 5</emphasis>
. Drawings of genital systems and details of distal portions. A.
<taxonomicName id="4C0598FB152C5F21FBFFF9AEFACCE128" box="[1055,1322,1556,1579]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="41" phylum="Mollusca" rank="species" species="bimbachensis">
<emphasis id="B9713F6A152C5F21FBFFF9AEFACCE128" box="[1055,1322,1556,1579]" italics="true" pageId="8" pageNumber="41">Canariella bimbachensis</emphasis>
</taxonomicName>
from Las Lajas (El Hierro; taken from Ibáñez
<emphasis id="B9713F6A152C5F21FD13F98DFCC7E14D" box="[755,801,1591,1614]" italics="true" pageId="8" pageNumber="41">et al</emphasis>
. 2002, Fig. 1B). B.
<taxonomicName id="4C0598FB152C5F21FC17F98DFB6EE14D" box="[1015,1160,1591,1614]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="41" phylum="Mollusca" rank="species" species="discobolus">
<emphasis id="B9713F6A152C5F21FC17F98DFB6EE14D" box="[1015,1160,1591,1614]" italics="true" pageId="8" pageNumber="41">C. discobolus</emphasis>
</taxonomicName>
from Barranco de La Rajita (La Gomera; taken from Ibáñez
<emphasis id="B9713F6A152C5F21FD05F9E0FCF5E172" box="[741,787,1626,1649]" italics="true" pageId="8" pageNumber="41">et al</emphasis>
. 1995, Fig. 34). C.
<taxonomicName id="4C0598FB152C5F21FC01F9E0FB84E172" box="[993,1122,1626,1649]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="41" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A152C5F21FC01F9E0FB84E172" box="[993,1122,1626,1649]" italics="true" pageId="8" pageNumber="41">C. hispidula</emphasis>
</taxonomicName>
from Tabaiba Alta (Tenerife; taken from Ibáñez
<emphasis id="B9713F6A152C5F21FDB1F9C6FD62E190" box="[593,644,1660,1683]" italics="true" pageId="8" pageNumber="41">et al</emphasis>
. 1995, Fig. 28). D.
<taxonomicName id="4C0598FB152C5F21FC8BF9C6FC37E190" box="[875,977,1660,1683]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="41" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B9713F6A152C5F21FC8BF9C6FC37E190" box="[875,977,1660,1683]" italics="true" pageId="8" pageNumber="41">C. giustii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2428211152C5F21FC3BF9C6FBD3E197" box="[987,1077,1660,1684]" pageId="8" pageNumber="41" rank="species">
<emphasis id="B9713F6A152C5F21FC3BF9C6FBD3E197" bold="true" box="[987,1077,1660,1684]" pageId="8" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
from La Punta (Teno massif, Tenerife). a, general appearance of the whole genital system; at, atrium; ag, albumen gland; b, detail of the distal female duct; bc, bursa copulatrix; c, detail of the distal male duct; d, epiphallus­penis diagram (without scale); e, vaginal cross­section diagram with arrangement of vaginal glands (without scale); ef, epiphallar fold; ep, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; p, penis; pi, pilaster; pp, penial papilla; r, retractor muscle; s, sheath; t, body tegument; v, vagina; vd, vas deferens; vg, vaginal gland.
</paragraph>
</caption>
<subSubSection id="C31FB0F3152D5F23FED8FEA1FB91E538" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42" type="discussion">
<paragraph id="8BBAE378152D5F20FED8FEA1FC7FE6F2" blockId="9.[264,1323,283,683]" pageId="9" pageNumber="42">
<emphasis id="B9713F6A152D5F20FED8FEA1FE4CE636" bold="true" box="[312,426,283,309]" pageId="9" pageNumber="42">Remarks</emphasis>
.
<taxonomicName id="4C0598FB152D5F20FE21FEA6FDD3E636" box="[449,565,284,309]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152D5F20FE21FEA6FDD3E636" box="[449,565,284,309]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
differs from all the other living
<taxonomicName id="4C0598FB152D5F20FC0DFEA6FB8AE636" box="[1005,1132,284,309]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A152D5F20FC0DFEA6FB8AE636" box="[1005,1132,284,309]" italics="true" pageId="9" pageNumber="42">Canariella</emphasis>
</taxonomicName>
species by its broadened penis. The penis diameter of
<taxonomicName id="4C0598FB152D5F20FD00FEFBFCADE659" box="[736,843,321,346]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152D5F20FD00FEFBFCADE659" box="[736,843,321,346]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
is twice than that of the epiphallus, with an eccentric, lateral­proximal epiphallar opening; the distal male duct broadens rapidly in the proximal part of penis. All the other living
<taxonomicName id="4C0598FB152D5F20FCA9FE36FC2EE6A6" box="[841,968,396,421]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A152D5F20FCA9FE36FC2EE6A6" box="[841,968,396,421]" italics="true" pageId="9" pageNumber="42">Canariella</emphasis>
</taxonomicName>
species have the penis about 11½ times broader than the epiphallus, with a central epiphallar opening, and the distal male duct broadens gently in the proximal part of penis.
</paragraph>
<paragraph id="8BBAE378152D5F20FED8FE46FE48E5A8" blockId="9.[264,1323,283,683]" pageId="9" pageNumber="42">
Moreover,
<taxonomicName id="4C0598FB152D5F20FE5BFE46FDCFE516" box="[443,553,508,533]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152D5F20FE5BFE46FDCFE516" box="[443,553,508,533]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
differs from all the other living
<taxonomicName id="4C0598FB152D5F20FC55FE46FBD2E516" box="[949,1076,508,533]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A152D5F20FC55FE46FBD2E516" box="[949,1076,508,533]" italics="true" pageId="9" pageNumber="42">Canariella</emphasis>
</taxonomicName>
species in its penial anatomy. It is the only
<taxonomicName id="4C0598FB152D5F20FDD8FD9BFD51E539" box="[568,695,545,570]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="B9713F6A152D5F20FDD8FD9BFD51E539" box="[568,695,545,570]" italics="true" pageId="9" pageNumber="42">Canariella</emphasis>
</taxonomicName>
species in which the two epiphallar folds extend together in the penis, tilting towards the proximal penial concavity opposite to the epiphallar insertion and ending very close but independent of each other, not merged into a penial papilla.
</paragraph>
<caption id="DF7AB3F0152D5F20FEE8F923FBE3E1D5" httpUri="https://zenodo.org/record/173132/files/figure.png" pageId="9" pageNumber="42" targetBox="[264,1323,737,1668]" targetPageId="9">
<paragraph id="8BBAE378152D5F20FEE8F923FBE3E1D5" blockId="9.[264,1322,1689,1750]" pageId="9" pageNumber="42">
<emphasis id="B9713F6A152D5F20FEE8F923FE62E1B2" bold="true" box="[264,388,1689,1713]" pageId="9" pageNumber="42">FIGURE 6</emphasis>
. Photographs of genital systems. A.
<taxonomicName id="4C0598FB152D5F20FCEFF923FC1EE1B3" box="[783,1016,1689,1712]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A152D5F20FCEFF923FC1EE1B3" box="[783,1016,1689,1712]" italics="true" pageId="9" pageNumber="42">Canariella pontelirae</emphasis>
</taxonomicName>
. B.
<taxonomicName id="4C0598FB152D5F20FBC5F923FB60E1B3" box="[1061,1158,1689,1712]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii" status="sp. nov.">
<emphasis id="B9713F6A152D5F20FBC5F923FB60E1B3" box="[1061,1158,1689,1712]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
<emphasis id="B9713F6A152D5F20FB6EF923FB02E1B2" bold="true" box="[1166,1252,1689,1713]" pageId="9" pageNumber="42">
<taxonomicNameLabel id="A2428211152D5F20FB6EF923FB02E1B2" box="[1166,1252,1689,1713]" pageId="9" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(both from La Punta, Teno massif, Tenerife). Abbreviations same as in Fig. 5.
</paragraph>
</caption>
<paragraph id="8BBAE378152D5F23FED8F8BDFB91E538" blockId="9.[264,1322,1799,1899]" lastBlockId="10.[264,1323,284,571]" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
The species most similar to
<taxonomicName id="4C0598FB152D5F20FD6CF8BDFD1FE023" box="[652,761,1799,1824]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152D5F20FD6CF8BDFD1FE023" box="[652,761,1799,1824]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
(
<figureCitation id="133EFFFD152D5F20FCECF8BDFCB3E022" box="[780,853,1799,1825]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="9" pageNumber="42">Fig. 4</figureCitation>
B) in shell dimensions are
<taxonomicName id="4C0598FB152D5F20FB79F8BDFACCE023" box="[1177,1322,1799,1824]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A152D5F20FB79F8BDFACCE023" box="[1177,1322,1799,1824]" italics="true" pageId="9" pageNumber="42">C. hispidula</emphasis>
</taxonomicName>
(
<figureCitation id="133EFFFD152D5F20FEF1F896FEBEE045" box="[273,344,1836,1862]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="9" pageNumber="42">Fig. 4</figureCitation>
C) and
<taxonomicName id="4C0598FB152D5F20FE52F896FDA8E046" box="[434,590,1836,1861]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A152D5F20FE52F896FDA8E046" box="[434,590,1836,1861]" italics="true" pageId="9" pageNumber="42">C. pontelirae</emphasis>
</taxonomicName>
(
<figureCitation id="133EFFFD152D5F20FDBFF896FD26E045" box="[607,704,1836,1862]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="9" pageNumber="42">Fig. 4A</figureCitation>
). The
<taxonomicName id="4C0598FB152D5F20FCECF896FC9FE046" box="[780,889,1836,1861]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="42" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152D5F20FCECF896FC9FE046" box="[780,889,1836,1861]" italics="true" pageId="9" pageNumber="42">C. giustii</emphasis>
</taxonomicName>
shell measurements (
<figureCitation id="133EFFFD152D5F20FB9CF896FB25E045" box="[1148,1219,1836,1862]" captionStart="FIGURE 3" captionStartId="6.[264,367,1577,1601]" captionTargetBox="[273,1312,335,1546]" captionTargetId="figure@6.[264,1324,326,1555]" captionTargetPageId="6" captionText="FIGURE 3. Scatter plots of some shell measurements for Canariella pontelirae, C. giustii sp. nov. and C. hispidula. Abbreviations same as in Fig. 2." httpUri="https://zenodo.org/record/173129/files/figure.png" pageId="9" pageNumber="42">Fig. 3</figureCitation>
) occupy an intermediate position between the other two, but with a broad overlap, although the shell height and frontal surface of
<taxonomicName id="4C0598FB152E5F23FD4BFEA6FCFEE636" box="[683,792,284,309]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152E5F23FD4BFEA6FCFEE636" box="[683,792,284,309]" italics="true" pageId="10" pageNumber="43">C. giustii</emphasis>
</taxonomicName>
are bigger than the others in relation to the maximum diameter and frontal perimeter, respectively.
<taxonomicName id="4C0598FB152E5F23FC72FEFBFC1BE659" box="[914,1021,321,346]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152E5F23FC72FEFBFC1BE659" box="[914,1021,321,346]" italics="true" pageId="10" pageNumber="43">C. giustii</emphasis>
</taxonomicName>
has a narrower umbilicus than those of the other two species (
<figureCitation id="133EFFFD152E5F23FD3BFEDDFCCDE682" box="[731,811,359,385]" captionStart="FIGURE 4. A" captionStartId="7.[264,367,1697,1721]" captionTargetBox="[264,1323,625,1676]" captionTargetId="figure@7.[264,1323,625,1676]" captionTargetPageId="7" captionText="FIGURE 4. A. Canariella pontelirae living specimen and shell, from La Punta (Teno massif, Tenerife). B. C. giustii sp. nov., holotype shell. C. C. hispidula shell, from Las Caletillas (Tenerife)." httpUri="https://zenodo.org/record/173130/files/figure.png" pageId="10" pageNumber="43">Fig. 4</figureCitation>
). The
<taxonomicName id="4C0598FB152E5F23FC62FEDDFC15E683" box="[898,1011,359,384]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152E5F23FC62FEDDFC15E683" box="[898,1011,359,384]" italics="true" pageId="10" pageNumber="43">C. giustii</emphasis>
</taxonomicName>
teleoconch is angular at periphery and densely, regularly, radially ribbed, whereas the
<taxonomicName id="4C0598FB152E5F23FC01FE36FB9AE6A6" box="[993,1148,396,421]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="pontelirae">
<emphasis id="B9713F6A152E5F23FC01FE36FB9AE6A6" box="[993,1148,396,421]" italics="true" pageId="10" pageNumber="43">C. pontelirae</emphasis>
</taxonomicName>
teleoconch is keeled and has prominent and regularly spaced radial ribs separated by 45 small ribs. The
<taxonomicName id="4C0598FB152E5F23FEE8FE6DFE7FE6F3" box="[264,409,471,496]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="hispidula">
<emphasis id="B9713F6A152E5F23FEE8FE6DFE7FE6F3" box="[264,409,471,496]" italics="true" pageId="10" pageNumber="43">C. hispidula</emphasis>
</taxonomicName>
shell is the most similar to that of
<taxonomicName id="4C0598FB152E5F23FCD9FE6DFC40E6F3" box="[825,934,471,496]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152E5F23FCD9FE6DFC40E6F3" box="[825,934,471,496]" italics="true" pageId="10" pageNumber="43">C. giustii</emphasis>
</taxonomicName>
in shape and ornamentation, but has a more angular periphery and it is shaggy, whereas that of
<taxonomicName id="4C0598FB152E5F23FC06FE46FBB7E516" box="[998,1105,508,533]" class="Gastropoda" family="Hygromiidae" genus="Canariella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="43" phylum="Mollusca" rank="species" species="giustii">
<emphasis id="B9713F6A152E5F23FC06FE46FBB7E516" box="[998,1105,508,533]" italics="true" pageId="10" pageNumber="43">C. giustii</emphasis>
</taxonomicName>
is almost hairless, the adult live specimens only having well­developed hairs at the periphery.
</paragraph>
</subSubSection>
</treatment>
</document>