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<mods:title id="4CDFFDF997E6F96F1E754FC7BF9FB27E">A new brachypterous species of Phoberus MacLeay (Coleoptera: Trogidae) and status changes of species in the East African P. nigrociliatus group</mods:title>
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<paragraph id="8BBAE341FF8CB03040D92632FD2EFBBB" blockId="3.[151,667,1123,1183]" box="[151,667,1123,1150]" pageId="3" pageNumber="295">
<heading id="D0F2542DFF8CB03040D92632FD2EFBBB" bold="true" box="[151,667,1123,1150]" fontSize="11" level="1" pageId="3" pageNumber="295" reason="1">
<emphasis id="B9713F53FF8CB03040D92632FD2EFBBB" bold="true" box="[151,667,1123,1150]" pageId="3" pageNumber="295">
<taxonomicName id="4C0598C2FF8CB03040D92632FDA5FBBB" ID-CoL="4GJWX" authority="Kolbe" authorityName="Kolbe" box="[151,528,1123,1150]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="nigrociliatus" status="stat. nov.">
<emphasis id="B9713F53FF8CB03040D92632FE1EFBB8" bold="true" box="[151,427,1123,1149]" italics="true" pageId="3" pageNumber="295">Phoberus nigrociliatus</emphasis>
(Kolbe)
</taxonomicName>
<taxonomicNameLabel id="A2428228FF8CB03042562635FD2EFBBB" box="[536,667,1124,1150]" pageId="3" pageNumber="295">new status</taxonomicNameLabel>
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<paragraph id="8BBAE341FF8CB03040D926D7FE95FB5A" blockId="3.[151,667,1123,1183]" box="[151,288,1158,1183]" pageId="3" pageNumber="295">
(
<figureCitation id="133EFFC4FF8CB03040D126D7FF43FB5A" box="[159,246,1158,1183]" captionStart="FIGURE 3" captionStartId="4.[151,250,922,944]" captionTargetBox="[432,1194,244,850]" captionTargetId="figure@4.[381,1206,195,901]" captionTargetPageId="4" captionText="FIGURE 3. A, Dorsal habitus of Phoberus nigrociliatus; B C, dorsal and lateral views of the aedeagus of P. nigrociliatus." httpUri="https://zenodo.org/record/231850/files/figure.png" pageId="3" pageNumber="295">Fig. 3A</figureCitation>
C)
</paragraph>
</subSubSection>
<subSubSection id="C31FB0CAFF8CB03040D92681FF5EFA02" pageId="3" pageNumber="295" type="reference_group">
<paragraph id="8BBAE341FF8CB03040D92681FD08FB21" blockId="3.[151,951,1230,1375]" box="[151,701,1230,1253]" pageId="3" pageNumber="295">
<treatmentCitationGroup id="AB15C46FFF8CB03040D92681FD08FB21" box="[151,701,1230,1253]" pageId="3" pageNumber="295">
<taxonomicName id="4C0598C2FF8CB03040D92681FDB1FB21" ID-CoL="59DNC" authority="Kolbe, 1904: 292" authorityName="Kolbe" authorityPageNumber="292" authorityYear="1904" box="[151,516,1230,1253]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="B9713F53FF8CB03040D92681FEF9FB21" box="[151,332,1230,1253]" italics="true" pageId="3" pageNumber="295">Trox nigrociliatus</emphasis>
<treatmentCitation id="0AA4C550FF8CB030411F269FFDB1FB21" author="Kolbe" box="[337,516,1230,1252]" page="292" pageId="3" pageNumber="295" year="1904">
<bibRefCitation id="EF949EB0FF8CB030411F269FFE64FB21" author="Kolbe" box="[337,465,1230,1252]" pageId="3" pageNumber="295" refString="Kolbe, H. J. (1904) Uber einige interessante Lamellicornier und Tenebrioniden Afrikas. Berliner Entomologische Zeitschrift, 49, 282 - 302." type="journal article" year="1904">Kolbe, 1904</bibRefCitation>
: 292
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AA4C550FF8CB0304241269EFD08FB21" author="Arrow" box="[527,701,1230,1252]" page="465" pageId="3" pageNumber="295" year="1927">
<bibRefCitation id="EF949EB0FF8CB0304241269EFD38FB21" author="Arrow" box="[527,653,1230,1252]" pageId="3" pageNumber="295" refString="Arrow, G. J. (1927) A note on the coleopterous genus Trox, with description of a few new Asiatic species. Annals and Magazine of Natural History, series 9, volume 19, 465 - 468." type="journal article" year="1927">Arrow 1927</bibRefCitation>
: 465
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</paragraph>
<paragraph id="8BBAE341FF8CB03040D926BFFDC8FAC6" blockId="3.[151,951,1230,1375]" box="[151,637,1261,1283]" pageId="3" pageNumber="295">
<treatmentCitationGroup id="AB15C46FFF8CB03040D926BFFDC8FAC6" box="[151,637,1261,1283]" pageId="3" pageNumber="295">
<taxonomicName id="4C0598C2FF8CB03040D926BFFDC8FAC6" ID-CoL="59DNC" authority="Haaf 1953: 328" authorityName="Haaf" authorityPageNumber="328" authorityYear="1953" box="[151,637,1261,1283]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subSpecies="nigrociliatus">
<emphasis id="B9713F53FF8CB03040D926BFFE67FAC6" box="[151,466,1261,1283]" italics="true" pageId="3" pageNumber="295">Trox nigrociliatus nigrociliatus</emphasis>
:
<treatmentCitation id="0AA4C550FF8CB030419126BCFDC8FAC6" author="Haaf" box="[479,637,1261,1283]" page="328" pageId="3" pageNumber="295" year="1953">
<bibRefCitation id="EF949EB0FF8CB030419126BCFDF8FAC6" author="Haaf" box="[479,589,1261,1283]" pageId="3" pageNumber="295" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey, 4, 309 - 346." type="journal article" year="1953">Haaf 1953</bibRefCitation>
: 328
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</taxonomicName>
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</paragraph>
<paragraph id="8BBAE341FF8CB03040D9275CFCDEFA9A" blockId="3.[151,951,1230,1375]" pageId="3" pageNumber="295">
<treatmentCitationGroup id="AB15C46FFF8CB03040D9275CFCDEFA9A" pageId="3" pageNumber="295">
<taxonomicName id="4C0598C2FF8CB03040D9275CFD67FAE7" ID-CoL="59DNC" authority="Scholtz 1980: 92" authorityName="Scholtz" authorityPageNumber="92" authorityYear="1980" box="[151,722,1292,1314]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subGenus="Trox" subSpecies="nigrociliatus">
<emphasis id="B9713F53FF8CB03040D9275CFF71FAE7" box="[151,196,1293,1314]" italics="true" pageId="3" pageNumber="295">Trox</emphasis>
(
<emphasis id="B9713F53FF8CB030409D275CFF4AFAE7" box="[211,255,1293,1314]" italics="true" pageId="3" pageNumber="295">Trox</emphasis>
)
<emphasis id="B9713F53FF8CB0304141275DFDA3FAE7" box="[271,534,1292,1314]" italics="true" pageId="3" pageNumber="295">nigrociliatus nigrociliatus</emphasis>
:
<treatmentCitation id="0AA4C550FF8CB030426A275DFD67FAE7" author="Scholtz" box="[548,722,1292,1314]" page="92" pageId="3" pageNumber="295" year="1980">
<bibRefCitation id="EF949EB0FF8CB030426A275DFD19FAE7" author="Scholtz" box="[548,684,1292,1314]" pageId="3" pageNumber="295" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of Subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980</bibRefCitation>
: 92
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AA4C550FF8CB0304293275DFCC9FAE7" author="Scholtz" box="[733,892,1292,1314]" page="5" pageId="3" pageNumber="295" year="1982">
<bibRefCitation id="EF949EB0FF8CB0304293275DFCD0FAE7" author="Scholtz" box="[733,869,1292,1314]" pageId="3" pageNumber="295" refString="Scholtz, C. H. (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries, Republic of South Africa, 54, 1 - 27." type="journal article" year="1982">Scholtz 1982</bibRefCitation>
: 5
</treatmentCitation>
<taxonomicName id="4C0598C2FF8CB03040D9277AFCA6FA85" ID-CoL="59DNC" authority="Scholtz 1986: 361" authorityName="Scholtz" authorityPageNumber="361" authorityYear="1986" box="[151,787,1322,1344]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subGenus="Phoberus" subSpecies="nigrociliatus">
<emphasis id="B9713F53FF8CB03040D9277AFF71FA85" box="[151,196,1323,1344]" italics="true" pageId="3" pageNumber="295">Trox</emphasis>
(
<emphasis id="B9713F53FF8CB030409D277BFE81FA85" box="[211,308,1322,1344]" italics="true" pageId="3" pageNumber="295">Phoberus</emphasis>
)
<emphasis id="B9713F53FF8CB030410D277BFDFFFA85" box="[323,586,1322,1344]" italics="true" pageId="3" pageNumber="295">nigrociliatus nigrociliatus</emphasis>
:
<treatmentCitation id="0AA4C550FF8CB0304219277BFCA6FA85" author="Scholtz" box="[599,787,1322,1344]" page="361" pageId="3" pageNumber="295" year="1986">
<bibRefCitation id="EF949EB0FF8CB0304219277BFD55FA85" author="Scholtz" box="[599,736,1322,1344]" pageId="3" pageNumber="295" refString="Scholtz, C. H. (1986) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology, 11, 355 - 363." type="journal article" year="1986">Scholtz 1986</bibRefCitation>
: 361
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AA4C550FF8CB0304353277BFC02FA85" author="Zidek" box="[797,951,1322,1344]" page="13" pageId="3" pageNumber="295" year="2013">
<bibRefCitation id="EF949EB0FF8CB0304353277BFC21FA85" author="Zidek" box="[797,916,1322,1344]" pageId="3" pageNumber="295" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek 2013</bibRefCitation>
: 13
</treatmentCitation>
<taxonomicName id="4C0598C2FF8CB03040D92718FDEBFA9A" ID-CoL="4GJWX" authority="Kolbe" authorityName="Kolbe" box="[151,606,1353,1375]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="subSpecies" species="nigrociliatus" subSpecies="nigrociliatus">
<emphasis id="B9713F53FF8CB03040D92718FDB3FA9A" box="[151,518,1353,1375]" italics="true" pageId="3" pageNumber="295">Phoberus nigrociliatus nigrociliatus</emphasis>
(Kolbe)
</taxonomicName>
:
<treatmentCitation id="0AA4C550FF8CB03042262718FCDEFA9A" author="Strumpher" box="[616,875,1353,1375]" page="78" pageId="3" pageNumber="295" year="2016">
<bibRefCitation id="EF949EB0FF8CB03042262718FCFDFA9A" author="Strumpher" box="[616,840,1353,1375]" pageId="3" pageNumber="295" refString="Strumpher, W. P., Villet, M. H., Sole, C. L. &amp; Scholtz, C. H. (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics and Evolution, 47, 53 - 82. https: // doi. org / 10.1163 / 1876312 X- 46052133" type="journal article" year="2016">
Strümpher
<emphasis id="B9713F53FF8CB0304296271BFCB0FA9A" box="[728,773,1353,1375]" italics="true" pageId="3" pageNumber="295">et al</emphasis>
. 2016
</bibRefCitation>
: 78
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</paragraph>
<paragraph id="8BBAE341FF8CB03040D927DDFF5EFA02" blockId="3.[151,1436,1420,1589]" pageId="3" pageNumber="295">
<emphasis id="B9713F53FF8CB03040D927DDFEB2FA60" bold="true" box="[151,263,1420,1445]" pageId="3" pageNumber="295">Synonym</emphasis>
.
<taxonomicName id="4C0598C2FF8CB030415B27DCFD0AFA61" ID-CoL="59DFY" authority="Balthasar, 1939: 32" authorityName="Balthasar" authorityPageNumber="32" authorityYear="1939" box="[277,703,1420,1445]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="asmaraensis">
<emphasis id="B9713F53FF8CB030415B27DCFE6EFA61" box="[277,475,1421,1444]" italics="true" pageId="3" pageNumber="295">Trox asmaraensis</emphasis>
<bibRefCitation id="EF949EB0FF8CB03041AC27DDFD20FA61" author="Balthasar" box="[482,661,1420,1445]" pageId="3" pageNumber="295" refString="Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico della Universita di Firenze. Redia, 25, 1 - 36." type="journal article" year="1939">Balthasar, 1939</bibRefCitation>
: 32
</taxonomicName>
; synonymy by
<bibRefCitation id="EF949EB0FF8CB030432527DDFC50FA60" author="Haaf" box="[875,997,1420,1445]" pageId="3" pageNumber="295" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). Entomologische Arbeiten aus dem Museum Georg Frey, 4, 309 - 346." type="journal article" year="1953">Haaf 1953</bibRefCitation>
: 328;
<bibRefCitation id="EF949EB0FF8CB030446727DDFB74FA61" author="Scholtz" box="[1065,1217,1420,1445]" pageId="3" pageNumber="295" refString="Scholtz, C. H. (1980) Monograph of the genus Trox F. (Coleoptera: Trogidae) of Subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980</bibRefCitation>
: 92, 1982: 5;
<bibRefCitation id="EF949EB0FF8CB030451527DDFF66FA0D" author="Zidek" pageId="3" pageNumber="295" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek 2013</bibRefCitation>
: 7
</paragraph>
</subSubSection>
<subSubSection id="C31FB0CAFF8CB03040892785FBA1F9F0" pageId="3" pageNumber="295" type="diagnosis">
<paragraph id="8BBAE341FF8CB03040892785FBA1F9F0" blockId="3.[151,1436,1420,1589]" pageId="3" pageNumber="295">
<emphasis id="B9713F53FF8CB03040892785FEF4FA28" bold="true" box="[199,321,1492,1517]" pageId="3" pageNumber="295">Diagnosis.</emphasis>
<taxonomicName id="4C0598C2FF8CB03041062785FDFDFA28" box="[328,584,1492,1517]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="B9713F53FF8CB03041062785FDFDFA28" box="[328,584,1492,1517]" italics="true" pageId="3" pageNumber="295">Phoberus nigrociliatus</emphasis>
</taxonomicName>
can be easily separated from other members in the group on the basis of the pronotum. It has the pronotum smoothly rounded, whereas the other members have the pronotum ridged with large tubercles.
<taxonomicName id="4C0598C2FF8CB0304144244CFE06F9F0" box="[266,435,1564,1589]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="nigrociliatus">
<emphasis id="B9713F53FF8CB0304144244CFE06F9F0" box="[266,435,1564,1589]" italics="true" pageId="3" pageNumber="295">P. nigrociliatus</emphasis>
</taxonomicName>
has only been recorded from the Ethiopian highlands.
</paragraph>
</subSubSection>
</treatment>
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