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<document id="8FB019B6AA226C9DAFAFFB0D0B834185" ID-CLB-Dataset="8153" ID-DOI="10.5252/z2009n4a5" ID-GBIF-Dataset="5ead431d-4736-4458-963f-4a8b8bdd603c" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520337" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612807457570" checkinUser="felipe" docAuthor="Duy-Jacquemin, Monique Nguyen" docDate="2009" docId="03AC1E75213E9771039DB871FEEAFC3E" docLanguage="en" docName="Zoosystema.31.4.829-848.pdf" docOrigin="Zoosystema 31 (4)" docSource="http://www.bioone.org/doi/abs/10.5252/z2009n4a5" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Macroxenodes navassaensis Duy-Jacquemin 2009, n. sp." docType="treatment" docVersion="8" lastPageNumber="841" masterDocId="FF95660D2134977D0357BA22FFD6FFB2" masterDocTitle="Revision of the genera of Polyxenidae bearing pseudoarticulated gnathochilarial sensilla (Diplopoda, Polyxenida, Polyxenidae), with the description of two new species" masterLastPageNumber="848" masterPageNumber="829" pageNumber="839" updateTime="1698928237598" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="360E1B6B158FE5C44542F0A7B47E761B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D779501B64701894CEFF0FCDB51E7C33">
<mods:title id="F1CB7397542011D210B2C4AA45D3D60B">Revision of the genera of Polyxenidae bearing pseudoarticulated gnathochilarial sensilla (Diplopoda, Polyxenida, Polyxenidae), with the description of two new species</mods:title>
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<mods:name id="337E1D47B2638C3C7919A5245A1A24B7" type="personal">
<mods:role id="A4F6B719BCFA91EEE6D4BE2D3D01C68B">
<mods:roleTerm id="AE6E7DC0DA58F8E4EDDD882DC9865B99">Author</mods:roleTerm>
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<mods:namePart id="1989DF42CF6A07743FE1BCBB9A08E456">Duy-Jacquemin, Monique Nguyen</mods:namePart>
<mods:affiliation id="73CED462E1D486096760B04BF9CC82B7">Muséum national dHistoire naturelle, Département Systématique et Évolution, Zoologie - Arthropodes, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) monguyen @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="D6961FAC27321BB958C0C4AF08D6C28D" type="email">monguyen@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="603A3319FB3F3382D7B983978F84720B">text</mods:typeOfResource>
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<mods:title id="FB46EEE5FD8585390672D73089A8B0DC">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="97618E4F1E85ACA9EE528826B619D53D">
<mods:date id="95A87BCD7DDCCF302E058190D13E4D28">2009</mods:date>
<mods:detail id="8979E545532BBE94E6756035E264C743" type="pubDate">
<mods:number id="3F1FAE1A3A21751ACD34B0CC56F680B5">2009-12-31</mods:number>
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<mods:number id="30AF251BEB562F95FAAFF5D3021D91E5">31</mods:number>
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<mods:identifier id="A856D4A55596A01AC3AA29659DF3A796" type="CLB-Dataset">8153</mods:identifier>
<mods:identifier id="7F949CF7087B01CB8102B384B8515A68" type="DOI">10.5252/z2009n4a5</mods:identifier>
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<treatment id="03AC1E75213E9771039DB871FEEAFC3E" ID-DOI="http://doi.org/10.5281/zenodo.4548355" ID-GBIF-Taxon="177423088" ID-Zenodo-Dep="4548355" LSID="urn:lsid:plazi:treatment:03AC1E75213E9771039DB871FEEAFC3E" httpUri="http://treatment.plazi.org/id/03AC1E75213E9771039DB871FEEAFC3E" lastPageId="12" lastPageNumber="841" pageId="10" pageNumber="839">
<subSubSection id="C31FFCE8213E9777039DB871FDF8FDDD" box="[202,558,595,623]" pageId="10" pageNumber="839" type="nomenclature">
<paragraph id="8BBAAF63213E9777039DB871FDF8FDDD" blockId="10.[202,558,595,655]" box="[202,558,595,623]" pageId="10" pageNumber="839">
<heading id="D0F2180F213E9777039DB871FDF8FDDD" box="[202,558,595,623]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="839" reason="2">
<taxonomicName id="4C05D4E0213E9777039DB871FE3AFDDC" authority="Duy-Jacquemin, 2009" authorityName="Duy-Jacquemin" authorityYear="2009" box="[202,492,595,622]" class="Diplopoda" family="Polyxenidae" genus="Macroxenodes" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="839" phylum="Arthropoda" rank="species" species="navassaensis" status="sp. nov.">
<emphasis id="B9717371213E9777039DB871FE3AFDDC" bold="true" box="[202,492,595,622]" italics="true" pageId="10" pageNumber="839">Macroxenodes navassaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242CE0A213E977702A4B876FDF8FDDD" box="[499,558,596,623]" pageId="10" pageNumber="839" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31FFCE8213E97770203B856FE72FD3D" box="[340,420,628,655]" pageId="10" pageNumber="839" type="description">
<paragraph id="8BBAAF63213E97770203B856FE72FD3D" blockId="10.[202,558,595,655]" box="[340,420,628,655]" pageId="10" pageNumber="839">
<heading id="D0F2180F213E97770203B856FE72FD3D" box="[340,420,628,655]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="839" reason="2">
(
<figureCitation id="133EB3E6213E9777020BB856FE4DFD3D" box="[348,411,628,655]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31FFCE8213E97770326B88FFF08FCA6" pageId="10" pageNumber="839" type="materials_examined">
<paragraph id="8BBAAF63213E97770326B88FFF08FCA6" blockId="10.[113,646,684,788]" pageId="10" pageNumber="839">
TYPE MATERIAL. —
<emphasis id="B9717371213E9777026CB88EFE32FD76" bold="true" box="[315,484,684,708]" pageId="10" pageNumber="839">Greater Antilles.</emphasis>
<collectingCountry id="F312EFF3213E977702BAB88FFD54FD77" box="[493,642,685,709]" name="Navassa Island" pageId="10" pageNumber="839">Navassa Island</collectingCountry>
, Clench coll.,
<date id="FFBB89A3213E97770256B8E5FE9DFD6D" box="[257,331,711,735]" pageId="10" pageNumber="839" value="1930-01">I.1930</date>
.,
<typeStatus id="54BE11C1213E9777020BB8E5FE60FD6D" box="[348,438,711,735]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
ad.
<specimenCount id="9D0364EA213E977702B2B8EAFE21FD6D" box="[485,503,712,735]" pageId="10" pageNumber="839" type="female"></specimenCount>
and
<typeStatus id="54BE11C1213E97770178B8E5FD50FD6D" box="[559,646,711,735]" pageId="10" pageNumber="839" type="paratype">paratype</typeStatus>
ad.
<specimenCount id="9D0364EA213E977703C0B8C1FF7EFD48" box="[151,168,739,762]" pageId="10" pageNumber="839" type="female"></specimenCount>
very near to moulting, both mounted on slides (
<collectionCode id="ED1437A6213E9777032DB8DEFF04FCA6" box="[122,210,764,788]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="839">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31FFCE8213E97770326B90EFE4CFCEC" pageId="10" pageNumber="839" type="etymology">
<paragraph id="8BBAAF63213E97770326B90EFE4CFCEC" blockId="10.[112,647,812,862]" pageId="10" pageNumber="839">
ETYMOLOGY. — The name refers to the island where the
<typeStatus id="54BE11C1213E97770327B964FF4CFCEC" box="[112,154,838,862]" pageId="10" pageNumber="839">type</typeStatus>
specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="C31FFCE8213E97710326B9B3FF34FE03" lastPageId="12" lastPageNumber="841" pageId="10" pageNumber="839" type="description">
<paragraph id="8BBAAF63213E97770326B9B3FE04FC1B" blockId="10.[111,649,913,1700]" box="[113,466,913,939]" pageId="10" pageNumber="839">DESCRIPTION OF FEMALE ADULTS</paragraph>
<paragraph id="8BBAAF63213E97770326B992FE94FB99" blockId="10.[111,649,913,1700]" pageId="10" pageNumber="839">
Measurements: body length (without caudal penicillus):
<quantity id="4CFD0286213E977703F3B9F2FED0FC59" box="[164,262,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="839" unit="mm" value="3.1">3.10 mm</quantity>
(
<typeStatus id="54BE11C1213E97770245B9F2FEA5FC59" box="[274,371,976,1003]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
) and
<quantity id="4CFD0286213E977702FBB9F2FDD8FC59" box="[428,526,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="839" unit="mm" value="3.5">3.50 mm</quantity>
(
<typeStatus id="54BE11C1213E9777014DB9F2FDAAFC59" box="[538,636,976,1003]" pageId="10" pageNumber="839" type="paratype">paratype</typeStatus>
). Tarsus II length of 13th leg:
<quantity id="4CFD0286213E977702C5B9D2FE33FBB9" box="[402,485,1008,1035]" metricMagnitude="-1" metricUnit="m" metricValue="1.43" pageId="10" pageNumber="839" unit="mm" value="143.0">143 µm</quantity>
(
<typeStatus id="54BE11C1213E977702A6B9D2FD87FBB9" box="[497,593,1008,1035]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
) and
<quantity id="4CFD0286213E97770338BE32FF12FB99" box="[111,196,1040,1067]" metricMagnitude="-1" metricUnit="m" metricValue="1.46" pageId="10" pageNumber="839" unit="mm" value="146.0">146 µm</quantity>
(
<typeStatus id="54BE11C1213E97770384BE32FEE0FB99" box="[211,310,1040,1067]" pageId="10" pageNumber="839" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8BBAAF63213E977703DFBE0DFED7FA9A" blockId="10.[111,649,913,1700]" pageId="10" pageNumber="839">
Head: Eight ocelli on each side. Vertex with one pair of posterior tufts, consisting of two rows: anterior row with 13-18, posterior row with 8 or 9 trichomes, the last in front of internal trichobothrium (
<figureCitation id="133EB3E6213E97770246BE8CFEB2FB7B" box="[273,356,1198,1225]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7E</figureCitation>
); the insertions of two last trichomes of posterior row are oriented perpendicularly to others (
<figureCitation id="133EB3E6213E97770268BECFFE47FABA" box="[319,401,1261,1288]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7E</figureCitation>
). The distance between tufts is small.
</paragraph>
<paragraph id="8BBAAF63213E977703DFBF0FFCB1FEE0" blockId="10.[111,649,913,1700]" lastBlockId="10.[675,1212,216,1700]" pageId="10" pageNumber="839">
Proportions of antennal articles as shown in
<figureCitation id="133EB3E6213E97770112BF0FFF47FAD5" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Figure 7A</figureCitation>
; length of article VI about twice its diameter. Antennal article VI with two groups of dorsal and postero-ventral sensilla: dorsal group composed of three thick sensilla basiconica: anterior
<emphasis id="B9717371213E9777011DBF8EFD81FA77" box="[586,599,1452,1477]" italics="true" pageId="10" pageNumber="839">a</emphasis>
, intermediate
<emphasis id="B9717371213E977703BBBFEEFF22FA57" box="[236,244,1484,1509]" italics="true" pageId="10" pageNumber="839">i</emphasis>
and posterior
<emphasis id="B9717371213E977702C5BFEEFE49FA57" box="[402,415,1484,1509]" italics="true" pageId="10" pageNumber="839">p</emphasis>
(
<emphasis id="B9717371213E977702E7BFEEFE6EFA57" box="[432,440,1484,1509]" italics="true" pageId="10" pageNumber="839">i</emphasis>
being shorter than
<emphasis id="B9717371213E97770326BFC9FFABF9B6" box="[113,125,1515,1540]" italics="true" pageId="10" pageNumber="839">a</emphasis>
and
<emphasis id="B9717371213E977703E1BFC9FF15F9B6" box="[182,195,1515,1540]" italics="true" pageId="10" pageNumber="839">p</emphasis>
), one posterior sensillum coeloconicum
<emphasis id="B9717371213E9777012ABFC9FD50F9B6" box="[637,646,1515,1540]" italics="true" pageId="10" pageNumber="839">c</emphasis>
and one trifid setiform sensillum
<emphasis id="B9717371213E9777028CBC29FE34F996" box="[475,482,1547,1572]" italics="true" pageId="10" pageNumber="839">s</emphasis>
between
<emphasis id="B9717371213E9777011DBC29FD80F996" box="[586,598,1547,1572]" italics="true" pageId="10" pageNumber="839">a</emphasis>
and
<emphasis id="B9717371213E97770326BC09FFAFF9F6" box="[113,121,1579,1604]" italics="true" pageId="10" pageNumber="839">i</emphasis>
(
<figureCitation id="133EB3E6213E977703D0BC08FF2BF9F7" box="[135,253,1578,1605]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7C, D</figureCitation>
); postero-ventral group composed of two similar, thick sensilla basiconica, shorter than
<emphasis id="B9717371213E97770326BC48FFABF931" box="[113,125,1642,1667]" italics="true" pageId="10" pageNumber="839">a</emphasis>
and
<emphasis id="B9717371213E977703E0BC48FF12F931" box="[183,196,1642,1667]" italics="true" pageId="10" pageNumber="839">p</emphasis>
. Antennal article VII with two subequal, thick, dorsal sensilla basiconica, one posterior sensillum coeloconicum and one trifid setiform sensillum between the two sensilla basiconica (
<figureCitation id="133EB3E6213E9777070DBADAFB7BFEA1" box="[1114,1197,248,275]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7B</figureCitation>
). Three trichobothria of equal size, with cylindrical funiculi (
<figureCitation id="133EB3E6213E97770050BB15FC8DFEE0" box="[775,859,311,338]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BBAAF63213E977701ECBB75FC72FD7C" blockId="10.[675,1212,216,1700]" pageId="10" pageNumber="839">
Labrum clothed with numerous small cuspidate papillae in addition to three rows (four towards lateral side) of spherical papillae (
<figureCitation id="133EB3E6213E97770773BBB4FBADFE03" box="[1060,1147,406,433]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7F</figureCitation>
). 6+6 (
<typeStatus id="54BE11C1213E977701F9BB94FCC2FE63" box="[686,788,438,465]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
) and 8+8 (
<figureCitation id="133EB3E6213E977700C2BB94FC3FFE63" box="[917,1001,438,465]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7F</figureCitation>
) lamellar teeth on anterior margin. Clypeo-labrum with 13 and 14 setae along posterior margin. In
<typeStatus id="54BE11C1213E9777075CBBD7FBBBFDA2" box="[1035,1133,501,528]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
, lateral expansions of gnathochilarium with 14 and 15 sensilla pseudoarticulated at apex (
<figureCitation id="133EB3E6213E9777077EB816FB57FDFD" box="[1065,1153,564,591]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7G</figureCitation>
) and middle palpus with 21 sensilla, comprising 12 long, pseudoarticulated sensilla and 9 short sensilla not articulated at apex; gnathochilarial sensilla cannot be counted in
<typeStatus id="54BE11C1213E97770017B891FC76FD7C" box="[832,928,691,718]" pageId="10" pageNumber="839" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8BBAAF63213E977701ECB8F1FCA0FB1B" blockId="10.[675,1212,216,1700]" pageId="10" pageNumber="839">
Trunk: trichomes are lacking; however, their orientation can be estimed thanks to their insertion. Except on collum and tergites IX and X, trichomes arranged in two lateral tufts with 16-33 trichomes connected by three trichomic rows: anterior row with 19-40 trichomes directed towards head; middle row with 7-21 trichomes and a sinuous posterior row with 42-74 trichomes directed towards telson; anterior and posterior rows uninterrupted. On some tergites the anterior and middle rows are slightly sinuous. Total number of tergal trichomes as follows: I (collum), 103 (
<typeStatus id="54BE11C1213E9777008EBE0DFBEAFBF8" box="[985,1084,1071,1098]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
) or 123; II to VIII, 109 or 178; IX and X, 85 or 76. Lateral protuberance of tergite I with 5 (
<typeStatus id="54BE11C1213E9777074FBE4DFBADFB38" box="[1048,1147,1135,1162]" pageId="10" pageNumber="839" type="holotype">holotype</typeStatus>
) or 6 trichomes in a row.
</paragraph>
<paragraph id="8BBAAF63213E977701ECBE8CFCB3F936" blockId="10.[675,1212,216,1700]" pageId="10" pageNumber="839">
Legs: all articles of legs bearing sensory setae, first tarsus excepted. Setae on coxa, trochanter and prefemur with an oval base, provided with an apical process (
<figureCitation id="133EB3E6213E9777003EBF2FFC60FA9A" box="[873,950,1293,1320]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7J</figureCitation>
). Chaetotaxy as follows: coxae II to XII with 2 setae (1 seta in six cases, 3 setae in one case); trochanters with 1 seta; prefemora leg I to XIII with 1 seta and 1 or 2 small, trifid setae (
<figureCitation id="133EB3E6213E97770026BFAEFC35FA15" box="[881,995,1420,1447]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7H, I</figureCitation>
); almost all femora and tibiae with same small seta as prefemur. Second tarsus with a small seta as shown in
<figureCitation id="133EB3E6213E97770723BFE9FD1CF9B4" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Figure 7K</figureCitation>
. Telotarsus bearing an anterior process with a spinous projection longer than claw, two subequal latero-anterior and posterior spiniform processes; posterior lamellar process thickened and basally pleated (
<figureCitation id="133EB3E6213E97770053BC4BFC8FF936" box="[772,857,1641,1668]" captionStart="FIG" captionStartId="11.[113,124,1544,1561]" captionTargetBox="[115,1201,216,1427]" captionTargetId="figure-13@11.[106,1225,199,1450]" captionTargetPageId="11" captionText="FIG. 7. — Macroxenodes navassaensis n. sp.: A, right antenna of paratype; B, C, sensilla on right antennal articles VII and VI of holotype; D, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation; E, left part of head of holotype with insertions of dorsal trichomes; F, labrum of paratype, only part of cuticular setae shown; G, right palpus of gnathochilarium of holotype; H-L, holotype; H, I, small setae of prefemora of right legs III and VII; J, coxal seta of right leg I; K, seta of tarsus II of right leg VI; L, telotarsus of right leg III. Abbreviations: a, anterior sensillum basiconicum; c, sensillum coeloconicum; i, intermediate sensillum basiconicum; p, posterior sensillum basiconicum; s, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm." figureDoi="http://doi.org/10.5281/zenodo.4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="10" pageNumber="839">Fig. 7L</figureCitation>
).
</paragraph>
<paragraph id="8BBAAF63213E977701ECBCABFB58F916" blockId="10.[675,1212,216,1700]" box="[699,1166,1673,1700]" pageId="10" pageNumber="839">
Four setae above the anal valve in
<typeStatus id="54BE11C1213E9777077DBCABFB5CF916" box="[1066,1162,1673,1700]" pageId="10" pageNumber="839" type="paratype">paratype</typeStatus>
.
</paragraph>
<caption id="DF7AFFEB213F97760326BC2AFEBDF913" ID-DOI="http://doi.org/10.5281/zenodo.4520363" ID-Zenodo-Dep="4520363" httpUri="https://zenodo.org/record/4520363/files/figure.png" pageId="11" pageNumber="840" startId="11.[113,124,1544,1561]" targetBox="[115,1201,216,1427]" targetPageId="11">
<paragraph id="8BBAAF63213F97760326BC2AFEBDF913" blockId="11.[113,1211,1544,1697]" pageId="11" pageNumber="840">
FIG. 7. —
<taxonomicName id="4C05D4E0213F97760390BC2AFE6FF9AB" authority="Duy-Jacquemin, 2009" authorityName="Duy-Jacquemin" authorityYear="2009" box="[199,441,1544,1561]" class="Diplopoda" family="Polyxenidae" genus="Macroxenodes" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="840" phylum="Arthropoda" rank="species" species="navassaensis" status="sp. nov.">
<emphasis id="B9717371213F97760390BC2AFE6FF9AB" box="[199,441,1544,1561]" italics="true" pageId="11" pageNumber="840">Macroxenodes navassaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242CE0A213F97760297BC2AFE22F9AB" box="[448,500,1544,1561]" pageId="11" pageNumber="840" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B9717371213F977602ABBC2AFDDFF9AB" bold="true" box="[508,521,1544,1561]" pageId="11" pageNumber="840">A</emphasis>
, right antenna of paratype;
<emphasis id="B9717371213F977601ACBC2AFCDEF9AB" bold="true" box="[763,776,1544,1561]" pageId="11" pageNumber="840">B</emphasis>
,
<emphasis id="B9717371213F97760043BC2AFCF4F9AB" bold="true" box="[788,802,1544,1561]" pageId="11" pageNumber="840">C</emphasis>
, sensilla on right antennal articles VII and VI of holotype;
<emphasis id="B9717371213F97760391BC3DFF02F982" bold="true" box="[198,212,1567,1584]" pageId="11" pageNumber="840">D</emphasis>
, sensilla on left antennal article VI of paratype, the new sensilla shown with dotted lines will appear after exuviation;
<emphasis id="B9717371213F97760324BC17FFA8F9F4" bold="true" box="[115,126,1589,1606]" pageId="11" pageNumber="840">E</emphasis>
, left part of head of holotype with insertions of dorsal trichomes;
<emphasis id="B9717371213F97760193BC17FD19F9F4" bold="true" box="[708,719,1589,1606]" pageId="11" pageNumber="840">F</emphasis>
, labrum of paratype, only part of cuticular setae shown;
<emphasis id="B9717371213F97760324BC6EFF56F9EF" bold="true" box="[115,128,1612,1629]" pageId="11" pageNumber="840">G</emphasis>
, right palpus of gnathochilarium of holotype;
<emphasis id="B9717371213F977602AABC6EFDCBF9EF" bold="true" box="[509,541,1612,1629]" pageId="11" pageNumber="840">H -L</emphasis>
, holotype;
<emphasis id="B9717371213F9776012CBC6EFD5FF9EF" bold="true" box="[635,649,1612,1629]" pageId="11" pageNumber="840">H</emphasis>
,
<emphasis id="B9717371213F977601C3BC6EFD4CF9EF" bold="true" box="[660,666,1612,1629]" pageId="11" pageNumber="840">I</emphasis>
, small setae of prefemora of right legs III and VII;
<emphasis id="B9717371213F9776076EBC6EFB95F9EF" bold="true" box="[1081,1091,1612,1629]" pageId="11" pageNumber="840">J</emphasis>
, coxal seta of right leg I;
<emphasis id="B9717371213F9776039CBC41FF0EF9C6" bold="true" box="[203,216,1635,1652]" pageId="11" pageNumber="840">K</emphasis>
, seta of tarsus II of right leg VI;
<emphasis id="B9717371213F977602B3BC41FE39F9C6" bold="true" box="[484,495,1635,1652]" pageId="11" pageNumber="840">L</emphasis>
, telotarsus of right leg III. Abbreviations:
<emphasis id="B9717371213F9776001EBC41FC82F9C6" bold="true" box="[841,852,1635,1652]" italics="true" pageId="11" pageNumber="840">a</emphasis>
, anterior sensillum basiconicum;
<emphasis id="B9717371213F9776073ABC41FBA1F9C6" bold="true" box="[1133,1143,1635,1652]" italics="true" pageId="11" pageNumber="840">c</emphasis>
, sensillum coeloconicum;
<emphasis id="B9717371213F9776024EBC5BFEC8F938" bold="true" box="[281,286,1657,1674]" italics="true" pageId="11" pageNumber="840">i</emphasis>
, intermediate sensillum basiconicum;
<emphasis id="B9717371213F97760130BC5BFDA4F938" bold="true" box="[615,626,1657,1674]" italics="true" pageId="11" pageNumber="840">p</emphasis>
, posterior sensillum basiconicum;
<emphasis id="B9717371213F977600C9BC5BFC7EF938" bold="true" box="[926,936,1657,1674]" italics="true" pageId="11" pageNumber="840">s</emphasis>
, setiform sensillum. Scale bars: A, E, 50 μm; B-D, F-L, 25 μm.
</paragraph>
</caption>
<paragraph id="8BBAAF632138977103DFBAFAFF34FE03" blockId="12.[112,647,216,433]" pageId="12" pageNumber="841">
Telson: dorsal face of telson with 13-16 barbate trichomes
<emphasis id="B97173712138977103B4BADAFF39FEA3" box="[227,239,248,273]" italics="true" pageId="12" pageNumber="841">a</emphasis>
; one trichome
<emphasis id="B97173712138977102C2BAD5FE74FEA3" box="[405,418,247,273]" italics="true" pageId="12" pageNumber="841">b</emphasis>
inner to group of 19 trichomes
<emphasis id="B97173712138977103B7BB3AFF3FFE83" box="[224,233,280,305]" italics="true" pageId="12" pageNumber="841">c</emphasis>
observed in
<typeStatus id="54BE11C121389771023ABB35FE1BFE80" box="[365,461,279,306]" pageId="12" pageNumber="841" type="paratype">paratype</typeStatus>
. Ventral face with a subrectilinear row of 14+14 barbate trichomes
<emphasis id="B971737121389771012EBB1AFD53FEE3" box="[633,645,312,337]" italics="true" pageId="12" pageNumber="841">v</emphasis>
in
<typeStatus id="54BE11C12138977103C7BB75FF39FEC0" box="[144,239,343,370]" pageId="12" pageNumber="841" type="holotype">holotype</typeStatus>
and 13+
<quantity id="4CFD0286213897710205BB75FE5DFEC0" box="[338,395,343,370]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="12" pageNumber="841" unit="in" value="15.0">15 in</quantity>
<typeStatus id="54BE11C12138977102C3BB75FE26FEC0" box="[404,496,343,370]" pageId="12" pageNumber="841" type="paratype">paratype</typeStatus>
(
<figureCitation id="133EB3E6213897710157BB75FD85FEC0" box="[512,595,343,370]" captionStart="FIG" captionStartId="9.[113,124,1590,1607]" captionTargetBox="[126,1173,217,1574]" captionTargetId="figure-13@9.[122,1198,186,1587]" captionTargetPageId="9" captionText="FIG. 6. — A-D, Macroxenodes meinerti (Silvestri, 1898); A, B, distal parts of barbate and hooked trichomes of caudal penicillus of male; C, ventral face of telson of male, dorsal trichomes shown with dotted lines; D, dorsal face of telson of male; E, M. navassaensis n. sp., caudal view of female paratype. All the biggest insertions belonging to barbate trichomes are shown; only part of insertions of hooked trichomes shown; dorsal trichomes a with dotted lines. Abbreviations: a1-a10, insertions of barbate trichomes a; b, insertion of trichome b; c, group of trichomes c; v1-vX, insertions of ventral barbate trichomes on edge of penicillus. Scale bars: A, B, D, 25μm; C, E, 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4520361" httpUri="https://zenodo.org/record/4520361/files/figure.png" pageId="12" pageNumber="841">Fig. 6E</figureCitation>
), located on the edge of the insertions of fallen hooked trichomes.
</paragraph>
</subSubSection>
<subSubSection id="C31FFCE8213897710326BBF5FEEAFC3E" pageId="12" pageNumber="841" type="discussion">
<paragraph id="8BBAAF63213897710326BBF5FF32FE5C" blockId="12.[111,648,471,908]" box="[113,228,471,497]" pageId="12" pageNumber="841">AFFINITIES</paragraph>
<paragraph id="8BBAAF63213897710326BBD7FEEAFC3E" blockId="12.[111,648,471,908]" pageId="12" pageNumber="841">
This species is close to
<taxonomicName id="4C05D4E0213897710225BBD7FD54FD82" authority="Ishii, Nguyen Duy-Jacquemin &amp; Conde, 1999" authorityName="Ishii, Nguyen Duy-Jacquemin &amp; Conde" authorityYear="1999" class="Diplopoda" family="Polyxenidae" genus="Macroxenodes" kingdom="Animalia" order="Polyxenida" pageId="12" pageNumber="841" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B9717371213897710225BBD7FD50FDBD" box="[370,646,501,527]" italics="true" pageId="12" pageNumber="841">Macroxenodes amazonicus</emphasis>
Ishii, Nguyen Duy-Jacquemin &amp; Condé, 1999
</taxonomicName>
. However, there are significant differences between them. The postero-ventral group of antennal sensilla of article VI has only two sensilla, as opposed to
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<taxonomicName id="4C05D4E0213897710394B8B6FEBCFD1F" authorityName="Ishii, Nguyen Duy-Jacquemin &amp; Conde" authorityYear="1999" box="[195,362,660,685]" class="Diplopoda" family="Polyxenidae" genus="Macroxenodes" kingdom="Animalia" order="Polyxenida" pageId="12" pageNumber="841" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B9717371213897710394B8B6FEBCFD1F" box="[195,362,660,685]" italics="true" pageId="12" pageNumber="841">M. amazonicus</emphasis>
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. Th ere are three anterior rows of spherical papillae on the labrum, whereas in
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<emphasis id="B97173712138977103C5B8F1FEECFD5E" box="[146,314,723,748]" italics="true" pageId="12" pageNumber="841">M. amazonicus</emphasis>
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there are only two. The new species has 15 sensilla at most on the lateral palp expansion, compared to
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<taxonomicName id="4C05D4E02138977102B7B931FDA9FC9E" authorityName="Ishii, Nguyen Duy-Jacquemin &amp; Conde" authorityYear="1999" box="[480,639,787,812]" class="Diplopoda" family="Polyxenidae" genus="Macroxenodes" kingdom="Animalia" order="Polyxenida" pageId="12" pageNumber="841" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B97173712138977102B7B931FDA9FC9E" box="[480,639,787,812]" italics="true" pageId="12" pageNumber="841">M. amazonicus</emphasis>
</taxonomicName>
. The median interruption of the anterior trichomic row of the tergites of
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<emphasis id="B9717371213897710204B970FE27FCD9" box="[339,497,850,875]" italics="true" pageId="12" pageNumber="841">M. amazonicus</emphasis>
</taxonomicName>
is not present in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>