113 lines
18 KiB
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113 lines
18 KiB
XML
<document id="043C6DF8EC4DF88CA65B237F131EDF91" ID-DOI="10.5281/zenodo.5312012" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312012" ID-ZooBank="D72D1DC9-87D1-4209-A01F-8185A4D21F88" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118846529" checkinUser="marcus" docAuthor="Sehnal, Richard, Král, David & Bezděk, Aleš" docDate="2014" docId="03A4923D7D79AD271557FD50F68A4F5C" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.139-171.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Canuschiza insularis Lacroix 1999" docType="treatment" docVersion="4" lastPageNumber="142" masterDocId="FF9DEA457D7AAD241413FFF8F470496C" masterDocTitle="Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group" masterLastPageNumber="171" masterPageNumber="139" pageNumber="142" updateTime="1699158890131" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4DE0746F0245972BB200232F2FF29C8C">Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group</mods:title>
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<mods:namePart id="03AD5C1407EDD8A32EC478915E6EA76C">Sehnal, Richard</mods:namePart>
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<mods:affiliation id="3080D4BBE7F0D4EEFDFA98D54794A5CA">Czech University of Life Sciences Prague, Faculty of Agrobiology, Food and Natural Resources, Department of Zoology and Fisheries, Kamýcká 129, CZ- 165 21 Praha 6, Czech Republic; e-mail: richard. sehnal @ seznam. cz</mods:affiliation>
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<mods:namePart id="8ED773ACE8D39B4F4EECF630BD3EFD6F">Král, David</mods:namePart>
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<mods:affiliation id="56348CE3779FE34FAF6BB93D17D0A2A7">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Praha 2, Czech Republic; e-mail: kral @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="89D782504F0B3575EEB6D36FF8007D93">Bezděk, Aleš</mods:namePart>
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<mods:affiliation id="57AF49898D65CF298A6920961DD7CAA0">Biology Centre ASCR, Institute of Entomology, Branišovská 31, CZ- 370 05 České Budějovice, Czech Republic; e-mail: bezdek @ entu. cas. cz</mods:affiliation>
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<mods:title id="5069782E7758504559A89E09F4B6463C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="2793303FFE5495C283DEDB85901D417F">2014</mods:date>
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<mods:number id="6270AD9A9788631693EA433087EBEDB9">2014-12-30</mods:number>
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<treatment id="03A4923D7D79AD271557FD50F68A4F5C" ID-DOI="http://doi.org/10.5281/zenodo.5449175" ID-Zenodo-Dep="5449175" LSID="urn:lsid:plazi:treatment:03A4923D7D79AD271557FD50F68A4F5C" httpUri="http://treatment.plazi.org/id/03A4923D7D79AD271557FD50F68A4F5C" lastPageNumber="142" pageId="3" pageNumber="142">
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<subSubSection id="C31770A07D79AD271557FD50F71E4BAC" box="[324,878,680,704]" pageId="3" pageNumber="142" type="nomenclature">
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<paragraph id="8BB2232B7D79AD271557FD50F71E4BAC" blockId="3.[324,878,680,704]" box="[324,878,680,704]" pageId="3" pageNumber="142">
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<heading id="D0FA94477D79AD271557FD50F71E4BAC" bold="true" box="[324,878,680,704]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="142" reason="3">
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<emphasis id="B979FF397D79AD271557FD50F71E4BAC" bold="true" box="[324,878,680,704]" pageId="3" pageNumber="142">
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<taxonomicName id="4C0D58A87D79AD271557FD50F6BE4BAC" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" box="[324,718,680,704]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B979FF397D79AD271557FD50F65C4BAC" bold="true" box="[324,556,680,704]" italics="true" pageId="3" pageNumber="142">Canuschiza insularis</emphasis>
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<bibRefCitation id="EF9C5EDA7D79AD271620FD51F6BE4BAC" author="LACROIX M." box="[563,718,681,704]" pageId="3" pageNumber="142" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
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species group
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</emphasis>
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</paragraph>
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<subSubSection id="C31770A07D79AD271462FD21F68A4F5C" pageId="3" pageNumber="142" type="diagnosis">
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<paragraph id="8BB2232B7D79AD271462FD21F6BF4C1C" blockId="3.[113,1090,729,1584]" pageId="3" pageNumber="142">
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<emphasis id="B979FF397D79AD271462FD21F4964B9C" bold="true" box="[113,230,729,752]" pageId="3" pageNumber="142">Diagnosis.</emphasis>
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Body elongate, almost parallel, weakly convex; colour chestnut brown to black, elytra often somewhat alutaceous, macrosetation pale. Dorsal surface of head, pronotum, scutellar shield and elytra covered with short recumbent scale-like macrosetae, ventral surface of thorax and abdomen with recumbent scale-like macrosetae. Labrum transverse, deeply bilobed, lobes rounded. Clypeus transverse, anterior angles rounded. Head with simple darkened fronto-clypeal suture (translucent endocarina) (
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<figureCitation id="13363FAE7D79AD2716E8FC81F7214AFC" box="[763,849,889,912]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="3" pageNumber="142">Fig. 1A</figureCitation>
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). Eye-canthus narrow, short, bare or with few macrosetae. Antenna with eight or ten antennomeres; club with three antennomeres, almost straight to regularly arcuate, sparsely and shortly macrosetaceous. Terminal maxillary palpomere elongate, truncate apically. Pronotum transverse, weakly convex, widest approximately at middle. Posterior angles approximately rectangular or even distinctly produced posteriad (e.g.
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<figureCitation id="13363FAE7D79AD2715F5FBE1F64C4D5C" box="[486,572,1049,1072]" captionStart="Figs 6" captionStartId="15.[113,150,1533,1552]" captionTargetBox="[121,1079,224,1420]" captionTargetId="figure-14@15.[113,1088,219,1440]" captionTargetPageId="15" captionText="Figs 6A–D. Canuschiza hagher sp. nov., male, holotype, body length 17.0 mm, dorsal view.A – habitus; B – protarsal claw; C – detail of laterobasal area of pronotum and elytron; D – head. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312030" httpUri="https://zenodo.org/record/5312030/files/figure.png" pageId="3" pageNumber="142">Figs 6C</figureCitation>
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,
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<figureCitation id="13363FAE7D79AD27165BFBE1F61A4D5C" box="[584,618,1049,1072]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="3" pageNumber="142">9C</figureCitation>
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). Scutellar shield large, almost equilaterally triangular. Elytra weakly convex, parallel-sided, rounded apically. Striae missing, except of feebly visible sutural stria. Surface finely microsculptured, alutaceous. Punctation coarse, each puncture bearing short, pale scale-like, almost recumbent macroseta. Macropterous or micropterous. Protibia narrow, tridentate, terminal calcar inserted against medial dent. Meso- and metatibia slightly expanded apicad, with two macrosetiferous longitudinal carinae. Upper terminal calcar of metatibia flattened. Claws bifid, with ventrobasal teeth (e.g.
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<figureCitation id="13363FAE7D79AD271005FB21F4E34C7C" captionStart="Figs 2" captionStartId="5.[113,150,1529,1548]" captionTargetBox="[130,1078,233,1493]" captionTargetId="figure-14@5.[121,1082,219,1495]" captionTargetPageId="5" captionText="Figs 2A–H. Canuschiza adah sp. nov., dorsal view: A–D – male, holotype, body length 13.7 mm; E–H – female, paratype No. 7, body length 13.0 mm. A, E – habitus; B, F – protarsal claw; C, G – detail of laterobasal area of pronotum and elytron; D, H – head. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312018" httpUri="https://zenodo.org/record/5312018/files/figure.png" pageId="3" pageNumber="142">Fig. 2B</figureCitation>
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). Pygidium slightly transverse, convex, all around bordered, broadly rounded apically, irregularly covered with coarse macrosetiferous punctures. Parameres symmetrical, shorter than phallobasis, regularly curved in lateral aspect, rounded apically in dorsal aspect; apex with tuft of long pale macrosetae (
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<figureCitation id="13363FAE7D79AD2715CCFAA1F6164C1C" box="[479,614,1369,1392]" captionStart="Figs 10" captionStartId="27.[113,150,1513,1532]" captionTargetBox="[121,1075,225,1464]" captionTargetId="figure-14@27.[113,1088,220,1476]" captionTargetPageId="27" captionText="Figs 10A–D.Aedeagus in dorsal (left) and lateral (right) view:A – Canuschiza adah sp. nov., holotype; B – C. croton sp. nov., holotype; C – C. dracaena sp. nov., holotype; D – C. firmihin sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312042" httpUri="https://zenodo.org/record/5312042/files/figure.png" pageId="3" pageNumber="142">Figs 10A–D</figureCitation>
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,
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<figureCitation id="13363FAE7D79AD271662FAA1F6B34C1C" box="[625,707,1369,1392]" captionStart="Figs 11" captionStartId="28.[150,187,1487,1506]" captionTargetBox="[159,1117,227,1393]" captionTargetId="figure-11@28.[151,1126,218,1396]" captionTargetPageId="28" captionText="Figs 11A–D. Aedeagus in dorsal (left) and lateral (right) view: A – Canuschiza hagher sp. nov., holotype; B – C. insularis Lacroix, 1999, holotype; C – C. jatropha sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312044" httpUri="https://zenodo.org/record/5312044/files/figure.png" pageId="3" pageNumber="142">11A–D</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB2232B7D79AD271462FA81F68A4F5C" blockId="3.[113,1090,729,1584]" pageId="3" pageNumber="142">
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<emphasis id="B979FF397D79AD271462FA81F5754CFC" bold="true" box="[113,261,1401,1424]" pageId="3" pageNumber="142">Comparison.</emphasis>
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Members of
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<taxonomicName id="4C0D58A87D79AD271580FA80F67E4CFC" authorityName="Lacroix" authorityYear="1999" box="[403,526,1400,1424]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B979FF397D79AD271580FA80F67E4CFC" box="[403,526,1400,1424]" italics="true" pageId="3" pageNumber="142">C. insularis</emphasis>
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</taxonomicName>
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species group differ from
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<taxonomicName id="4C0D58A87D79AD27173BFA80F7E14CFC" authorityName="Lacroix" authorityYear="1999" box="[808,913,1400,1424]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="minuta">
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<emphasis id="B979FF397D79AD27173BFA80F7E14CFC" box="[808,913,1400,1424]" italics="true" pageId="3" pageNumber="142">C. minuta</emphasis>
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</taxonomicName>
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mainly in larger size (9.0–
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<quantity id="4CF58ECE7D79AD2714CBFA61F5484CDC" box="[216,312,1433,1456]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="142" unit="mm" value="17.2">17.2 mm</quantity>
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), weakly convex body, simple darkened fronto-clypeal suture (translucent endocarina) (
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<figureCitation id="13363FAE7D79AD2714ECFA41F5244CBC" box="[255,340,1465,1488]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="3" pageNumber="142">Fig. 1A</figureCitation>
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), and the rectangular shape of posterior angles of protonum, in some species even distinctly produced posteriad (e.g.
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<figureCitation id="13363FAE7D79AD271679FA21F6CE4C9C" box="[618,702,1497,1520]" captionStart="Figs 6" captionStartId="15.[113,150,1533,1552]" captionTargetBox="[121,1079,224,1420]" captionTargetId="figure-14@15.[113,1088,219,1440]" captionTargetPageId="15" captionText="Figs 6A–D. Canuschiza hagher sp. nov., male, holotype, body length 17.0 mm, dorsal view.A – habitus; B – protarsal claw; C – detail of laterobasal area of pronotum and elytron; D – head. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312030" httpUri="https://zenodo.org/record/5312030/files/figure.png" pageId="3" pageNumber="142">Figs 6C</figureCitation>
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,
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<figureCitation id="13363FAE7D79AD2716DBFA21F69A4C9C" box="[712,746,1497,1520]" captionStart="Figs 9" captionStartId="26.[151,188,1513,1532]" captionTargetBox="[164,1117,226,1448]" captionTargetId="figure-11@26.[156,1121,219,1457]" captionTargetPageId="26" captionText="Figs 9A–I. Canuschiza skand sp. nov., dorsal view: A–E – male, holotype, body length 12.5 mm; F–I – female, paratype No. 33, body length 14.2 mm. A, F – habitus; B, G – protarsal claw; C, H – detail of laterobasal area of pronotum and elytron; D, I – head; E – detail of lateroapical area of pronotum. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312038" httpUri="https://zenodo.org/record/5312038/files/figure.png" pageId="3" pageNumber="142">9C</figureCitation>
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), while
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<taxonomicName id="4C0D58A87D79AD27172EFA20F7D54C9C" authorityName="Lacroix" authorityYear="1999" box="[829,933,1496,1520]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="minuta">
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<emphasis id="B979FF397D79AD27172EFA20F7D54C9C" box="[829,933,1496,1520]" italics="true" pageId="3" pageNumber="142">C. minuta</emphasis>
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</taxonomicName>
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is smaller (
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<quantity id="4CF58ECE7D79AD27100DFA21F4EE4F7C" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="142" unit="mm" value="7.5">7.5 mm</quantity>
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), with distinctly convex body, head with well visible elevated double fronto-clypeal carina (
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<figureCitation id="13363FAE7D79AD27146AF9E1F4BA4F5C" box="[121,202,1561,1584]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="3" pageNumber="142">Fig. 1B</figureCitation>
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) and broadly rounded posterior angles of protonum.
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</paragraph>
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</treatment>
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