treatments-xml/data/03/A4/87/03A487E6D645D332FF3A7DF6FA75FBFC.xml
2024-06-21 12:22:17 +02:00

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<document id="27D282FDF265250702F9D44E6BA5FE02" ID-DOI="10.11646/palaeoentomology.6.2.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="7928885" ID-ZooBank="1E80FE4B-F846-4243-ABB9-15CF7F8341F4" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1683887594507" checkinUser="plazi" docAuthor="GOROCHOV, ANDREI V. &amp; CORAM, ROBERT A." docDate="2023" docId="03A487E6D645D332FF3A7DF6FA75FBFC" docLanguage="en" docName="palaeoentomology.6.2.11.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Protogryllinae Zeuner 1937" docType="treatment" docVersion="1" lastPageNumber="199" masterDocId="FF9DFF9ED644D333FFB27E7DFFD7FF86" masterDocTitle="New and little known taxa of the order Orthoptera (Insecta) from the Upper Triassic and Lower Jurassic of England" masterLastPageNumber="204" masterPageNumber="198" pageNumber="199" updateTime="1685380773502" updateUser="valdenar">
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<mods:title id="19897D140830545B61F07E5B94B43CBC">New and little known taxa of the order Orthoptera (Insecta) from the Upper Triassic and Lower Jurassic of England</mods:title>
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<mods:namePart id="0CFF063200657E03755F8E1C6FE7F2D7">CORAM, ROBERT A.</mods:namePart>
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<mods:date id="BE840DF633CF7F4424263A68E356BD79">2023</mods:date>
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<subSubSection id="C317657BD645D332FF3A7DF6FD96FC25" box="[136,577,907,931]" pageId="1" pageNumber="199" type="nomenclature">
<paragraph id="8BB236F0D645D332FF3A7DF6FD96FC25" blockId="1.[136,613,798,931]" box="[136,577,907,931]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FF3A7DF6FD96FC25" bold="true" box="[136,577,907,931]" pageId="1" pageNumber="199">
Subfamily
<taxonomicName id="4C0D4D73D645D332FEB67DF6FD96FC25" ID-CoL="8PRZY" authority="Zeuner, 1937" authorityName="Zeuner" authorityYear="1937" box="[260,577,907,931]" class="Insecta" family="Protogryllidae" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="subFamily" subFamily="Protogryllinae">Protogryllinae Zeuner, 1937</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C317657BD645D332FF3A7DAFFA75FBFC" pageId="1" pageNumber="199" type="discussion">
<paragraph id="8BB236F0D645D332FF3A7DAFFD5DF8D5" blockId="1.[136,778,978,2092]" pageId="1" pageNumber="199">
<emphasis id="B979EAE2D645D332FF3A7DAFFF2BFC6A" bold="true" box="[136,252,978,1004]" pageId="1" pageNumber="199">Remarks.</emphasis>
The generic composition of this subfamily is unclear, because its type genus,
<taxonomicName id="4C0D4D73D645D332FE417D8BFF14FBB3" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FE417D8BFD57FB96" box="[499,640,1014,1040]" italics="true" pageId="1" pageNumber="199">Protogryllus</emphasis>
Handlirsch, 1906
</taxonomicName>
, is poorly understood. Designation of the type species of this genus (
<taxonomicName id="4C0D4D73D645D332FE297A42FF17FBFB" authority="Geinitz, 1880" authorityName="Geinitz" authorityYear="1880" class="Insecta" family="Gryllidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="dobbertinensis">
<emphasis id="B979EAE2D645D332FE297A42FD75FBDF" box="[411,674,1087,1113]" italics="true" pageId="1" pageNumber="199">Gryllus dobbertinensis</emphasis>
<bibRefCitation id="EF9C4B01D645D332FD027A42FF17FBFB" author="Geinitz, F. E." pageId="1" pageNumber="199" pagination="510 - 535" refId="ref4869" refString="Geinitz, F. E. (1880) Der Jura von Dobbertin in Mecklenburg und seine Versteinerungen. Zeitschrift der Deutschen geologischen Gesellschaft, 32, 510 - 535." type="journal article" year="1880">Geinitz, 1880</bibRefCitation>
</taxonomicName>
from the Lower Jurassic of
<collectingCountry id="F31A7660D645D332FE4C7A1EFDBFFBFB" box="[510,616,1123,1149]" name="Germany" pageId="1" pageNumber="199">Germany</collectingCountry>
) was made by
<bibRefCitation id="EF9C4B01D645D332FF3A7AF5FEFBFB24" author="Zeuner, F. E." box="[136,300,1160,1186]" pageId="1" pageNumber="199" refId="ref5166" refString="Zeuner, F. E. (1939) Fossil Orthoptera Ensifera. British Museum (Natural History), London, 321 pp." type="book" year="1939">Zeuner (1939)</bibRefCitation>
based only on an inadequate description and picture by
<bibRefCitation id="EF9C4B01D645D332FE867AD1FE02FB40" author="Geinitz, F. E." box="[308,469,1196,1222]" pageId="1" pageNumber="199" pagination="510 - 535" refId="ref4869" refString="Geinitz, F. E. (1880) Der Jura von Dobbertin in Mecklenburg und seine Versteinerungen. Zeitschrift der Deutschen geologischen Gesellschaft, 32, 510 - 535." type="journal article" year="1880">Geinitz (1880</bibRefCitation>
: fig. XXII, 6),
<emphasis id="B979EAE2D645D332FD307AD0FD5DFB40" box="[642,650,1197,1222]" italics="true" pageId="1" pageNumber="199">i</emphasis>
.
<emphasis id="B979EAE2D645D332FD237AD0FD4AFB40" box="[657,669,1197,1222]" italics="true" pageId="1" pageNumber="199">e</emphasis>
., without restudy of its male
<typeStatus id="54B68852D645D332FED77AADFE1CFB6C" box="[357,459,1232,1258]" pageId="1" pageNumber="199">holotype</typeStatus>
. At the same time and later, several other specimens, included in this species, were described and illustrated; however, they probably belong to several different species. The
<specimenCount id="9D0BFD79D645D332FE457B40FDBEFAD1" box="[503,617,1341,1367]" pageId="1" pageNumber="199" type="male">two males</specimenCount>
from the type locality have a rather long harp area and weakly arcuate chords in the tegmina (
<bibRefCitation id="EF9C4B01D645D332FE397BFBFDF1FA26" author="Zeuner, F. E." box="[395,550,1414,1440]" pageId="1" pageNumber="199" refId="ref5166" refString="Zeuner, F. E. (1939) Fossil Orthoptera Ensifera. British Museum (Natural History), London, 321 pp." type="book" year="1939">Zeuner, 1939</bibRefCitation>
: figs XXXIX, 1 and LVIII, 2) whereas the other male from the same locality (
<bibRefCitation id="EF9C4B01D645D332FF227BB3FEFCFA6E" author="Zeuner, F. E." box="[144,299,1486,1512]" pageId="1" pageNumber="199" refId="ref5166" refString="Zeuner, F. E. (1939) Fossil Orthoptera Ensifera. British Museum (Natural History), London, 321 pp." type="book" year="1939">Zeuner, 1939</bibRefCitation>
: p. 193 and fig. LXXII, 3) and
<specimenCount id="9D0BFD79D645D332FD247BB3FCDEFA6E" box="[662,777,1486,1512]" pageId="1" pageNumber="199" type="male">two males</specimenCount>
from another German locality of the same age (
<bibRefCitation id="EF9C4B01D645D332FD117B8FFF13F9B6" author="Ansorge, J." pageId="1" pageNumber="199" pagination="1 - 132" refId="ref4692" refString="Ansorge, J. (1996) Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland). Neue Palaontologische Abhandlungen, 2, 1 - 132." type="journal article" year="1996">Ansorge, 1996</bibRefCitation>
: fig. 14A, B) have a distinctly shorter harp area and more strongly curved chords in the tegmina. Two further males from the latter locality also have a short harp area, but have less strongly curved chords with a clearly wider area between them and the anterior part of the diagonal vein (moreover, the areas between the chords as well as between the chords and CuA2 differ in shape in these additional males, which may therefore also belong to separate species;
<bibRefCitation id="EF9C4B01D645D332FEFB7944FE26F8D5" author="Ansorge, J." box="[329,497,1849,1875]" pageId="1" pageNumber="199" pagination="1 - 132" refId="ref4692" refString="Ansorge, J. (1996) Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland). Neue Palaontologische Abhandlungen, 2, 1 - 132." type="journal article" year="1996">Ansorge, 1996</bibRefCitation>
: fig. 14C, D).
</paragraph>
<paragraph id="8BB236F0D645D332FF0F7920FA73FB8B" blockId="1.[136,778,978,2092]" lastBlockId="1.[810,1452,173,1146]" pageId="1" pageNumber="199">
Synonymy of
<taxonomicName id="4C0D4D73D645D332FEEE7920FCDEF8F1" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[348,777,1885,1911]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="cantans">
<emphasis id="B979EAE2D645D332FEEE7920FD95F8F1" box="[348,578,1885,1911]" italics="true" pageId="1" pageNumber="199">Protogryllus cantans</emphasis>
Handlirsch, 1939
</taxonomicName>
and
<taxonomicName id="4C0D4D73D645D332FF0B79FFFDD7F81D" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[185,512,1921,1948]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="muzicus">
<emphasis id="B979EAE2D645D332FF0B79FFFF1DF81D" box="[185,202,1922,1947]" italics="true" pageId="1" pageNumber="199">P</emphasis>
.
<emphasis id="B979EAE2D645D332FF6879FFFEE3F81D" box="[218,308,1922,1947]" italics="true" pageId="1" pageNumber="199">muzicus</emphasis>
Handlirsch, 1939
</taxonomicName>
with
<taxonomicName id="4C0D4D73D645D332FDF679FFFCDDF81D" authorityName="Geinitz" authorityYear="1880" box="[580,778,1921,1948]" class="Insecta" family="Gryllidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="dobbertinensis">
<emphasis id="B979EAE2D645D332FDF679FFFD82F81D" box="[580,597,1922,1947]" italics="true" pageId="1" pageNumber="199">P</emphasis>
.
<emphasis id="B979EAE2D645D332FDD779FCFCDDF81D" box="[613,778,1921,1947]" italics="true" pageId="1" pageNumber="199">dobbertinensis</emphasis>
</taxonomicName>
is not evident, because the first species has male tegmina more or less similar to those of the males with strongly curved tegminal chords mentioned above, but the male tegmina of the second species have less strongly curved chords (very similar to some other males attributed to
<taxonomicName id="4C0D4D73D645D332FC987EAEFC3BFF6A" authorityName="Geinitz" authorityYear="1880" box="[810,1004,210,236]" class="Insecta" family="Gryllidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="dobbertinensis">
<emphasis id="B979EAE2D645D332FC987EAEFCECFF6A" box="[810,827,211,236]" italics="true" pageId="1" pageNumber="199">P</emphasis>
.
<emphasis id="B979EAE2D645D332FCF57EAFFC3BFF6A" box="[839,1004,210,236]" italics="true" pageId="1" pageNumber="199">dobbertinensis</emphasis>
</taxonomicName>
). Synonymy of this Early Jurassic genus with the enigmatic genus
<taxonomicName id="4C0D4D73D645D332FBEC7E8BFA70FE96" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[1118,1447,246,272]" class="Insecta" family="Protogryllidae" genus="Achaetites" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FBEC7E8BFB04FE96" box="[1118,1235,246,272]" italics="true" pageId="1" pageNumber="199">Achaetites</emphasis>
Handlirsch, 1906
</taxonomicName>
, based on a female from the Early Cretaceous,
<taxonomicName id="4C0D4D73D645D332FAEC7F67FBE4FEDC" authority="Brodie, 1845" authorityName="Brodie" authorityYear="1845" class="Insecta" family="Gryllidae" genus="Acheta" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="species" species="sedgwicki">
<emphasis id="B979EAE2D645D332FAEC7F67FC4FFEDF" italics="true" pageId="1" pageNumber="199">Acheta sedgwicki</emphasis>
Brodie, 1845
</taxonomicName>
(
<bibRefCitation id="EF9C4B01D645D332FBF07F42FB0BFEDF" author="Zeuner, F. E." box="[1090,1244,319,345]" pageId="1" pageNumber="199" refId="ref5166" refString="Zeuner, F. E. (1939) Fossil Orthoptera Ensifera. British Museum (Natural History), London, 321 pp." type="book" year="1939">Zeuner, 1939</bibRefCitation>
: p. 188), is almost undoubtedly erroneous. Differences of
<taxonomicName id="4C0D4D73D645D332FAAD7F1EFA7BFEFB" authorityName="Handlirsch" authorityYear="1906" box="[1311,1452,355,381]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FAAD7F1EFA7BFEFB" box="[1311,1452,355,381]" italics="true" pageId="1" pageNumber="199">Protogryllus</emphasis>
</taxonomicName>
from the other genera of this subfamily (
<taxonomicName id="4C0D4D73D645D332FB417FF5FC0AFE40" authority="Haughton, 1924" authorityName="Haughton" authorityYear="1924" class="Insecta" family="Protogryllidae" genus="Archaegryllodes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FB417FF5FA7CFE24" box="[1267,1451,392,418]" italics="true" pageId="1" pageNumber="199">Archaegryllodes</emphasis>
Haughton, 1924
</taxonomicName>
and
<taxonomicName id="4C0D4D73D645D332FBA17FD1FAACFE40" authority="Gorochov, 1984" authorityName="Gorochov" authorityYear="1984" box="[1043,1403,428,454]" class="Insecta" family="Protogryllidae" genus="Bacharogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FBA17FD1FB17FE40" box="[1043,1216,428,454]" italics="true" pageId="1" pageNumber="199">Bacharogryllus</emphasis>
Gorochov, 1984
</taxonomicName>
) are also rather problematic:
<taxonomicName id="4C0D4D73D645D332FBF57FADFB28FE6C" authorityName="Haughton" authorityYear="1924" box="[1095,1279,464,490]" class="Insecta" family="Protogryllidae" genus="Archaegryllodes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FBF57FADFB28FE6C" box="[1095,1279,464,490]" italics="true" pageId="1" pageNumber="199">Archaegryllodes</emphasis>
</taxonomicName>
was described based on a Triassic male tegmen from
<collectingCountry id="F31A7660D645D332FAA77F88FA7BFD89" box="[1301,1452,501,527]" name="South Africa" pageId="1" pageNumber="199">South Africa</collectingCountry>
having a long and rather wide lancet-like area with only one crossvein in the apical part;
<taxonomicName id="4C0D4D73D645D332FB3B7C43FAE1FDDE" authorityName="Gorochov" authorityYear="1984" box="[1161,1334,574,600]" class="Insecta" family="Protogryllidae" genus="Bacharogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FB3B7C43FAE1FDDE" box="[1161,1334,574,600]" italics="true" pageId="1" pageNumber="199">Bacharogryllus</emphasis>
</taxonomicName>
was based on male tegmina of two species from the Middle Jurassic of Mongolia which have a clearly smaller (shorter and narrower) lancet-like area with two or more crossveins; in the diverse European males from the Lower Jurassic, usually attributed to
<taxonomicName id="4C0D4D73D645D332FBA77C89FB75FC88" authorityName="Handlirsch" authorityYear="1906" box="[1045,1186,756,782]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FBA77C89FB75FC88" box="[1045,1186,756,782]" italics="true" pageId="1" pageNumber="199">Protogryllus</emphasis>
</taxonomicName>
, this lancet-like area is often unknown or rather long and narrow (but sometimes rather wide) as well as without distinct crossveins. Numerous species from the Lower Jurassic of Europe, included in
<taxonomicName id="4C0D4D73D645D332FC0C7DF8FB9CFC19" authorityName="Handlirsch" authorityYear="1906" box="[958,1099,901,927]" class="Insecta" family="Protogryllidae" genus="Protogryllus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAE2D645D332FC0C7DF8FB9CFC19" box="[958,1099,901,927]" italics="true" pageId="1" pageNumber="199">Protogryllus</emphasis>
</taxonomicName>
, are not suitable for generic taxonomy (if they are described after female tegmina which do not provide suitably good characters) or are in need of restudy and may belong to more than one genus.
</paragraph>
<paragraph id="8BB236F0D645D332FCED7A6AFA75FBFC" blockId="1.[810,1452,173,1146]" pageId="1" pageNumber="199">Thus, the male tegmen considered below, having the lancet-like area somewhat different from that of the taxa discussed above, most probably belongs to a new genus.</paragraph>
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