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<document id="BE30D3ED926721DB151F5DA5A1CA336D" ID-CLB-Dataset="42346" ID-DOI="10.11646/zootaxa.3911.2.5" ID-GBIF-Dataset="b0c15d7f-7b3b-44b8-9743-1c1caef29d27" ID-ISSN="1175-5326" ID-Zenodo-Dep="240391" ID-ZooBank="22BEC736-D607-4997-8B76-90C03149E751" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460941215999" checkinUser="plazi" docAuthor="Hancock, E. Geoffrey, Hewitt, Stephen M., Horsfield, David, Lyszkowsi, Richard M., Macgowan, Iain, Ricarte, Antonio, Rotheray, Graham E. &amp; Watt, Kenneth" docDate="2015" docId="03A4879BFFF80F1FFF32A09A62D86898" docLanguage="en" docName="zt03911p244.pdf" docOrigin="Zootaxa 3911 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prionolabis pjotri Hancock, Hewitt, Horsfield, Lyszkowsi, Macgowan, Ricarte, Rotheray &amp; Watt, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="236" masterDocId="FF9DFFE3FFFD0F1AFFA5A35A62386F69" masterDocTitle="Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae" masterLastPageNumber="244" masterPageNumber="231" pageNumber="236" updateTime="1698584370164" updateUser="ExternalLinkService">
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<mods:title id="4287EC6784CDC90D02E4F7C918AB7EE2">Nematocera flies recorded in Serra do Courel, northwest Spain, May 2012 (Diptera: Anisopodidae, Blepharoceridae, Cylindrotomidae, Limoniidae, Pediciidae, Tipulidae and Trichoceridae) including descriptions of two new species of Limoniidae</mods:title>
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<mods:namePart id="4BD7DA0F5461F99B7FBAF3DEAB82CBCA">Hancock, E. Geoffrey</mods:namePart>
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<mods:namePart id="E2B56321AF7449C539234857338C5D80">Hewitt, Stephen M.</mods:namePart>
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<mods:namePart id="420EF02F3B97F2CE1FF5DF6EFD586014">Horsfield, David</mods:namePart>
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<mods:namePart id="7EBBE701E91C96F7E0CA2CB8A9FF62C6">Lyszkowsi, Richard M.</mods:namePart>
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<mods:namePart id="F1F335B2BC90A6724B120868D641DA19">Macgowan, Iain</mods:namePart>
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<mods:namePart id="20B216D9DD3725550514538D569A12DF">Ricarte, Antonio</mods:namePart>
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<mods:namePart id="A0208EA66885528B0D02E55B88611654">Rotheray, Graham E.</mods:namePart>
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<mods:namePart id="770121068A96AFC686FA416637E1FF2B">Watt, Kenneth</mods:namePart>
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<mods:date id="A3564A7E484322DDAA654CFE5B90307F">2015</mods:date>
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<mods:identifier id="9F38C50768B7D9003BBE438739C4C179" type="DOI">10.11646/zootaxa.3911.2.5</mods:identifier>
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<paragraph id="8BB2368DFFF80F1FFF32A09A607F6CB3" blockId="5.[151,583,960,986]" box="[151,583,960,986]" pageId="5" pageNumber="236">
<heading id="D0FA81E1FFF80F1FFF32A09A607F6CB3" bold="true" box="[151,583,960,986]" fontSize="11" level="1" pageId="5" pageNumber="236" reason="1">
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<taxonomicName id="4C0D4D0EFFF80F1FFF32A09A63536CB3" ID-CoL="8GZGF" box="[151,363,960,986]" class="Insecta" family="Limoniidae" genus="Prionolabis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="pjotri" status="sp. nov.">
<emphasis id="B979EA9FFFF80F1FFF32A09A63536CB3" bold="true" box="[151,363,960,986]" italics="true" pageId="5" pageNumber="236">Prionolabis pjotri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A57E4FFF80F1FFED6A09A63E86CB3" box="[371,464,960,986]" pageId="5" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
Hancock
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</paragraph>
</subSubSection>
<subSubSection id="C3176506FFF80F1FFF32A75F66F769B1" pageId="5" pageNumber="236" type="description">
<paragraph id="8BB2368DFFF80F1FFF32A75F612B6B98" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
<emphasis id="B979EA9FFFF80F1FFF32A75F63116B77" bold="true" box="[151,297,1029,1054]" pageId="5" pageNumber="236">Description.</emphasis>
Head and entire body shining black, quite strongly dusted (pruinose) especially on the pleurae, less so on scutum and dorsally on abdomen, sparsely pale haired; apices of coxae, trochanters and bases of femora brown. Antennae 16 segmented. Wing membrane pale grey, stigma dark, wing veins yellow on basal third; medial vein four-branched (i.e., upper branch forked and so cell M1 present). Halteres yellow. Male wing length 8.00mm, ranging from 6.6 to 9.0mm (sample size = 19). Male genitalia (
<figureCitation id="13362A08FFF80F1FFCF7A7C961F96BC5" box="[850,961,1171,1196]" captionStart="FIGURE 15" captionStartId="8.[151,250,1245,1267]" captionText="FIGURE 15. Prionolabis pjotri n. sp., hypopygium of male." pageId="5" pageNumber="236">Figure 15</figureCitation>
) with outer dististyle bifurcate at tip, inner one angled after swollen medial section resembling a birds beak. Parameres are flattened and only lightly sclerotised, expanding before a squared off truncated tip.
</paragraph>
<paragraph id="8BB2368DFFF80F1FFF62A7A461556A50" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
Female same colouration as male but with apex of 8th tergite, cerci and sternal valves of ovipositor yellowbrown; wing reduced (
<figureCitation id="13362A08FFF80F1FFE36A67A603B6A50" box="[403,515,1312,1337]" captionStart="FIGURE 16" captionStartId="8.[151,250,2011,2033]" captionText="FIGURE 16. Prionolabis pjotri n. sp., wing of female (total length 2 mm)" pageId="5" pageNumber="236">Figure 16</figureCitation>
), length
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(sample size = 1).
</paragraph>
<paragraph id="8BB2368DFFF80F1FFF62A61966166A16" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
<typeStatus id="54B6882FFFF80F1FFF62A619630C6A35" box="[199,308,1347,1372]" pageId="5" pageNumber="236" type="holotype">Holotype</typeStatus>
: male, Alto do Couto, Lugo, Galicia,
<collectingCountry id="F31A761DFFF80F1FFCACA61961746A35" box="[777,844,1347,1372]" name="Spain" pageId="5" pageNumber="236">Spain</collectingCountry>
,
<date id="FFB3104DFFF80F1FFCFBA619663D6A35" box="[862,1029,1347,1372]" pageId="5" pageNumber="236" value="2012-05-25">25 May 2012</date>
, E.G. Hancock. Deposited in The Hunterian Zoology Museum, University of Glasgow (Accession number
<date id="FFB3104DFFF80F1FFC6FA63C661B6A16" box="[970,1059,1382,1407]" pageId="5" pageNumber="236" value="30-06-15">150630</date>
).
</paragraph>
<paragraph id="8BB2368DFFF80F1FFF62A6D360086926" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
<typeStatus id="54B6882FFFF80F1FFF62A6D363016AC8" box="[199,313,1417,1441]" pageId="5" pageNumber="236" type="paratype">Paratypes</typeStatus>
: five males swept on woodland edge, nr Moreda,
<date id="FFB3104DFFF80F1FFCDEA6D266216AC8" box="[891,1049,1416,1441]" pageId="5" pageNumber="236" value="2012-05-26">26 May 2012</date>
; eleven males and one female, nr Mercurin,
<date id="FFB3104DFFF80F1FFEA8A6F6639D6AAD" box="[269,421,1452,1476]" pageId="5" pageNumber="236" value="2012-05-27">27 May 2012</date>
; one male by Rio Lor, Ferrería Nova, Seoane do Courel,
<date id="FFB3104DFFF80F1FFB81A6F666856AAD" box="[1060,1213,1452,1476]" pageId="5" pageNumber="236" value="2012-05-30">30 May 2012</date>
; E.G. Hancock, The Hunterian Zoology Museum, University of Glasgow (Entry No. 897). Three additional specimens also designated as
<typeStatus id="54B6882FFFF80F1FFF1DA6AB631B6960" box="[184,291,1521,1545]" pageId="5" pageNumber="236" type="paratype">paratypes</typeStatus>
in the collection of J. Starý, Palacký University, Olomouc,
<collectingCountry id="F31A761DFFF80F1FFC72A6AA66A86960" box="[983,1168,1520,1545]" name="Czech Republic" pageId="5" pageNumber="236">Czech Republic</collectingCountry>
: one male, San Martin, Lugo,
<date id="FFB3104DFFF80F1FFF44A54E63A86945" box="[225,400,1555,1580]" pageId="5" pageNumber="236" value="1984-03-22">22 March 1984</date>
, E. Eiroa; two males,
<quantity id="4CF59B68FFF80F1FFD2FA54E60EA6945" box="[650,722,1555,1580]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="236" unit="km" value="23.0">23 km</quantity>
south of Betanzos, A Coruna, in wet deciduous forest,
<date id="FFB3104DFFF80F1FFAE0A54962EA6926" pageId="5" pageNumber="236" value="1994-05-16">16 May 1994</date>
, P. Oosterbroek &amp; C. Hartveld.
</paragraph>
<paragraph id="8BB2368DFFF80F1FFF62A50266F769B1" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
Details in the male genitalia are distinctive, particularly the inner dististyle with its strong resemblance to the profile of a bird and the truncate apex to the pale and flattened parameres (
<figureCitation id="13362A08FFF80F1FFC7FA521667269FD" box="[986,1098,1659,1684]" captionStart="FIGURE 15" captionStartId="8.[151,250,1245,1267]" captionText="FIGURE 15. Prionolabis pjotri n. sp., hypopygium of male." pageId="5" pageNumber="236">Figure 15</figureCitation>
). The females wings are long enough to retain recognisable venation. The colouration seen in the live habitus photographs (
<figureCitation id="13362A08FFF80F1FFB66A5C4670669DE" box="[1219,1342,1694,1719]" captionStart="FIGURE 17" captionStartId="9.[151,250,913,935]" captionTargetBox="[412,1180,193,881]" captionTargetId="figure@9.[412,1180,193,882]" captionTargetPageId="9" captionText="FIGURE 17. Prionolabis pjotri n. sp., photograph of male, Galicia, 2012 (Courtesy of Belen, Diptera. Info website)" httpUri="https://zenodo.org/record/240406/files/figure.png" pageId="5" pageNumber="236">Figures 17</figureCitation>
and
<figureCitation id="13362A08FFF80F1FFAD2A5C567AC69DE" box="[1399,1428,1695,1719]" captionStart="FIGURE 18" captionStartId="9.[151,250,1973,1995]" captionTargetBox="[446,1144,988,1938]" captionTargetId="figure@9.[446,1144,988,1939]" captionTargetPageId="9" captionText="FIGURE 18. Prionolabis pjotri n. sp., photograph of female, Galicia, 2012 (Courtesy of Belen, Diptera. Info website)." httpUri="https://zenodo.org/record/240407/files/figure.png" pageId="5" pageNumber="236">18</figureCitation>
) shows the strong contrast between the yellow wing bases and the rest of the insect in both sexes.
</paragraph>
</subSubSection>
<subSubSection id="C3176506FFF80F1FFF62A5B966986876" pageId="5" pageNumber="236" type="etymology">
<paragraph id="8BB2368DFFF80F1FFF62A5B966986876" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
<emphasis id="B979EA9FFFF80F1FFF62A5B963766995" bold="true" box="[199,334,1763,1788]" pageId="5" pageNumber="236">Etymology.</emphasis>
The expertise of Pjotr Oosterbroek, managing the collections under his care and the creation and development of the Cranefly Catalog of the World is commemorated in naming this species.
</paragraph>
</subSubSection>
<subSubSection id="C3176506FFF80F1FFF62A47262D86898" pageId="5" pageNumber="236" type="discussion">
<paragraph id="8BB2368DFFF80F1FFF62A47262D86898" blockId="5.[151,1437,1029,2033]" pageId="5" pageNumber="236">
<emphasis id="B979EA9FFFF80F1FFF62A47263746828" bold="true" box="[199,332,1832,1857]" pageId="5" pageNumber="236">Discussion.</emphasis>
There is one other known Spanish species in this genus,
<taxonomicName id="4C0D4D0EFFF80F1FFC4AA47267146828" authority="Egger" authorityName="Egger" box="[1007,1324,1832,1857]" class="Insecta" family="Limoniidae" genus="Prionolabis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="B979EA9FFFF80F1FFC4AA47266F16828" box="[1007,1225,1832,1857]" italics="true" pageId="5" pageNumber="236">Prionolabis hospes</emphasis>
(Egger)
</taxonomicName>
, which is widespread in the western Palaearctic. It cannot be confused with
<taxonomicName id="4C0D4D0EFFF80F1FFC25A41661E2680A" box="[896,986,1868,1891]" class="Insecta" family="Limoniidae" genus="Prionolabis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="pjotri">
<emphasis id="B979EA9FFFF80F1FFC25A41661E2680A" box="[896,986,1868,1891]" italics="true" pageId="5" pageNumber="236">P. pjotri</emphasis>
</taxonomicName>
in its lack of a branch to vein M
<quantity id="4CF59B68FFF80F1FFAE8A400674A680D" box="[1357,1394,1867,1896]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="236" unit="in" value="1.0">1 in</quantity>
the male, the female having even smaller wings with much reduced venation and is not black in colour. One other species
<taxonomicName id="4C0D4D0EFFF80F1FFF55A4C960B268C5" authority="Strobl" authorityName="Strobl" box="[240,650,1939,1964]" class="Insecta" family="Limoniidae" genus="Prionolabis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="longeantennata">
<emphasis id="B979EA9FFFF80F1FFF55A4C9601268C5" box="[240,554,1939,1964]" italics="true" pageId="5" pageNumber="236">Prionolabis longeantennata</emphasis>
(Strobl)
</taxonomicName>
does have four branches to M but the antennae are almost as long as the body as does
<emphasis id="B979EA9FFFF80F1FFEF9A4ED63F468A7" box="[348,460,1974,1999]" italics="true" pageId="5" pageNumber="236">P. s c h m i d i</emphasis>
Starý, but which has even longer antennae, as long as or longer than the body (
<bibRefCitation id="EF9C4B7CFFF80F1FFAFCA4EC62EC6898" author="Stary" pageId="5" pageNumber="236" refString="Stary, J. (2006) New information resulting from a study of Strobl's types of Limoniidae. Acta universitatis Carolinae biologica, 49, 187 - 203." type="journal article" year="2006">Starý, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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