180 lines
28 KiB
XML
180 lines
28 KiB
XML
<document id="7AC16095A1B141818789D3AB67EF52DE" ID-CLB-Dataset="9458" ID-DOI="10.11646/zootaxa.4889.1.1" ID-GBIF-Dataset="11a122a9-8f3e-4a1c-a8d3-5ef3bea09b59" ID-ISSN="1175-5326" ID-Zenodo-Dep="4297816" ID-ZooBank="0B8630F6-2EF0-44E6-9D3A-7386BF949FD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606731381730" checkinUser="plazi" docAuthor="Grzymala, Traci L. & Leschen, Richard A. B." docDate="2020" docId="03A48794FFC8FFF66F854ABC734BFCE1" docLanguage="en" docName="zootaxa.4889.1.1.pdf" docOrigin="Zootaxa 4889 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudozena Grzymala & Leschen 2020, gen. n." docType="treatment" docUuid="0FB818D0-77B8-4D42-98D4-C7AB35354363" docUuidSource="ZooBank" docVersion="11" lastPageNumber="24" masterDocId="FF9DFFECFFDEFFE16F124B66712FFFBC" masterDocTitle="Sexual Dimorphism of New Zealand Puppet Beetles (Aderidae, Coleoptera, Tenebrionoidea): Systematic Revision, Description of Three New Genera, and Phylogeny for Zenascus, gen. n." masterLastPageNumber="59" masterPageNumber="1" pageNumber="23" updateTime="1698846900633" updateUser="ExternalLinkService">
|
||
<mods:mods id="6A93B78367AC1150A384855674DCB9D8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8BEEF58CB7F09AAC54C674196356B74F">
|
||
<mods:title id="AFB5DACEE6C47B3DCFD70A03B77A5C32">Sexual Dimorphism of New Zealand Puppet Beetles (Aderidae, Coleoptera, Tenebrionoidea): Systematic Revision, Description of Three New Genera, and Phylogeny for Zenascus, gen. n.</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FC445F4F7642019D69A5BBE1D6ED0C32" type="personal">
|
||
<mods:role id="8673C5F1F2B3B6D21A91F59534234DAA">
|
||
<mods:roleTerm id="460552BE79E3719376BE3AEF623FA082">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3A0D1F5A7DF1E0E16EDFA1CFD1B550CA">Grzymala, Traci L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D9115E8A0D8FDBC1A7D1C4C703435E0B" type="personal">
|
||
<mods:role id="D7F1BE208CB5EC40DAC74F9473DEFC61">
|
||
<mods:roleTerm id="E9EB7CCB52327BFA7AE2510C61A9E8F8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5B3537840FB0C0BFD05E9077CF18C776">Leschen, Richard A. B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4EFA68AE42F90833D4CC3E5DD6C80266">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F8D3111430AC49635EF07B3777BF34BE" type="host">
|
||
<mods:titleInfo id="60F0095311ABF3BD984058A67F14F6FE">
|
||
<mods:title id="218C08792DE8CAE5491E2C1219310696">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="137082C598603F4A443F43C8E230228B">
|
||
<mods:date id="0E8821E321621B7E1CEE520C020CA47F">2020</mods:date>
|
||
<mods:detail id="BAE4BE19DD858EB45F042BADFE95D1A0" type="pubDate">
|
||
<mods:number id="EDA16E1D50DB94E7B2D4CA0E727B1B20">2020-11-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="348779B6E73B7D85B264DABBC5C50A37" type="volume">
|
||
<mods:number id="026DB941E3C7D9ABB8A788EE59975FCF">4889</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="65A332BD5B006B479EA097AE5619045D" type="issue">
|
||
<mods:number id="50F493111BFEC6B52D91555A6EDEC093">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="47057890A6B6389908AE6BD47F225BD8" unit="page">
|
||
<mods:start id="A71C61FA6E4E2A25F3EF12A5D4BAA9BA">1</mods:start>
|
||
<mods:end id="F19F7B87AFF9ADFFA8F5B2B25D9843DA">59</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="19C871B017D7186BDE279D7FB12429E5">journal article</mods:classification>
|
||
<mods:identifier id="6168E7C246C98585BF7814021BEA45B5" type="CLB-Dataset">9458</mods:identifier>
|
||
<mods:identifier id="4AB12E790D44F89FDB727B876950DDA0" type="DOI">10.11646/zootaxa.4889.1.1</mods:identifier>
|
||
<mods:identifier id="E3675D23D8895015727F492424515391" type="GBIF-Dataset">11a122a9-8f3e-4a1c-a8d3-5ef3bea09b59</mods:identifier>
|
||
<mods:identifier id="0BE9FE1460D2A1262996DEC96D762B79" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="63F415662054AFA35CAB4EE551873828" type="Zenodo-Dep">4297816</mods:identifier>
|
||
<mods:identifier id="361967487C2D1D93B09CBE1DC1ADF9A2" type="ZooBank">0B8630F6-2EF0-44E6-9D3A-7386BF949FD0</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A48794FFC8FFF66F854ABC734BFCE1" ID-DOI="http://doi.org/10.5281/zenodo.4334374" ID-GBIF-Taxon="170606625" ID-Zenodo-Dep="4334374" LSID="urn:lsid:zoobank.org:act:0FB818D0-77B8-4D42-98D4-C7AB35354363" httpUri="http://treatment.plazi.org/id/03A48794FFC8FFF66F854ABC734BFCE1" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<subSubSection id="C3176509FFC8FFF76F854ABC70AFFE49" box="[151,384,474,501]" pageId="22" pageNumber="23" type="nomenclature">
|
||
<paragraph id="8BB23682FFC8FFF76F854ABC70AFFE49" blockId="22.[151,384,474,501]" box="[151,384,474,501]" pageId="22" pageNumber="23">
|
||
<heading id="D0FA81EEFFC8FFF76F854ABC70AFFE49" bold="true" box="[151,384,474,501]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
|
||
<emphasis id="B979EA90FFC8FFF76F854ABC70AFFE49" bold="true" box="[151,384,474,501]" pageId="22" pageNumber="23">
|
||
<taxonomicName id="4C0D4D01FFC8FFF76F854ABC7009FE49" authority="Grzymala & Leschen, 2020" authorityName="Grzymala & Leschen" authorityYear="2020" box="[151,294,474,501]" class="Insecta" family="Aderidae" genus="Pseudozena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B979EA90FFC8FFF76F854ABC7009FE49" bold="true" box="[151,294,474,501]" italics="true" pageId="22" pageNumber="23">Pseudozena</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A57EBFFC8FFF76E3C4ABD70AFFE49" box="[302,384,475,501]" pageId="22" pageNumber="23" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC8FFF76F8549797220FD8B" box="[151,783,543,567]" pageId="22" pageNumber="23" type="type_taxon">
|
||
<paragraph id="8BB23682FFC8FFF76F8549797220FD8B" blockId="22.[151,783,543,567]" box="[151,783,543,567]" pageId="22" pageNumber="23">
|
||
<emphasis id="B979EA90FFC8FFF76F8549797009FD8B" bold="true" box="[151,294,543,567]" pageId="22" pageNumber="23">
|
||
<typeStatus id="54B68820FFC8FFF76F85497971E3FD8B" box="[151,204,543,567]" pageId="22" pageNumber="23">Type</typeStatus>
|
||
species:
|
||
</emphasis>
|
||
<taxonomicName id="4C0D4D01FFC8FFF76E3E49797332FD8B" authority="Grzymala & Leschen, 2020" authorityName="Grzymala & Leschen" authorityYear="2020" box="[300,541,543,567]" class="Insecta" family="Aderidae" genus="Pseudozena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
|
||
<emphasis id="B979EA90FFC8FFF76E3E49797332FD8B" box="[300,541,543,567]" italics="true" pageId="22" pageNumber="23">Pseudozena denticulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B979EA90FFC8FFF76D3649797372FD8B" bold="true" box="[548,605,543,567]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A24A57EBFFC8FFF76D3649797372FD8B" box="[548,605,543,567]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, here designated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC8FFF76F85490373CEFD74" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="8BB23682FFC8FFF76F85490373CEFD74" blockId="22.[151,1437,613,2008]" pageId="22" pageNumber="23">
|
||
<emphasis id="B979EA90FFC8FFF76F8549037031FDC3" bold="true" box="[151,286,613,639]" pageId="22" pageNumber="23">Etymology.</emphasis>
|
||
The genus is named with reference to the deceptively similar overall dorsal habitus to the other more speciose
|
||
<collectingCountry id="F31A7612FFC8FFF76FEC49EC70BBFD1F" box="[254,404,649,676]" name="New Zealand" pageId="22" pageNumber="23">New Zealand</collectingCountry>
|
||
genus,
|
||
<taxonomicName id="4C0D4D01FFC8FFF76EFE49EC737BFD1F" authorityName="Grzymala & Leschen" authorityYear="2020" box="[492,596,650,675]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EA90FFC8FFF76EFE49EC737BFD1F" box="[492,596,650,675]" italics="true" pageId="22" pageNumber="23">Zenascus</emphasis>
|
||
</taxonomicName>
|
||
. The name is formed from the Greek
|
||
<emphasis id="B979EA90FFC8FFF76B1349EF757FFD1F" box="[1025,1104,649,675]" italics="true" pageId="22" pageNumber="23">pseudo</emphasis>
|
||
(misrepresent, practice trickery) combined with
|
||
<emphasis id="B979EA90FFC8FFF76E6B49C87083FD7B" box="[377,428,686,711]" italics="true" pageId="22" pageNumber="23">zena</emphasis>
|
||
as a reference to
|
||
<taxonomicName id="4C0D4D01FFC8FFF76D6349C873F6FD7B" authorityName="Grzymala & Leschen" authorityYear="2020" box="[625,729,686,711]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EA90FFC8FFF76D6349C873F6FD7B" box="[625,729,686,711]" italics="true" pageId="22" pageNumber="23">Zenascus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC8FFF76FD549B773E8FC1C" pageId="22" pageNumber="23" type="diagnosis">
|
||
<paragraph id="8BB23682FFC8FFF76FD549B773E8FC1C" blockId="22.[151,1437,613,2008]" pageId="22" pageNumber="23">
|
||
<emphasis id="B979EA90FFC8FFF76FD549B7706DFD57" bold="true" box="[199,322,721,747]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
|
||
This genus is distinguished by the subglobular scape and pedicel, the anntennomere 3 subequal in length and width to antennomere 4, the biseriate elytra, and the metafemur featuring a characteristic ctenidium accompanied by a row of sclerotized ridges present on the postero-ventral margin of both males and females (
|
||
<figureCitation id="13362A07FFC8FFF76A50487F71FAFCEB" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="22" pageNumber="23">Figs 9D & 9F</figureCitation>
|
||
). The genus is superficially very similar to the genus
|
||
<taxonomicName id="4C0D4D01FFC8FFF76C3F485872BAFCEB" authorityName="Grzymala & Leschen" authorityYear="2020" box="[813,917,830,855]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EA90FFC8FFF76C3F485872BAFCEB" box="[813,917,830,855]" italics="true" pageId="22" pageNumber="23">Zenascus</emphasis>
|
||
</taxonomicName>
|
||
, but can be distinguished by the subequal size and shape of antennomeres 3 & 4, the distinct modifications to the metafemora, and the interstitial setae of the elytra subequal in length and width to the primary setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC8FFF66FD548CF702EFD39" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="8BB23682FFC8FFF66FD548CF70EFFD81" blockId="22.[151,1437,613,2008]" lastBlockId="23.[151,1437,151,861]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<emphasis id="B979EA90FFC8FFF76FD548CF7076FC7F" bold="true" box="[199,345,937,963]" pageId="22" pageNumber="23">Description.</emphasis>
|
||
Average size for family, length
|
||
<quantity id="4CF59B67FFC8FFF76DD548CF7241FC78" box="[711,878,937,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.635" metricValueMax="1.67" metricValueMin="1.6" pageId="22" pageNumber="23" unit="mm" value="1.635" valueMax="1.67" valueMin="1.6">1.60–1.67 mm</quantity>
|
||
. Head abruptly constricted posteriorly, forming a neck, concealed within pronotum. Eyes moderately protuberant, coarsely faceted, with moderate anterior emargination, with short interfacetal setae. Antennal insertions anterad of eyes and eye emargination, produced on slight ridge. Frontoclypeal suture present. Mandible bidentate. Pronotum (
|
||
<figureCitation id="13362A07FFC8FFF76C9F4F7372CAFB93" box="[909,997,1045,1071]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="22" pageNumber="23">Fig. 8A</figureCitation>
|
||
) subquadrate, width subequal to slightly greater than length; pronotal width 0.5× elytral width; sides sinuate, anterior angles rounded, posterior angles rounded; disc with two strongly impressed basal fovea, and a distinct transverse sulcus just anterad of center; surface with deep, elongate punctures, unevenly spaced from one another, separated by an average of two punctural lengths; punctures confused on central disc. Prosternal process short, extremely narrowed posteriorly, apex acute. Mesonotum with slight central anterior notch; suture between mesonotum and scutellar shield distinct, centrally emarginate; scutellar shield (
|
||
<figureCitation id="13362A07FFC8FFF76EC94F8B731CFABB" box="[475,563,1261,1287]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="22" pageNumber="23">Fig. 8D</figureCitation>
|
||
) subquadrate, length approximately equal to width, posterior margin straight, lateral apices broadly rounded. Elytron (
|
||
<figureCitation id="13362A07FFC8FFF76D444E7773F6FA97" box="[598,729,1297,1323]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="22" pageNumber="23">Figs 8E, 8F</figureCitation>
|
||
) with moderately, obliquely impressed subscutellar depression; surface with elongate, elliptical punctures, separated by an average of three punctural lengths, pubescence biseriate, primary setae decumbent, thin, length about equal to two punctural lengths; secondary setae subdecumbent, subequal in width and length to primary, 2–3 present between each approximate pair of deep, elongate punctures; posterior margin rounded, without protrusions. Mesoventrite without procoxal rests. Mesanepitsternum with deep, punctation. Mesepimeron with deep punctation. Mesanepisternal-mesepimeral suture faintly indicated by absence of punctures. Mesoventral process (
|
||
<figureCitation id="13362A07FFC8FFF76D374E8F7353F9BF" box="[549,636,1513,1539]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="22" pageNumber="23">Fig. 8G</figureCitation>
|
||
) moderately broad, about 0.5× width of mesocoxa, posterior margin inserted below metaventral process. Metaventrite (
|
||
<figureCitation id="13362A07FFC8FFF76DA44D6B7222F99B" box="[694,781,1549,1575]" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
) with deep punctures, approximately evenly spaced across surface except medially and at margins anterad of metacoxae; discrimen indistinct, without invagination. Metendosternite with moderately long lateral arms; anterior process absent; laminae absent. Proleg with femur (
|
||
<figureCitation id="13362A07FFC8FFF76A144D337474F9D3" box="[1286,1371,1621,1647]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="22" pageNumber="23">Fig. 8C</figureCitation>
|
||
) simple, width subequal basally, medially, and apically; tibia gradually expanded in width apically, without apical spine; tarsomeres 1–3 subequal in width, each ventrally with spatulate, adhesive setae, equally concentrated on each; 1
|
||
<superScript id="7C789BCAFFC8FFF76A834DFB74B3F917" attach="none" box="[1425,1436,1693,1707]" fontSize="6" pageId="22" pageNumber="23">st</superScript>
|
||
protarsomere 2.5× longer than protarsomere 2; 2
|
||
<superScript id="7C789BCAFFC8FFF76DAD4DA773E0F973" attach="left" box="[703,719,1729,1743]" fontSize="6" pageId="22" pageNumber="23">nd</superScript>
|
||
protarsomere 1.5× longer than protarsomere 3; protarsomere 3 expanded ventrally; protarsomere 4 reduced; 5
|
||
<superScript id="7C789BCAFFC8FFF76DB14D83739FF94F" attach="left" box="[675,688,1765,1779]" fontSize="6" pageId="22" pageNumber="23">th</superScript>
|
||
protarsomere 2.5× longer than wide. Midleg with femur (
|
||
<figureCitation id="13362A07FFC8FFF76A2F4D8374BBF943" box="[1341,1428,1765,1791]" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="22" pageNumber="23">Fig. 9C</figureCitation>
|
||
) simple, width subequal basally, medially, and apically; tibia straight from base to apex, gradually expanded apically; tarsomeres 1–3 subequal in width, each ventrally with spatulate, adhesive setae, concentrated on mesotarsomere 3; 1
|
||
<superScript id="7C789BCAFFC8FFF76FB74C37719FF8E3" attach="left" box="[165,176,1873,1887]" fontSize="6" pageId="22" pageNumber="23">st</superScript>
|
||
mesotarsomere 2.5× longer than mesotarsomere 2; 2
|
||
<superScript id="7C789BCAFFC8FFF76C114C37723CF8E3" attach="left" box="[771,787,1873,1887]" fontSize="6" pageId="22" pageNumber="23">nd</superScript>
|
||
mesotarsomere 1.5× longer than mesotarsomere 3; mesotarsomere 3 expanded ventrally; mesotarsomere 4 reduced; 5
|
||
<superScript id="7C789BCAFFC8FFF76C534C137261F83F" attach="left" box="[833,846,1909,1923]" fontSize="6" pageId="22" pageNumber="23">th</superScript>
|
||
mesotarsomere 2.5× longer than wide. Hindleg with coxa (
|
||
<figureCitation id="13362A07FFC8FFF76FCE4CFF701CF80F" box="[220,307,1945,1971]" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="22" pageNumber="23">Fig. 9B</figureCitation>
|
||
) having deep punctures confined to posterior margin, with suture anterad of punctures; femur (
|
||
<figureCitation id="13362A07FFC8FFF76A794CFF71CAF86B" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="22" pageNumber="23">Figs 9C, 9F</figureCitation>
|
||
) with distinct line of thickened setae on basal half of ventro-posterior margin, with row of eight sclerotized ridges, confined to basal half, of varying thickness and height, gradually expanded in width medially, medial width approximately 1.8× width basally and apically; tibia (
|
||
<figureCitation id="13362A07FFC9FFF66DFE4BDD726FFF69" box="[748,832,187,213]" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="23" pageNumber="24">Fig. 9E</figureCitation>
|
||
) gradually expanded in width from base to apex, apex inner face without triangular expansion; tarsomere 1 elongate, length approximately 8× width; metatarsomere 2 length subequal to width, expanded ventrally, with spatulate, adhesive setae ventrally; metatarsomere 3 reduced; metatarsomere 4 length approximately 3.5× width. Abdomen with ventrites 1 and 2 connate (
|
||
<figureCitation id="13362A07FFC9FFF66BA94A41743AFEFD" box="[1211,1301,295,321]" captionStart="FIGURES 8" captionStartId="24.[151,264,1959,1983]" captionTargetBox="[179,1409,181,1935]" captionTargetId="figure-28@24.[179,1409,181,1935]" captionTargetPageId="24" captionText="FIGURES 8A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) pronotum; (B) antenna; (C) proleg; (D) scutellum; (E) elytron; (F) elytral integument close-up; (G) mesoventral and metaventral processes." figureDoi="http://doi.org/10.5281/zenodo.4297834" httpUri="https://zenodo.org/record/4297834/files/figure.png" pageId="23" pageNumber="24">Fig. 8G</figureCitation>
|
||
), suture not indicated, without appressed thick field of pubescence, length of ventrites 1 and 2 combined 2.5× length of ventrite 3; abdominal process rounded; length of ventrites 3 and 4 subequal; deep punctures confined to abdominal process and medial area of ventrite I, present medially on ventrites 2–4, absent laterally; small, shallow micropunctures present on ventrite 5; ventrite 5 without medial impression. Phallobase (
|
||
<figureCitation id="13362A07FFC9FFF66CD34AD17537FE6D" box="[961,1048,439,465]" captionStart="FIGURES 9" captionStartId="25.[151,264,1911,1935]" captionTargetBox="[151,1436,181,1888]" captionTargetId="figure-17@25.[151,1436,181,1888]" captionTargetPageId="25" captionText="FIGURES 9A–G. Pseudozena denticulata gen n. sp. n., scanning electron micrographs. (A) metaventrite; (B) metacoxa; (C) midleg (D) metafemur; (E) metatibia; (F) close-up of metafemur ventro-posterior margin; (G) abdomen." figureDoi="http://doi.org/10.5281/zenodo.4297836" httpUri="https://zenodo.org/record/4297836/files/figure.png" pageId="23" pageNumber="24">Fig. 9C</figureCitation>
|
||
) broadly rounded anteriorly; phallobase and apicale clearly separated, not fused; apicale posteriorly narrowly acute, accessory lobes present with four setae, one long and two short setae at apex, one long seta ventrally below apex; penis with anterior struts barely extending past phallobase.
|
||
</paragraph>
|
||
<paragraph id="8BB23682FFC9FFF66FD54921702EFD39" blockId="23.[151,1437,151,861]" pageId="23" pageNumber="24">
|
||
<emphasis id="B979EA90FFC9FFF66FD54921701FFDDD" bold="true" box="[199,304,583,609]" pageId="23" pageNumber="24">Females.</emphasis>
|
||
Ventrite 5 without medial setiferous sex patch. Apex of last abdominal ventrite and tergite without serration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC9FFF66FD549E8732AFD15" box="[199,517,654,681]" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8BB23682FFC9FFF66FD549E8732AFD15" blockId="23.[151,1437,151,861]" box="[199,517,654,681]" pageId="23" pageNumber="24">
|
||
<emphasis id="B979EA90FFC9FFF66FD549E8704EFD14" bold="true" box="[199,353,654,680]" pageId="23" pageNumber="24">Distribution.</emphasis>
|
||
<collectingCountry id="F31A7612FFC9FFF66E7549E9732EFD15" box="[359,513,655,681]" name="New Zealand" pageId="23" pageNumber="24">New Zealand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176509FFC9FFF66FD549D5734BFCE1" pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph id="8BB23682FFC9FFF66FD549D5734BFCE1" blockId="23.[151,1437,151,861]" pageId="23" pageNumber="24">
|
||
<emphasis id="B979EA90FFC9FFF66FD549D57014FD71" bold="true" box="[199,315,691,717]" pageId="23" pageNumber="24">Remarks.</emphasis>
|
||
The phylogenetic relationship of
|
||
<taxonomicName id="4C0D4D01FFC9FFF66DA549D57212FD71" authorityName="Grzymala & Leschen" authorityYear="2020" box="[695,829,691,717]" class="Insecta" family="Aderidae" genus="Pseudozena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EA90FFC9FFF66DA549D57212FD71" box="[695,829,691,717]" italics="true" pageId="23" pageNumber="24">Pseudozena</emphasis>
|
||
</taxonomicName>
|
||
to other aderid genera is currently unknown. The size and shape of the pronotum and male genitalia characteristics indicate a possible close relationship with the newly described
|
||
<taxonomicName id="4C0D4D01FFC9FFF66E18499D705DFCA8" authorityName="Grzymala & Leschen" authorityYear="2020" box="[266,370,763,788]" class="Insecta" family="Aderidae" genus="Zenascus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EA90FFC9FFF66E18499D705DFCA8" box="[266,370,763,788]" italics="true" pageId="23" pageNumber="24">Zenascus</emphasis>
|
||
</taxonomicName>
|
||
. The specific
|
||
<typeStatus id="54B68820FFC9FFF66D00499D736DFCA9" box="[530,578,763,789]" pageId="23" pageNumber="24">type</typeStatus>
|
||
of biseriate elytral pubescence and particular modifications to the metafemora are unique characteristics amongst currently described aderid taxa and a phylogenetic analysis is required to determine the relationships of this genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |