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<document ID-DOI="10.11646/zootaxa.5276.1.1" ID-GBIF-Dataset="590e0e3a-f3ea-4420-b976-b668987049e6" ID-ISSN="1175­5334" ID-Zenodo-Dep="7907294" ID-ZooBank="92300500-BB24-45B0-8ADD-977C3220A069" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1682929863379" checkinUser="plazi" docAuthor="Camargo, Alexssandro, Vieira, Rodrigo &amp; Rafael, José Albertino" docDate="2023" docId="03A4878EEE443842BEE1FA98F126FE03" docLanguage="en" docName="zootaxa.5276.1.1.pdf" docOrigin="Zootaxa 5276" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.2:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="2" docTitle="Ctenodontina mochica Lamas 1973" docType="treatment" docVersion="2" lastPageNumber="11" masterDocId="FF9DFFF6EE423848BE76FFD5F253FFD2" masterDocTitle="Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="7" updateTime="1683543292131" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Camargo, Alexssandro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vieira, Rodrigo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rafael, José Albertino</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5276</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5276.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">590e0e3a-f3ea-4420-b976-b668987049e6</mods:identifier>
<mods:identifier type="ISSN">1175­5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">7907294</mods:identifier>
<mods:identifier type="ZooBank">92300500-BB24-45B0-8ADD-977C3220A069</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7907304" ID-Zenodo-Dep="7907304" LSID="urn:lsid:plazi:treatment:03A4878EEE443842BEE1FA98F126FE03" httpUri="http://treatment.plazi.org/id/03A4878EEE443842BEE1FA98F126FE03" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7">
<subSubSection box="[151,592,1357,1384]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph blockId="6.[151,592,1357,1420]" box="[151,592,1357,1384]" pageId="6" pageNumber="7">
<heading bold="true" box="[151,592,1357,1384]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName ID-CoL="325RR" authority="Lamas, 1973" authorityName="Lamas" authorityYear="1973" box="[151,592,1357,1384]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mochica">
<emphasis bold="true" box="[151,592,1357,1384]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[151,426,1357,1384]" italics="true" pageId="6" pageNumber="7">Ctenodontina mochica</emphasis>
<bibRefCitation author="Lamas, G. M." box="[433,592,1358,1384]" pageId="6" pageNumber="7" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas, 1973</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,295,1394,1420]" pageId="6" pageNumber="7" type="description">
<paragraph blockId="6.[151,592,1357,1420]" box="[151,295,1394,1420]" pageId="6" pageNumber="7">
(
<figureCitation box="[159,260,1394,1420]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="7.[152,255,1943,1967]" captionStartId-1="8.[151,250,1874,1898]" captionStartId-2="9.[151,250,1943,1967]" captionTargetBox-0="[203,1385,181,1919]" captionTargetBox-1="[227,1350,189,1848]" captionTargetBox-2="[203,1385,181,1908]" captionTargetId-0="figure-17@7.[203,1385,181,1919]" captionTargetId-1="figure-21@8.[227,1361,181,1851]" captionTargetId-2="figure-17@9.[203,1385,181,1919]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." captionText-1="FIGURE 2. Ctenodontina mochica Lamas, 1973, male. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Sternite 8, ventral; E. Hypandrium, ventral; F. Gonopods, external, lateral; G. Same, internal, lateral; H. Phallus, lateral. I. Cercus and subepandrial sclerite, dorsal; J. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." captionText-2="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907300" figureDoi-1="http://doi.org/10.5281/zenodo.7907302" figureDoi-2="http://doi.org/10.5281/zenodo.7907306" httpUri-0="https://zenodo.org/record/7907300/files/figure.png" httpUri-1="https://zenodo.org/record/7907302/files/figure.png" httpUri-2="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">Figs 1–3</figureCitation>
,
<figureCitation box="[272,286,1394,1420]" captionStart="FIGURE 8" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[151,1435,813,1946]" captionTargetId="figure-286@15.[151,1435,813,1946]" captionTargetPageId="15" captionText="FIGURE 8. Distribution map of Ctenodontina Enderlein, 1914 with the distribution records of its two species: Ctenodontina mochica Lamas, 1973 and Ctenodontina pectinatipes Enderlein, 1914." figureDoi="http://doi.org/10.5281/zenodo.7907328" httpUri="https://zenodo.org/record/7907328/files/figure.png" pageId="6" pageNumber="7">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="reference_group">
<paragraph blockId="6.[151,1436,1457,1542]" pageId="6" pageNumber="7">
<treatmentCitationGroup pageId="6" pageNumber="7">
<treatmentCitation author="Lamas, G. M." box="[151,568,1457,1481]" page="276" pageId="6" pageNumber="7" year="1973">
<taxonomicName ID-CoL="325RR" authority="Lamas, 1973: 276" authorityName="Lamas" authorityPageNumber="276" authorityYear="1973" box="[151,568,1457,1481]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mochica">
<emphasis box="[151,380,1457,1481]" italics="true" pageId="6" pageNumber="7">Ctenodontina mochica</emphasis>
<bibRefCitation author="Lamas, G. M." box="[386,568,1457,1481]" pageId="6" pageNumber="7" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas, 1973: 276</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 277, figs. 4–7;
<treatmentCitation author="Artigas, J. N. &amp; Papavero, N." box="[726,1025,1457,1481]" page="36" pageId="6" pageNumber="7" year="1995">
<bibRefCitation author="Artigas, J. N. &amp; Papavero, N." box="[726,1025,1457,1481]" pageId="6" pageNumber="7" pagination="33 - 56" refId="ref34602" refString="Artigas, J. N. &amp; Papavero, N. (1995) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX. 10. Subfamily Asilinae Leach - Lecania - Group, with a catalogue of Neotropical species. Theoria, 4, 33 - 56." type="journal article" year="1995">Artigas &amp; Papavero, 1995: 36</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation author="Vieira, R." box="[1157,1307,1457,1481]" page="2" pageId="6" pageNumber="7" year="2012">
<bibRefCitation author="Vieira, R." box="[1157,1307,1457,1481]" pageId="6" pageNumber="7" pagination="1 - 6" refId="ref35861" refString="Vieira, R. (2012) Ctenodontina Enderlein, 1914 (Diptera, Asilidae, Asilinae): First record for Brazil and description of a new species. International Scholarly Research Notices, 2012, 1 - 6. https: // doi. org / 10.5402 / 2012 / 517410" type="journal article" year="2012">Vieira, 2012: 2</bibRefCitation>
</treatmentCitation>
, fig. 1 (key);
<treatmentCitation author="Vieira, R." box="[199,376,1488,1512]" page="314" pageId="6" pageNumber="7" year="2014">
<bibRefCitation author="Vieira, R." box="[199,376,1488,1512]" pageId="6" pageNumber="7" pagination="313 - 315" refId="ref35914" refString="Vieira, R. (2014) Description of the female of Ctenodontina nairae Vieira (Diptera, Asilidae, Asilinae), with new distribution records. Revista Brasileira de Entmologia, 58 (4), 313 - 315. https: // doi. org / 10.1590 / S 0085 - 56262014005000005" type="journal article" year="2014">Vieira, 2014: 314</bibRefCitation>
</treatmentCitation>
(comments);
<treatmentCitation author="Vieira, R. &amp; Ayala-Landa, J. M. &amp; Rafael, J. A." box="[516,934,1488,1512]" page="290" pageId="6" pageNumber="7" year="2017">
<bibRefCitation author="Vieira, R. &amp; Ayala-Landa, J. M. &amp; Rafael, J. A." box="[516,934,1488,1512]" pageId="6" pageNumber="7" pagination="279 - 291" refId="ref35969" refString="Vieira, R., Ayala-Landa, J. M. &amp; Rafael, J. A. (2017) Ctenodontina Enderlein, 1914 (Diptera, Asilidae, Asilinae): new combinations, synonyms, new species and new records for Argentina. Zootaxa, 4216 (3), 279 - 291. https: // doi. org / 10.11646 / zootaxa. 4216.3.5" type="journal article" year="2017">Vieira, Ayala-Landa &amp; Rafael, 2017: 290</bibRefCitation>
</treatmentCitation>
, figs. 1, 4 (key);
<treatmentCitation ID-DOI="http://doi.org/10.5281/zenodo.5492780" ID-Zenodo-Dep="5492780" author="Sanchez, P. &amp; Camargo, A." box="[1106,1432,1488,1512]" httpUri="http://treatment.plazi.org/id/03A187A3403F2E4FFF09FC9A5266FF6A" page="270" pageId="6" pageNumber="7" year="2021">
<bibRefCitation author="Sanchez, P. &amp; Camargo, A." box="[1106,1432,1488,1512]" pageId="6" pageNumber="7" pagination="269 - 281" refId="ref35747" refString="Sanchez, P. &amp; Camargo, A. (2021) Description of the female of Ctenodontina mochica Lamas, revalidation of Ctenodontina carrerai (Hull) stat. rev., and comments on the taxonomy and distribution of Peruvian species of Ctenodontina Enderlein (Diptera: Asilidae: Asilinae). Zootaxa, 5027 (2), 269 - 281. https: // doi. org / 10.11646 / zootaxa. 5027.2.7" type="journal article" year="2021">Sánchez &amp; Camargo, 2021: 270</bibRefCitation>
</treatmentCitation>
, 278, figs. 1–3 (female description, key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="diagnosis">
<paragraph blockId="6.[151,1436,1583,2041]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[151,274,1583,1609]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
Face silvery pruinose; mystacal macrosetae white (
<figureCitation box="[849,942,1583,1609]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
,
<figureCitation box="[953,991,1583,1609]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">3A</figureCitation>
); scape and pedicel dark yellow-brown; postpedicel and stylus brownish (
<figureCitation box="[525,618,1619,1645]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
,
<figureCitation box="[630,668,1619,1645]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">3A</figureCitation>
); thorax light yellow-brown; paramedian stripes, pre and postsutural spots and prescutellar spot dark; legs yellow brown with apex of last tarsomeres dark brown almost black (
<figureCitation captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
,
<figureCitation box="[202,240,1691,1717]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">3A</figureCitation>
); hind femur with a conspicuous sack-like swelling on distal forth ventrally, bearing a number of short white spines on its apex (
<figureCitation box="[443,565,1727,1753]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1B–C</figureCitation>
); an elongated concavity on proximal sixth of ventral surface of hind tibiae, opposite the femoral swelling (
<figureCitation box="[504,626,1763,1789]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1B–C</figureCitation>
); abdomen yellow brown with silvery and golden pruinosity (
<figureCitation box="[1331,1421,1763,1789]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
); male terminalia reddish brown (
<figureCitation box="[506,626,1799,1825]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1D–E</figureCitation>
), S8 with a mid-posterior projection bilobed apically (
<figureCitation box="[1226,1313,1799,1825]" captionStart="FIGURE 2" captionStartId="8.[151,250,1874,1898]" captionTargetBox="[227,1350,189,1848]" captionTargetId="figure-21@8.[227,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 2. Ctenodontina mochica Lamas, 1973, male. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Sternite 8, ventral; E. Hypandrium, ventral; F. Gonopods, external, lateral; G. Same, internal, lateral; H. Phallus, lateral. I. Cercus and subepandrial sclerite, dorsal; J. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907302" httpUri="https://zenodo.org/record/7907302/files/figure.png" pageId="6" pageNumber="7">Fig. 2D</figureCitation>
) and white setose apically (
<figureCitation box="[335,462,1835,1861]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Figs 1D–E</figureCitation>
,
<figureCitation box="[476,514,1835,1861]" captionStart="FIGURE 2" captionStartId="8.[151,250,1874,1898]" captionTargetBox="[227,1350,189,1848]" captionTargetId="figure-21@8.[227,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 2. Ctenodontina mochica Lamas, 1973, male. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Sternite 8, ventral; E. Hypandrium, ventral; F. Gonopods, external, lateral; G. Same, internal, lateral; H. Phallus, lateral. I. Cercus and subepandrial sclerite, dorsal; J. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907302" httpUri="https://zenodo.org/record/7907302/files/figure.png" pageId="6" pageNumber="7">2A</figureCitation>
); epandrium with rounded apex, in lateral view (
<figureCitation box="[1080,1170,1835,1861]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1D</figureCitation>
), with black and white setae (
<figureCitation box="[223,345,1871,1897]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="6" pageNumber="7">Fig. 1D–E</figureCitation>
); gonocoxite subtriangular and rounded apically in lateral view (
<figureCitation box="[1072,1192,1871,1897]" captionStart="FIGURE 2" captionStartId="8.[151,250,1874,1898]" captionTargetBox="[227,1350,189,1848]" captionTargetId="figure-21@8.[227,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 2. Ctenodontina mochica Lamas, 1973, male. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Sternite 8, ventral; E. Hypandrium, ventral; F. Gonopods, external, lateral; G. Same, internal, lateral; H. Phallus, lateral. I. Cercus and subepandrial sclerite, dorsal; J. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907302" httpUri="https://zenodo.org/record/7907302/files/figure.png" pageId="6" pageNumber="7">Fig. 2F–G</figureCitation>
); hypandrium convex anteriorly and with a triangular indentation at posterior margin (
<figureCitation box="[885,974,1907,1933]" captionStart="FIGURE 2" captionStartId="8.[151,250,1874,1898]" captionTargetBox="[227,1350,189,1848]" captionTargetId="figure-21@8.[227,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 2. Ctenodontina mochica Lamas, 1973, male. A. Terminalia, lateral; B. Same, dorsal; C. Same, ventral; D. Sternite 8, ventral; E. Hypandrium, ventral; F. Gonopods, external, lateral; G. Same, internal, lateral; H. Phallus, lateral. I. Cercus and subepandrial sclerite, dorsal; J. Same, ventral. Abbreviations: cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907302" httpUri="https://zenodo.org/record/7907302/files/figure.png" pageId="6" pageNumber="7">Fig. 2E</figureCitation>
); female terminalia compressed laterally and shining reddish brown (
<figureCitation box="[461,581,1943,1969]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">Fig. 3B–C</figureCitation>
); T8 as long as T6 and T7 combined (
<figureCitation box="[997,1119,1943,1969]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">Fig. 3B–D</figureCitation>
); spermatheca with two oval capsules (
<figureCitation box="[260,346,1979,2005]" captionStart="FIGURE 3" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1908]" captionTargetId="figure-17@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 3. Ctenodontina mochica Lamas, 1973, female. A. Habitus, lateral; B. Terminalia, dorsal; C. Same, lateral; D. Same, lateral, dissected; E. Spermathecal capsule. Abbreviations: cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
). This diagnosis is based on the original description of
<bibRefCitation author="Lamas, G. M." box="[954,1109,1979,2006]" pageId="6" pageNumber="7" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas (1973)</bibRefCitation>
and an additional description of the male terminalia and female by
<bibRefCitation author="Sanchez, P. &amp; Camargo, A." box="[570,886,2015,2041]" pageId="6" pageNumber="7" pagination="269 - 281" refId="ref35747" refString="Sanchez, P. &amp; Camargo, A. (2021) Description of the female of Ctenodontina mochica Lamas, revalidation of Ctenodontina carrerai (Hull) stat. rev., and comments on the taxonomy and distribution of Peruvian species of Ctenodontina Enderlein (Diptera: Asilidae: Asilinae). Zootaxa, 5027 (2), 269 - 281. https: // doi. org / 10.11646 / zootaxa. 5027.2.7" type="journal article" year="2021">Sánchez &amp; Camargo (2021)</bibRefCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7907300" ID-Zenodo-Dep="7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="7" pageNumber="8" startId="7.[152,255,1943,1967]" targetBox="[203,1385,181,1919]" targetPageId="7" targetType="figure">
<paragraph blockId="7.[151,1437,1943,2040]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[152,280,1943,1967]" pageId="7" pageNumber="8">FIGURE 1.</emphasis>
<taxonomicName authority="Lamas, 1973" authorityName="Lamas" authorityYear="1973" box="[286,654,1944,1968]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mochica">
<emphasis box="[286,515,1944,1968]" italics="true" pageId="7" pageNumber="8">Ctenodontina mochica</emphasis>
<bibRefCitation author="Lamas, G. M." box="[521,654,1944,1968]" pageId="7" pageNumber="8" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas, 1973</bibRefCitation>
</taxonomicName>
, male.
<emphasis bold="true" box="[725,749,1943,1967]" pageId="7" pageNumber="8">A.</emphasis>
Habitus, lateral;
<emphasis bold="true" box="[922,945,1944,1968]" pageId="7" pageNumber="8">B.</emphasis>
Hind femora, anterior;
<emphasis bold="true" box="[1183,1208,1943,1967]" pageId="7" pageNumber="8">C.</emphasis>
Same, detail, anterior;
<emphasis bold="true" box="[151,176,1980,2004]" pageId="7" pageNumber="8">D.</emphasis>
Terminalia, lateral;
<emphasis bold="true" box="[380,404,1980,2004]" pageId="7" pageNumber="8">E.</emphasis>
Same, dorsal.
<emphasis bold="true" box="[553,714,1979,2003]" pageId="7" pageNumber="8">Abbreviations:</emphasis>
cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7907302" ID-Zenodo-Dep="7907302" httpUri="https://zenodo.org/record/7907302/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1874,1898]" targetBox="[227,1350,189,1848]" targetPageId="8" targetType="figure">
<paragraph blockId="8.[151,1437,1874,2043]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[151,276,1874,1898]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
<taxonomicName authority="Lamas, 1973" authorityName="Lamas" authorityYear="1973" box="[283,656,1875,1899]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mochica">
<emphasis box="[283,514,1875,1899]" italics="true" pageId="8" pageNumber="9">Ctenodontina mochica</emphasis>
<bibRefCitation author="Lamas, G. M." box="[522,656,1875,1899]" pageId="8" pageNumber="9" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas, 1973</bibRefCitation>
</taxonomicName>
, male.
<emphasis bold="true" box="[730,755,1874,1898]" pageId="8" pageNumber="9">A.</emphasis>
Terminalia, lateral;
<emphasis bold="true" box="[962,986,1875,1899]" pageId="8" pageNumber="9">B.</emphasis>
Same, dorsal;
<emphasis bold="true" box="[1139,1164,1874,1898]" pageId="8" pageNumber="9">C.</emphasis>
Same, ventral;
<emphasis bold="true" box="[1326,1350,1875,1899]" pageId="8" pageNumber="9">D.</emphasis>
Sternite 8, ventral;
<emphasis bold="true" box="[261,285,1911,1935]" pageId="8" pageNumber="9">E.</emphasis>
Hypandrium, ventral;
<emphasis bold="true" box="[517,536,1911,1935]" pageId="8" pageNumber="9">F.</emphasis>
Gonopods, external, lateral;
<emphasis bold="true" box="[832,858,1910,1934]" pageId="8" pageNumber="9">G.</emphasis>
Same, internal, lateral;
<emphasis bold="true" box="[1102,1129,1911,1935]" pageId="8" pageNumber="9">H.</emphasis>
Phallus, lateral.
<emphasis bold="true" box="[1299,1315,1911,1935]" pageId="8" pageNumber="9">I.</emphasis>
Cercus and subepandrial sclerite, dorsal;
<emphasis bold="true" box="[449,468,1947,1971]" pageId="8" pageNumber="9">J.</emphasis>
Same, ventral.
<emphasis bold="true" box="[627,788,1946,1970]" pageId="8" pageNumber="9">Abbreviations:</emphasis>
cerc: cercus; ej apod: ejaculatory apodeme; goncx: gonocoxite; goncx apod: gonocoxal apodeme; gonst: gonostylus; hypd: hypandrium; ph: phallus; st: sternite; subepand scl: subepandrial sclerite; tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7907306" ID-Zenodo-Dep="7907306" httpUri="https://zenodo.org/record/7907306/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1943,1967]" targetBox="[203,1385,181,1908]" targetPageId="9" targetType="figure">
<paragraph blockId="9.[151,1437,1943,2040]" pageId="9" pageNumber="10">
<emphasis bold="true" box="[151,275,1943,1967]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
<taxonomicName authority="Lamas, 1973" authorityName="Lamas" authorityYear="1973" box="[281,650,1944,1968]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mochica">
<emphasis box="[281,511,1944,1968]" italics="true" pageId="9" pageNumber="10">Ctenodontina mochica</emphasis>
<bibRefCitation author="Lamas, G. M." box="[517,650,1944,1968]" pageId="9" pageNumber="10" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas, 1973</bibRefCitation>
</taxonomicName>
, female.
<emphasis bold="true" box="[742,766,1943,1967]" pageId="9" pageNumber="10">A.</emphasis>
Habitus, lateral;
<emphasis bold="true" box="[940,963,1944,1968]" pageId="9" pageNumber="10">B.</emphasis>
Terminalia, dorsal;
<emphasis bold="true" box="[1167,1191,1943,1967]" pageId="9" pageNumber="10">C.</emphasis>
Same, lateral;
<emphasis bold="true" box="[1343,1368,1944,1968]" pageId="9" pageNumber="10">D.</emphasis>
Same, lateral, dissected;
<emphasis bold="true" box="[332,355,1980,2004]" pageId="9" pageNumber="10">E.</emphasis>
Spermathecal capsule.
<emphasis bold="true" box="[591,752,1979,2003]" pageId="9" pageNumber="10">Abbreviations:</emphasis>
cerc: cercus; gen fk: genital fork; spe du: spermathecal duct; spmth: spermathecae; st: sternite; tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph blockId="10.[151,1437,150,465]" pageId="10" pageNumber="11">
<emphasis bold="true" box="[199,469,150,176]" pageId="10" pageNumber="11">Taxonomic discussion.</emphasis>
<bibRefCitation author="Lamas, G. M." box="[479,638,151,177]" pageId="10" pageNumber="11" pagination="275 - 280" refId="ref35408" refString="Lamas, G. M. (1973) Some comments on the genus Ctenodontina Enderlein (Diptera, Asilidae), with the description of a new species from coastal Peru. Papeis Avulsos de Zoologia, 26 (23), 275 - 280." type="journal article" year="1973">Lamas (1973)</bibRefCitation>
described this species based on a single male specimen that seemed to him to be a newly hatched imago and that the colour-pattern could be somewhat darker in an older specimen. Recently
<bibRefCitation author="Sanchez, P. &amp; Camargo, A." box="[257,566,223,249]" pageId="10" pageNumber="11" pagination="269 - 281" refId="ref35747" refString="Sanchez, P. &amp; Camargo, A. (2021) Description of the female of Ctenodontina mochica Lamas, revalidation of Ctenodontina carrerai (Hull) stat. rev., and comments on the taxonomy and distribution of Peruvian species of Ctenodontina Enderlein (Diptera: Asilidae: Asilinae). Zootaxa, 5027 (2), 269 - 281. https: // doi. org / 10.11646 / zootaxa. 5027.2.7" type="journal article" year="2021">Sánchez &amp; Camargo (2021)</bibRefCitation>
described the female and provided an addendum to the description of the male terminalia of this species.
</paragraph>
<paragraph blockId="10.[151,1437,150,465]" pageId="10" pageNumber="11">
This species has a very conspicuous sack-like swelling on the distal fourth ventrally of the hind femora of males and the hind tibia is curved on the anterior third with a ventral hollow which seems to accommodate the femoral projection while in resting position (
<figureCitation box="[557,681,367,393]" captionStart="FIGURE 1" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 1. Ctenodontina mochica Lamas, 1973, male. A. Habitus, lateral; B. Hind femora, anterior; C. Same, detail, anterior; D. Terminalia, lateral; E. Same, dorsal. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907300" httpUri="https://zenodo.org/record/7907300/files/figure.png" pageId="10" pageNumber="11">Fig. 1A–C</figureCitation>
). In addition, the S8 has a mid-posterior bilobed projection (Fig. D) which helps to separate it from
<taxonomicName authorityName="Enderlein" authorityYear="1914" box="[504,799,403,429]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pectinatipes">
<emphasis box="[504,799,403,429]" italics="true" pageId="10" pageNumber="11">Ctenodontina pectinatipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,885,438,465]" pageId="10" pageNumber="11" type="distribution">
<paragraph blockId="10.[151,1437,150,465]" box="[199,885,438,465]" pageId="10" pageNumber="11">
<emphasis bold="true" box="[199,445,438,464]" pageId="10" pageNumber="11">
Distribution (
<figureCitation box="[360,430,438,464]" captionStart="FIGURE 8" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[151,1435,813,1946]" captionTargetId="figure-286@15.[151,1435,813,1946]" captionTargetPageId="15" captionText="FIGURE 8. Distribution map of Ctenodontina Enderlein, 1914 with the distribution records of its two species: Ctenodontina mochica Lamas, 1973 and Ctenodontina pectinatipes Enderlein, 1914." figureDoi="http://doi.org/10.5281/zenodo.7907328" httpUri="https://zenodo.org/record/7907328/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
).
</emphasis>
<collectingCountry box="[452,503,439,465]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
(
<collectingRegion box="[518,612,439,465]" country="Peru" name="Tumbes" pageId="10" pageNumber="11">Tumbes</collectingRegion>
,
<collectingRegion box="[623,682,439,465]" country="Peru" name="Piura" pageId="10" pageNumber="11">Piura</collectingRegion>
and
<collectingRegion box="[736,873,439,465]" country="Peru" name="La Libertad" pageId="10" pageNumber="11">La Libertad</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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