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<document id="44DCC4258BD0D29A98F99BBE36240105" ID-CLB-Dataset="8690" ID-DOI="10.11646/zootaxa.4827.1.1" ID-GBIF-Dataset="c93b6c76-aaac-4fcd-acca-3dc3e0fd62d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402308" ID-ZooBank="2CE528C2-DE07-4155-B37D-BE2C342A3901" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609343086441" checkinUser="plazi" docAuthor="Raines, Bret K." docDate="2020" docId="03A3DF49FF9DFF03FF44F9F6FA8F395A" docLanguage="en" docName="zootaxa.4827.1.1.pdf" docOrigin="Zootaxa 4827 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Caecum draperi Raines 2020, species novum" docType="treatment" docVersion="12" lastPageNumber="131" masterDocId="FF9AA731FFE2FF81FFD3FFD7FF8D381B" masterDocTitle="A Rosetta Stone for eastern Pacific Caecidae (Gastropoda: Caenogastropoda)" masterLastPageNumber="146" masterPageNumber="1" pageNumber="128" updateTime="1698849882115" updateUser="ExternalLinkService">
<mods:mods id="953EC98CE0628E4BC6F41463CE03E058" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="604944C985A5E16A595A2660969FF227">
<mods:title id="4BF530EE094ABE2C5949CEE9EE9B30DC">A Rosetta Stone for eastern Pacific Caecidae (Gastropoda: Caenogastropoda)</mods:title>
</mods:titleInfo>
<mods:name id="784358BECB388C439EDE48936E3DD5D3" type="personal">
<mods:role id="56FB79189E528DB3B3C92CB3B542D892">
<mods:roleTerm id="F6ED3F7C246CD297E72B0857287CD79A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D06B23CF1E9DA3A27A90875D726DC47">Raines, Bret K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C4BEFE09F67AB80F2BC9733905326773">text</mods:typeOfResource>
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<mods:titleInfo id="036C38E17D49A08BB60A3682205FCEF3">
<mods:title id="AFA9EAAC98F5C3DE08504F4DCD67733A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AE9C93834397E460FFE7F3585214BF07">
<mods:date id="E20AB2BB347297A1142A54E9456F6E7B">2020</mods:date>
<mods:detail id="F739469B36F835DBC6D30FB71369FF2C" type="pubDate">
<mods:number id="B6FFA5E6ADC696F87671AFC5C53D8E7A">2020-08-11</mods:number>
</mods:detail>
<mods:detail id="6E88DDD5F49FB10EB581055DD591A739" type="volume">
<mods:number id="EB845890E23A910E43A8086E44E7F29D">4827</mods:number>
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<mods:number id="A868C9E0D76DEDEA23B5893D2707D02C">1</mods:number>
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<mods:classification id="B804F9A84D47372493665B0317E6C08B">journal article</mods:classification>
<mods:identifier id="4EB5267068F73033066BD69F7B18BC62" type="CLB-Dataset">8690</mods:identifier>
<mods:identifier id="B63FB2863B763328C43A6B866161CD21" type="DOI">10.11646/zootaxa.4827.1.1</mods:identifier>
<mods:identifier id="199956B4971636F03DE0B9BCAC1C66FB" type="GBIF-Dataset">c93b6c76-aaac-4fcd-acca-3dc3e0fd62d4</mods:identifier>
<mods:identifier id="A245C121C49F3BB99297AA3666EEEB99" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="342F44E279411A1D6EA3A2E2DBE94AC8" type="Zenodo-Dep">4402308</mods:identifier>
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</mods:mods>
<treatment id="03A3DF49FF9DFF03FF44F9F6FA8F395A" ID-DOI="http://doi.org/10.5281/zenodo.4457839" ID-GBIF-Taxon="177057001" ID-Zenodo-Dep="4457839" LSID="urn:lsid:plazi:treatment:03A3DF49FF9DFF03FF44F9F6FA8F395A" httpUri="http://treatment.plazi.org/id/03A3DF49FF9DFF03FF44F9F6FA8F395A" lastPageId="130" lastPageNumber="131" pageId="127" pageNumber="128">
<subSubSection id="C3103DD4FF9DFFFEFF44F9F6FD983E20" box="[151,533,1568,1595]" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="8BB56E5FFF9DFFFEFF44F9F6FD983E20" blockId="127.[151,619,1568,1631]" box="[151,533,1568,1595]" pageId="127" pageNumber="128">
<heading id="D0FDD933FF9DFFFEFF44F9F6FD983E20" bold="true" box="[151,533,1568,1595]" fontSize="11" level="2" pageId="127" pageNumber="128" reason="4">
<emphasis id="B97EB24DFF9DFFFEFF44F9F6FD983E20" bold="true" box="[151,533,1568,1595]" pageId="127" pageNumber="128">
<taxonomicName id="4C0A15DCFF9DFFFEFF44F9F6FED73E20" ID-CoL="7QJ29" authority="Raines, 2020" authorityName="Raines" authorityYear="2020" box="[151,346,1568,1595]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="draperi" status="sp. nov.">
<emphasis id="B97EB24DFF9DFFFEFF44F9F6FED73E20" bold="true" box="[151,346,1568,1595]" italics="true" pageId="127" pageNumber="128">Caecum draperi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D0F36FF9DFFFEFEB2F9F6FD983E20" box="[353,533,1569,1595]" pageId="127" pageNumber="128" rank="species">species novum</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF9DFFFEFF44F992FDE63E44" box="[151,619,1605,1631]" pageId="127" pageNumber="128" type="description">
<paragraph id="8BB56E5FFF9DFFFEFF44F992FDE63E44" blockId="127.[151,619,1568,1631]" box="[151,619,1605,1631]" pageId="127" pageNumber="128">
<figureCitation id="133172DAFF9DFFFEFF44F992FE943E44" box="[151,281,1605,1631]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="127" pageNumber="128">Figures 111</figureCitation>
A–G, 112A–J, 113A–B, 126B
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF9DFFFEFF44F95EFD2B3EB9" box="[151,678,1673,1698]" pageId="127" pageNumber="128" type="reference_group">
<paragraph id="8BB56E5FFF9DFFFEFF44F95EFD2B3EB9" blockId="127.[151,678,1673,1698]" box="[151,678,1673,1698]" pageId="127" pageNumber="128">
<treatmentCitationGroup id="AB1A4971FF9DFFFEFF44F95EFD2B3EB9" box="[151,678,1673,1698]" pageId="127" pageNumber="128">
<taxonomicName id="4C0A15DCFF9DFFFEFF44F95EFEE83EBA" authority="(Bartsch, 1920)" box="[151,357,1673,1697]" class="Gastropoda" family="Caecidae" genus="Brochina" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="127" pageNumber="128" phylum="Mollusca" rank="species" species="carpenteri">
<emphasis id="B97EB24DFF9DFFFEFF44F95EFEE83EBA" box="[151,357,1673,1697]" italics="true" pageId="127" pageNumber="128">Brochina carpenteri</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AAB484EFF9DFFFEFEADF95EFDDE3EB9" author="Lightfoot, J." box="[382,595,1673,1698]" page="82" pageId="127" pageNumber="128" year="1993">
<bibRefCitation id="EF9B13AEFF9DFFFEFEADF95EFDDE3EB9" author="Lightfoot, J." box="[382,595,1673,1698]" pageId="127" pageNumber="128" pagination="75 - 87" refId="ref98203" refString="Lightfoot, J. (1993 b) Caecidae of the Panamic Province. Part II. Of Sea &amp; Shore, 16, 75 - 87." type="journal article" year="1993">Lightfoot, 1993b: 82</bibRefCitation>
</treatmentCitation>
, fig. 30.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF9DFF01FF44F907FC2038AA" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="8BB56E5FFF9DFFFEFF44F907FDBE3F14" blockId="127.[151,1437,1743,2022]" pageId="127" pageNumber="128">
<emphasis id="B97EB24DFF9DFFFEFF44F907FEC93EF2" bold="true" box="[151,324,1743,1770]" pageId="127" pageNumber="128">Type material.</emphasis>
<materialsCitation id="3B626402FF9DFFFEFE99F907FDBE3F14" ID-GBIF-Occurrence="3023454645" collectionCode="SBMNH, LACM" pageId="127" pageNumber="128" specimenCode="SBMNH 637748, SBMNH 637747, SBMNH 637749, LACM 3756" specimenCount="2" typeStatus="holotype">
<typeStatus id="54B1D0FDFF9DFFFEFE99F907FE383EF1" box="[330,437,1744,1770]" pageId="127" pageNumber="128" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBACC624FF9DFFFEFE13F907FD0F3EF1" box="[448,642,1744,1770]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH 637748</specimenCode>
;
<typeStatus id="54B1D0FDFF9DFFFEFD5FF907FD773EF1" box="[652,762,1744,1770]" pageId="127" pageNumber="128" type="paratype">paratypes</typeStatus>
,
<specimenCode id="DBACC624FF9DFFFEFCD7F907FC4F3EF1" box="[772,962,1744,1770]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH 637747</specimenCode>
(4 figured);
<specimenCode id="DBACC624FF9DFFFEFB9DF907FA813EF1" box="[1102,1292,1744,1770]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH 637749</specimenCode>
(3 figured, 1 unfigured);
<specimenCode id="DBACC624FF9DFFFEFECFF923FE213F15" box="[284,428,1779,1806]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="127" pageNumber="128" type="Museum">LACM 3756</specimenCode>
(2 figured).
</materialsCitation>
</paragraph>
<paragraph id="8BB56E5FFF9DFFFEFF14F8C0FD993FFD" blockId="127.[151,1437,1743,2022]" pageId="127" pageNumber="128">
<emphasis id="B97EB24DFF9DFFFEFF14F8C0FD483F2A" bold="true" box="[199,709,1815,1841]" pageId="127" pageNumber="128">Additional material examined. NHMLAC:</emphasis>
<collectionCode id="ED1BF69AFF9DFFFEFD1FF8CFFC903F29" box="[716,797,1816,1842]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="127" pageNumber="128" type="Museum">LACM</collectionCode>
1977-144.15, Chame Point,
<collectingCountry id="F31D2ECFFF9DFFFEFBB2F8CFFB373F29" box="[1121,1210,1816,1842]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Bay,
<collectingCountry id="F31D2ECFFF9DFFFEFB28F8CFFAD93F29" box="[1275,1364,1816,1842]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFF44F8EBFF7E3F4D" box="[151,243,1852,1878]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 6 sh.
<emphasis id="B97EB24DFF9DFFFEFE97F8ECFE393F4E" bold="true" box="[324,436,1851,1877]" pageId="127" pageNumber="128">
<collectionCode id="ED1BF69AFF9DFFFEFE97F8ECFE223F4E" box="[324,431,1851,1877]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
:
</emphasis>
<collectionCode id="ED1BF69AFF9DFFFEFE6DF8EBFDAF3F4D" box="[446,546,1852,1878]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
452501, Punta Chame,
<collectingCountry id="F31D2ECFFF9DFFFEFCEEF8EBFC1B3F4D" box="[829,918,1852,1878]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Bay,
<collectingCountry id="F31D2ECFFF9DFFFEFC0EF8EBFBBB3F4D" box="[989,1078,1852,1878]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFB58F8EBFB6A3F4D" box="[1163,1255,1852,1878]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 6 sh;
<collectionCode id="ED1BF69AFF9DFFFEFAEAF8EBFA103F4D" box="[1337,1437,1852,1878]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
452539, Pier 18, Balboa,
<collectingCountry id="F31D2ECFFF9DFFFEFE60F8B7FD813F61" box="[435,524,1888,1914]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFD89F8B7FD3B3F61" box="[602,694,1888,1914]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 2 sh;
<collectionCode id="ED1BF69AFF9DFFFEFD2FF8B7FCED3F61" box="[764,864,1888,1914]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
452440, Pier 18, Balboa,
<collectingCountry id="F31D2ECFFF9DFFFEFB52F8B7FB573F61" box="[1153,1242,1888,1914]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFAFBF8B7FA093F61" box="[1320,1412,1888,1914]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 2 sh;
<collectionCode id="ED1BF69AFF9DFFFEFF6DF853FEAF3F85" box="[190,290,1924,1950]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
452441, Pier 18, Balboa,
<collectingCountry id="F31D2ECFFF9DFFFEFD90F853FD113F85" box="[579,668,1924,1950]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFD39F853FCCB3F85" box="[746,838,1924,1950]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 3 sh;
<collectionCode id="ED1BF69AFF9DFFFEFC58F853FC623F85" box="[907,1007,1924,1950]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
620531, Playa Venado,
<collectingCountry id="F31D2ECFFF9DFFFEFB2FF853FAD83F85" box="[1276,1365,1924,1950]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFF44F87FFF7E3FD9" box="[151,243,1960,1986]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 20 sh;
<collectionCode id="ED1BF69AFF9DFFFEFE95F87FFE273FD9" box="[326,426,1960,1986]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
637751, Fuerte Amador,
<collectingCountry id="F31D2ECFFF9DFFFEFD17F87FFCAD3FD9" box="[708,800,1960,1986]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, intertidal, 1 sh;
<collectionCode id="ED1BF69AFF9DFFFEFC04F87FFBB63FD9" box="[983,1083,1960,1986]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="127" pageNumber="128" type="Museum">SBMNH</collectionCode>
637746, Punta Chame,
<collectingCountry id="F31D2ECFFF9DFFFEFA97F87FFA103FD9" box="[1348,1437,1960,1986]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Bay,
<collectingCountry id="F31D2ECFFF9DFFFEFF03F81BFEA43FFD" box="[208,297,1996,2022]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF9DFFFEFEABF81BFE593FFD" box="[376,468,1996,2022]" name="Panama" pageId="127" pageNumber="128">Panama</collectingCountry>
, 3 sh.
</paragraph>
<paragraph id="8BB56E5FFF62FF01FF14FF40FC2038AA" blockId="128.[151,1437,150,861]" box="[199,941,150,177]" pageId="128" pageNumber="129">
<materialsCitation id="3B626402FF62FF01FF14FF40FDE838AA" ID-GBIF-Occurrence="3023454827" box="[199,613,150,177]" country="Panama" location="Chame Point" pageId="128" pageNumber="129" specimenCount="1" typeStatus="holotype">
<emphasis id="B97EB24DFF62FF01FF14FF40FEEB38AB" bold="true" box="[199,358,150,177]" pageId="128" pageNumber="129">
<typeStatus id="54B1D0FDFF62FF01FF14FF40FE8F38AA" box="[199,258,151,177]" pageId="128" pageNumber="129">Type</typeStatus>
locality.
</emphasis>
<location id="8ED53884FF62FF01FEBFFF40FD8C38AA" LSID="urn:lsid:plazi:treatment:03A3DF49FF9DFF03FF44F9F6FA8F395A:8ED53884FF62FF01FEBFFF40FD8C38AA" box="[364,513,151,177]" country="Panama" name="Chame Point" pageId="128" pageNumber="129">Chame Point</location>
,
<collectingCountry id="F31D2ECFFF62FF01FDDFFF40FDE838AA" box="[524,613,151,177]" name="Panama" pageId="128" pageNumber="129">Panama</collectingCountry>
</materialsCitation>
Bay,
<collectingCountry id="F31D2ECFFF62FF01FD76FF40FD7338AA" box="[677,766,151,177]" name="Panama" pageId="128" pageNumber="129">Panama</collectingCountry>
Prov.,
<collectingCountry id="F31D2ECFFF62FF01FC9FFF40FC2538AA" box="[844,936,151,177]" name="Panama" pageId="128" pageNumber="129">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF62FF01FF14FF6DFE6D395A" pageId="128" pageNumber="129" type="diagnosis">
<paragraph id="8BB56E5FFF62FF01FF14FF6DFE6D395A" blockId="128.[151,1437,150,861]" pageId="128" pageNumber="129">
<emphasis id="B97EB24DFF62FF01FF14FF6DFECF38CF" bold="true" box="[199,322,186,212]" pageId="128" pageNumber="129">Diagnosis.</emphasis>
Teleoconch subcylindrical, with slight increase in diameter from posterior end to aperture. Surface smooth with 12–16 large, irregularly spaced, axial ribs, which increase in strength aperturally, semi-translucent to creamy white in color. Septum mucronate, moderately inflated, with low blunt mucro. Aperture simple, rather thin. Periostracum thin, yellowish.
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF62FF01FF14FE9DFCE73B46" pageId="128" pageNumber="129" type="description">
<paragraph id="8BB56E5FFF62FF01FF14FE9DFCE73B46" blockId="128.[151,1437,150,861]" pageId="128" pageNumber="129">
<emphasis id="B97EB24DFF62FF01FF14FE9DFED4397F" bold="true" box="[199,345,330,356]" pageId="128" pageNumber="129">Description.</emphasis>
Protoconch and teleoconch I unknown. Teleoconch II surface smooth (
<figureCitation id="133172DAFF62FF01FB5BFE9CFB73397E" box="[1160,1278,331,357]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112D</figureCitation>
), transition to teleoconch III marked by slight incised axial line (
<figureCitation id="133172DAFF62FF01FD19FEB8FCCD3992" box="[714,832,367,393]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112A</figureCitation>
). Teleoconch III smooth with irregularly spaced, axial ribs near apertural end (
<figureCitation id="133172DAFF62FF01FE63FE44FDBF39B6" box="[432,562,403,429]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Figs. 112B</figureCitation>
), transition to teleoconch IV marked by suture (
<figureCitation id="133172DAFF62FF01FBB6FE44FB5239B6" box="[1125,1247,403,429]" captionStart="FIGURE 113" captionStartId="130.[151,250,1175,1199]" captionTargetBox="[151,1435,345,1151]" captionTargetId="figure-111@130.[151,1435,345,1151]" captionTargetPageId="130" captionText="FIGURE 113. Caecum draperi sp. nov. A. Conceptual reconstruction of growth stages. B. Species distribution map." figureDoi="http://doi.org/10.5281/zenodo.4402601" httpUri="https://zenodo.org/record/4402601/files/figure.png" pageId="128" pageNumber="129">Fig. 113A</figureCitation>
). Teleoconch IV with identical characters as previous stage, transition to teleoconch V marked by suture (
<figureCitation id="133172DAFF62FF01FBAAFE60FB6239CA" box="[1145,1263,439,465]" captionStart="FIGURE 113" captionStartId="130.[151,250,1175,1199]" captionTargetBox="[151,1435,345,1151]" captionTargetId="figure-111@130.[151,1435,345,1151]" captionTargetPageId="130" captionText="FIGURE 113. Caecum draperi sp. nov. A. Conceptual reconstruction of growth stages. B. Species distribution map." figureDoi="http://doi.org/10.5281/zenodo.4402601" httpUri="https://zenodo.org/record/4402601/files/figure.png" pageId="128" pageNumber="129">Fig. 113A</figureCitation>
). Teleoconch V smooth with large, irregularly spaced axial ribs (
<figureCitation id="133172DAFF62FF01FD6BFE0CFCAF39EE" box="[696,802,475,501]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Figs. 112</figureCitation>
G–H), transition to teleoconch VI marked by suture, followed by increase in diameter (
<figureCitation id="133172DAFF62FF01FE28FE28FDEA3A02" box="[507,615,511,537]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112I</figureCitation>
). Teleoconch VI (last growth stage) large for genus [Tol
<quantity id="4CF2C3BAFF62FF01FB3BFE29FA1C3A02" box="[1256,1425,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="5.04" metricValueMax="5.18" metricValueMin="4.9" pageId="128" pageNumber="129" unit="mm" value="5.04" valueMax="5.18" valueMin="4.9">4.90–5.18 mm</quantity>
], tubular, rather thin, mildly and regularly arched [Larc: 1.81–2.00 mm; Arc:
<quantity id="4CF2C3BAFF62FF01FC2DFDF5FB253A26" box="[1022,1192,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="4.65" metricValueMax="5.3" metricValueMin="4.0" pageId="128" pageNumber="129" unit="mm" value="0.465" valueMax="0.53" valueMin="0.4">0.40–0.53 mm</quantity>
], subcylindrical, with slight increase in diameter from posterior to aperture, semi-translucent to creamy white (
<figureCitation id="133172DAFF62FF01FB54FD90FA8B3A7A" box="[1159,1286,583,609]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Figs. 111A</figureCitation>
,
<figureCitation id="133172DAFF62FF01FAC0FD90FAC63A7A" box="[1299,1355,583,609]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">112J</figureCitation>
). Axial sculpture of 12–16 large, rather irregularly spaced axial ribs, increasing in strength aperturally (
<figureCitation id="133172DAFF62FF01FB14FDBCFAB93A9E" box="[1223,1332,619,645]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112J</figureCitation>
). Surface smooth with thin axial lines overall (
<figureCitation id="133172DAFF62FF01FDECFD58FD3A3AB2" box="[575,695,655,681]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112G</figureCitation>
). Posterior [Dpe:
<quantity id="4CF2C3BAFF62FF01FC50FD58FBA63AB2" box="[899,1067,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.2" metricValueMin="8.0" pageId="128" pageNumber="129" unit="mm" value="0.81" valueMax="0.82" valueMin="0.8">0.80–0.82 mm</quantity>
] with rather thick edge, squared shoulder (
<figureCitation id="133172DAFF62FF01FED5FD64FEFB3AD6" box="[262,374,691,717]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111E</figureCitation>
). Septum mucronate, moderately inflated (
<figureCitation id="133172DAFF62FF01FC9FFD64FC353AD6" box="[844,952,691,717]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112J</figureCitation>
). Mucro conical, low, blunt, positioned near dorsal margin (
<figureCitation id="133172DAFF62FF01FEECFD00FE3E3AEA" box="[319,435,727,753]" captionStart="FIGURE 112" captionStartId="129.[151,250,1798,1822]" captionTargetBox="[151,1435,181,1773]" captionTargetId="figure-15@129.[151,1435,181,1773]" captionTargetPageId="129" captionText="FIGURE 112. Caecum draperi sp. nov. shells at different growth stages. A. Teleoconchs II and III (SBMNH 637747). B. Teleoconch III (SBMNH 637747). C. Teleoconch IV (SBMNH 637747). D. Surface of teleoconchs II to IV (SBMNH 637747). E. Axial ribs of teleoconch IV (SBMNH 637747). F. Surface of teleoconchs V to VI (SBMNH 637747). G. Axial ribs of teleoconchs V to VI (SBMNH 637748). H. Teleoconch V (LACM 3756). I. Teleoconchs V to VI (SBMNH 637747). J. Teleoconch VI (SBMNH 637748)." figureDoi="http://doi.org/10.5281/zenodo.4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="128" pageNumber="129">Fig. 112C</figureCitation>
). Aperture [Da:
<quantity id="4CF2C3BAFF62FF01FDB6FD01FC873AEA" box="[613,778,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="128" pageNumber="129" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.90–0.95 mm</quantity>
] simple, rather thin (
<figureCitation id="133172DAFF62FF01FC3CFD00FBED3AEA" box="[1007,1120,727,753]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111E</figureCitation>
). No varix. Lip smooth (
<figureCitation id="133172DAFF62FF01FABCFD00FF593B0E" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111E</figureCitation>
). Periostracum thin, smooth, yellowish (
<figureCitation id="133172DAFF62FF01FD70FD2CFC963B0E" box="[675,795,763,789]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111A</figureCitation>
). Operculum thick, multi-layered, tapering, light brown, interior surface with central subcylindrical cusp (
<figureCitation id="133172DAFF62FF01FD6AFCC8FCA43B22" box="[697,809,799,825]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111F</figureCitation>
); exterior surface with 6–7 concentric rings (
<figureCitation id="133172DAFF62FF01FACFFCC8FA1D3B22" box="[1308,1424,799,825]" captionStart="FIGURE 111" captionStartId="128.[151,250,1716,1740]" captionTargetBox="[151,1435,885,1693]" captionTargetId="figure-322@128.[151,1435,885,1693]" captionTargetPageId="128" captionText="FIGURE 111. Caecum draperi sp. nov. A. Holotype, profile view without periostracum (SBMNH 637748). B. Paratype, profile view with periostracum (LACM 3756). C. Paratype, profile view (SBMNH 637747). D. Paratype, profile view (SBMNH 637747). E. Paratype, ventral view (LACM 3756). F. Operculum, profile view (SBMNH 637747). G. Operculum, exterior surface (SBMNH 637747)." figureDoi="http://doi.org/10.5281/zenodo.4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129">Fig. 111G</figureCitation>
). Conceptual reconstruction of growth stages shown in
<figureCitation id="133172DAFF62FF01FD21FC94FCEB3B46" box="[754,870,835,861]" captionStart="FIGURE 113" captionStartId="130.[151,250,1175,1199]" captionTargetBox="[151,1435,345,1151]" captionTargetId="figure-111@130.[151,1435,345,1151]" captionTargetPageId="130" captionText="FIGURE 113. Caecum draperi sp. nov. A. Conceptual reconstruction of growth stages. B. Species distribution map." figureDoi="http://doi.org/10.5281/zenodo.4402601" httpUri="https://zenodo.org/record/4402601/files/figure.png" pageId="128" pageNumber="129">Fig. 113A</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF753ED7FF62FF01FF44F963FE263F23" ID-DOI="http://doi.org/10.5281/zenodo.4402595" ID-Zenodo-Dep="4402595" httpUri="https://zenodo.org/record/4402595/files/figure.png" pageId="128" pageNumber="129" startId="128.[151,250,1716,1740]" targetBox="[151,1435,885,1693]" targetPageId="128">
<paragraph id="8BB56E5FFF62FF01FF44F963FE263F23" blockId="128.[151,1437,1716,1848]" pageId="128" pageNumber="129">
<emphasis id="B97EB24DFF62FF01FF44F963FEA63ED7" bold="true" box="[151,299,1716,1740]" pageId="128" pageNumber="129">FIGURE 111.</emphasis>
<taxonomicName id="4C0A15DCFF62FF01FEE2F962FE5A3ED7" authority="Raines, 2020" authorityName="Raines" authorityYear="2020" box="[305,471,1716,1740]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="draperi" status="sp. nov.">
<emphasis id="B97EB24DFF62FF01FEE2F962FE5A3ED7" box="[305,471,1716,1740]" italics="true" pageId="128" pageNumber="129">Caecum draperi</emphasis>
</taxonomicName>
<emphasis id="B97EB24DFF62FF01FE0DF963FDC53ED7" bold="true" box="[478,584,1716,1740]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A24D0F36FF62FF01FE0DF963FDA23ED7" box="[478,559,1716,1740]" pageId="128" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. Holotype, profile view without periostracum (SBMNH 637748).
<emphasis id="B97EB24DFF62FF01FB3DF963FB723ED7" bold="true" box="[1262,1279,1716,1740]" pageId="128" pageNumber="129">B</emphasis>
. Paratype, profile view with periostracum (LACM 3756).
<emphasis id="B97EB24DFF62FF01FD89F90FFDE13EEB" bold="true" box="[602,620,1752,1776]" pageId="128" pageNumber="129">C</emphasis>
. Paratype, profile view (SBMNH 637747).
<emphasis id="B97EB24DFF62FF01FBE3F90FFBCF3EEB" bold="true" box="[1072,1090,1752,1776]" pageId="128" pageNumber="129">D</emphasis>
. Paratype, profile view (SBMNH 637747).
<emphasis id="B97EB24DFF62FF01FF28F92BFE813F0F" bold="true" box="[251,268,1788,1812]" pageId="128" pageNumber="129">E</emphasis>
. Paratype, ventral view (LACM 3756).
<emphasis id="B97EB24DFF62FF01FD61F92BFD4C3F0F" bold="true" box="[690,705,1788,1812]" pageId="128" pageNumber="129">F</emphasis>
. Operculum, profile view (SBMNH 637747).
<emphasis id="B97EB24DFF62FF01FB7AF92BFB303F0F" bold="true" box="[1193,1213,1788,1812]" pageId="128" pageNumber="129">G</emphasis>
. Operculum, exterior surface (SBMNH 637747).
</paragraph>
</caption>
<subSubSection id="C3103DD4FF62FF01FF14F8B1FB4E3FBF" pageId="128" pageNumber="129" type="etymology">
<paragraph id="8BB56E5FFF62FF01FF14F8B1FB4E3FBF" blockId="128.[151,1436,1894,2029]" pageId="128" pageNumber="129">
<emphasis id="B97EB24DFF62FF01FF14F8B1FEC33F9B" bold="true" box="[199,334,1894,1920]" pageId="128" pageNumber="129">Etymology.</emphasis>
The species is named in honor of the late Bertram “Bert” C. Draper (1904
<emphasis id="B97EB24DFF62FF01FB41F8B1FB2D3F9B" bold="true" box="[1170,1184,1894,1920]" pageId="128" pageNumber="129">–</emphasis>
2000), for his work on
<taxonomicName id="4C0A15DCFF62FF01FF44F85DFF733FBF" authorityName="Gray" authorityYear="1850" box="[151,254,1930,1956]" class="Gastropoda" family="Caecidae" kingdom="Animalia" order="Littorinimorpha" pageId="128" pageNumber="129" phylum="Mollusca" rank="family">Caecidae</taxonomicName>
while a Museum Associate with the Los Angeles County Museum of Natural History.
</paragraph>
</subSubSection>
<subSubSection id="C3103DD4FF62FF01FF14F879FC653FF7" pageId="128" pageNumber="129" type="distribution">
<paragraph id="8BB56E5FFF62FF01FF14F879FC653FF7" blockId="128.[151,1436,1894,2029]" pageId="128" pageNumber="129">
<emphasis id="B97EB24DFF62FF01FF14F879FE773FD3" bold="true" box="[199,506,1966,1992]" pageId="128" pageNumber="129">Distribution and Habitat.</emphasis>
From Chame Point,
<collectingCountry id="F31D2ECFFF62FF01FD39F878FCCE3FD2" box="[746,835,1967,1993]" name="Panama" pageId="128" pageNumber="129">Panama</collectingCountry>
Bay,
<collectingCountry id="F31D2ECFFF62FF01FC56F878FC533FD2" box="[901,990,1967,1993]" name="Panama" pageId="128" pageNumber="129">Panama</collectingCountry>
to Carrillo Peninsula de Nicoya,
<collectingCountry id="F31D2ECFFF62FF01FA8FF879FF433FF7" name="Costa Rica" pageId="128" pageNumber="129">Costa Rica</collectingCountry>
. Intertidal to subtidal. Uncommon on algae and coarse sand substrates.
</paragraph>
</subSubSection>
<caption id="DF753ED7FF63FF00FF44F8D1FEF03FB5" ID-DOI="http://doi.org/10.5281/zenodo.4402597" ID-Zenodo-Dep="4402597" httpUri="https://zenodo.org/record/4402597/files/figure.png" pageId="129" pageNumber="130" startId="129.[151,250,1798,1822]" targetBox="[151,1435,181,1773]" targetPageId="129">
<paragraph id="8BB56E5FFF63FF00FF44F8D1FEF03FB5" blockId="129.[151,1437,1798,1966]" pageId="129" pageNumber="130">
<emphasis id="B97EB24DFF63FF00FF44F8D1FEA13F05" bold="true" box="[151,300,1798,1822]" pageId="129" pageNumber="130">FIGURE 112.</emphasis>
<taxonomicName id="4C0A15DCFF63FF00FEE0F8D1FE543F05" authority="Raines, 2020" authorityName="Raines" authorityYear="2020" box="[307,473,1798,1822]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="129" pageNumber="130" phylum="Mollusca" rank="species" species="draperi" status="sp. nov.">
<emphasis id="B97EB24DFF63FF00FEE0F8D1FE543F05" box="[307,473,1798,1822]" italics="true" pageId="129" pageNumber="130">Caecum draperi</emphasis>
</taxonomicName>
<emphasis id="B97EB24DFF63FF00FE33F8D1FDBC3F05" bold="true" box="[480,561,1798,1822]" pageId="129" pageNumber="130">
<taxonomicNameLabel id="A24D0F36FF63FF00FE33F8D1FDBC3F05" box="[480,561,1798,1822]" pageId="129" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shells at different growth stages.
<emphasis id="B97EB24DFF63FF00FC59F8D1FC113F05" bold="true" box="[906,924,1798,1822]" pageId="129" pageNumber="130">A</emphasis>
. Teleoconchs II and III (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FA8EF8D1FAE33F05" bold="true" box="[1373,1390,1798,1822]" pageId="129" pageNumber="130">B</emphasis>
. Teleoconch III (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FE34F8FDFE743F59" bold="true" box="[487,505,1834,1858]" pageId="129" pageNumber="130">C</emphasis>
. Teleoconch IV (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FCA2F8FDFC0E3F59" bold="true" box="[881,899,1834,1858]" pageId="129" pageNumber="130">D</emphasis>
. Surface of teleoconchs II to IV (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FF44F899FF253F7D" bold="true" box="[151,168,1870,1894]" pageId="129" pageNumber="130">E</emphasis>
. Axial ribs of teleoconch IV (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FD4BF899FD2A3F7D" bold="true" box="[664,679,1870,1894]" pageId="129" pageNumber="130">F</emphasis>
. Surface of teleoconchs V to VI (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FB6EF899FB5C3F7D" bold="true" box="[1213,1233,1870,1894]" pageId="129" pageNumber="130">G</emphasis>
. Axial ribs of teleoconchs V to VI (SBMNH 637748).
<emphasis id="B97EB24DFF63FF00FDD1F8A5FD9B3F91" bold="true" box="[514,534,1906,1930]" pageId="129" pageNumber="130">H</emphasis>
. Teleoconch V (LACM 3756).
<emphasis id="B97EB24DFF63FF00FC85F8A5FCED3F91" bold="true" box="[854,864,1906,1930]" pageId="129" pageNumber="130">I</emphasis>
. Teleoconchs V to VI (SBMNH 637747).
<emphasis id="B97EB24DFF63FF00FADCF8A5FA913F91" bold="true" box="[1295,1308,1906,1930]" pageId="129" pageNumber="130">J</emphasis>
. Teleoconch VI (SBMNH 637748).
</paragraph>
</caption>
<subSubSection id="C3103DD4FF60FF03FF14FF40FA8F395A" pageId="130" pageNumber="131" type="discussion">
<paragraph id="8BB56E5FFF60FF03FF14FF40FA8F395A" blockId="130.[151,1437,151,321]" pageId="130" pageNumber="131">
<emphasis id="B97EB24DFF60FF03FF14FF40FEB638AA" bold="true" box="[199,315,151,177]" pageId="130" pageNumber="131">Remarks.</emphasis>
<bibRefCitation id="EF9B13AEFF60FF03FE91FF40FD8138A9" author="Lightfoot, J." box="[322,524,151,178]" pageId="130" pageNumber="131" pagination="75 - 87" refId="ref98203" refString="Lightfoot, J. (1993 b) Caecidae of the Panamic Province. Part II. Of Sea &amp; Shore, 16, 75 - 87." type="journal article" year="1993">Lightfoot (1993b)</bibRefCitation>
misidentified
<taxonomicName id="4C0A15DCFF60FF03FD7DFF40FCE838AA" authorityName="Raines" authorityYear="2020" box="[686,869,151,177]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="draperi">
<emphasis id="B97EB24DFF60FF03FD7DFF40FCE838AA" box="[686,869,151,177]" italics="true" pageId="130" pageNumber="131">Caecum draperi</emphasis>
</taxonomicName>
as
<taxonomicName id="4C0A15DCFF60FF03FC58FF40FBAC38AB" box="[907,1057,151,177]" class="Gastropoda" family="Caecidae" genus="Brochina" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="carpenteri">
<emphasis id="B97EB24DFF60FF03FC58FF40FBAC38AB" box="[907,1057,151,177]" italics="true" pageId="130" pageNumber="131">C. carpenteri</emphasis>
</taxonomicName>
=
<taxonomicName id="4C0A15DCFF60FF03FBECFF40FB3C38AA" authorityName="Raines" authorityYear="2020" box="[1087,1201,151,177]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="adamsi">
<emphasis id="B97EB24DFF60FF03FBECFF40FB3C38AA" box="[1087,1201,151,177]" italics="true" pageId="130" pageNumber="131">C. adamsi</emphasis>
</taxonomicName>
from
<collectingCountry id="F31D2ECFFF60FF03FB25FF40FADF38AA" box="[1270,1362,151,177]" name="Panama" pageId="130" pageNumber="131">Panama</collectingCountry>
. However,
<taxonomicName id="4C0A15DCFF60FF03FF06FF6CFEC638CE" authorityName="Raines" authorityYear="2020" box="[213,331,187,213]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="draperi">
<emphasis id="B97EB24DFF60FF03FF06FF6CFEC638CE" box="[213,331,187,213]" italics="true" pageId="130" pageNumber="131">C. draperi</emphasis>
</taxonomicName>
can be distinguished by its well-defined undulating axial ribs, lack of microsculpture, and unique multi-layered operculum. And as previously noted, the range of
<taxonomicName id="4C0A15DCFF60FF03FCB0FF08FC5938E2" authorityName="Raines" authorityYear="2020" box="[867,980,223,249]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="adamsi">
<emphasis id="B97EB24DFF60FF03FCB0FF08FC5938E2" box="[867,980,223,249]" italics="true" pageId="130" pageNumber="131">C. adamsi</emphasis>
</taxonomicName>
is restricted to southern California south to the Gulf of California,
<collectingCountry id="F31D2ECFFF60FF03FE63FED4FD843906" box="[432,521,259,285]" name="Mexico" pageId="130" pageNumber="131">Mexico</collectingCountry>
. Lightfoot also noted examining three specimens from
<collectingCountry id="F31D2ECFFF60FF03FBB9FED4FB6A3906" box="[1130,1255,259,285]" name="Costa Rica" pageId="130" pageNumber="131">Costa Rica</collectingCountry>
. Given the close proximity to the
<typeStatus id="54B1D0FDFF60FF03FE80FEF0FE0E395A" box="[339,387,295,321]" pageId="130" pageNumber="131">type</typeStatus>
locality, it is reasonable to deduce these three specimens are
<taxonomicName id="4C0A15DCFF60FF03FBE3FEF0FB28395A" authorityName="Raines" authorityYear="2020" box="[1072,1189,295,321]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="130" pageNumber="131" phylum="Mollusca" rank="species" species="draperi">
<emphasis id="B97EB24DFF60FF03FBE3FEF0FB28395A" box="[1072,1189,295,321]" italics="true" pageId="130" pageNumber="131">C. draperi</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
</treatment>
</document>
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