978 lines
123 KiB
XML
978 lines
123 KiB
XML
<document id="EA072A5AF816A54D5A56B4769BFD640E" ID-CLB-Dataset="2402" ID-DOI="10.5852/ejt.2022.789.1627" ID-GBIF-Dataset="3b07e8e7-5768-4279-9266-e962ae1315b0" ID-ISSN="2118-9773" ID-Zenodo-Dep="5906438" ID-ZooBank="B9AB4431-7FD3-4733-9FCA-F0DDDC7CE5A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643204463038" checkinUser="carolina" docAuthor="Souza, Maysa Fernanda Villela Rezende & Ferreira, Rodrigo Lopes" docDate="2022" docId="03A37718FF86357DFDC7FA89FEF0FCC1" docLanguage="en" docName="ejt-789_11-48.pdf" docOrigin="European Journal of Taxonomy 789" docSource="http://dx.doi.org/10.5852/ejt.2022.789.1627" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Allokoenenia canhembora Souza & Ferreira 2022, sp. nov." docType="treatment" docUuid="F6B9B9E0-6B19-4FCB-82CF-A8B4F754F492" docUuidSource="ZooBank" docVersion="8" lastPageNumber="26" masterDocId="FF9A0F60FF80356DFF80FFE5FFD9FF97" masterDocTitle="Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description" masterLastPageNumber="48" masterPageNumber="11" pageNumber="17" updateTime="1699305254263" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F4E3731E589B006CD4C92235C39F42E9">Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description</mods:title>
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<mods:roleTerm id="F2AED12F4433C0BBDCE0A7AFC39BAA83">Author</mods:roleTerm>
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<mods:namePart id="52DA739B55277E1CEF6C69CCE084FAAF">Souza, Maysa Fernanda Villela Rezende</mods:namePart>
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<mods:affiliation id="A23AF124C10E7313E1E98CAEFC3D9D2A">Centro de Estudos em Biologia Subterrânea, Departamento de Ecologia e Conservação, Universidade Federal de Lavras, Lavras, MG. CEP 37200 - 000, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="24D1293D4ECE4BBD0095A57B700D7CA9" type="email">mvillelabio@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="D1E496693ECB2A8FB40886C684FF876D">Ferreira, Rodrigo Lopes</mods:namePart>
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<mods:affiliation id="EBD7FD6F4FFF4CB04A3E5E224F3A9E7C">Centro de Estudos em Biologia Subterrânea, Departamento de Ecologia e Conservação, Universidade Federal de Lavras, Lavras, MG. CEP 37200 - 000, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="2579D36E087534072D16CA9DE5C35E30" type="email">drops@ufla.br</mods:nameIdentifier>
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<mods:title id="6ABA317139F46AA9D5A4C3177025461C">European Journal of Taxonomy</mods:title>
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<mods:date id="5D5299CA93E491F692A733FD387B7FC9">2022</mods:date>
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<treatment id="03A37718FF86357DFDC7FA89FEF0FCC1" ID-DOI="http://doi.org/10.5281/zenodo.5907244" ID-GBIF-Taxon="192629977" ID-Zenodo-Dep="5907244" LSID="urn:lsid:zoobank.org:act:F6B9B9E0-6B19-4FCB-82CF-A8B4F754F492" httpUri="http://treatment.plazi.org/id/03A37718FF86357DFDC7FA89FEF0FCC1" lastPageId="16" lastPageNumber="26" pageId="6" pageNumber="17">
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<subSubSection id="C3109585FF86356BFDC7FA89FC34FA10" box="[583,1005,1388,1415]" pageId="6" pageNumber="17" type="nomenclature">
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<paragraph id="8BB5C60EFF86356BFDC7FA89FC34FA10" blockId="6.[367,1221,1388,1485]" box="[583,1005,1388,1415]" pageId="6" pageNumber="17">
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<heading id="D0FD7162FF86356BFDC7FA89FC34FA10" box="[583,1005,1388,1415]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="17" reason="2">
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<taxonomicName id="4C0ABD8DFF86356BFDC7FA89FC52FA10" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[583,907,1388,1415]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
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<emphasis id="B97E1A1CFF86356BFDC7FA89FC52FA10" bold="true" box="[583,907,1388,1415]" italics="true" pageId="6" pageNumber="17">Allokoenenia canhembora</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24DA767FF86356BFC12FA88FC34FA10" box="[914,1005,1389,1415]" pageId="6" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C3109585FF86356BFEEFFA75FC95FA5A" pageId="6" pageNumber="17" type="description">
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urn:lsid:zoobank.org:act:
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<uuid id="FFACFCDBFF86356BFD10FA75FB1CFA3C" box="[656,1221,1424,1451]" pageId="6" pageNumber="17">F6B9B9E0-6B19-4FCB-82CF-A8B4F754F492</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BB5C60EFF86356BFD67FA56FC95FA5A" blockId="6.[367,1221,1388,1485]" box="[743,844,1459,1485]" pageId="6" pageNumber="17">
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<figureCitation id="1331DA8BFF86356BFD67FA56FC95FA5A" box="[743,844,1459,1485]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="7.[189,232,1892,1918]" captionStartId-1="8.[189,232,1928,1954]" captionStartId-2="9.[189,232,1963,1989]" captionStartId-3="10.[189,232,1928,1954]" captionStartId-4="11.[189,232,1892,1918]" captionStartId-5="12.[189,232,1928,1954]" captionStartId-6="14.[189,232,1749,1775]" captionTargetBox-0="[222,1368,282,1855]" captionTargetBox-1="[189,1398,786,1888]" captionTargetBox-2="[197,1383,782,1888]" captionTargetBox-3="[197,1388,288,1858]" captionTargetBox-4="[196,1355,474,1841]" captionTargetBox-5="[189,1398,1009,1888]" captionTargetBox-6="[299,1296,275,1709]" captionTargetId-0="figure-11@7.[209,1378,265,1855]" captionTargetId-1="figure-133@8.[189,1398,786,1888]" captionTargetId-2="figure-179@9.[189,1398,767,1924]" captionTargetId-3="figure-15@10.[189,1398,277,1888]" captionTargetId-4="figure-64@11.[189,1398,466,1853]" captionTargetId-5="figure-380@12.[189,1398,1009,1888]" captionTargetId-6="figure-14@14.[290,1296,265,1712]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="14" captionText-0="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." captionText-1="Fig. 4. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Deutotritosternal setae. B. Propeltidial chaetotaxy. C. Proximal setae (p 1–p 6) on basal segment of chelicera (p 5 out of focus). Abbreviations: see Material and methods. Scale bars: A= 20 μm; B = 50 μm; C = 45 μm." captionText-2="Fig. 5. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-femur of right pedipalp. B. Tibia-tarsus 2 of left pedipalp.Abbreviations: see Material and methods. Scale bar = 100 μm." captionText-3="Fig. 6. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-patella of right leg I. B. Tibia-basitarsus 2 of left leg I. C. Basitarsus 3-tarsus 2 of left leg I. D. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm." captionText-4="Fig. 7. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Tarsus 3 of left pedipalp. B. Basitarsus of right leg IV. C. First lobe of female genitalia. D. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods." captionText-5="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." captionText-6="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi-0="http://doi.org/10.5281/zenodo.5906444" figureDoi-1="http://doi.org/10.5281/zenodo.5906446" figureDoi-2="http://doi.org/10.5281/zenodo.5906448" figureDoi-3="http://doi.org/10.5281/zenodo.5906450" figureDoi-4="http://doi.org/10.5281/zenodo.5906452" figureDoi-5="http://doi.org/10.5281/zenodo.5906454" figureDoi-6="http://doi.org/10.5281/zenodo.5906456" httpUri-0="https://zenodo.org/record/5906444/files/figure.png" httpUri-1="https://zenodo.org/record/5906446/files/figure.png" httpUri-2="https://zenodo.org/record/5906448/files/figure.png" httpUri-3="https://zenodo.org/record/5906450/files/figure.png" httpUri-4="https://zenodo.org/record/5906452/files/figure.png" httpUri-5="https://zenodo.org/record/5906454/files/figure.png" httpUri-6="https://zenodo.org/record/5906456/files/figure.png" pageId="6" pageNumber="17">Figs 3–9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3109585FF86356BFF3DFA11FCB7F8C0" pageId="6" pageNumber="17" type="diagnosis">
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<paragraph id="8BB5C60EFF86356BFF3DFA11FEEEF999" blockId="6.[189,311,1524,1550]" box="[189,311,1524,1550]" pageId="6" pageNumber="17">
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<heading id="D0FD7162FF86356BFF3DFA11FEEEF999" bold="true" box="[189,311,1524,1550]" fontSize="11" level="3" pageId="6" pageNumber="17" reason="3">
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<emphasis id="B97E1A1CFF86356BFF3DFA11FEEEF999" bold="true" box="[189,311,1524,1550]" pageId="6" pageNumber="17">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB5C60EFF86356BFF3DF9C6FCB7F8C0" blockId="6.[188,1399,1571,1879]" pageId="6" pageNumber="17">
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Frontal organ formed by 2 reticulated and expanded branches; 5 blades finely reticulated in the lateral organs; 5 setae on deutotritosternum; 7 pairs of short setae on propeltidium; 3 pairs of setae with similar length on metapeltidium; cheliceral fingers with 9 teeth each; coxae II–IV with 3, 3, 1 thick setae, respectively; 5 setae (
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<emphasis id="B97E1A1CFF86356BFE44F968FE3EF931" box="[452,487,1677,1702]" italics="true" pageId="6" pageNumber="17">grt</emphasis>
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,
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<emphasis id="B97E1A1CFF86356BFE77F968FDDBF931" box="[503,514,1677,1702]" italics="true" pageId="6" pageNumber="17">r</emphasis>
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,
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<emphasis id="B97E1A1CFF86356BFD94F968FDE2F931" box="[532,571,1677,1702]" italics="true" pageId="6" pageNumber="17">esp</emphasis>
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and 2
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<emphasis id="B97E1A1CFF86356BFD12F968FD60F930" box="[658,697,1677,1703]" italics="true" pageId="6" pageNumber="17">esd</emphasis>
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) on basitarsus of leg IV; opisthosomal tergites II–VI with two pairs of setae
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<emphasis id="B97E1A1CFF86356BFE16F954FE47F95D" box="[406,414,1713,1738]" italics="true" pageId="6" pageNumber="17">t</emphasis>
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between one setae
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<emphasis id="B97E1A1CFF86356BFD05F954FD49F95D" box="[645,656,1713,1738]" italics="true" pageId="6" pageNumber="17">s</emphasis>
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on each side; opisthosomal sternites IV–VI with three pairs of
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<emphasis id="B97E1A1CFF86356BFF3EF931FF14F97A" box="[190,205,1748,1773]" italics="true" pageId="6" pageNumber="17">a</emphasis>
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setae (
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<emphasis id="B97E1A1CFF86356BFEA1F931FEE0F963" box="[289,313,1748,1780]" italics="true" pageId="6" pageNumber="17">
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a
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<subScript id="178EC44BFF86356BFEB0F903FEE0F963" attach="left" box="[304,313,1766,1780]" fontSize="6" pageId="6" pageNumber="17">1</subScript>
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</emphasis>
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shorter than
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<emphasis id="B97E1A1CFF86356BFE56F931FE34F963" box="[470,493,1748,1780]" italics="true" pageId="6" pageNumber="17">
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a
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<subScript id="178EC44BFF86356BFE64F903FE34F963" attach="left" box="[484,493,1766,1780]" fontSize="6" pageId="6" pageNumber="17">2</subScript>
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</emphasis>
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and
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<emphasis id="B97E1A1CFF86356BFDA8F931FD99F963" box="[552,576,1748,1780]" italics="true" pageId="6" pageNumber="17">
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a
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<subScript id="178EC44BFF86356BFDB7F903FD99F963" attach="left" box="[567,576,1766,1780]" fontSize="6" pageId="6" pageNumber="17">3</subScript>
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</emphasis>
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) flanked by one
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<emphasis id="B97E1A1CFF86356BFC88F931FCCAF97A" box="[776,787,1748,1773]" italics="true" pageId="6" pageNumber="17">s</emphasis>
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seta on each side; opisthosomal segment XI with 2 pairs of ventral setae arranged in 2 rows; first lobe of female genitalia with 10 pairs of setae; flagellum formed by 14 moniliform segments with setae of different lengths, including very long setae inserted in a V arrangement from the sixth flagellar segment onwards.
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</paragraph>
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</subSubSection>
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<caption id="DF759686FF87356AFF3DF881FE75F87E" ID-DOI="http://doi.org/10.5281/zenodo.5906444" ID-Zenodo-Dep="5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="7" pageNumber="18" startId="7.[189,232,1892,1918]" targetBox="[222,1368,282,1855]" targetPageId="7">
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<paragraph id="8BB5C60EFF87356AFF3DF881FE75F87E" blockId="7.[189,1399,1892,2025]" pageId="7" pageNumber="18">
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<emphasis id="B97E1A1CFF87356AFF3DF881FED5F8E8" bold="true" box="[189,268,1892,1919]" pageId="7" pageNumber="18">Fig. 3.</emphasis>
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<taxonomicName id="4C0ABD8DFF87356AFE95F880FD96F8E8" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[277,591,1893,1919]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
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<emphasis id="B97E1A1CFF87356AFE95F880FD96F8E8" box="[277,591,1893,1919]" italics="true" pageId="7" pageNumber="18">Allokoenenia canhembora</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24DA767FF87356AFDD7F880FD6FF8E8" box="[599,694,1893,1919]" pageId="7" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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, ♀, holotype (ISLA 50394).
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<emphasis id="B97E1A1CFF87356AFB8DF881FBFBF8E9" bold="true" box="[1037,1058,1892,1918]" pageId="7" pageNumber="18">A</emphasis>
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. Frontal organ.
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<emphasis id="B97E1A1CFF87356AFB63F880FB2CF8E8" bold="true" box="[1251,1269,1893,1919]" pageId="7" pageNumber="18">B</emphasis>
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. Blades of lateral organ.
|
||
<emphasis id="B97E1A1CFF87356AFEDFF86DFEADF835" bold="true" box="[351,372,1928,1954]" pageId="7" pageNumber="18">C</emphasis>
|
||
. Coxa of left pedipalp.
|
||
<emphasis id="B97E1A1CFF87356AFD0CF86DFD70F835" bold="true" box="[652,681,1928,1954]" pageId="7" pageNumber="18">D.</emphasis>
|
||
Coxa of left leg I.
|
||
<emphasis id="B97E1A1CFF87356AFC08F86DFC42F835" bold="true" box="[904,923,1928,1954]" pageId="7" pageNumber="18">E</emphasis>
|
||
. Coxa of right leg II.
|
||
<emphasis id="B97E1A1CFF87356AFB1EF86DFB69F835" bold="true" box="[1182,1200,1928,1954]" pageId="7" pageNumber="18">F</emphasis>
|
||
. Coxa of left leg III.
|
||
<emphasis id="B97E1A1CFF87356AFF6AF84EFED8F852" bold="true" box="[234,257,1963,1989]" pageId="7" pageNumber="18">G</emphasis>
|
||
. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3109585FF883565FF3DFEECFDB5FE2C" pageId="8" pageNumber="19" type="etymology">
|
||
<paragraph id="8BB5C60EFF883565FF3DFEECFE9CFEB4" blockId="8.[189,325,265,291]" box="[189,325,265,291]" pageId="8" pageNumber="19">
|
||
<heading id="D0FD7162FF883565FF3DFEECFE9CFEB4" bold="true" box="[189,325,265,291]" fontSize="11" level="3" pageId="8" pageNumber="19" reason="3">
|
||
<emphasis id="B97E1A1CFF883565FF3DFEECFE9CFEB4" bold="true" box="[189,325,265,291]" pageId="8" pageNumber="19">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF883565FF3DFED2FDB5FE2C" blockId="8.[187,1399,311,443]" pageId="8" pageNumber="19">The species ephitet is derived from a word of the Tupi language meaning ʻfugitiveʼ. The species received this name due to the difficulty of capturing the specimens, which quickly ran away and hid under the calcite plates on the cave floor as soon as they were discovered (see: Habitats and threats). The name is to be treated as a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3109585FF883565FF3DFE07FBADFD41" pageId="8" pageNumber="19" type="materials_examined">
|
||
<paragraph id="8BB5C60EFF883565FF3DFE07FE74FE6B" blockId="8.[189,429,482,508]" box="[189,429,482,508]" pageId="8" pageNumber="19">
|
||
<heading id="D0FD7162FF883565FF3DFE07FE74FE6B" bold="true" box="[189,429,482,508]" fontSize="11" level="3" pageId="8" pageNumber="19" reason="3">
|
||
<emphasis id="B97E1A1CFF883565FF3DFE07FE74FE6B" bold="true" box="[189,429,482,508]" pageId="8" pageNumber="19">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF883565FF6AFDF4FCC2FDE5" blockId="8.[189,1399,529,626]" pageId="8" pageNumber="19">
|
||
<materialsCitation id="3B62CC53FF883565FF6AFDF4FCC2FDE5" ID-GBIF-Occurrence="3457762304" collectingDate="2012-06-11" collectionCode="R" collectorName="R. L. Ferreira" country="Brazil" elevation="527" latitude="-10.511373" location="Toca do Goncalo Cave" longLatPrecision="1" longitude="-40.89503" municipality="Campo Formoso" pageId="8" pageNumber="19" specimenCode="ISLA 50394" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="holotype">
|
||
<emphasis id="B97E1A1CFF883565FF6AFDF4FE84FDBC" bold="true" box="[234,349,529,555]" pageId="8" pageNumber="19">
|
||
<typeStatus id="54B178ACFF883565FF6AFDF4FE84FDBC" box="[234,349,529,555]" pageId="8" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F31D869EFF883565FF3DFDD1FEFFFDD9" box="[189,294,564,590]" name="Brazil" pageId="8" pageNumber="19">BRAZIL</collectingCountry>
|
||
•
|
||
<specimenCount id="9D0C0D87FF883565FEBEFDD1FE8CFDD9" box="[318,341,564,590]" pageId="8" pageNumber="19" type="female">♀</specimenCount>
|
||
;
|
||
<collectingRegion id="49CE08ECFF883565FEE3FDD1FE73FDD9" box="[355,426,564,590]" country="Brazil" name="Bahia" pageId="8" pageNumber="19">Bahia</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BD15C74FF883565FE36FDD1FDA5FDD9" box="[438,636,564,590]" pageId="8" pageNumber="19">Campo Formoso</collectingMunicipality>
|
||
,
|
||
<location id="8ED590D5FF883565FD07FDD1FC4FFDD9" LSID="urn:lsid:plazi:treatment:03A37718FF86357DFDC7FA89FEF0FCC1:8ED590D5FF883565FD07FDD1FC4FFDD9" box="[647,918,564,590]" country="Brazil" latitude="-10.511373" longLatPrecision="1" longitude="-40.89503" municipality="Campo Formoso" name="Toca do Goncalo Cave" pageId="8" pageNumber="19" stateProvince="Bahia">Toca do Gonçalo Cave</location>
|
||
;
|
||
<geoCoordinate id="EE3EA0C9FF883565FC21FDD1FB8CFDD9" box="[929,1109,562,590]" degrees="10" direction="south" minutes="30" orientation="latitude" pageId="8" pageNumber="19" precision="1" seconds="40.94" value="-10.511373">10°30′40.94″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3EA0C9FF883565FBE2FDD1FAF8FDD9" box="[1122,1313,562,591]" degrees="40" direction="west" minutes="53" orientation="longitude" pageId="8" pageNumber="19" precision="1" seconds="42.10" value="-40.89503">40°53′42.10″ W</geoCoordinate>
|
||
;
|
||
<elevation id="0027213DFF883565FAADFDD1FF2CFDE6" metricMagnitude="2" metricUnit="m" metricValue="5.27" pageId="8" pageNumber="19" unit="m" value="527.0">
|
||
<quantity id="4CF26BEBFF883565FAADFDD1FAAEFDD9" box="[1325,1399,564,591]" metricMagnitude="2" metricUnit="m" metricValue="5.27" pageId="8" pageNumber="19" unit="m" value="527.0">527 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFB4E0CEFF883565FE83FDB2FE45FDE6" box="[259,412,599,626]" pageId="8" pageNumber="19" value="2012-06-11">
|
||
<collectingDate id="EFF01926FF883565FE83FDB2FE45FDE6" box="[259,412,599,626]" pageId="8" pageNumber="19" value="2012-06-11">11 Jun. 2012</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26FFA3D8FF883565FE28FDBDFD98FDE6" box="[424,577,599,626]" pageId="8" pageNumber="19">R.L. Ferreira</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBAC6E75FF883565FD02FDB2FCCFFDE5" box="[642,790,599,626]" pageId="8" pageNumber="19">ISLA 50394</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF883565FF6AFD7DFE71FD25" blockId="8.[189,1140,664,726]" box="[234,424,664,690]" pageId="8" pageNumber="19">
|
||
<emphasis id="B97E1A1CFF883565FF6AFD7DFE71FD25" bold="true" box="[234,424,664,690]" pageId="8" pageNumber="19">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF883565FF3DFD59FBADFD41" blockId="8.[189,1140,664,726]" box="[189,1140,699,726]" pageId="8" pageNumber="19">
|
||
<materialsCitation id="3B62CC53FF883565FF3DFD59FBADFD41" ID-GBIF-Occurrence="3457762302" box="[189,1140,699,726]" collectingDate="2014-03-16" collectionCode="R" collectorName="L. Ferreira" country="Brazil" elevation="527" latitude="-10.511373" location="BRAZIL" longLatPrecision="1" longitude="-40.89503" municipality="Campo Formoso" pageId="8" pageNumber="19" specimenCode="ISLA 50395" specimenCount="1" specimenCount-female="1" stateProvince="Bahia">
|
||
<collectingCountry id="F31D869EFF883565FF3DFD59FEFFFD41" box="[189,294,700,726]" name="Brazil" pageId="8" pageNumber="19">BRAZIL</collectingCountry>
|
||
•
|
||
<specimenCount id="9D0C0D87FF883565FEBFFD59FEB5FD41" box="[319,364,700,726]" pageId="8" pageNumber="19" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<date id="FFB4E0CEFF883565FCB0FD5EFC08FD41" box="[816,977,699,726]" pageId="8" pageNumber="19" value="2014-03-16">
|
||
<collectingDate id="EFF01926FF883565FCB0FD5EFC08FD41" box="[816,977,699,726]" pageId="8" pageNumber="19" value="2014-03-16">16 Mar. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="DBAC6E75FF883565FC5DFD59FBB4FD41" box="[989,1133,699,726]" pageId="8" pageNumber="19">ISLA 50395</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF759686FF883565FF3DF86DFB0DF87E" ID-DOI="http://doi.org/10.5281/zenodo.5906446" ID-Zenodo-Dep="5906446" httpUri="https://zenodo.org/record/5906446/files/figure.png" pageId="8" pageNumber="19" startId="8.[189,232,1928,1954]" targetBox="[189,1398,786,1888]" targetPageId="8">
|
||
<paragraph id="8BB5C60EFF883565FF3DF86DFB0DF87E" blockId="8.[189,1399,1928,2025]" pageId="8" pageNumber="19">
|
||
<emphasis id="B97E1A1CFF883565FF3DF86DFEC8F835" bold="true" box="[189,273,1928,1954]" pageId="8" pageNumber="19">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF883565FE9EF86DFD82F835" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[286,603,1928,1954]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF883565FE9EF86DFD82F835" box="[286,603,1928,1954]" italics="true" pageId="8" pageNumber="19">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF883565FDE8F86DFD12F835" box="[616,715,1928,1954]" pageId="8" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, holotype (ISLA 50394).
|
||
<emphasis id="B97E1A1CFF883565FBB7F86DFB95F835" bold="true" box="[1079,1100,1928,1954]" pageId="8" pageNumber="19">A</emphasis>
|
||
. Deutotritosternal setae.
|
||
<emphasis id="B97E1A1CFF883565FF3DF84EFF08F852" bold="true" box="[189,209,1963,1989]" pageId="8" pageNumber="19">B</emphasis>
|
||
. Propeltidial chaetotaxy.
|
||
<emphasis id="B97E1A1CFF883565FD84F84EFDCEF852" bold="true" box="[516,535,1963,1989]" pageId="8" pageNumber="19">C</emphasis>
|
||
. Proximal setae (
|
||
<emphasis id="B97E1A1CFF883565FD6CF849FCF3F85B" box="[748,810,1964,1996]" italics="true" pageId="8" pageNumber="19">
|
||
p
|
||
<subScript id="178EC44BFF883565FD7AF85BFCDAF85B" attach="left" box="[762,771,1982,1996]" fontSize="6" pageId="8" pageNumber="19">1</subScript>
|
||
–p
|
||
<subScript id="178EC44BFF883565FCA1F85BFCF3F85B" attach="left" box="[801,810,1982,1996]" fontSize="6" pageId="8" pageNumber="19">6</subScript>
|
||
</emphasis>
|
||
) on basal segment of chelicera (
|
||
<emphasis id="B97E1A1CFF883565FB34F849FB15F85B" box="[1204,1228,1964,1996]" italics="true" pageId="8" pageNumber="19">
|
||
p
|
||
<subScript id="178EC44BFF883565FB43F85BFB15F85B" attach="left" box="[1219,1228,1982,1996]" fontSize="6" pageId="8" pageNumber="19">5</subScript>
|
||
</emphasis>
|
||
out of focus). Abbreviations: see Material and methods. Scale bars: A= 20 μm; B = 50 μm; C = 45 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3109585FF893560FF3DFEECFEF3FA42" lastPageId="13" lastPageNumber="24" pageId="9" pageNumber="20" type="description">
|
||
<paragraph id="8BB5C60EFF893564FF3DFEECFE96FEB4" blockId="9.[189,335,265,291]" box="[189,335,265,291]" pageId="9" pageNumber="20">
|
||
<heading id="D0FD7162FF893564FF3DFEECFE96FEB4" bold="true" box="[189,335,265,291]" fontSize="11" level="3" pageId="9" pageNumber="20" reason="3">
|
||
<emphasis id="B97E1A1CFF893564FF3DFEECFE96FEB4" bold="true" box="[189,335,265,291]" pageId="9" pageNumber="20">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF893564FF6AFED2FE55FEC6" blockId="9.[189,657,311,373]" box="[234,396,311,337]" pageId="9" pageNumber="20">
|
||
<emphasis id="B97E1A1CFF893564FF6AFED2FE55FEC6" bold="true" box="[234,396,311,337]" pageId="9" pageNumber="20">Adult female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF893564FF3DFEBEFD48FEE2" blockId="9.[189,657,311,373]" box="[189,657,347,373]" pageId="9" pageNumber="20">
|
||
MEASUREMENTS AND RATIOS. See
|
||
<tableCitation id="C688F3B5FF893564FDB3FEBEFD48FEE2" box="[563,657,347,373]" captionStart="Table 2" captionStartId="4.[189,259,265,291]" captionTargetPageId="4" captionText="Table 2. Measurements and ratios of morphological characters of the three Brazilian Allokoenenia Silvestri, 1913 and the few measurements obtained from the adult females of the type series of A. afra Silvestri, 1913. The asterisks indicate the absence of the gla seta on basitarsus IV; the dashes indicate structures that could not be accurately measured. The body length (L) of A. afra was obtained from Silvestri (1913)." httpUri="http://table.plazi.org/id/DF759686FF843569FF3DFEECFEA5FE27" pageId="9" pageNumber="20" tableUuid="DF759686FF843569FF3DFEECFEA5FE27">Table 2.</tableCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF893564FF3DFE78FD62FE20" blockId="9.[189,699,412,439]" box="[189,699,412,439]" pageId="9" pageNumber="20">BODY LENGTH. Without flagellum: 1420 μm.</paragraph>
|
||
<paragraph id="8BB5C60EFF893566FF3DFE3BFCB4FEFE" blockId="9.[187,1399,477,722]" lastBlockId="11.[188,1398,265,362]" lastPageId="11" lastPageNumber="22" pageId="9" pageNumber="20">
|
||
PROSOMA. Frontal organ formed by two reticulated and expanded branches (42 long), with blunt tips (
|
||
<figureCitation id="1331DA8BFF893564FF46FDE4FEFEFD8C" box="[198,295,513,539]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="9" pageNumber="20">Fig. 3A</figureCitation>
|
||
). Lateral organ with 5 blades pointed-lanceolate (32 long) and finely reticulated (
|
||
<figureCitation id="1331DA8BFF893564FA89FDE4FAB1FD8C" box="[1289,1384,513,539]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="9" pageNumber="20">Fig. 3B</figureCitation>
|
||
). Propeltidium with 7 +7 short setae (
|
||
<figureCitation id="1331DA8BFF893564FDEDFDC1FD15FDA9" box="[621,716,548,574]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[189,1398,786,1888]" captionTargetId="figure-133@8.[189,1398,786,1888]" captionTargetPageId="8" captionText="Fig. 4. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Deutotritosternal setae. B. Propeltidial chaetotaxy. C. Proximal setae (p 1–p 6) on basal segment of chelicera (p 5 out of focus). Abbreviations: see Material and methods. Scale bars: A= 20 μm; B = 50 μm; C = 45 μm." figureDoi="http://doi.org/10.5281/zenodo.5906446" httpUri="https://zenodo.org/record/5906446/files/figure.png" pageId="9" pageNumber="20">Fig. 4B</figureCitation>
|
||
). Setae
|
||
<emphasis id="B97E1A1CFF893564FCAEFDC0FCE4FDD2" box="[814,829,549,581]" italics="true" pageId="9" pageNumber="20">
|
||
t
|
||
<subScript id="178EC44BFF893564FCB4FDD2FCE4FDD2" attach="left" box="[820,829,567,581]" fontSize="6" pageId="9" pageNumber="20">1</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF893564FCCEFDC0FC86FDD2" box="[846,863,549,581]" italics="true" pageId="9" pageNumber="20">
|
||
t
|
||
<subScript id="178EC44BFF893564FCD6FDD2FC86FDD2" attach="left" box="[854,863,567,581]" fontSize="6" pageId="9" pageNumber="20">2</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B97E1A1CFF893564FC1EFDC0FC74FDD2" box="[926,941,549,581]" italics="true" pageId="9" pageNumber="20">
|
||
t
|
||
<subScript id="178EC44BFF893564FC24FDD2FC74FDD2" attach="left" box="[932,941,567,581]" fontSize="6" pageId="9" pageNumber="20">3</subScript>
|
||
</emphasis>
|
||
of metapeltidium 77, 75 and 77 long, respectively. Deuto-tritosternum with 5 setae in U-shaped arrangement (
|
||
<figureCitation id="1331DA8BFF893564FC7BFDA2FB8EFDF6" box="[1019,1111,583,609]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[189,1398,786,1888]" captionTargetId="figure-133@8.[189,1398,786,1888]" captionTargetPageId="8" captionText="Fig. 4. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Deutotritosternal setae. B. Propeltidial chaetotaxy. C. Proximal setae (p 1–p 6) on basal segment of chelicera (p 5 out of focus). Abbreviations: see Material and methods. Scale bars: A= 20 μm; B = 50 μm; C = 45 μm." figureDoi="http://doi.org/10.5281/zenodo.5906446" httpUri="https://zenodo.org/record/5906446/files/figure.png" pageId="9" pageNumber="20">Fig. 4A</figureCitation>
|
||
). Labrum with 5+5 short setae. Basal segment of chelicera 195 long (dorsal length), with 6 proximal setae (
|
||
<emphasis id="B97E1A1CFF893564FB14FD8EFB75FD1C" box="[1172,1196,619,651]" italics="true" pageId="9" pageNumber="20">
|
||
p
|
||
<subScript id="178EC44BFF893564FB23FD98FB75FD1C" attach="left" box="[1187,1196,637,651]" fontSize="6" pageId="9" pageNumber="20">4</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B97E1A1CFF893564FB69FD8EFAD9FD1C" box="[1257,1280,619,651]" italics="true" pageId="9" pageNumber="20">
|
||
p
|
||
<subScript id="178EC44BFF893564FB77FD98FAD9FD1C" attach="left" box="[1271,1280,637,651]" fontSize="6" pageId="9" pageNumber="20">6</subScript>
|
||
</emphasis>
|
||
thickened and densely barbed;
|
||
<emphasis id="B97E1A1CFF893564FE2FFD6BFE1EFD39" box="[431,455,654,686]" italics="true" pageId="9" pageNumber="20">
|
||
p
|
||
<subScript id="178EC44BFF893564FE3EFD45FE1EFD39" attach="left" box="[446,455,672,686]" fontSize="6" pageId="9" pageNumber="20">1</subScript>
|
||
</emphasis>
|
||
slightly thinner and barbed) (
|
||
<figureCitation id="1331DA8BFF893564FCA3FD68FCA6FD3F" box="[803,895,653,680]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[189,1398,786,1888]" captionTargetId="figure-133@8.[189,1398,786,1888]" captionTargetPageId="8" captionText="Fig. 4. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Deutotritosternal setae. B. Propeltidial chaetotaxy. C. Proximal setae (p 1–p 6) on basal segment of chelicera (p 5 out of focus). Abbreviations: see Material and methods. Scale bars: A= 20 μm; B = 50 μm; C = 45 μm." figureDoi="http://doi.org/10.5281/zenodo.5906446" httpUri="https://zenodo.org/record/5906446/files/figure.png" pageId="9" pageNumber="20">Fig. 4C</figureCitation>
|
||
), and 3 distal setae:
|
||
<emphasis id="B97E1A1CFF893564FBEDFD68FB5CFD39" box="[1133,1157,653,686]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FBFCFD45FB5CFD39" attach="left" box="[1148,1157,672,686]" fontSize="6" pageId="9" pageNumber="20">3</subScript>
|
||
</emphasis>
|
||
(107) longer than
|
||
<emphasis id="B97E1A1CFF893564FADFFD68FAAEFD39" box="[1375,1399,653,686]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FAEEFD45FAAEFD39" attach="left" box="[1390,1399,672,686]" fontSize="6" pageId="9" pageNumber="20">1</subScript>
|
||
</emphasis>
|
||
(60) and
|
||
<emphasis id="B97E1A1CFF893564FEA9FD54FE99FD45" box="[297,320,689,722]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FEB7FD21FE99FD45" attach="left" box="[311,320,708,722]" fontSize="6" pageId="9" pageNumber="20">2</subScript>
|
||
</emphasis>
|
||
(62);
|
||
<emphasis id="B97E1A1CFF893564FE09FD54FE79FD45" box="[393,416,689,722]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FE17FD21FE79FD45" attach="left" box="[407,416,708,722]" fontSize="6" pageId="9" pageNumber="20">3</subScript>
|
||
</emphasis>
|
||
smooth near base and barbed in its distal half,
|
||
<emphasis id="B97E1A1CFF893564FC4DFD54FC3CFD45" box="[973,997,689,722]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FC5CFD21FC3CFD45" attach="left" box="[988,997,708,722]" fontSize="6" pageId="9" pageNumber="20">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B97E1A1CFF893564FB9FFD54FBEFFD45" box="[1055,1078,689,722]" italics="true" pageId="9" pageNumber="20">
|
||
d
|
||
<subScript id="178EC44BFF893564FBADFD21FBEFFD45" attach="left" box="[1069,1078,708,722]" fontSize="6" pageId="9" pageNumber="20">2</subScript>
|
||
</emphasis>
|
||
thin, flexible and with tiny projections in apex; and one apical seta. Hand of chelicera with 7 setae: 4 dorsal setae, 2 setae in its outer portion (1 close to articulation of movable finger and 1 on a tubercle close to teeth of fixed finger) and 1 seta inserted in its inner portion. Fingers with 9 teeth each.
|
||
</paragraph>
|
||
<caption id="DF759686FF893564FF3DF84EFAAEF87E" ID-DOI="http://doi.org/10.5281/zenodo.5906448" ID-Zenodo-Dep="5906448" httpUri="https://zenodo.org/record/5906448/files/figure.png" pageId="9" pageNumber="20" startId="9.[189,232,1963,1989]" targetBox="[197,1383,782,1888]" targetPageId="9">
|
||
<paragraph id="8BB5C60EFF893564FF3DF84EFAAEF87E" blockId="9.[189,1399,1963,2025]" pageId="9" pageNumber="20">
|
||
<emphasis id="B97E1A1CFF893564FF3DF84EFED7F852" bold="true" box="[189,270,1963,1989]" pageId="9" pageNumber="20">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF893564FE98F84EFD8AF852" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[280,595,1963,1989]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF893564FE98F84EFD8AF852" box="[280,595,1963,1989]" italics="true" pageId="9" pageNumber="20">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF893564FDDEF849FD67F851" box="[606,702,1964,1990]" pageId="9" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, holotype (ISLA 50394).
|
||
<emphasis id="B97E1A1CFF893564FB9DF84EFBEBF852" bold="true" box="[1053,1074,1963,1989]" pageId="9" pageNumber="20">A</emphasis>
|
||
. Trochanter-femur of right pedipalp.
|
||
<emphasis id="B97E1A1CFF893564FEAEF82BFE9BF87F" bold="true" box="[302,322,1998,2024]" pageId="9" pageNumber="20">B</emphasis>
|
||
. Tibia-tarsus 2 of left pedipalp.Abbreviations: see Material and methods. Scale bar = 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF759686FF8A3567FF3DF86DFBA2F87E" ID-DOI="http://doi.org/10.5281/zenodo.5906450" ID-Zenodo-Dep="5906450" httpUri="https://zenodo.org/record/5906450/files/figure.png" pageId="10" pageNumber="21" startId="10.[189,232,1928,1954]" targetBox="[197,1388,288,1858]" targetPageId="10">
|
||
<paragraph id="8BB5C60EFF8A3567FF3DF86DFBA2F87E" blockId="10.[189,1399,1928,2025]" pageId="10" pageNumber="21">
|
||
<emphasis id="B97E1A1CFF8A3567FF3DF86DFED4F835" bold="true" box="[189,269,1928,1954]" pageId="10" pageNumber="21">Fig. 6.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF8A3567FE97F86DFD8BF835" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[279,594,1928,1954]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8A3567FE97F86DFD8BF835" box="[279,594,1928,1954]" italics="true" pageId="10" pageNumber="21">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8A3567FDDBF86DFD63F835" box="[603,698,1928,1954]" pageId="10" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, holotype (ISLA 50394).
|
||
<emphasis id="B97E1A1CFF8A3567FB97F86DFBF5F835" bold="true" box="[1047,1068,1928,1954]" pageId="10" pageNumber="21">A</emphasis>
|
||
. Trochanter-patella of right leg I.
|
||
<emphasis id="B97E1A1CFF8A3567FE83F84EFECFF852" bold="true" box="[259,278,1963,1989]" pageId="10" pageNumber="21">B</emphasis>
|
||
. Tibia-basitarsus 2 of left leg I.
|
||
<emphasis id="B97E1A1CFF8A3567FD14F84EFD70F852" bold="true" box="[660,681,1963,1989]" pageId="10" pageNumber="21">C</emphasis>
|
||
. Basitarsus 3-tarsus 2 of left leg I.
|
||
<emphasis id="B97E1A1CFF8A3567FBCBF84EFBB9F852" bold="true" box="[1099,1120,1963,1989]" pageId="10" pageNumber="21">D</emphasis>
|
||
. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF759686FF8B3566FF3DF881FE28F87F" ID-DOI="http://doi.org/10.5281/zenodo.5906452" ID-Zenodo-Dep="5906452" httpUri="https://zenodo.org/record/5906452/files/figure.png" pageId="11" pageNumber="22" startId="11.[189,232,1892,1918]" targetBox="[196,1355,474,1841]" targetPageId="11">
|
||
<paragraph id="8BB5C60EFF8B3566FF3DF881FE28F87F" blockId="11.[189,1399,1892,2025]" pageId="11" pageNumber="22">
|
||
<emphasis id="B97E1A1CFF8B3566FF3DF881FED7F8E8" bold="true" box="[189,270,1892,1919]" pageId="11" pageNumber="22">Fig. 7.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF8B3566FE99F880FD8DF8E8" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[281,596,1893,1919]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8B3566FE99F880FD8DF8E8" box="[281,596,1893,1919]" italics="true" pageId="11" pageNumber="22">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8B3566FDDFF880FD66F8E8" box="[607,703,1893,1919]" pageId="11" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, holotype (ISLA 50394).
|
||
<emphasis id="B97E1A1CFF8B3566FB9FF881FBEDF8E9" bold="true" box="[1055,1076,1892,1918]" pageId="11" pageNumber="22">A</emphasis>
|
||
. Tarsus 3 of left pedipalp.
|
||
<emphasis id="B97E1A1CFF8B3566FF3DF86DFF08F835" bold="true" box="[189,209,1928,1954]" pageId="11" pageNumber="22">B</emphasis>
|
||
. Basitarsus of right leg IV.
|
||
<emphasis id="B97E1A1CFF8B3566FD90F86DFDFCF835" bold="true" box="[528,549,1928,1954]" pageId="11" pageNumber="22">C</emphasis>
|
||
. First lobe of female genitalia.
|
||
<emphasis id="B97E1A1CFF8B3566FC13F86DFC71F835" bold="true" box="[915,936,1928,1954]" pageId="11" pageNumber="22">D</emphasis>
|
||
. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB5C60EFF8C3561FF3DFEECFC66FE1A" blockId="12.[189,1400,265,397]" pageId="12" pageNumber="23">
|
||
COXAL CHAETOTAXY. Pedipalp coxa with 18 setae (
|
||
<figureCitation id="1331DA8BFF8C3561FCA2FEECFC5AFEB4" box="[802,899,265,291]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="12" pageNumber="23">Fig. 3C</figureCitation>
|
||
); coxa I with 13 ordinary setae and two microsetae (
|
||
<figureCitation id="1331DA8BFF8C3561FECDFEC9FE74FED1" box="[333,429,300,326]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="12" pageNumber="23">Fig. 3D</figureCitation>
|
||
); coxa II with 3 thick setae, 2 macrosetae and 8 ordinary setae (
|
||
<figureCitation id="1331DA8BFF8C3561FB26FEC9FADAFED1" box="[1190,1283,300,326]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="12" pageNumber="23">Fig. 3E</figureCitation>
|
||
); coxa III with 3 thick setae, 1 macroseta, and 8 ordinary setae (including 1 small seta adjacent to macroseta) (
|
||
<figureCitation id="1331DA8BFF8C3561FF46FE97FEC5FE1A" box="[198,284,370,397]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="12" pageNumber="23">Fig. 3F</figureCitation>
|
||
) and coxa IV with 1 thick and 8 ordinary setae (
|
||
<figureCitation id="1331DA8BFF8C3561FCD3FE97FC6BFE1A" box="[851,946,370,397]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="12" pageNumber="23">Fig. 3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8C3561FF3DFE50FCD5FDAF" blockId="12.[188,1399,436,569]" pageId="12" pageNumber="23">
|
||
PEDIPALP.
|
||
<emphasis id="B97E1A1CFF8C3561FEB0FE50FE9CFE59" box="[304,325,437,462]" italics="true" pageId="12" pageNumber="23">tc</emphasis>
|
||
with 8 setae (2 considerably smaller than others);
|
||
<emphasis id="B97E1A1CFF8C3561FC1AFE51FC76FE59" box="[922,943,436,462]" italics="true" pageId="12" pageNumber="23">fe</emphasis>
|
||
with 8 setae (
|
||
<figureCitation id="1331DA8BFF8C3561FBD4FE51FB6AFE58" box="[1108,1203,436,463]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[197,1383,782,1888]" captionTargetId="figure-179@9.[189,1398,767,1924]" captionTargetPageId="9" captionText="Fig. 5. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-femur of right pedipalp. B. Tibia-tarsus 2 of left pedipalp.Abbreviations: see Material and methods. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906448" httpUri="https://zenodo.org/record/5906448/files/figure.png" pageId="12" pageNumber="23">Fig. 5A</figureCitation>
|
||
);
|
||
<emphasis id="B97E1A1CFF8C3561FB4AFE50FB03FE59" box="[1226,1242,437,462]" italics="true" pageId="12" pageNumber="23">ti</emphasis>
|
||
with 8 setae;
|
||
<emphasis id="B97E1A1CFF8C3561FF3DFE32FF28FE66" box="[189,241,471,497]" italics="true" pageId="12" pageNumber="23">bta1</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FEB2FE32FE82FE66" box="[306,347,471,497]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="23" unit="m" value="2.0">
|
||
2
|
||
<emphasis id="B97E1A1CFF8C3561FEC6FE3DFE82FE66" box="[326,347,472,497]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
and 1 normal seta;
|
||
<emphasis id="B97E1A1CFF8C3561FDBDFE32FDA8FE66" box="[573,625,471,497]" italics="true" pageId="12" pageNumber="23">bta2</emphasis>
|
||
with 2 normal setae and
|
||
<quantity id="4CF26BEBFF8C3561FC14FE32FC64FE66" box="[916,957,471,497]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="23" unit="m" value="4.0">
|
||
4
|
||
<emphasis id="B97E1A1CFF8C3561FC28FE3DFC64FE66" box="[936,957,472,497]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
;
|
||
<emphasis id="B97E1A1CFF8C3561FC4DFE32FC2BFE66" box="[973,1010,471,497]" italics="true" pageId="12" pageNumber="23">ta1</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FBB3FE32FB85FE66" box="[1075,1116,471,497]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="23" unit="m" value="2.0">
|
||
2
|
||
<emphasis id="B97E1A1CFF8C3561FBC7FE3DFB85FE66" box="[1095,1116,472,497]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
;
|
||
<emphasis id="B97E1A1CFF8C3561FBECFE32FB4BFE66" box="[1132,1170,471,497]" italics="true" pageId="12" pageNumber="23">ta2</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FB53FE32FB25FE66" box="[1235,1276,471,497]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="23" unit="m" value="6.0">
|
||
6
|
||
<emphasis id="B97E1A1CFF8C3561FB67FE3DFB25FE66" box="[1255,1276,472,497]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
(
|
||
<figureCitation id="1331DA8BFF8C3561FA8CFE32FAB1FE65" box="[1292,1384,471,498]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[197,1383,782,1888]" captionTargetId="figure-179@9.[189,1398,767,1924]" captionTargetPageId="9" captionText="Fig. 5. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-femur of right pedipalp. B. Tibia-tarsus 2 of left pedipalp.Abbreviations: see Material and methods. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906448" httpUri="https://zenodo.org/record/5906448/files/figure.png" pageId="12" pageNumber="23">Fig. 5B</figureCitation>
|
||
);
|
||
<emphasis id="B97E1A1CFF8C3561FF3CFE1EFF3BFD82" box="[188,226,507,533]" italics="true" pageId="12" pageNumber="23">ta3</emphasis>
|
||
with 1 long
|
||
<emphasis id="B97E1A1CFF8C3561FE02FE1FFE4FFD83" box="[386,406,506,532]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
, 1
|
||
<emphasis id="B97E1A1CFF8C3561FE41FE1EFE03FD83" box="[449,474,507,532]" italics="true" pageId="12" pageNumber="23">cs</emphasis>
|
||
with a conspicuous spine, 2
|
||
<emphasis id="B97E1A1CFF8C3561FCBDFE1EFC91FD83" box="[829,840,507,532]" italics="true" pageId="12" pageNumber="23">r</emphasis>
|
||
,
|
||
<quantity id="4CF26BEBFF8C3561FCDAFE1EFC4FFD83" box="[858,918,507,533]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="m" value="10.0">
|
||
10
|
||
<emphasis id="B97E1A1CFF8C3561FC01FE1EFC4FFD83" box="[897,918,507,532]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
(one macroseta with basal denticle and conspicuous spine) and 10 normal setae (
|
||
<figureCitation id="1331DA8BFF8C3561FD20FDFBFD26FDAF" box="[672,767,542,568]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[196,1355,474,1841]" captionTargetId="figure-64@11.[189,1398,466,1853]" captionTargetPageId="11" captionText="Fig. 7. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Tarsus 3 of left pedipalp. B. Basitarsus of right leg IV. C. First lobe of female genitalia. D. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5906452" httpUri="https://zenodo.org/record/5906452/files/figure.png" pageId="12" pageNumber="23">Fig. 7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8C3561FF3DFD85FB60FCBD" blockId="12.[189,1400,607,816]" pageId="12" pageNumber="23">
|
||
LEG I.
|
||
<emphasis id="B97E1A1CFF8C3561FE90FD85FEFCFDEE" box="[272,293,608,633]" italics="true" pageId="12" pageNumber="23">tc</emphasis>
|
||
with 12 normal setae (1 considerably smaller than others);
|
||
<emphasis id="B97E1A1CFF8C3561FC7BFDBAFBC9FDEE" box="[1019,1040,607,633]" italics="true" pageId="12" pageNumber="23">fe</emphasis>
|
||
with 9 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FA9BFD85FAE1FDEE" box="[1307,1336,608,633]" italics="true" pageId="12" pageNumber="23">pa</emphasis>
|
||
with 9 normal setae and 1
|
||
<emphasis id="B97E1A1CFF8C3561FE41FD67FE01FD0B" box="[449,472,642,668]" italics="true" pageId="12" pageNumber="23">tb</emphasis>
|
||
(
|
||
<figureCitation id="1331DA8BFF8C3561FE69FD67FD91FD0A" box="[489,584,642,669]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[197,1388,288,1858]" captionTargetId="figure-15@10.[189,1398,277,1888]" captionTargetPageId="10" captionText="Fig. 6. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-patella of right leg I. B. Tibia-basitarsus 2 of left leg I. C. Basitarsus 3-tarsus 2 of left leg I. D. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm." figureDoi="http://doi.org/10.5281/zenodo.5906450" httpUri="https://zenodo.org/record/5906450/files/figure.png" pageId="12" pageNumber="23">Fig. 6A</figureCitation>
|
||
);
|
||
<emphasis id="B97E1A1CFF8C3561FDE2FD66FDABFD0B" box="[610,626,643,668]" italics="true" pageId="12" pageNumber="23">ti</emphasis>
|
||
with 9 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FCF8FD67FC75FD0B" box="[888,940,642,668]" italics="true" pageId="12" pageNumber="23">bta1</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FC71FD67FBC7FD0B" box="[1009,1054,642,668]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="m" value="1.0">
|
||
1
|
||
<emphasis id="B97E1A1CFF8C3561FB8AFD66FBC7FD0B" box="[1034,1054,643,668]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
, 1 normal seta, 2
|
||
<emphasis id="B97E1A1CFF8C3561FB79FD67FAC9FD0B" box="[1273,1296,642,668]" italics="true" pageId="12" pageNumber="23">tb</emphasis>
|
||
and 1
|
||
<emphasis id="B97E1A1CFF8C3561FAE4FD67FAA1FD0B" box="[1380,1400,642,668]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
(with inner branch shorter than outer branch);
|
||
<emphasis id="B97E1A1CFF8C3561FD5AFD40FCD7FD28" box="[730,782,677,703]" italics="true" pageId="12" pageNumber="23">bta2</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FCD1FD43FCA5FD28" box="[849,892,678,704]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="23" unit="m" value="4.0">
|
||
4
|
||
<emphasis id="B97E1A1CFF8C3561FCE7FD43FCA5FD28" box="[871,892,678,703]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
, 2
|
||
<emphasis id="B97E1A1CFF8C3561FC20FD40FC6EFD28" box="[928,951,677,703]" italics="true" pageId="12" pageNumber="23">tb</emphasis>
|
||
and 1 long
|
||
<emphasis id="B97E1A1CFF8C3561FBC2FD43FB8FFD57" box="[1090,1110,678,704]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
(
|
||
<figureCitation id="1331DA8BFF8C3561FBE5FD40FB18FD57" box="[1125,1217,677,704]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[197,1388,288,1858]" captionTargetId="figure-15@10.[189,1398,277,1888]" captionTargetPageId="10" captionText="Fig. 6. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-patella of right leg I. B. Tibia-basitarsus 2 of left leg I. C. Basitarsus 3-tarsus 2 of left leg I. D. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm." figureDoi="http://doi.org/10.5281/zenodo.5906450" httpUri="https://zenodo.org/record/5906450/files/figure.png" pageId="12" pageNumber="23">Fig. 6B</figureCitation>
|
||
);
|
||
<emphasis id="B97E1A1CFF8C3561FB57FD40FAD5FD28" box="[1239,1292,677,703]" italics="true" pageId="12" pageNumber="23">bta3</emphasis>
|
||
with 1
|
||
<emphasis id="B97E1A1CFF8C3561FAE3FD43FAB7FD28" box="[1379,1390,678,703]" italics="true" pageId="12" pageNumber="23">r</emphasis>
|
||
, 1
|
||
<emphasis id="B97E1A1CFF8C3561FF53FD2CFF2FFD75" box="[211,246,713,738]" italics="true" pageId="12" pageNumber="23">grt</emphasis>
|
||
and 1 short normal seta;
|
||
<emphasis id="B97E1A1CFF8C3561FD9DFD2CFD88FD74" box="[541,593,713,739]" italics="true" pageId="12" pageNumber="23">bta4</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FD14FD2DFD67FD75" box="[660,702,712,739]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="23" unit="m" value="5.0">
|
||
5
|
||
<emphasis id="B97E1A1CFF8C3561FD29FD2CFD67FD75" box="[681,702,713,738]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
, 1
|
||
<emphasis id="B97E1A1CFF8C3561FD63FD2CFD23FD74" box="[739,762,713,739]" italics="true" pageId="12" pageNumber="23">tb</emphasis>
|
||
and 1 long
|
||
<emphasis id="B97E1A1CFF8C3561FC04FD2CFC41FD74" box="[900,920,713,739]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
;
|
||
<emphasis id="B97E1A1CFF8C3561FC27FD2CFC14FD74" box="[935,973,713,739]" italics="true" pageId="12" pageNumber="23">ta1</emphasis>
|
||
with 5 normal setae (2 considerably smaller than others);
|
||
<emphasis id="B97E1A1CFF8C3561FE39FD09FE07FC91" box="[441,478,748,774]" italics="true" pageId="12" pageNumber="23">ta2</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FDA2FD09FD97FC91" box="[546,590,748,775]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="23" unit="m" value="5.0">
|
||
5
|
||
<emphasis id="B97E1A1CFF8C3561FDB9FD08FD97FC91" box="[569,590,749,774]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
, 1
|
||
<emphasis id="B97E1A1CFF8C3561FDF5FD09FD55FC91" box="[629,652,748,774]" italics="true" pageId="12" pageNumber="23">tb</emphasis>
|
||
and 1 long
|
||
<emphasis id="B97E1A1CFF8C3561FC99FD09FCF4FC91" box="[793,813,748,774]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
(
|
||
<figureCitation id="1331DA8BFF8C3561FCBEFD09FC42FC90" box="[830,923,748,775]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[197,1388,288,1858]" captionTargetId="figure-15@10.[189,1398,277,1888]" captionTargetPageId="10" captionText="Fig. 6. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-patella of right leg I. B. Tibia-basitarsus 2 of left leg I. C. Basitarsus 3-tarsus 2 of left leg I. D. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm." figureDoi="http://doi.org/10.5281/zenodo.5906450" httpUri="https://zenodo.org/record/5906450/files/figure.png" pageId="12" pageNumber="23">Fig. 6C</figureCitation>
|
||
);
|
||
<emphasis id="B97E1A1CFF8C3561FC32FD09FC01FC91" box="[946,984,748,774]" italics="true" pageId="12" pageNumber="23">ta3</emphasis>
|
||
with 5
|
||
<emphasis id="B97E1A1CFF8C3561FBB3FD09FB9EFC91" box="[1075,1095,748,774]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
(with subequal branches) arranged as
|
||
<emphasis id="B97E1A1CFF8C3561FECBFCEAFEB1FCA7" box="[331,360,783,816]" italics="true" pageId="12" pageNumber="23">
|
||
fs
|
||
<subScript id="178EC44BFF8C3561FEDFFCC7FEB1FCA7" attach="left" box="[351,360,802,816]" fontSize="6" pageId="12" pageNumber="23">1</subScript>
|
||
</emphasis>
|
||
/
|
||
<emphasis id="B97E1A1CFF8C3561FEEFFCEAFE55FCA7" box="[367,396,783,816]" italics="true" pageId="12" pageNumber="23">
|
||
fs
|
||
<subScript id="178EC44BFF8C3561FE03FCC7FE55FCA7" attach="left" box="[387,396,802,816]" fontSize="6" pageId="12" pageNumber="23">2</subScript>
|
||
</emphasis>
|
||
/
|
||
<emphasis id="B97E1A1CFF8C3561FE14FCEAFE68FCA7" box="[404,433,783,816]" italics="true" pageId="12" pageNumber="23">
|
||
fs
|
||
<subScript id="178EC44BFF8C3561FE28FCC7FE68FCA7" attach="left" box="[424,433,802,816]" fontSize="6" pageId="12" pageNumber="23">3</subScript>
|
||
</emphasis>
|
||
/
|
||
<emphasis id="B97E1A1CFF8C3561FE39FCEAFE14FCBE" box="[441,461,783,809]" italics="true" pageId="12" pageNumber="23">fs</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8C3561FE78FCF5FDD6FCBE" box="[504,527,784,809]" italics="true" pageId="12" pageNumber="23">rs</emphasis>
|
||
(
|
||
<emphasis id="B97E1A1CFF8C3561FDA0FCF5FDEEFCBE" box="[544,567,784,809]" italics="true" pageId="12" pageNumber="23">rs</emphasis>
|
||
/
|
||
<emphasis id="B97E1A1CFF8C3561FDC0FCEAFD85FCA7" box="[576,604,783,816]" italics="true" pageId="12" pageNumber="23">
|
||
fs
|
||
<subScript id="178EC44BFF8C3561FDD3FCC7FD85FCA7" attach="left" box="[595,604,802,816]" fontSize="6" pageId="12" pageNumber="23">1</subScript>
|
||
</emphasis>
|
||
=2.2), 2
|
||
<emphasis id="B97E1A1CFF8C3561FD43FCF5FD17FCBE" box="[707,718,784,809]" italics="true" pageId="12" pageNumber="23">r</emphasis>
|
||
, 1
|
||
<emphasis id="B97E1A1CFF8C3561FD77FCF5FCC9FCBE" box="[759,784,784,809]" italics="true" pageId="12" pageNumber="23">cs</emphasis>
|
||
,
|
||
<quantity id="4CF26BEBFF8C3561FC9FFCEAFC81FCBE" box="[799,856,783,809]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="23" unit="m" value="13.0">
|
||
13
|
||
<emphasis id="B97E1A1CFF8C3561FCC3FCF5FC81FCBE" box="[835,856,784,809]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
and 5 normal setae (
|
||
<figureCitation id="1331DA8BFF8C3561FBCDFCEAFB72FCBD" box="[1101,1195,783,810]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[197,1388,288,1858]" captionTargetId="figure-15@10.[189,1398,277,1888]" captionTargetPageId="10" captionText="Fig. 6. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Trochanter-patella of right leg I. B. Tibia-basitarsus 2 of left leg I. C. Basitarsus 3-tarsus 2 of left leg I. D. Tarsus 3 of right leg I. Abbreviations: see Material and methods. Scale bars: A–C = 100 μm; D = 40 μm." figureDoi="http://doi.org/10.5281/zenodo.5906450" httpUri="https://zenodo.org/record/5906450/files/figure.png" pageId="12" pageNumber="23">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8C3561FE4CFCC7FE30FCA7" blockId="12.[189,1400,607,816]" box="[460,489,802,816]" pageId="12" pageNumber="23">
|
||
<emphasis id="B97E1A1CFF8C3561FE4CFCC7FE30FCA7" box="[460,489,802,816]" italics="true" pageId="12" pageNumber="23">
|
||
<subScript id="178EC44BFF8C3561FE4CFCC7FE30FCA7" attach="left" box="[460,489,802,816]" fontSize="6" pageId="12" pageNumber="23">4+5</subScript>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8C3561FF3DFCB4FAAEFC19" blockId="12.[189,1399,848,910]" pageId="12" pageNumber="23">
|
||
LEG II.
|
||
<emphasis id="B97E1A1CFF8C3561FE96FCB4FEF2FCFD" box="[278,299,849,874]" italics="true" pageId="12" pageNumber="23">tc</emphasis>
|
||
with 3 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FDACFCB4FD98FCFC" box="[556,577,849,875]" italics="true" pageId="12" pageNumber="23">fe</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FD05FCB4FD68FCFD" box="[645,689,849,875]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="m" value="1.0">
|
||
1
|
||
<emphasis id="B97E1A1CFF8C3561FD1CFCB4FD68FCFD" box="[668,689,849,874]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
and 4 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FC28FCB4FC1CFCFD" box="[936,965,849,874]" italics="true" pageId="12" pageNumber="23">pa</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8C3561FB89FCB4FBECFCFD" box="[1033,1077,849,875]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="23" unit="m" value="2.0">
|
||
2
|
||
<emphasis id="B97E1A1CFF8C3561FBA0FCB4FBECFCFD" box="[1056,1077,849,874]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
, 1 thick seta, and 2 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FE86FC91FECFFC1A" box="[262,278,884,909]" italics="true" pageId="12" pageNumber="23">ti</emphasis>
|
||
with 1 thick seta and 4 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FD45FC91FD33FC19" box="[709,746,884,910]" italics="true" pageId="12" pageNumber="23">bta</emphasis>
|
||
with 6 normal setae;
|
||
<emphasis id="B97E1A1CFF8C3561FC61FC91FC21FC1A" box="[993,1016,884,909]" italics="true" pageId="12" pageNumber="23">ta</emphasis>
|
||
with 1
|
||
<emphasis id="B97E1A1CFF8C3561FBCCFC91FB8EFC1A" box="[1100,1111,884,909]" italics="true" pageId="12" pageNumber="23">r</emphasis>
|
||
,
|
||
<quantity id="4CF26BEBFF8C3561FBE6FC91FB56FC1A" box="[1126,1167,884,910]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="23" unit="m" value="1.0">
|
||
1
|
||
<emphasis id="B97E1A1CFF8C3561FBFAFC91FB56FC1A" box="[1146,1167,884,909]" italics="true" pageId="12" pageNumber="23">m</emphasis>
|
||
</quantity>
|
||
and 9 normal setae.
|
||
</paragraph>
|
||
<caption id="DF759686FF8C3561FF3DF86DFB42F87E" ID-DOI="http://doi.org/10.5281/zenodo.5906454" ID-Zenodo-Dep="5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="12" pageNumber="23" startId="12.[189,232,1928,1954]" targetBox="[189,1398,1009,1888]" targetPageId="12">
|
||
<paragraph id="8BB5C60EFF8C3561FF3DF86DFB42F87E" blockId="12.[187,1399,1928,2025]" pageId="12" pageNumber="23">
|
||
<emphasis id="B97E1A1CFF8C3561FF3DF86DFED5F835" bold="true" box="[189,268,1928,1954]" pageId="12" pageNumber="23">Fig. 8.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF8C3561FE94F86DFD94F835" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[276,589,1928,1954]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="12" pageNumber="23" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8C3561FE94F86DFD94F835" box="[276,589,1928,1954]" italics="true" pageId="12" pageNumber="23">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8C3561FDD5F86DFD6AF835" box="[597,691,1928,1954]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, holotype (ISLA 50394).
|
||
<emphasis id="B97E1A1CFF8C3561FB87F86DFBC5F835" bold="true" box="[1031,1052,1928,1954]" pageId="12" pageNumber="23">A</emphasis>
|
||
. Chaetotaxy of opisthosomal tergite III.
|
||
<emphasis id="B97E1A1CFF8C3561FEB6F84EFE93F852" bold="true" box="[310,330,1963,1989]" pageId="12" pageNumber="23">B</emphasis>
|
||
. Chaetotaxy of opisthosomal sternite VI.
|
||
<emphasis id="B97E1A1CFF8C3561FCA9F84EFCE7F852" bold="true" box="[809,830,1963,1989]" pageId="12" pageNumber="23">C</emphasis>
|
||
. Opisthosomal segments VIII–XI.
|
||
<emphasis id="B97E1A1CFF8C3561FB56F84EFB32F852" bold="true" box="[1238,1259,1963,1989]" pageId="12" pageNumber="23">D</emphasis>
|
||
. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFEEFFD71FED0" blockId="13.[188,1398,265,327]" pageId="13" pageNumber="24">
|
||
LEG III.
|
||
<emphasis id="B97E1A1CFF8D3560FE9DFEEFFEEBFEB4" box="[285,306,266,291]" italics="true" pageId="13" pageNumber="24">tc</emphasis>
|
||
with 2 normal setae;
|
||
<emphasis id="B97E1A1CFF8D3560FDA9FEECFDE7FEB4" box="[553,574,265,291]" italics="true" pageId="13" pageNumber="24">fe</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FDCDFEEFFDB3FEB4" box="[589,618,266,291]" italics="true" pageId="13" pageNumber="24">pa</emphasis>
|
||
, and
|
||
<emphasis id="B97E1A1CFF8D3560FD28FEEFFD61FEB4" box="[680,696,266,291]" italics="true" pageId="13" pageNumber="24">ti</emphasis>
|
||
with 1 thick seta and 4 normal setae each;
|
||
<emphasis id="B97E1A1CFF8D3560FB27FEECFB15FEB4" box="[1191,1228,265,291]" italics="true" pageId="13" pageNumber="24">bta</emphasis>
|
||
with 6 normal setae;
|
||
<emphasis id="B97E1A1CFF8D3560FE87FEC8FEC7FED1" box="[263,286,301,326]" italics="true" pageId="13" pageNumber="24">ta</emphasis>
|
||
with 1
|
||
<emphasis id="B97E1A1CFF8D3560FEF7FEC8FE5BFED1" box="[375,386,301,326]" italics="true" pageId="13" pageNumber="24">r</emphasis>
|
||
,
|
||
<quantity id="4CF26BEBFF8D3560FE11FEC9FE65FED1" box="[401,444,300,326]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="24" unit="m" value="1.0">
|
||
1
|
||
<emphasis id="B97E1A1CFF8D3560FE27FEC8FE65FED1" box="[423,444,301,326]" italics="true" pageId="13" pageNumber="24">m</emphasis>
|
||
</quantity>
|
||
and 9 normal setae.
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFE8BFB70FE3C" blockId="13.[188,1399,366,427]" pageId="13" pageNumber="24">
|
||
LEG IV.
|
||
<emphasis id="B97E1A1CFF8D3560FE9BFE8BFEE9FE10" box="[283,304,366,391]" italics="true" pageId="13" pageNumber="24">tc</emphasis>
|
||
with 3 normal setae;
|
||
<emphasis id="B97E1A1CFF8D3560FDAAFE8BFDE6FE1F" box="[554,575,366,392]" italics="true" pageId="13" pageNumber="24">fe</emphasis>
|
||
with
|
||
<quantity id="4CF26BEBFF8D3560FD02FE8BFD75FE10" box="[642,684,366,392]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="24" unit="m" value="1.0">
|
||
1
|
||
<emphasis id="B97E1A1CFF8D3560FD17FE8BFD75FE10" box="[663,684,366,391]" italics="true" pageId="13" pageNumber="24">m</emphasis>
|
||
</quantity>
|
||
and 2 normal setae;
|
||
<emphasis id="B97E1A1CFF8D3560FC1DFE8BFC63FE10" box="[925,954,366,391]" italics="true" pageId="13" pageNumber="24">pa</emphasis>
|
||
and
|
||
<emphasis id="B97E1A1CFF8D3560FC72FE8BFBDBFE10" box="[1010,1026,366,391]" italics="true" pageId="13" pageNumber="24">ti</emphasis>
|
||
with 1 thick and 4 normal setae each;
|
||
<emphasis id="B97E1A1CFF8D3560FE82FE74FEFEFE3C" box="[258,295,401,427]" italics="true" pageId="13" pageNumber="24">bta</emphasis>
|
||
with
|
||
<emphasis id="B97E1A1CFF8D3560FEEAFE77FE54FE3C" box="[362,397,402,427]" italics="true" pageId="13" pageNumber="24">grt</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FE1CFE77FE7EFE3C" box="[412,423,402,427]" italics="true" pageId="13" pageNumber="24">r</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FE36FE77FE04FE3C" box="[438,477,402,427]" italics="true" pageId="13" pageNumber="24">esp</emphasis>
|
||
, and 2
|
||
<emphasis id="B97E1A1CFF8D3560FDB4FE74FD82FE3C" box="[564,603,401,427]" italics="true" pageId="13" pageNumber="24">esd</emphasis>
|
||
;
|
||
<emphasis id="B97E1A1CFF8D3560FDEBFE74FD48FE3C" box="[619,657,401,427]" italics="true" pageId="13" pageNumber="24">ta1</emphasis>
|
||
with 4 normal setae;
|
||
<emphasis id="B97E1A1CFF8D3560FC0EFE74FC6DFE3C" box="[910,948,401,427]" italics="true" pageId="13" pageNumber="24">ta2</emphasis>
|
||
with 8 normal setae.
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFE36FC02FD87" blockId="13.[189,1398,466,528]" pageId="13" pageNumber="24">
|
||
IVBTA. 7.2 × longer than wide and with 5 setae (
|
||
<emphasis id="B97E1A1CFF8D3560FD77FE36FCC3FE7B" box="[759,794,467,492]" italics="true" pageId="13" pageNumber="24">grt</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FCA9FE36FCEDFE7B" box="[809,820,467,492]" italics="true" pageId="13" pageNumber="24">r</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FCC4FE36FCB2FE7B" box="[836,875,467,492]" italics="true" pageId="13" pageNumber="24">esp</emphasis>
|
||
and 2
|
||
<emphasis id="B97E1A1CFF8D3560FC3CFE37FC3AFE7B" box="[956,995,466,492]" italics="true" pageId="13" pageNumber="24">esd</emphasis>
|
||
). Seta
|
||
<emphasis id="B97E1A1CFF8D3560FBB7FE36FB9BFE7B" box="[1079,1090,467,492]" italics="true" pageId="13" pageNumber="24">r</emphasis>
|
||
inserted in distal half (
|
||
<emphasis id="B97E1A1CFF8D3560FAD4FE37FAB7FE7B" box="[1364,1390,466,492]" italics="true" pageId="13" pageNumber="24">dr</emphasis>
|
||
/
|
||
<emphasis id="B97E1A1CFF8D3560FF3DFE10FF26FD98" box="[189,255,501,527]" italics="true" pageId="13" pageNumber="24">IVbta</emphasis>
|
||
=0.64) and
|
||
<emphasis id="B97E1A1CFF8D3560FE09FE13FE75FD98" box="[393,428,502,527]" italics="true" pageId="13" pageNumber="24">grt</emphasis>
|
||
inserted in proximal half of segment (
|
||
<figureCitation id="1331DA8BFF8D3560FCF1FE10FC17FD87" box="[881,974,501,528]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[196,1355,474,1841]" captionTargetId="figure-64@11.[189,1398,466,1853]" captionTargetPageId="11" captionText="Fig. 7. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Tarsus 3 of left pedipalp. B. Basitarsus of right leg IV. C. First lobe of female genitalia. D. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5906452" httpUri="https://zenodo.org/record/5906452/files/figure.png" pageId="13" pageNumber="24">Fig. 7B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFDD2FB44FC19" blockId="13.[189,1399,567,910]" pageId="13" pageNumber="24">
|
||
OPISTHOSOMA. Tergites II–VI with 3 +3 dorsal setae, two pairs of
|
||
<emphasis id="B97E1A1CFF8D3560FC3FFDDDFC1EFDC6" box="[959,967,568,593]" italics="true" pageId="13" pageNumber="24">t</emphasis>
|
||
setae (
|
||
<emphasis id="B97E1A1CFF8D3560FB9BFDDDFBF4FDCF" box="[1051,1069,568,600]" italics="true" pageId="13" pageNumber="24">
|
||
t
|
||
<subScript id="178EC44BFF8D3560FBA4FDAFFBF4FDCF" attach="left" box="[1060,1069,586,600]" fontSize="6" pageId="13" pageNumber="24">1</subScript>
|
||
</emphasis>
|
||
=35‒41 μm,
|
||
<emphasis id="B97E1A1CFF8D3560FB4CFDDDFB04FDCF" box="[1228,1245,568,600]" italics="true" pageId="13" pageNumber="24">
|
||
t
|
||
<subScript id="178EC44BFF8D3560FB54FDAFFB04FDCF" attach="left" box="[1236,1245,586,600]" fontSize="6" pageId="13" pageNumber="24">2</subScript>
|
||
</emphasis>
|
||
= 37‒42 μm) between a pair of slender setae (
|
||
<emphasis id="B97E1A1CFF8D3560FDBFFDBEFD93FDE3" box="[575,586,603,628]" italics="true" pageId="13" pageNumber="24">s</emphasis>
|
||
=27‒35 μm) (
|
||
<figureCitation id="1331DA8BFF8D3560FD76FDBFFC8CFDE3" box="[758,853,602,628]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">Fig. 8A</figureCitation>
|
||
). Sternite III with 2+ 2 setae. Sternites IV–VI each with 3+ 3 thickened setae (
|
||
<emphasis id="B97E1A1CFF8D3560FDC2FD9BFD83FD09" box="[578,602,638,670]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FDD1FD75FD83FD09" attach="left" box="[593,602,656,670]" fontSize="6" pageId="13" pageNumber="24">1</subScript>
|
||
</emphasis>
|
||
=30 μm,
|
||
<emphasis id="B97E1A1CFF8D3560FD50FD9BFD31FD09" box="[720,744,638,670]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FD5FFD75FD31FD09" attach="left" box="[735,744,656,670]" fontSize="6" pageId="13" pageNumber="24">2</subScript>
|
||
</emphasis>
|
||
=41‒45 μm,
|
||
<emphasis id="B97E1A1CFF8D3560FC0AFD9BFC7BFD09" box="[906,930,638,670]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FC19FD75FC7BFD09" attach="left" box="[921,930,656,670]" fontSize="6" pageId="13" pageNumber="24">3</subScript>
|
||
</emphasis>
|
||
=48‒53 μm) between a pair of slender setae (
|
||
<emphasis id="B97E1A1CFF8D3560FE89FD44FECDFD2D" box="[265,276,673,698]" italics="true" pageId="13" pageNumber="24">s</emphasis>
|
||
= 36‒40 μm) (inserted caudal to thick setae); pair of pores present between
|
||
<emphasis id="B97E1A1CFF8D3560FB0FFD44FB7FFD56" box="[1167,1190,673,705]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FB1DFD56FB7FFD56" attach="left" box="[1181,1190,691,705]" fontSize="6" pageId="13" pageNumber="24">1</subScript>
|
||
</emphasis>
|
||
setae on sternites IV‒VI (
|
||
<figureCitation id="1331DA8BFF8D3560FE9AFD21FEACFD49" box="[282,373,708,734]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">Fig. 8B</figureCitation>
|
||
). Presence of inconspicuous paired cavities in intersegmental furrows between sternites III–IV, IV–V, V–VI and VI–VII, one at each side of opisthosoma. Segments VII–X with 8 setae each (4 dorsal and 4 ventral). Segment XI elongated (1.15 × as long as wide), with dorsal row of 2+2 long setae (inner pair=160 μm; outer pair =117‒125 μm) inserted in its distal half and 2 pairs of ventral setae, 1 inserted around middle of segment (60 μm) and other inserted in apical region (75 μm) (
|
||
<figureCitation id="1331DA8BFF8D3560FA8AFCB5FAB0FCFC" box="[1290,1385,848,875]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">Fig. 8C</figureCitation>
|
||
). Intermediate ring of flagellum reduced in size (11 μm) and bears 2 tiny setae (8 μm).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFC53FBB0FBCB" blockId="13.[188,1399,949,1116]" pageId="13" pageNumber="24">
|
||
FEMALE GENITALIA. First lobe with 10 +10 setae in 5 transverse rows: 2+ 2 sternal setae (
|
||
<emphasis id="B97E1A1CFF8D3560FB37FC53FB0DFC41" box="[1207,1236,950,982]" italics="true" pageId="13" pageNumber="24">
|
||
st
|
||
<subScript id="178EC44BFF8D3560FB4BFC2DFB0DFC41" attach="left" box="[1227,1236,968,982]" fontSize="6" pageId="13" pageNumber="24">1</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FB61FC53FB27FC41" box="[1249,1278,950,982]" italics="true" pageId="13" pageNumber="24">
|
||
st
|
||
<subScript id="178EC44BFF8D3560FB75FC2DFB27FC41" attach="left" box="[1269,1278,968,982]" fontSize="6" pageId="13" pageNumber="24">2</subScript>
|
||
</emphasis>
|
||
) followed by 2+2, 1+1, 1 +1 and 4 +4 distal setae (
|
||
<emphasis id="B97E1A1CFF8D3560FD1EFC3CFD6CFC6E" box="[670,693,985,1017]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FD2CFC0EFD6CFC6E" attach="left" box="[684,693,1003,1017]" fontSize="6" pageId="13" pageNumber="24">1</subScript>
|
||
</emphasis>
|
||
=11 μm;
|
||
<emphasis id="B97E1A1CFF8D3560FCA4FC3CFCE5FC6E" box="[804,828,985,1017]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FCB3FC0EFCE5FC6E" attach="left" box="[819,828,1003,1017]" fontSize="6" pageId="13" pageNumber="24">2</subScript>
|
||
</emphasis>
|
||
= 12 μm;
|
||
<emphasis id="B97E1A1CFF8D3560FC2CFC3CFC1DFC6E" box="[940,964,985,1017]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FC3BFC0EFC1DFC6E" attach="left" box="[955,964,1003,1017]" fontSize="6" pageId="13" pageNumber="24">3</subScript>
|
||
</emphasis>
|
||
=20 μm;
|
||
<emphasis id="B97E1A1CFF8D3560FBB4FC3CFB95FC6E" box="[1076,1100,985,1017]" italics="true" pageId="13" pageNumber="24">
|
||
a
|
||
<subScript id="178EC44BFF8D3560FBC3FC0EFB95FC6E" attach="left" box="[1091,1100,1003,1017]" fontSize="6" pageId="13" pageNumber="24">4</subScript>
|
||
</emphasis>
|
||
=30 μm); interior surface of the first lobe with a group of 3 orifices on either side and a medial pair of small orifices (
|
||
<figureCitation id="1331DA8BFF8D3560FA8BFC1EFAB0FB81" box="[1291,1385,1019,1046]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[196,1355,474,1841]" captionTargetId="figure-64@11.[189,1398,466,1853]" captionTargetPageId="11" captionText="Fig. 7. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Tarsus 3 of left pedipalp. B. Basitarsus of right leg IV. C. First lobe of female genitalia. D. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5906452" httpUri="https://zenodo.org/record/5906452/files/figure.png" pageId="13" pageNumber="24">Fig. 7C</figureCitation>
|
||
). Second lobe with 3+2 setae, asymmetry caused by lack of 1
|
||
<emphasis id="B97E1A1CFF8D3560FC14FBC5FC78FBAE" box="[916,929,1056,1081]" italics="true" pageId="13" pageNumber="24">y</emphasis>
|
||
seta (
|
||
<emphasis id="B97E1A1CFF8D3560FC68FBC5FBD4FBAE" box="[1000,1037,1056,1081]" italics="true" pageId="13" pageNumber="24">x =</emphasis>
|
||
15 μm;
|
||
<emphasis id="B97E1A1CFF8D3560FBEDFBC5FB4BFBAE" box="[1133,1170,1056,1081]" italics="true" pageId="13" pageNumber="24">y =</emphasis>
|
||
29 μm;
|
||
<emphasis id="B97E1A1CFF8D3560FB72FBC5FACCFBAE" box="[1266,1301,1056,1081]" italics="true" pageId="13" pageNumber="24">z =</emphasis>
|
||
27 μm), and with cuticular spines; presence of group of 4 orifices on each half (
|
||
<figureCitation id="1331DA8BFF8D3560FC7DFBA7FB85FBCB" box="[1021,1116,1090,1116]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[196,1355,474,1841]" captionTargetId="figure-64@11.[189,1398,466,1853]" captionTargetPageId="11" captionText="Fig. 7. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Tarsus 3 of left pedipalp. B. Basitarsus of right leg IV. C. First lobe of female genitalia. D. Second lobe of female genitalia. Scale bars = 40 μm. Dotted circles and ellipses in figures C and D represent cuticular orifices. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5906452" httpUri="https://zenodo.org/record/5906452/files/figure.png" pageId="13" pageNumber="24">Fig. 7D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFB61FADDFAE6" blockId="13.[189,1399,1155,1393]" pageId="13" pageNumber="24">
|
||
FLAGELLUM. Arched and formed by 14 short and moniliform segments. First flagellar segment with 4 short setae (21–44 μm); second with 2 short setae (27–30 μm); third with 2 long setae (75 μm) and 4 short setae (10 – 30 μm); fourth with 3 short setae (15–31 μm); fifth with 5 short setae (25–33 μm); sixth with 2 very long setae; seventh with 1 short seta (15 μm) and 4 very long setae; eighth with 3 short setae (10–15 μm) and 2 very long setae; remaining segments bear 1 short seta (10–23 μm) and 4 very long setae each (
|
||
<figureCitation id="1331DA8BFF8D3560FEC3FAD6FE79FADA" box="[323,416,1331,1357]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">Fig. 8D</figureCitation>
|
||
). Very long setae inserted laterally on segments, being arranged in V along flagellum (
|
||
<figureCitation id="1331DA8BFF8D3560FF46FAB3FEFDFAE6" box="[198,292,1366,1393]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="13" pageNumber="24">Fig. 9H</figureCitation>
|
||
). Length (μm) and total number of setae (N) of flagellar articles available in
|
||
<tableCitation id="C688F3B5FF8D3560FB27FAB3FB26FAE6" box="[1191,1279,1366,1393]" captionStart="Table 1" captionStartId="3.[189,259,265,291]" captionTargetPageId="3" captionText="Table 1. Length (μm) and total number of setae (N) of the flagellar articles of Allokoenenia canhembora sp. nov. and length of the flagellar articles II–XIV of Allokoenenia afra Silvestri, 1913." httpUri="http://table.plazi.org/id/DF759686FF83356EFF3DFEECFB65FED1" pageId="13" pageNumber="24" tableUuid="DF759686FF83356EFF3DFEECFB65FED1">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF6AFA7DFE36FA25" blockId="13.[189,495,1432,1493]" box="[234,495,1432,1458]" pageId="13" pageNumber="24">
|
||
<emphasis id="B97E1A1CFF8D3560FF6AFA7DFE36FA25" bold="true" box="[234,495,1432,1458]" pageId="13" pageNumber="24">Male and immatures</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFA5EFEF3FA42" blockId="13.[189,495,1432,1493]" box="[189,298,1467,1493]" pageId="13" pageNumber="24">Unkown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3109585FF8D3560FF3DFA19FD80F9D2" box="[189,601,1532,1606]" pageId="13" pageNumber="24" type="distribution">
|
||
<paragraph id="8BB5C60EFF8D3560FF3DFA19FE81F981" blockId="13.[189,344,1532,1558]" box="[189,344,1532,1558]" pageId="13" pageNumber="24">
|
||
<heading id="D0FD7162FF8D3560FF3DFA19FE81F981" bold="true" box="[189,344,1532,1558]" fontSize="11" level="3" pageId="13" pageNumber="24" reason="3">
|
||
<emphasis id="B97E1A1CFF8D3560FF3DFA19FE81F981" bold="true" box="[189,344,1532,1558]" pageId="13" pageNumber="24">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3560FF3DF9C9FD80F9D2" blockId="13.[189,601,1579,1606]" box="[189,601,1579,1606]" pageId="13" pageNumber="24">
|
||
Known only from the
|
||
<typeStatus id="54B178ACFF8D3560FE42F9C9FE2CF9D1" box="[450,501,1580,1606]" pageId="13" pageNumber="24">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3109585FF8D3562FF3DF988FCC4FD8D" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24" type="discussion">
|
||
<paragraph id="8BB5C60EFF8D3560FF3DF988FEE9F910" blockId="13.[189,304,1645,1671]" box="[189,304,1645,1671]" pageId="13" pageNumber="24">
|
||
<heading id="D0FD7162FF8D3560FF3DF988FEE9F910" bold="true" box="[189,304,1645,1671]" fontSize="11" level="3" pageId="13" pageNumber="24" reason="3">
|
||
<emphasis id="B97E1A1CFF8D3560FF3DF988FEE9F910" bold="true" box="[189,304,1645,1671]" pageId="13" pageNumber="24">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8D3562FF3DF97EFCC4FD8D" blockId="13.[188,1400,1691,1999]" lastBlockId="15.[188,1399,265,538]" lastPageId="15" lastPageNumber="26" pageId="13" pageNumber="24">
|
||
<taxonomicName id="4C0ABD8DFF8D3560FF3DF97EFE2FF922" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[189,502,1691,1717]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8D3560FF3DF97EFE2FF922" box="[189,502,1691,1717]" italics="true" pageId="13" pageNumber="24">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8D3560FE7FF979FD82F921" box="[511,603,1692,1718]" pageId="13" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the genus
|
||
<taxonomicName id="4C0ABD8DFF8D3560FCE1F97EFC24F922" authorityName="Silvestri" authorityYear="1913" box="[865,1021,1691,1717]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="13" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E1A1CFF8D3560FCE1F97EFC24F922" box="[865,1021,1691,1717]" italics="true" pageId="13" pageNumber="24">Allokoenenia</emphasis>
|
||
</taxonomicName>
|
||
mainly because of the shape of the last opisthosomal segments and flagellum (
|
||
<figureCitation id="1331DA8BFF8D3560FD6CF95BFCA9F94F" box="[748,880,1726,1752]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">Figs 8C–D</figureCitation>
|
||
,
|
||
<figureCitation id="1331DA8BFF8D3560FCFEF95BFC1FF94F" box="[894,966,1726,1752]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="13" pageNumber="24">9F–H</figureCitation>
|
||
). It shares with
|
||
<taxonomicName id="4C0ABD8DFF8D3560FB06F95AFB00F94F" authorityName="Silvestri" authorityYear="1913" box="[1158,1241,1726,1753]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="afra">
|
||
<emphasis id="B97E1A1CFF8D3560FB06F95AFB00F94F" box="[1158,1241,1726,1753]" italics="true" pageId="13" pageNumber="24">A. afra</emphasis>
|
||
</taxonomicName>
|
||
the presence of a short and arched flagellum formed by 14 moniliform segments, the chaetotaxy of opisthosomal sternites IV–VI (1
|
||
<emphasis id="B97E1A1CFF8D3560FE1AF8E3FE7CF888" box="[410,421,1798,1823]" italics="true" pageId="13" pageNumber="24">s</emphasis>
|
||
+3
|
||
<emphasis id="B97E1A1CFF8D3560FE4CF8E3FE02F888" box="[460,475,1798,1823]" italics="true" pageId="13" pageNumber="24">a</emphasis>
|
||
+3
|
||
<emphasis id="B97E1A1CFF8D3560FD81F8E3FDC9F888" box="[513,528,1798,1823]" italics="true" pageId="13" pageNumber="24">a</emphasis>
|
||
+1
|
||
<emphasis id="B97E1A1CFF8D3560FDB6F8E3FD98F888" box="[566,577,1798,1823]" italics="true" pageId="13" pageNumber="24">s</emphasis>
|
||
), and the arrangement of the ventral setae of the last opisthosomal segment (in two rows) (
|
||
<figureCitation id="1331DA8BFF8D3560FE4FF8CDFDC3F8D5" box="[463,538,1832,1858]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[406,1188,1247,1911]" captionTargetId="figure-58@5.[382,1206,1224,1925]" captionTargetPageId="5" captionText="Fig. 2. Allokoenenia afra Silvestri, 1913, immature, cotypus, ventral view of opisthosomal segment XI (the insertions of the dorsal setae are represented in dashed line). Scale bar = 40 μm." figureDoi="http://doi.org/10.5281/zenodo.5906442" httpUri="https://zenodo.org/record/5906442/files/figure.png" pageId="13" pageNumber="24">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="1331DA8BFF8D3560FDA5F8CDFD93F8D5" box="[549,586,1832,1858]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,1009,1888]" captionTargetId="figure-380@12.[189,1398,1009,1888]" captionTargetPageId="12" captionText="Fig. 8. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Chaetotaxy of opisthosomal tergite III.B. Chaetotaxy of opisthosomal sternite VI.C. Opisthosomal segments VIII–XI. D. Flagellum. Abbreviations: see Material and methods. Scale bars: A–B = 20 μm; C–D = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5906454" httpUri="https://zenodo.org/record/5906454/files/figure.png" pageId="13" pageNumber="24">8C</figureCitation>
|
||
). However, they can be distinguished by the shape of the frontal organ (
|
||
<figureCitation id="1331DA8BFF8D3560FF44F8AEFEFDF8F2" box="[196,292,1867,1893]" captionStart="Fig" captionStartId="5.[189,232,1003,1029]" captionTargetBox="[189,1398,265,966]" captionTargetId="figure-11@5.[189,1398,265,966]" captionTargetPageId="5" captionText="Fig. 1. Allokoenenia afra Silvestri, 1913. A. Opisthosomal segments VIII–XI and first flagellar segment, ♀, typus. B. Flagellum (second to fourteenth flagellar segments), ♀, typus. C. Frontal organ, immature, cotypus. Scale bars: A–B = 100 μm; C = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5906440" httpUri="https://zenodo.org/record/5906440/files/figure.png" pageId="13" pageNumber="24">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation id="1331DA8BFF8D3560FEB2F8AEFE80F8F2" box="[306,345,1867,1893]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[222,1368,282,1855]" captionTargetId="figure-11@7.[209,1378,265,1855]" captionTargetPageId="7" captionText="Fig. 3. Allokoenenia canhembora sp. nov., ♀, holotype (ISLA 50394). A. Frontal organ. B. Blades of lateral organ. C. Coxa of left pedipalp. D. Coxa of left leg I. E. Coxa of right leg II. F. Coxa of left leg III. G. Coxa of right leg IV. Scale bars: A–B = 20 μm. C–G = 40 μm. Thickened setae on coxae II–IV indicated by arrows." figureDoi="http://doi.org/10.5281/zenodo.5906444" httpUri="https://zenodo.org/record/5906444/files/figure.png" pageId="13" pageNumber="24">3A</figureCitation>
|
||
), and by the number of blades on lateral organs (5 vs 1) and deutotritosternal setae (5 vs 1) (
|
||
<bibRefCitation id="EF9BBBFFFF8D3560FF68F88BFE45F81F" author="Silvestri F." box="[232,412,1902,1928]" pageId="13" pageNumber="24" pagination="211 - 217" refId="ref15863" refString="Silvestri F. 1913. Novi generi e specie de Koeneniidae (Arachnida, Palpigradi). Bollettino di Laboratorio di Entomologia Agraria Portici 7: 211 - 217." type="journal article" year="1913">Silvestri, 1913</bibRefCitation>
|
||
). Furthermore, we observed that the immature specimen of
|
||
<taxonomicName id="4C0ABD8DFF8D3560FBEFF88AFB1DF81F" authorityName="Silvestri" authorityYear="1913" box="[1135,1220,1902,1929]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="afra">
|
||
<emphasis id="B97E1A1CFF8D3560FBEFF88AFB1DF81F" box="[1135,1220,1902,1929]" italics="true" pageId="13" pageNumber="24">A. afra</emphasis>
|
||
</taxonomicName>
|
||
has 6 setae on basitarsus IV (
|
||
<emphasis id="B97E1A1CFF8D3560FEE6F877FE50F83C" box="[358,393,1938,1963]" italics="true" pageId="13" pageNumber="24">grt</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FE17F877FE7BF83C" box="[407,418,1938,1963]" italics="true" pageId="13" pageNumber="24">r</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FE30F874FE0FF83C" box="[432,470,1937,1963]" italics="true" pageId="13" pageNumber="24">gla</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8D3560FE64F877FDD2F83C" box="[484,523,1938,1963]" italics="true" pageId="13" pageNumber="24">esp</emphasis>
|
||
and 2
|
||
<emphasis id="B97E1A1CFF8D3560FDD8F874FDA6F83C" box="[600,639,1937,1963]" italics="true" pageId="13" pageNumber="24">esd</emphasis>
|
||
). This feature was not visible in the adults and was not described by
|
||
<bibRefCitation id="EF9BBBFFFF8D3560FF61F850FE42F858" author="Silvestri F." box="[225,411,1973,1999]" pageId="13" pageNumber="24" pagination="211 - 217" refId="ref15863" refString="Silvestri F. 1913. Novi generi e specie de Koeneniidae (Arachnida, Palpigradi). Bollettino di Laboratorio di Entomologia Agraria Portici 7: 211 - 217." type="journal article" year="1913">Silvestri (1913)</bibRefCitation>
|
||
, but they probably have at least 6 setae, since in general the setation of this article is already complete in immatures of this stage or they have fewer setae than the adults. In the new species, the basitarsus IV bears only 5 setae (
|
||
<emphasis id="B97E1A1CFF8F3562FDEEFEC8FD48FED1" box="[622,657,301,326]" italics="true" pageId="15" pageNumber="26">grt</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8F3562FD20FEC8FD72FED1" box="[672,683,301,326]" italics="true" pageId="15" pageNumber="26">r</emphasis>
|
||
,
|
||
<emphasis id="B97E1A1CFF8F3562FD3BFEC8FD3BFED1" box="[699,738,301,326]" italics="true" pageId="15" pageNumber="26">esp</emphasis>
|
||
and 2
|
||
<emphasis id="B97E1A1CFF8F3562FCB3FEC9FC83FED1" box="[819,858,300,326]" italics="true" pageId="15" pageNumber="26">esd</emphasis>
|
||
). Some flagellar characters can also be useful to distinguish these two species: the flagellar segments 1–3, 5, 7 and 9 of
|
||
<taxonomicName id="4C0ABD8DFF8F3562FB9BFEB5FBB4FEFE" authorityName="Silvestri" authorityYear="1913" box="[1051,1133,335,362]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="afra">
|
||
<emphasis id="B97E1A1CFF8F3562FB9BFEB5FBB4FEFE" box="[1051,1133,335,362]" italics="true" pageId="15" pageNumber="26">A. afra</emphasis>
|
||
</taxonomicName>
|
||
bear the typical apical cuticular spines present in the flagellum of other palpigrades, as well as setae of uniform shape despite showing some variation in their lengths, as represented in
|
||
<bibRefCitation id="EF9BBBFFFF8F3562FCE1FE73FB92FE27" author="Silvestri F." box="[865,1099,406,432]" pageId="15" pageNumber="26" pagination="211 - 217" refId="ref15863" refString="Silvestri F. 1913. Novi generi e specie de Koeneniidae (Arachnida, Palpigradi). Bollettino di Laboratorio di Entomologia Agraria Portici 7: 211 - 217." type="journal article" year="1913">Silvestri (1913: 216</bibRefCitation>
|
||
, fig. VI, 5) and confirmed by the study of the specimens (
|
||
<figureCitation id="1331DA8BFF8F3562FDABFE5CFD5FFE44" box="[555,646,441,467]" captionStart="Fig" captionStartId="5.[189,232,1003,1029]" captionTargetBox="[189,1398,265,966]" captionTargetId="figure-11@5.[189,1398,265,966]" captionTargetPageId="5" captionText="Fig. 1. Allokoenenia afra Silvestri, 1913. A. Opisthosomal segments VIII–XI and first flagellar segment, ♀, typus. B. Flagellum (second to fourteenth flagellar segments), ♀, typus. C. Frontal organ, immature, cotypus. Scale bars: A–B = 100 μm; C = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.5906440" httpUri="https://zenodo.org/record/5906440/files/figure.png" pageId="15" pageNumber="26">Fig. 1B</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C0ABD8DFF8F3562FD3DFE5CFCABFE44" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[701,882,441,467]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8F3562FD3DFE5CFCABFE44" box="[701,882,441,467]" italics="true" pageId="15" pageNumber="26">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8F3562FCFAFE5CFC0EFE44" box="[890,983,441,467]" pageId="15" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the cuticular spines are absent and the flagellar segments carry setae of very different lengths and shapes, including very short (around 10–15 μm) to very long setae (longer than 100 μm).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF759686FF8E3563FF3DF930FB77F871" ID-DOI="http://doi.org/10.5281/zenodo.5906456" ID-Zenodo-Dep="5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="14" pageNumber="25" startId="14.[189,232,1749,1775]" targetBox="[299,1296,275,1709]" targetPageId="14">
|
||
<paragraph id="8BB5C60EFF8E3563FF3DF930FB77F871" blockId="14.[189,1399,1749,2023]" pageId="14" pageNumber="25">
|
||
<emphasis id="B97E1A1CFF8E3563FF3DF930FED7F978" bold="true" box="[189,270,1749,1775]" pageId="14" pageNumber="25">Fig. 9.</emphasis>
|
||
<taxonomicName id="4C0ABD8DFF8E3563FE98F930FD8AF978" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[280,595,1749,1775]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8E3563FE98F930FD8AF978" box="[280,595,1749,1775]" italics="true" pageId="14" pageNumber="25">Allokoenenia canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8E3563FDDDF933FD62F967" box="[605,699,1750,1776]" pageId="14" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B97E1A1CFF8E3563FD44F930FD00F978" bold="true" box="[708,729,1749,1775]" pageId="14" pageNumber="25">A</emphasis>
|
||
. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of
|
||
<taxonomicName id="4C0ABD8DFF8E3563FB41F91CFAAFF884" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[1217,1398,1785,1811]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8E3563FB41F91CFAAFF884" box="[1217,1398,1785,1811]" italics="true" pageId="14" pageNumber="25">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8E3563FF3DF8F8FECEF8A0" box="[189,279,1821,1847]" pageId="14" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B97E1A1CFF8E3563FE9EF8F9FEEBF8A1" bold="true" box="[286,306,1820,1846]" pageId="14" pageNumber="25">B</emphasis>
|
||
. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate.
|
||
<emphasis id="B97E1A1CFF8E3563FEDAF8DAFEB7F8CE" bold="true" box="[346,366,1855,1881]" pageId="14" pageNumber="25">C</emphasis>
|
||
. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of
|
||
<taxonomicName id="4C0ABD8DFF8E3563FF5DF886FE4AF8EB" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[221,403,1890,1917]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="14" pageNumber="25" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8E3563FF5DF886FE4AF8EB" box="[221,403,1890,1917]" italics="true" pageId="14" pageNumber="25">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8E3563FE1BF886FE21F8EA" box="[411,504,1891,1917]" pageId="14" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were observed.
|
||
<emphasis id="B97E1A1CFF8E3563FD3AF887FD16F8EB" bold="true" box="[698,719,1890,1916]" pageId="14" pageNumber="25">D</emphasis>
|
||
. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface.
|
||
<emphasis id="B97E1A1CFF8E3563FDF0F863FD5DF837" bold="true" box="[624,644,1926,1952]" pageId="14" pageNumber="25">E</emphasis>
|
||
. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts.
|
||
<emphasis id="B97E1A1CFF8E3563FDBDF84DFDADF855" bold="true" box="[573,628,1960,1986]" pageId="14" pageNumber="25">F–G</emphasis>
|
||
. Live specimens photographed on a tray before being captured.
|
||
<emphasis id="B97E1A1CFF8E3563FF3DF829FF0DF871" bold="true" box="[189,212,1996,2022]" pageId="14" pageNumber="25">H</emphasis>
|
||
. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3109585FF8F357DFF3DFDA5FEF0FCC1" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26" type="biology_ecology">
|
||
<paragraph id="8BB5C60EFF8F3562FF3DFDA5FE6CFDCC" blockId="15.[189,437,576,603]" box="[189,437,576,603]" pageId="15" pageNumber="26">
|
||
<heading id="D0FD7162FF8F3562FF3DFDA5FE6CFDCC" bold="true" box="[189,437,576,603]" fontSize="11" level="3" pageId="15" pageNumber="26" reason="3">
|
||
<emphasis id="B97E1A1CFF8F3562FF3DFDA5FE6CFDCC" bold="true" box="[189,437,576,603]" pageId="15" pageNumber="26">Habitat and threats</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8F3562FF3DFD8AFB9FFB2B" blockId="15.[188,1400,623,1213]" pageId="15" pageNumber="26">
|
||
The Toca do Gonçalo Cave is associated to carbonatic rocks from the Caatinga geological group, which comprises relatively young rocks deposited around three million years ago under a lacustrine condition. Such rocks are closely inserted to old carbonatic rocks from the Una geological group, from the Neoproterozoic. The cave is located in a semiarid biome (the Caatinga Formation), in the Campo Formoso Municipality,
|
||
<collectingRegion id="49CE08ECFF8F3562FDAFFD19FD60FC81" box="[559,697,764,790]" country="Brazil" name="Bahia" pageId="15" pageNumber="26">Bahia State</collectingRegion>
|
||
, Northeastern
|
||
<collectingCountry id="F31D869EFF8F3562FCE4FD19FC75FC81" box="[868,940,764,790]" name="Brazil" pageId="15" pageNumber="26">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="1331DA8BFF8F3562FC3CFD19FBC5FC81" box="[956,1052,764,790]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9A</figureCitation>
|
||
). It presents around
|
||
<quantity id="4CF26BEBFF8F3562FA8CFD19FA8FFC80" box="[1292,1366,764,791]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="26" unit="m" value="500.0">500 m</quantity>
|
||
of topographed conduits and a single horizontal lenticular-shaped entrance (
|
||
<figureCitation id="1331DA8BFF8F3562FB9BFCFAFBAEFCAE" box="[1051,1143,799,825]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9B</figureCitation>
|
||
), which connects to a descending conduit. This cave presents two distinct levels: the upper level (which corresponds to the left branch of the cave), is predominantly dry, and the lower level (corresponding to the right branch) is quite humid (
|
||
<figureCitation id="1331DA8BFF8F3562FE98FC6CFEACFC34" box="[280,373,905,931]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9C</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EF9BBBFFFF8F3562FE0EFC6CFD37FC33" author="Souza-Silva M. & Ferreira R. L." box="[398,750,905,932]" pageId="15" pageNumber="26" pagination="1 - 21" refId="ref16231" refString="Souza-Silva M. & Ferreira R. L. 2016. The first two hotspots of subterranean biodiversity in South America. Subterranean Biology 19: 1 - 21. https: // doi. org / 10.3897 / subtbiol. 19.8207" type="journal article" year="2016">Souza-Silva & Ferreira 2016</bibRefCitation>
|
||
). The lower level used to be mostly submerged by the water table, but it is currently accessible, since the water table reduced dramatically, as will be discussed further on. The main sources of organic matter for both terrestrial and aquatic species is the dissolved and particulate organic matter imported from the external habitats during strong rains, but also the small guano deposits and some root mats that can be used as food by invertebrates. The Toca do Gonçalo Cave is considered a hotspot of subterranean biodiversity (only three hotspots are currently known to South America), presenting 22 troglobitic species (
|
||
<bibRefCitation id="EF9BBBFFFF8F3562FD55FBB9FBEEFBE0" author="Souza-Silva M. & Ferreira R. L." box="[725,1079,1116,1143]" pageId="15" pageNumber="26" pagination="1 - 21" refId="ref16231" refString="Souza-Silva M. & Ferreira R. L. 2016. The first two hotspots of subterranean biodiversity in South America. Subterranean Biology 19: 1 - 21. https: // doi. org / 10.3897 / subtbiol. 19.8207" type="journal article" year="2016">Souza-Silva & Ferreira 2016</bibRefCitation>
|
||
). Hostpots of subterranean biodiversity are those caves (or cave systems) with 20 or more cave-restricted species (
|
||
<bibRefCitation id="EF9BBBFFFF8F3562FB49FB9AFF27FB2A" author="Culver D. C. & Sket B." pageId="15" pageNumber="26" pagination="11 - 17" refId="ref15420" refString="Culver D. C. & Sket B. 2000. Hotspots of subterranean biodiversity in caves and wells. Journal of Cave and Karst Studies 62: 11 - 17." type="journal article" year="2000">Culver & Sket 2000</bibRefCitation>
|
||
). However, most troglobitic species from this cave remain undescribed.
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8F3562FF3DFB01FAD8F933" blockId="15.[188,1401,1252,1701]" pageId="15" pageNumber="26">
|
||
All individuals of
|
||
<taxonomicName id="4C0ABD8DFF8F3562FE1CFB01FD8AFB69" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[412,595,1252,1278]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8F3562FE1CFB01FD8AFB69" box="[412,595,1252,1278]" italics="true" pageId="15" pageNumber="26">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8F3562FDDEFB00FD62FB68" box="[606,699,1253,1279]" pageId="15" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were found at the lower cave level, always in very damp areas. In 2012, several specimens were observed, but only a single exemplar was captured since the cave substrata in the area prevented us to collect most of them. The water in karst systems usually presents high contents of dissolved calcium carbonates, which sometimes can form conspicuous deposits in the water surface inside caves. Such formations are usually called “calcite rafts” and are considered to be a kind of speleothem (
|
||
<figureCitation id="1331DA8BFF8F3562FE2FFA71FDD7FA39" box="[431,526,1428,1454]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9D</figureCitation>
|
||
). Since the water table in Toca do Gonçalo Cave used to fluctuate in time, the areas devoid of water usually present a muddy substrate covered by many layers of calcite rafts (
|
||
<figureCitation id="1331DA8BFF8F3562FF46FA3FFEFBFA63" box="[198,290,1498,1524]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9E</figureCitation>
|
||
). Most specimens of
|
||
<taxonomicName id="4C0ABD8DFF8F3562FDA0FA3EFD0CFA63" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[544,725,1498,1525]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF8F3562FDA0FA3EFD0CFA63" box="[544,725,1498,1525]" italics="true" pageId="15" pageNumber="26">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF8F3562FD5DFA3EFCE1FA62" box="[733,824,1499,1525]" pageId="15" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were found below those calcite rafts, so in most cases, as soon as we turned a piece of calcite, the specimen used to run away and hide in the uncountable small spaces between the layers, thus making their capture unfeasible. This is the reason why we decided to name the species “canhembora”, meaning ʻfugitiveʼ as explained in the etymology. A visit in 2013 revealed specimens (although in smaler number in comparison to 2012), but none could be captured. Finally, in 2014, we could find a single specimen that was successfully sampled (
|
||
<figureCitation id="1331DA8BFF8F3562FBF6F96FFB2CF933" box="[1142,1269,1674,1700]" captionStart="Fig" captionStartId="14.[189,232,1749,1775]" captionTargetBox="[299,1296,275,1709]" captionTargetId="figure-14@14.[290,1296,265,1712]" captionTargetPageId="14" captionText="Fig. 9. Allokoenenia canhembora sp. nov. A. Map of Brazil showing the placement of the Caatinga Biome, delimited in yellow, and of Toca do Gonçalo Cave (red dot), the type locality of A. canhembora sp. nov. B. Entrance of Toca do Gonçalo Cave and its surrounding landscape, characterized by the semiarid climate. C. Gallery of the lower level (right branch) of Toca do Gonçalo Cave, where individuals of A. canhembora sp. nov. were observed. D. Puddle of water inside the Gruta do Gonçalo Cave, with numerous calcite rafts on its surface. E. Area of the cave devoid of water with muddy substrate covered by many layers of calcite rafts. F–G. Live specimens photographed on a tray before being captured. H. Habitus of the specimen (ISLA 50395) captured in 2014 under a stereo microcope." figureDoi="http://doi.org/10.5281/zenodo.5906456" httpUri="https://zenodo.org/record/5906456/files/figure.png" pageId="15" pageNumber="26">Fig. 9F–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB5C60EFF8F357DFF3DF929FEF0FCC1" blockId="15.[188,1401,1740,2013]" lastBlockId="16.[188,1401,265,854]" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26">
|
||
All the cave visits from 2015 onward resulted in no further observations of specimens. Since the lower level of the cave was extremely dry, we believe that the specimens had probably migrated to inaccessible lower compartments. However, it is important to emphasize the fact that this species might be seriously threatened, as are other cave-restricted species from the Toca do Gonçalo Cave. This cave has been the target of several human impacts, which were highly intensified over the last years. In 1997, one of the authors (RLF) could attest that most part of the lower level was inaccessible due to the high level of the water table. In that period, the villagers of Gonçalo settlement used to draw water from the cave for their subsistence and this practice occurred during decades, by means of a diesel pump installed inside the cave. The villagers used to remove water from the cave once a week, thus the water table was persisting (although varying along the time), since the amount of water pumped out from the cave was negligible. However, an electric pump used by a local farmer for irrigation was installed in the cave in 2010 (
|
||
<bibRefCitation id="EF9BBBFFFF90357DFEA9FE97FDE0FE1A" author="Prevorcnik S. & Ferreira R. L. & Sket B." box="[297,569,370,397]" pageId="16" pageNumber="27" pagination="47 - 65" refId="ref15753" refString="Prevorcnik S., Ferreira R. L. & Sket B. 2012. Brasileirinidae, a new isopod family (Crustacea: Isopoda) from the cave in Bahia (Brazil) with a discussion on its taxonomic position. Zootaxa 3452: 47 - 65. https: // doi. org / 10.11646 / zootaxa. 3452.1.2" type="journal article" year="2012">
|
||
Prevorčnik
|
||
<emphasis id="B97E1A1CFF90357DFE35FE96FE2BFE1B" box="[437,498,370,396]" italics="true" pageId="16" pageNumber="27">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9BBBFFFF90357DFDC7FE97FC7CFE1A" author="Souza-Silva M. & Ferreira R. L." box="[583,933,370,397]" pageId="16" pageNumber="27" pagination="1 - 21" refId="ref16231" refString="Souza-Silva M. & Ferreira R. L. 2016. The first two hotspots of subterranean biodiversity in South America. Subterranean Biology 19: 1 - 21. https: // doi. org / 10.3897 / subtbiol. 19.8207" type="journal article" year="2016">Souza-Silva & Ferreira 2016</bibRefCitation>
|
||
). According to the villagers, this pump remained active during the whole day. In one of our surveys in 2010, a pronounced decrease of the water table (at least two meters) was observed. In 2012, the equipments were removed from the cave since the National Cave Research and Conservation
|
||
<collectingRegion id="49CE08ECFF90357DFD69FE39FCEEFE61" box="[745,823,476,502]" country="France" name="Centre" pageId="16" pageNumber="27">Center</collectingRegion>
|
||
intervened and asked the municipal government to dig an artesian well for the villagers. In
|
||
<date id="FFB4E0CEFF90357DFD45FE1AFC5CFD8E" box="[709,901,511,537]" pageId="16" pageNumber="27" value="2018-12">December 2018</date>
|
||
, a cave visit revealed that the water table was no longer observed. Hence, previously inaccessible areas could be explored, revealing the cave to be much longer than thought. However, from the 22 troglobitic species occurring in this cave, only two (an isopod and a springtail) were observed in this visit, in some moist areas located deep inside the cave. No specimens of
|
||
<taxonomicName id="4C0ABD8DFF90357DFE50FD68FD5CFD31" authority="Souza & Ferreira, 2022" authorityName="Souza & Ferreira" authorityYear="2022" box="[464,645,652,679]" class="Arachnida" family="Eukoeneniidae" genus="Allokoenenia" kingdom="Animalia" order="Palpigradi" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="canhembora" status="sp. nov.">
|
||
<emphasis id="B97E1A1CFF90357DFE50FD68FD5CFD31" box="[464,645,652,679]" italics="true" pageId="16" pageNumber="27">A. canhembora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DA767FF90357DFD0DFD68FD31FD30" box="[653,744,653,679]" pageId="16" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were observed, despite the intense efforts employed by the team in searching for them. The villagers mentioned that the building of a reservoir some dozens of kilometers from the cave, has altered the water balance, so that eventual flooding that used to reach the area never occurred again. Therefore, the Toca do Gonçalo Cave is seriously threatened and it is crucial that the Brazilian government intervene, in order to ensure the protection of this unique habitat in South America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |