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<document id="DB0471EC4668CBD1241D759C193B0D31" ID-CLB-Dataset="30106" ID-DOI="10.11646/zootaxa.4418.5.2" ID-GBIF-Dataset="76924ecc-2833-4231-97e9-fcbaf6bfcda7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1244980" ID-ZooBank="7E160945-52D3-460D-A1AA-B5890C09C10E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525974547493" checkinUser="plazi" docAuthor="Qian, Shuonan &amp; Li, Houhun" docDate="2018" docId="03A240729172FF8EFF72FEE7FD9ED0DF" docLanguage="en" docName="zootaxa.4418.5.2.pdf" docOrigin="Zootaxa 4418 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Commatarcha hamata Qian &amp; Li 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="444" masterDocId="FF9B380A917EFF82FFE5FFABFFAFD63D" masterDocTitle="Taxonomic review of the genus Commatarcha Meyrick (Lepidoptera: Carposinoidea: Carposinidae) from China, with descriptions of four new species" masterLastPageNumber="448" masterPageNumber="432" pageNumber="444" updateTime="1698543693336" updateUser="ExternalLinkService">
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<mods:title id="05E4692AAE9BCFDE0B8BD4B553F127FB">Taxonomic review of the genus Commatarcha Meyrick (Lepidoptera: Carposinoidea: Carposinidae) from China, with descriptions of four new species</mods:title>
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<mods:namePart id="B2E2959A07035622330EB5ECEA31C503">Qian, Shuonan</mods:namePart>
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<mods:namePart id="87C0E718C6D2A4B41706F73EB7CB3FCD">Li, Houhun</mods:namePart>
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<mods:title id="C7314D4B7F3F8FC9A17CFE8D2B16597F">Zootaxa</mods:title>
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<mods:date id="CB6B650FE6839EC9039DDFD1A0689987">2018</mods:date>
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<treatment id="03A240729172FF8EFF72FEE7FD9ED0DF" ID-DOI="http://doi.org/10.5281/zenodo.5979299" ID-GBIF-Taxon="143929667" ID-Zenodo-Dep="5979299" LSID="urn:lsid:plazi:treatment:03A240729172FF8EFF72FEE7FD9ED0DF" httpUri="http://treatment.plazi.org/id/03A240729172FF8EFF72FEE7FD9ED0DF" lastPageNumber="444" pageId="12" pageNumber="444">
<subSubSection id="C311A2EF9172FF8EFF72FEE7FE89D7BA" pageId="12" pageNumber="444" type="nomenclature">
<paragraph id="8BB4F1649172FF8EFF72FEE7FD9BD75B" blockId="12.[151,564,332,391]" box="[151,564,332,358]" pageId="12" pageNumber="444">
<heading id="D0FC46089172FF8EFF72FEE7FD9BD75B" bold="true" box="[151,564,332,358]" fontSize="11" level="1" pageId="12" pageNumber="444" reason="1">
<emphasis id="B97F2D769172FF8EFF72FEE7FD9BD75B" bold="true" box="[151,564,332,358]" pageId="12" pageNumber="444">
<taxonomicName id="4C0B8AE79172FF8EFF72FEE7FE66D75B" ID-CoL="BKSXY" authority="Li" authorityName="Qian &amp; Li" authorityYear="2018" box="[151,457,332,358]" class="Insecta" family="Carposinidae" genus="Commatarcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="hamata" status="sp. nov.">
<emphasis id="B97F2D769172FF8EFF72FEE7FE0AD75B" bold="true" box="[151,421,332,358]" italics="true" pageId="12" pageNumber="444">Commatarcha hamata</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="A24C900D9172FF8EFE33FEE7FD9BD75B" box="[470,564,332,358]" pageId="12" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4F1649172FF8EFF72FEC5FE89D7BA" blockId="12.[151,564,332,391]" box="[151,294,366,391]" pageId="12" pageNumber="444">
(
<figureCitation id="1330EDE19172FF8EFF7AFEC5FF5BD7BA" box="[159,244,366,391]" captionStart="FIGURES 714" captionStartId="4.[151,264,1868,1890]" captionTargetBox="[164,1419,217,1831]" captionTargetId="figure@4.[151,1436,193,1847]" captionTargetPageId="4" captionText="FIGURES 714. Adults of Commatarcha spp. 7, C. acidodes Diakonoff, ♂; 8, C. convoluta sp. nov., holotype, ♀; 9, C. fanjingshana Li, ♀; 10, C. guangxiensis Li, ♂; 11, C. hamata sp. nov., holotype, ♂; 12, C. palaeosema Meyrick, ♂; 13, C. rotundivalvasp. nov., holotype, ♂; 14, C. setiferaedeaga sp. nov., holotype, ♂ (Scale bars = 2.0 mm)." httpUri="https://zenodo.org/record/1244986/files/figure.png" pageId="12" pageNumber="444">Figs 11</figureCitation>
,
<figureCitation id="1330EDE19172FF8EFEE5FEC4FEB3D7BA" box="[256,284,367,391]" captionStart="FIGURES1520" captionStartId="5.[151,264,1911,1933]" captionTargetBox="[221,1351,209,1873]" captionTargetId="figure@5.[180,1374,193,1878]" captionTargetPageId="5" captionText="FIGURES1520. Malegenitalia of Commatarchaspp.15, C. acidodes Diakonoff, slideNo.QSN17469;16, C. guangxiensis Li, slide No. QSN17515; 17, C. hamata sp. nov., paratype, slide No. QSN17773; 18, C. palaeosema Meyrick, slide No. QSN17502; 19, C. rotundivalva sp. nov., paratype, slide No. QSN17503; 20, C. setiferaedeaga sp. nov., paratype, slide No. QSN17487 (Scale bars = 0.5 mm)." httpUri="https://zenodo.org/record/1244988/files/figure.png" pageId="12" pageNumber="444">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311A2EF9172FF8EFF72FE1DFEB3D462" pageId="12" pageNumber="444" type="materials_examined">
<paragraph id="8BB4F1649172FF8EFF72FE1DFD61D7CE" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<materialsCitation id="3B63FB399172FF8EFF72FE1DFD61D7CE" ID-GBIF-Occurrence="1844422386" collectingDate="2015-04-05" collectorName="Kaijian Teng &amp; No." country="China" county="Taiyanghe National Forest Park" elevation="1450" location="Pu'er City" municipality="Taiyanghe National Forest" pageId="12" pageNumber="444" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="B97F2D769172FF8EFF72FE1DFE15D7F2" bold="true" box="[151,442,438,463]" pageId="12" pageNumber="444">
<typeStatus id="54B04FC69172FF8EFF72FE1DFF7DD7F2" box="[151,210,438,463]" pageId="12" pageNumber="444">Type</typeStatus>
material.
<collectingCountry id="F31CB1F49172FF8EFEB1FE1DFE1AD7F2" box="[340,437,438,463]" name="China" pageId="12" pageNumber="444">CHINA</collectingCountry>
:
</emphasis>
<typeStatus id="54B04FC69172FF8EFE20FE1DFD82D7F2" box="[453,557,438,463]" pageId="12" pageNumber="444" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0D3AED9172FF8EFDDCFE1CFDFFD7F3" box="[569,592,439,462]" pageId="12" pageNumber="444" type="male"></specimenCount>
,
<collectingCounty id="62D589E89172FF8EFD85FE1DFC7DD7F2" box="[608,978,438,463]" pageId="12" pageNumber="444">
<collectingMunicipality id="6BD06B1E9172FF8EFD85FE1DFC3ED7F3" box="[608,913,438,463]" pageId="12" pageNumber="444">Taiyanghe National Forest</collectingMunicipality>
Park
</collectingCounty>
,
<location id="8ED4A7BF9172FF8EFC07FE1CFBF9D7F2" LSID="urn:lsid:plazi:treatment:03A240729172FF8EFF72FEE7FD9ED0DF:8ED4A7BF9172FF8EFC07FE1CFBF9D7F2" box="[994,1110,438,463]" country="China" county="Taiyanghe National Forest Park" municipality="Taiyanghe National Forest" name="Pu'er City" pageId="12" pageNumber="444" stateProvince="Yunnan Province">Pu'er City</location>
,
<collectingRegion id="49CF3F869172FF8EFB83FE1CFA81D7F2" box="[1126,1326,438,463]" country="China" name="Yunnan" pageId="12" pageNumber="444">Yunnan Province</collectingRegion>
,
<elevation id="002616579172FF8EFAD9FE1DFA38D7F3" box="[1340,1431,438,463]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="444" unit="m" value="1450.0">
<quantity id="4CF35C819172FF8EFAD9FE1DFA38D7F3" box="[1340,1431,438,463]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="444" unit="m" value="1450.0">1450 m</quantity>
</elevation>
,
<collectingDate id="EFF12E4C9172FF8EFF72FE71FEAED7CE" box="[151,257,474,499]" pageId="12" pageNumber="444" value="2015-04-05">
<date id="FFB5D7A49172FF8EFF72FE71FEAED7CE" box="[151,257,474,499]" pageId="12" pageNumber="444" value="2015-04-05">5.iv.2015</date>
</collectingDate>
, leg.
<collectorName id="26FE94B29172FF8EFED8FE71FE7FD7CE" box="[317,464,474,499]" pageId="12" pageNumber="444">Kaijian Teng</collectorName>
, slide
<collectorName id="26FE94B29172FF8EFDFDFE70FDEDD7CF" box="[536,578,475,498]" pageId="12" pageNumber="444">No.</collectorName>
QSN17499.
</materialsCitation>
</paragraph>
<paragraph id="8BB4F1649172FF8EFF22FE54FEB3D462" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<materialsCitation id="3B63FB399172FF8EFF22FE54FEB7D406" ID-GBIF-Occurrence="1844422381" collectingDate="2015-04-04" collectingDateMax="2015-04-08" collectingDateMin="2015-04-04" collectorName="Nos." country="China" county="Taiyanghe National Forest Park" elevation="1450" location="Pu'er City" municipality="Taiyanghe National Forest" pageId="12" pageNumber="444" specimenCount="5" specimenCount-male="5" stateProvince="Yunnan Province" typeStatus="paratype">
<typeStatus id="54B04FC69172FF8EFF22FE54FE9BD42A" box="[199,308,511,535]" pageId="12" pageNumber="444" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D0D3AED9172FF8EFEA2FE54FED8D42A" box="[327,375,511,535]" pageId="12" pageNumber="444" type="male">3 ♂</specimenCount>
):
<emphasis id="B97F2D769172FF8EFE6AFE55FDC7D42A" bold="true" box="[399,616,510,535]" pageId="12" pageNumber="444">
<collectingRegion id="49CF3F869172FF8EFE6AFE55FDCDD42A" box="[399,610,510,535]" country="China" name="Yunnan" pageId="12" pageNumber="444">Yunnan Province</collectingRegion>
.
</emphasis>
<specimenCount id="9D0D3AED9172FF8EFD97FE54FD0ED42B" box="[626,673,511,535]" pageId="12" pageNumber="444" type="male">2 ♂</specimenCount>
,
<collectingDate id="EFF12E4C9172FF8EFD55FE55FC99D42A" box="[688,822,510,535]" pageId="12" pageNumber="444" value="2015-04-04" valueMax="2015-04-08" valueMin="2015-04-04">
<date id="FFB5D7A49172FF8EFD55FE55FC99D42A" box="[688,822,510,535]" pageId="12" pageNumber="444" value="2015-04-04" valueMax="2015-04-08" valueMin="2015-04-04">48.iv.2015</date>
</collectingDate>
, other same data as holotype, slide
<collectorName id="26FE94B29172FF8EFB3CFE54FAA1D42B" box="[1241,1294,511,534]" pageId="12" pageNumber="444">Nos.</collectorName>
QSN17523, QSN17773
</materialsCitation>
;
<materialsCitation id="3B63FB399172FF8EFEC6FD88FEB7D462" ID-GBIF-Occurrence="1844422387" collectingDate="2016-07-13" collectorName="Kaijian Teng &amp; Ga-Eun Lee &amp; Tao Wang &amp; No." country="China" elevation="1450" location="Taiyanghe Reserve" pageId="12" pageNumber="444" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount id="9D0D3AED9172FF8EFEC6FD88FEE0D407" box="[291,335,547,571]" pageId="12" pageNumber="444" type="male">1 ♂</specimenCount>
,
<location id="8ED4A7BF9172FF8EFEBEFD89FD9CD407" LSID="urn:lsid:plazi:treatment:03A240729172FF8EFF72FEE7FD9ED0DF:8ED4A7BF9172FF8EFEBEFD89FD9CD407" box="[347,563,546,571]" country="China" name="Taiyanghe Reserve" pageId="12" pageNumber="444" stateProvince="Yunnan Province">Taiyanghe Reserve</location>
,
<elevation id="002616579172FF8EFDDBFD89FD3BD407" box="[574,660,546,571]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="444" unit="m" value="1450.0">
<quantity id="4CF35C819172FF8EFDDBFD89FD3BD407" box="[574,660,546,571]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="444" unit="m" value="1450.0">1450 m</quantity>
</elevation>
,
<collectingDate id="EFF12E4C9172FF8EFD45FD89FC8ED406" box="[672,801,546,571]" pageId="12" pageNumber="444" value="2016-07-13">
<date id="FFB5D7A49172FF8EFD45FD89FC8ED406" box="[672,801,546,571]" pageId="12" pageNumber="444" value="2016-07-13">13.vii.2016</date>
</collectingDate>
, leg.
<collectorName id="26FE94B29172FF8EFCB8FD89FC5ED406" box="[861,1009,546,571]" pageId="12" pageNumber="444">Kaijian Teng</collectorName>
,
<collectorName id="26FE94B29172FF8EFC1EFD88FB2CD407" box="[1019,1155,547,571]" pageId="12" pageNumber="444">Ga-Eun Lee</collectorName>
and
<collectorName id="26FE94B29172FF8EFB5CFD88FA82D406" box="[1209,1325,547,571]" pageId="12" pageNumber="444">Tao Wang</collectorName>
, slide
<collectorName id="26FE94B29172FF8EFA96FD88FA32D407" box="[1395,1437,547,570]" pageId="12" pageNumber="444">No.</collectorName>
QSN17507
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311A2EF9172FF8EFF22FDC1FEE9D52E" pageId="12" pageNumber="444" type="diagnosis">
<paragraph id="8BB4F1649172FF8EFF22FDC1FEE9D52E" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<emphasis id="B97F2D769172FF8EFF22FDC1FEEED4BE" bold="true" box="[199,321,618,643]" pageId="12" pageNumber="444">Diagnosis.</emphasis>
This species is similar to
<taxonomicName id="4C0B8AE79172FF8EFD8BFDC0FCB9D4BE" authorityName="Meyrick" authorityYear="1935" box="[622,790,619,643]" class="Insecta" family="Carposinidae" genus="Commatarcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="palaeosema">
<emphasis id="B97F2D769172FF8EFD8BFDC0FD2ED4BF" box="[622,641,619,642]" italics="true" pageId="12" pageNumber="444">C</emphasis>
.
<emphasis id="B97F2D769172FF8EFD74FDC0FCB9D4BE" box="[657,790,619,643]" italics="true" pageId="12" pageNumber="444">palaeosema</emphasis>
</taxonomicName>
in the forewing pattern and the male genitalia. It can be differentiated by the forewing lacking an ochreous yellow spot at basal 1/3 of the costal margin, and in the male genitalia by the saccus about 3/4 the length of the valva, and the ectophallus as long as height of the tegumen. In
<taxonomicName id="4C0B8AE79172FF8EFA67FD18FEB2D4D3" authorityName="Meyrick" authorityYear="1935" class="Insecta" family="Carposinidae" genus="Commatarcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="444" phylum="Arthropoda" rank="species" species="palaeosema">
<emphasis id="B97F2D769172FF8EFA67FD18FA3AD4F7" box="[1410,1429,691,714]" italics="true" pageId="12" pageNumber="444">C</emphasis>
.
<emphasis id="B97F2D769172FF8EFF72FD7DFEB2D4D3" box="[151,285,726,750]" italics="true" pageId="12" pageNumber="444">palaeosema</emphasis>
</taxonomicName>
, the saccus is about 4/5 the length of the valva, and the length of the ectophallus is about 3/4 the height of the tegumen.
</paragraph>
</subSubSection>
<subSubSection id="C311A2EF9172FF8EFF22FCB5FE3FD04B" pageId="12" pageNumber="444" type="description">
<paragraph id="8BB4F1649172FF8EFF22FCB5FB65D333" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<emphasis id="B97F2D769172FF8EFF22FCB5FEF6D50A" bold="true" box="[199,345,798,823]" pageId="12" pageNumber="444">Description.</emphasis>
Adult (
<figureCitation id="1330EDE19172FF8EFE57FCB5FDABD50A" box="[434,516,798,823]" captionStart="FIGURES 714" captionStartId="4.[151,264,1868,1890]" captionTargetBox="[164,1419,217,1831]" captionTargetId="figure@4.[151,1436,193,1847]" captionTargetPageId="4" captionText="FIGURES 714. Adults of Commatarcha spp. 7, C. acidodes Diakonoff, ♂; 8, C. convoluta sp. nov., holotype, ♀; 9, C. fanjingshana Li, ♀; 10, C. guangxiensis Li, ♂; 11, C. hamata sp. nov., holotype, ♂; 12, C. palaeosema Meyrick, ♂; 13, C. rotundivalvasp. nov., holotype, ♂; 14, C. setiferaedeaga sp. nov., holotype, ♂ (Scale bars = 2.0 mm)." httpUri="https://zenodo.org/record/1244986/files/figure.png" pageId="12" pageNumber="444">Fig. 11</figureCitation>
). Wingspan 10.014.0 mm. Head black, mixed with greyish white-tipped scales. Labial palpus black, extending forward; second segment greyish white on inner surface, slightly longer than diameter of eye; third segment about 1/3 length of second segment, greyish white at apex. Antenna with scape black on dorsal surface, greyish white on ventral surface; flagellum brown, darker on basal several flagellomeres, cilia on ventral surface same length as diameter of flagellum. Thorax and tegula black, with purplish blue reflection. Forewing relatively broad, widened from base to basal 1/4, parallel from basal 1/4 to distal 1/5, then narrowed triangularly to rounded apex; ground coloration greyish brown, with silvery grey scales; costal margin straight, black, with seven greyish white dots evenly spaced from basal 1/3 to apex; termen obliquely straight, black; with nine raised black scale tufts: three scale tufts arranged in a row, running from basal 1/6 below costal margin obliquely inward to basal 1/8 of 1A+2A above dorsum, other scale tufts scattered on median 1/3 of wing, tufts at upper and lower angles of cell edged with ochreous yellow scales on outer side, forming a large ochreous yellow stripe and a small ochreous yellow dot respectively, other tufts tipped with blue; black band from middle of R3 curved to tornus, sinuate along both outer and inner margins; fringe black, mixed with greyish white-tipped scales. Hindwing greyish brown; apex narrowly rounded, termen obliquely straight; fringe grey.
</paragraph>
<paragraph id="8BB4F1649172FF8EFF22FAB2FDBDD06F" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<emphasis id="B97F2D769172FF8EFF22FAB2FEDCD30F" bold="true" box="[199,371,1305,1330]" pageId="12" pageNumber="444">Male genitalia</emphasis>
(
<figureCitation id="1330EDE19172FF8EFE66FAB2FE77D30F" box="[387,472,1305,1330]" captionStart="FIGURES1520" captionStartId="5.[151,264,1911,1933]" captionTargetBox="[221,1351,209,1873]" captionTargetId="figure@5.[180,1374,193,1878]" captionTargetPageId="5" captionText="FIGURES1520. Malegenitalia of Commatarchaspp.15, C. acidodes Diakonoff, slideNo.QSN17469;16, C. guangxiensis Li, slide No. QSN17515; 17, C. hamata sp. nov., paratype, slide No. QSN17773; 18, C. palaeosema Meyrick, slide No. QSN17502; 19, C. rotundivalva sp. nov., paratype, slide No. QSN17503; 20, C. setiferaedeaga sp. nov., paratype, slide No. QSN17487 (Scale bars = 0.5 mm)." httpUri="https://zenodo.org/record/1244988/files/figure.png" pageId="12" pageNumber="444">Fig. 17</figureCitation>
). Uncus broadly sub-triangular, broadly rounded apically. Tegumen narrowly banded entirely. Valva sub-triangular, wide basally, abruptly narrowed to hooked apex; dorsobasal process elongately clubbed, extending obliquely outward, with long sparse setae; ventrobasal process wedge-shaped, wide at base, narrowed to rounded apex, longer than dorsobasal process and subparallel to it. Transtilla widely banded. Vinculum with length of lateral arm about two times height of tegumen; saccus short, about 3/4 length of valva, rounded at apex. Juxta broad and short, digitate, wider at base. Ectophallus triangular, about same length as height of tegumen, narrowed from base to approximately basal 2/3, distal 1/3 uniformly slender, acute apically. Aedeagus shorter than half length of saccus, rounded basally, dilated before apex; cornuti shaped as a cluster of differently sized large spines, placed distally.
</paragraph>
<paragraph id="8BB4F1649172FF8EFF22F9F6FE3FD04B" blockId="12.[151,1437,438,1762]" box="[199,400,1629,1654]" pageId="12" pageNumber="444">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C311A2EF9172FF8EFF22F92AFDB9D0A4" box="[199,534,1665,1690]" pageId="12" pageNumber="444" type="distribution">
<paragraph id="8BB4F1649172FF8EFF22F92AFDB9D0A4" blockId="12.[151,1437,438,1762]" box="[199,534,1665,1690]" pageId="12" pageNumber="444">
<emphasis id="B97F2D769172FF8EFF22F92AFECFD0A7" bold="true" box="[199,352,1665,1690]" pageId="12" pageNumber="444">Distribution.</emphasis>
<collectingCountry id="F31CB1F49172FF8EFE82F92AFE01D0A7" box="[359,430,1665,1690]" name="China" pageId="12" pageNumber="444">China</collectingCountry>
:
<collectingRegion id="49CF3F869172FF8EFE5CF929FDBDD0A4" box="[441,530,1666,1689]" country="China" name="Yunnan" pageId="12" pageNumber="444">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C311A2EF9172FF8EFF22F90EFD9ED0DF" pageId="12" pageNumber="444" type="etymology">
<paragraph id="8BB4F1649172FF8EFF22F90EFD9ED0DF" blockId="12.[151,1437,438,1762]" pageId="12" pageNumber="444">
<emphasis id="B97F2D769172FF8EFF22F90EFEE1D083" bold="true" box="[199,334,1701,1726]" pageId="12" pageNumber="444">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="B97F2D769172FF8EFC99F90EFC74D080" box="[892,987,1701,1725]" italics="true" pageId="12" pageNumber="444">hamatus</emphasis>
(hooked) referring to the valva hookshaped distally in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>