161 lines
20 KiB
XML
161 lines
20 KiB
XML
<document id="678B324BC6FBBE23F84FF413BFC0E497" ID-CLB-Dataset="56750" ID-DOI="10.5852/ejt.2022.821.1795" ID-GBIF-Dataset="98e75316-f8a2-4557-9e65-2bae83d36768" ID-ISSN="2118-9773" ID-Zenodo-Dep="6580782" ID-ZooBank="6DD16DFE-028A-4454-8DF7-64150836813E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653484400412" checkinUser="felipe" docAuthor="Sonar, Mohan A., Pawar, Ravi V. & Wayal, Dnyaneshwar V." docDate="2022" docId="039E8160DE5BFFE6FD3CF315FE6C4F3B" docLanguage="en" docName="ejt-821_16-39.pdf" docOrigin="European Journal of Taxonomy 821" docSource="http://zoobank.org/6dd16dfe-028a-4454-8df7-64150836813e" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Licornia undefined-1" docType="treatment" docVersion="6" lastPageNumber="21" masterDocId="FFA7F918DE5EFFE0FF82F25FFF924C5D" masterDocTitle="Newly discovered species of cheilostomatid Bryozoa from the Miocene of western Kachchh, Gujarat, India" masterLastPageNumber="39" masterPageNumber="16" pageNumber="21" updateTime="1699338043508" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="55BDDAC298D22255A057E010223C9B4D">Newly discovered species of cheilostomatid Bryozoa from the Miocene of western Kachchh, Gujarat, India</mods:title>
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<mods:name id="69C34EFC619894567D14037FBA7F312D" type="personal">
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<mods:namePart id="C9017C51B29570101B989FD971D341F1">Sonar, Mohan A.</mods:namePart>
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<mods:affiliation id="7330946850C0FA284912CD99661EF2A3">Department of Geology, Government Institute of Science, Nipat Niranjan Nagar, Caves Road, Aurangabad, Maharashtra 431004, India.</mods:affiliation>
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<mods:nameIdentifier id="760D4C2825044ABA9F1ADA5B768BD52E" type="email">drmohansonar398@gmail.com</mods:nameIdentifier>
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<mods:namePart id="717D854CA497A874AE1030C9AD8E2808">Pawar, Ravi V.</mods:namePart>
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<mods:affiliation id="6981F3E1932A0A4908B35A0239466AD3">Department of Geology, Government Institute of Science, Nipat Niranjan Nagar, Caves Road, Aurangabad, Maharashtra 431004, India.</mods:affiliation>
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<mods:nameIdentifier id="5DA0A91411A037640AB784AC4C4F76E1" type="email">ravi4geology@gmail.com</mods:nameIdentifier>
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<mods:namePart id="885E4B23C75C4F229E2C5A1496353DFF">Wayal, Dnyaneshwar V.</mods:namePart>
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<mods:affiliation id="489FE4E12931D08BD0A31D05448E6483">Department of Geology, Government Institute of Science, Nipat Niranjan Nagar, Caves Road, Aurangabad, Maharashtra 431004, India.</mods:affiliation>
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<mods:nameIdentifier id="0E5805E4E2FBC9A7BD8613E31819EC97" type="email">wayaldnyaneshwar@gmail.com</mods:nameIdentifier>
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<mods:title id="32E6E6044E6CEB015BD5280878D70E20">European Journal of Taxonomy</mods:title>
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<mods:part id="F71153D8B8A2689285BB65BA675B4392">
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<mods:date id="2EFE846C5CB542110254506FE39DFCE8">2022</mods:date>
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<mods:number id="68775E2CF3E749AC5CDDEF8CE7D40C26">2022-05-24</mods:number>
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<treatment id="039E8160DE5BFFE6FD3CF315FE6C4F3B" ID-DOI="http://doi.org/10.5281/zenodo.6581841" ID-GBIF-Taxon="195629175" ID-Zenodo-Dep="6581841" LSID="urn:lsid:zoobank.org:act:1A669348-F122-4A0C-AB7C-185D0E1BA860" httpUri="http://treatment.plazi.org/id/039E8160DE5BFFE6FD3CF315FE6C4F3B" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="21">
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<subSubSection id="C32D63FDDE5BFFE5FD3CF315FCE44D38" box="[702,886,330,357]" pageId="5" pageNumber="21" type="nomenclature">
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<paragraph id="8B883076DE5BFFE5FD3CF315FCE44D38" blockId="5.[702,886,330,392]" box="[702,886,330,357]" pageId="5" pageNumber="21">
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<emphasis id="B943EC64DE5BFFE5FD3CF315FD5F4D39" bold="true" box="[702,717,330,356]" pageId="5" pageNumber="21">?</emphasis>
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<taxonomicName id="4C374BF5DE5BFFE5FD4FF315FCE44D38" box="[717,886,330,357]" class="Gymnolaemata" family="Candidae" genus="Licornia" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="21" phylum="Bryozoa" rank="species" species="undefined-1">
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<emphasis id="B943EC64DE5BFFE5FD4FF315FCA44D39" bold="true" box="[717,822,330,356]" italics="true" pageId="5" pageNumber="21">Licornia</emphasis>
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sp. 1
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D63FDDE5BFFE5FD47F331FCFD4DD5" box="[709,879,366,392]" pageId="5" pageNumber="21" type="description">
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<paragraph id="8B883076DE5BFFE5FD47F331FCFD4DD5" blockId="5.[702,886,330,392]" box="[709,879,366,392]" pageId="5" pageNumber="21">
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<figureCitation id="130C2CF3DE5BFFE5FD47F331FC994DD5" box="[709,779,366,392]" captionStart="Fig" captionStartId="5.[191,234,1928,1954]" captionTargetBox="[189,1398,1190,1888]" captionTargetId="figure-237@5.[189,1398,1190,1888]" captionTargetPageId="5" captionText="Fig. 3.?Licornia sp. 1 (GIS/B 0539). A. Frontal view of the colony autozooids, ooecium and avicularia. B. Enlargement of the colony showing autozooids and ooecium. C. Close-up of a single autozooid with ooecium having the ectooecium perforated by unequal sized pores." figureDoi="http://doi.org/10.5281/zenodo.6580790" httpUri="https://zenodo.org/record/6580790/files/figure.png" pageId="5" pageNumber="21">Fig. 3</figureCitation>
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;
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<tableCitation id="C6B505CDDE5BFFE5FC9AF331FCFD4DD5" box="[792,879,366,392]" captionStart="Table 2" captionStartId="6.[189,259,265,291]" captionTargetPageId="6" captionText="Table 2. Measurements in mm of?Licornia sp. 1 (GIS/B 0439), Burdigalian, lower Miocene, Chhasra Formation, western Kachchh, Gujarat, India. Abbreviations: see Material and methods." pageId="5" pageNumber="21">Table 2</tableCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32D63FDDE5BFFE5FF3FF3F0FAD34E46" pageId="5" pageNumber="21" type="materials_examined">
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<paragraph id="8B883076DE5BFFE5FF3FF3F0FE3E4D94" blockId="5.[189,428,431,457]" box="[189,428,431,457]" pageId="5" pageNumber="21">
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<heading id="D0C0871ADE5BFFE5FF3FF3F0FE3E4D94" bold="true" box="[189,428,431,457]" fontSize="11" level="3" pageId="5" pageNumber="21" reason="3">
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<emphasis id="B943EC64DE5BFFE5FF3FF3F0FE3E4D94" bold="true" box="[189,428,431,457]" pageId="5" pageNumber="21">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B883076DE5BFFE5FF3FF381FAD34E46" blockId="5.[189,1399,478,539]" pageId="5" pageNumber="21">
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<materialsCitation id="3B5F3A2BDE5BFFE5FF3FF381FAAF4E46" ID-GBIF-Occurrence="3783537302" collectedFrom="fossiliferous limestone at cliff section" collectingDate="2012-01" collectingDateMax="2012-02" collectingDateMin="2012-01" collectionCode="GIS/B" collectorName="Sonar" country="India" latitude="23.371944" location="Lakdi Nadi" longLatPrecision="21" longitude="68.969444" pageId="5" pageNumber="21" specimenCode="0439" specimenCount="1">
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<collectingCountry id="F32070E6DE5BFFE5FF3FF381FE9C4DA5" box="[189,270,478,504]" name="India" pageId="5" pageNumber="21">INDIA</collectingCountry>
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•
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<collectedFrom id="382B738BDE5BFFE5FEACF381FD674DA5" box="[302,757,478,504]" pageId="5" pageNumber="21">fossiliferous limestone at cliff section</collectedFrom>
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of
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<location id="8EE866ADDE5BFFE5FCA1F381FC394DA5" LSID="urn:lsid:plazi:treatment:039E8160DE5BFFE6FD3CF315FE6C4F3B:8EE866ADDE5BFFE5FCA1F381FC394DA5" box="[803,939,478,504]" country="India" county="Chhasra Formation" latitude="23.371944" longLatPrecision="21" longitude="68.969444" municipality="lower Miocene" name="Lakdi Nadi" pageId="5" pageNumber="21">Lakdi Nadi</location>
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near
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<location id="8EE866ADDE5BFFE5FC70F381FB1A4DA5" LSID="urn:lsid:plazi:treatment:039E8160DE5BFFE6FD3CF315FE6C4F3B:8EE866ADDE5BFFE5FC70F381FB1A4DA5" box="[1010,1160,478,504]" country="India" county="Chhasra Formation" latitude="23.371944" longLatPrecision="21" longitude="68.969444" municipality="lower Miocene" name="Tera Village" pageId="5" pageNumber="21">Tera Village</location>
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||
;
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<geologicalTimeScale id="F917B0DDDE5BFFE5FB15F381FAB74DA5" box="[1175,1317,478,504]" pageId="5" pageNumber="21">Burdigalian</geologicalTimeScale>
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,
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<geologicalTimeScale id="F917B0DDDE5BFFE5FAB6F381FEB54E46" pageId="5" pageNumber="21">lower Miocene</geologicalTimeScale>
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, Chhasra Formation;
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<geoCoordinate id="EE0356B1DE5BFFE5FDA1F3A0FD2B4E46" box="[547,697,511,539]" degrees="23" direction="north" minutes="22" orientation="latitude" pageId="5" pageNumber="21" precision="15" seconds="19" value="23.371944">23°22′19″ N</geoCoordinate>
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,
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<geoCoordinate id="EE0356B1DE5BFFE5FD47F3A0FCCA4E46" box="[709,856,511,539]" degrees="68" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="21" precision="15" seconds="10" value="68.969444">68°58′10″ E</geoCoordinate>
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||
;
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<date id="FF8916B6DE5BFFE5FCE6F05EFB884E46" box="[868,1050,513,539]" pageId="5" pageNumber="21" value="2012-01" valueMax="2012-02-31" valueMin="2012-01-00">
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<collectingDate id="EFCDEF5EDE5BFFE5FCE6F05EFB884E46" box="[868,1050,513,539]" pageId="5" pageNumber="21" value="2012-01" valueMax="2012-02-31" valueMin="2012-01-00">Jan.–Feb. 2012</collectingDate>
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</date>
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||
;
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<collectorName id="26C255A0DE5BFFE5FBA4F05EFBF84E46" box="[1062,1130,513,539]" pageId="5" pageNumber="21">Sonar</collectorName>
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leg.;
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<collectionCode id="ED26A8B3DE5BFFE5FB2FF05EFB654E46" box="[1197,1271,513,539]" pageId="5" pageNumber="21">GIS/B</collectionCode>
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<specimenCode id="DB91980DDE5BFFE5FB7DF05EFAAF4E46" box="[1279,1341,513,539]" pageId="5" pageNumber="21">0439</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D63FDDE5BFFE5FF3FF01DFBDA4FDC" pageId="5" pageNumber="21" type="description">
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<paragraph id="8B883076DE5BFFE5FF3FF01DFEDD4E01" blockId="5.[189,335,578,604]" box="[189,335,578,604]" pageId="5" pageNumber="21">
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<heading id="D0C0871ADE5BFFE5FF3FF01DFEDD4E01" bold="true" box="[189,335,578,604]" fontSize="11" level="3" pageId="5" pageNumber="21" reason="3">
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<emphasis id="B943EC64DE5BFFE5FF3FF01DFEDD4E01" bold="true" box="[189,335,578,604]" pageId="5" pageNumber="21">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B883076DE5BFFE5FF3FF02EFBDA4FDC" blockId="5.[187,1400,625,898]" pageId="5" pageNumber="21">
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Colony erect biserial, branching. Autozooids club-shaped, arranged in alternate longitudinal rows (
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<figureCitation id="130C2CF3DE5BFFE5FF44F0CBFEB54EF3" box="[198,295,660,686]" captionStart="Fig" captionStartId="5.[191,234,1928,1954]" captionTargetBox="[189,1398,1190,1888]" captionTargetId="figure-237@5.[189,1398,1190,1888]" captionTargetPageId="5" captionText="Fig. 3.?Licornia sp. 1 (GIS/B 0539). A. Frontal view of the colony autozooids, ooecium and avicularia. B. Enlargement of the colony showing autozooids and ooecium. C. Close-up of a single autozooid with ooecium having the ectooecium perforated by unequal sized pores." figureDoi="http://doi.org/10.5281/zenodo.6580790" httpUri="https://zenodo.org/record/6580790/files/figure.png" pageId="5" pageNumber="21">Fig. 3A</figureCitation>
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). Cryptocyst barely developed, when present descending into opesia. Gymnocyst extended for ⅓ of frontal surface, occupied by ooecia of the proximal zooid if fertile (
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<figureCitation id="130C2CF3DE5BFFE5FBCBF0E8FB344E8C" box="[1097,1190,695,721]" captionStart="Fig" captionStartId="5.[191,234,1928,1954]" captionTargetBox="[189,1398,1190,1888]" captionTargetId="figure-237@5.[189,1398,1190,1888]" captionTargetPageId="5" captionText="Fig. 3.?Licornia sp. 1 (GIS/B 0539). A. Frontal view of the colony autozooids, ooecium and avicularia. B. Enlargement of the colony showing autozooids and ooecium. C. Close-up of a single autozooid with ooecium having the ectooecium perforated by unequal sized pores." figureDoi="http://doi.org/10.5281/zenodo.6580790" httpUri="https://zenodo.org/record/6580790/files/figure.png" pageId="5" pageNumber="21">Fig. 3B</figureCitation>
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). Opesia rounded to oval. Scutum not preserved. Distal oral spine bases stout, usually two internal, distal spine bases obscured by the ovicells (
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<figureCitation id="130C2CF3DE5BFFE5FE63F0A2FDAE4F45" box="[481,572,765,792]" captionStart="Fig" captionStartId="5.[191,234,1928,1954]" captionTargetBox="[189,1398,1190,1888]" captionTargetId="figure-237@5.[189,1398,1190,1888]" captionTargetPageId="5" captionText="Fig. 3.?Licornia sp. 1 (GIS/B 0539). A. Frontal view of the colony autozooids, ooecium and avicularia. B. Enlargement of the colony showing autozooids and ooecium. C. Close-up of a single autozooid with ooecium having the ectooecium perforated by unequal sized pores." figureDoi="http://doi.org/10.5281/zenodo.6580790" httpUri="https://zenodo.org/record/6580790/files/figure.png" pageId="5" pageNumber="21">Fig. 3C</figureCitation>
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). Lateral avicularia large, on outer distolateral corner of each autozooid, laterally directed, with acute rostrum, with somewhat raised tip. A frontal vibracula obliquely placed distolaterally with small rootlet foramen. Ooecia broad, conspicuous, slightly tilted away from axis of branch, ectooecium smooth, perforated by 12–15 unequal sized pseudopores.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D63FDDE5BFFE6FF3FF1F6FE6C4F3B" lastPageId="6" lastPageNumber="22" pageId="5" pageNumber="21" type="discussion">
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||
<paragraph id="8B883076DE5BFFE5FF3FF1F6FEA24F9E" blockId="5.[189,304,937,963]" box="[189,304,937,963]" pageId="5" pageNumber="21">
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||
<heading id="D0C0871ADE5BFFE5FF3FF1F6FEA24F9E" bold="true" box="[189,304,937,963]" fontSize="11" level="3" pageId="5" pageNumber="21" reason="3">
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<emphasis id="B943EC64DE5BFFE5FF3FF1F6FEA24F9E" bold="true" box="[189,304,937,963]" pageId="5" pageNumber="21">Remarks</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B883076DE5BFFE6FF3FF188FE6C4F3B" blockId="5.[189,1399,983,1116]" lastBlockId="6.[189,1400,738,870]" lastPageId="6" lastPageNumber="22" pageId="5" pageNumber="21">
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A single, weathered specimen was available to study, which does not allow to confirm the presence or details of a scutum or oral spine bases, the presence of frontal avicularia or axial vibracula. Hence, the species is left in open nomenclature until better preserved material is available. The present species differs from the Recent Indo-Pacific species
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<taxonomicName id="4C374BF5DE5BFFE5FD50F61DFBD94801" authority="(Busk, 1852)" baseAuthorityName="Busk" baseAuthorityYear="1852" box="[722,1099,1089,1116]" class="Gymnolaemata" family="Candidae" genus="Licornia" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="21" phylum="Bryozoa" rank="species" species="diadema">
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<emphasis id="B943EC64DE5BFFE5FD50F61DFC354806" box="[722,935,1089,1115]" italics="true" pageId="5" pageNumber="21">Licornia diadema</emphasis>
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(
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<bibRefCitation id="EFA64D87DE5BFFE5FC3BF61EFBD14801" author="Busk G." box="[953,1091,1089,1116]" pageId="5" pageNumber="21" pagination="343 - 402" refId="ref6953" refString="Busk G. 1852. An account of the Polyzoa, and sertularian zoophytes, collected in the Voyage of the Rattlesnake, on the coasts of Australia and the Louisiade Archipelago. In: MacGillivray J. (ed.) Narrative of the Voyage of the H. M. S. Rattlesnake 1: 343 - 402." type="journal article" year="1852">Busk, 1852</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the absence of frontal avicularia, comparatively large lateral avicularia and the absence of a raised ooecial rim (see
|
||
<bibRefCitation id="EFA64D87DE58FFE6FB6DF0BDFE1D4F7D" author="Vieira L. M. & Spencer J. M. & Winston J. E." pageId="6" pageNumber="22" pagination="1911 - 1921" refId="ref8871" refString="Vieira L. M., Spencer J. M. & Winston J. E. 2013. Resurrection of the genus Licornia for Scrupocellaria jolloisii (Bryozoa) and related species, with documentation of L. jolloisii as a non-indigenous species in the western Atlantic. Journal of the Marine Biological Association of the United Kingdom 93 (7): 1911 - 1921. https: // doi. org / 10.1017 / S 0025315413000301" type="journal article" year="2013">
|
||
Vieira
|
||
<emphasis id="B943EC64DE58FFE6FAB9F0BCFAE44EA1" box="[1339,1398,738,764]" italics="true" pageId="6" pageNumber="22">et al.</emphasis>
|
||
2013: 1914–1915
|
||
</bibRefCitation>
|
||
, fig. 3e–f). The Recent south Pacific Australian species
|
||
<taxonomicName id="4C374BF5DE58FFE6FBA6F159FEC14F1E" authority="Tilbrook & Vieira, 2012" authorityName="Tilbrook & Vieira" authorityYear="2012" class="Gymnolaemata" family="Candidae" genus="Licornia" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="prolata">
|
||
<emphasis id="B943EC64DE58FFE6FBA6F159FB7B4F42" box="[1060,1257,773,799]" italics="true" pageId="6" pageNumber="22">Licornia prolata</emphasis>
|
||
<bibRefCitation id="EFA64D87DE58FFE6FB71F15AFEC14F1E" author="Tilbrook K. J. & Vieira L. M." pageId="6" pageNumber="22" pagination="29 - 48" refId="ref8832" refString="Tilbrook K. J. & Vieira L. M. 2012. Scrupocellaria (Bryozoa: Cheilostomata) from the Queensland coast, with the description of three new species. Zootaxa 3528: 29 - 48." type="journal article" year="2012">Tilbrook & Vieira, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
differs in the presence of frontal avicularia, small lateral avicularia and the number of frontal pores on the ooecia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |