treatments-xml/data/03/9D/F1/039DF108000E242FF5C144C374DFFF46.xml

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<mods:title id="7E8004E00007CE85FD9FC63A4C029DF7">Labuanium vitatum (Crustacea: Decapoda: Brachyura: Sesarmidae), a new Indo-West Pacific species of arboreal crab</mods:title>
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<mods:namePart id="2D52BB79A0B453C318565BB8DD827FE9">Ng, Peter K. L.</mods:namePart>
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<mods:date id="0276FC01814785380F6EE7E27B81C641">2011</mods:date>
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<emphasis id="B9409C0C000E2424F5C144C37571FA24" bold="true" box="[151,524,1437,1462]" pageId="2" pageNumber="37">
<taxonomicName id="4C343B9D000E2424F5C144C37601FA24" ID-CoL="7WTQY" box="[151,380,1437,1462]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000E2424F5C144C37601FA24" bold="true" box="[151,380,1437,1462]" italics="true" pageId="2" pageNumber="37">Labuanium vitatum</emphasis>
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(
<figureCitation id="130F5C9B000E2424F5C9449F7791FA4B" box="[159,236,1473,1497]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1968]" captionTargetBox="[261,1325,193,1924]" captionTargetId="figure@3.[261,1325,193,1925]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life of Labuanium vitatum new species. A, B, female (35.2 × 33.6 mm) (WAM C 258 - 89), Greta Beach, Christmas I (courtesy Hugh Yorkston).; C, male (39.5 × 38.5 mm) (WAM C 257 - 89), The Dales, Christmas I (courtesy Peter Greenaway)." httpUri="https://zenodo.org/record/206069/files/figure.png" pageId="2" pageNumber="37">Figs. 1</figureCitation>
, 
<figureCitation id="130F5C9B000E2424F5AC449F7675FA4B" box="[250,264,1473,1497]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="2" pageNumber="37">2</figureCitation>
A, 3A, 4A, 5A, B, 6A, 7, 8, 10A–G)
</paragraph>
</subSubSection>
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<paragraph id="8B8B401E000E2424F5C14754762DF9AC" blockId="2.[151,1436,1545,1752]" pageId="2" pageNumber="37">
<treatmentCitationGroup id="AB246730000E2424F5C14754762DF9AC" pageId="2" pageNumber="37">
<taxonomicName id="4C343B9D000E2424F5C147547619F98D" box="[151,356,1546,1567]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="oceanicum">
<emphasis id="B9409C0C000E2424F5C147547619F98D" box="[151,356,1546,1567]" italics="true" pageId="2" pageNumber="37">Sesarma oceanicum</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0A95660F000E2424F42847577546F98D" author="Alcock" box="[382,571,1545,1567]" page="423" pageId="2" pageNumber="37" year="1900">
<bibRefCitation id="EFA53DEF000E2424F4284757757BF98D" author="Alcock" box="[382,518,1545,1567]" pageId="2" pageNumber="37" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (3), 279 - 486." type="journal article" year="1900">Alcock 1900</bibRefCitation>
: 423
</treatmentCitation>
; 
<treatmentCitation id="0A95660F000E2424F71147547588F98D" author="George" box="[583,757,1546,1567]" page="14" pageId="2" pageNumber="37" year="1978">
<bibRefCitation id="EFA53DEF000E2424F711475475ADF98D" author="George" box="[583,720,1546,1567]" pageId="2" pageNumber="37" refString="George, R. W. (1978) The Land and Freshwater Crabs of Christmas Island. Unpublished report commissioned by the Australian National Parks and Wildlife Service, 22 pp." type="book" year="1978">George 1978</bibRefCitation>
: 14
</treatmentCitation>
[not 
<taxonomicName id="4C343B9D000E2424F666475473F6F98D" ID-CoL="7Z29T" authority="De Man, 1889" authorityName="De Man" authorityYear="1889" box="[816,1163,1546,1567]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="oceanica">
<emphasis id="B9409C0C000E2424F66647547496F98D" box="[816,1003,1546,1567]" italics="true" pageId="2" pageNumber="37">Sesarma oceanica</emphasis>
<collectingCountry id="F323008E000E2424F6A24754736CF98D" box="[1012,1041,1546,1567]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="37">De</collectingCountry>
<bibRefCitation id="EFA53DEF000E2424F14D475473F6F98D" author="Man" box="[1051,1163,1546,1567]" pageId="2" pageNumber="37" refString="Man, J. G. De (1889) Ueber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452, pls. 9, 10." type="journal article" year="1889">
<collectingCountry id="F323008E000E2424F14D47547331F98D" box="[1051,1100,1546,1567]" name="Isle of Man" pageId="2" pageNumber="37">Man</collectingCountry>
, 1889
</bibRefCitation>
</taxonomicName>
= 
<taxonomicName id="4C343B9D000E2424F1FD47577636F9AC" ID-CoL="7VK9K" authority="Hess, 1865" authorityName="Hess" authorityYear="1865" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B9409C0C000E2424F1FD475772E1F98D" box="[1195,1436,1545,1567]" italics="true" pageId="2" pageNumber="37">Labuanium rotundatum</emphasis>
(
<bibRefCitation id="EFA53DEF000E2424F5984776763EF9AC" author="Hess" box="[206,323,1576,1598]" pageId="2" pageNumber="37" refString="Hess, W. (1865) Beitrage zur Kenntnis der Decapoden-Krebse Ost-Australiens. Archiv fur Naturgeschicht e, 36, 127 - 173, pls. 1, 2." type="journal article" year="1865">Hess, 1865</bibRefCitation>
)
</taxonomicName>
]
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<treatmentCitationGroup id="AB246730000E2424F5C1471976C1F9E9" pageId="2" pageNumber="37">
<taxonomicName id="4C343B9D000E2424F5C147197548F9CF" box="[151,565,1607,1629]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="rotundata" subGenus="Episesarma" subSpecies="papuo">
<emphasis id="B9409C0C000E2424F5C147197793F9CE" box="[151,238,1607,1628]" italics="true" pageId="2" pageNumber="37">Sesarma</emphasis>
(
<emphasis id="B9409C0C000E2424F4564716760AF9CF" box="[256,375,1608,1629]" italics="true" pageId="2" pageNumber="37">Episesarma</emphasis>
) 
<emphasis id="B9409C0C000E2424F4DF47197548F9CF" box="[393,565,1607,1629]" italics="true" pageId="2" pageNumber="37">rotundata papuo</emphasis>
</taxonomicName>
-malesiaca— 
<treatmentCitation id="0A95660F000E2424F7E84718740CF9CE" author="Nobili" box="[702,881,1606,1628]" page="510" pageId="2" pageNumber="37" year="1900">
<bibRefCitation id="EFA53DEF000E2424F7E847187442F9CE" author="Nobili" box="[702,831,1606,1628]" pageId="2" pageNumber="37" refString="Nobili, G. (1900) Decapodi e Stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale &quot; G. Doria &quot;, 40, 473 - 523." type="journal article" year="1900">Nobili 1900</bibRefCitation>
: 510
</treatmentCitation>
(part). [not 
<taxonomicName id="4C343B9D000E2424F6A3471972EEF9CF" box="[1013,1427,1607,1629]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subSpecies" species="rotundata" subGenus="Episesarma" subSpecies="papuo">
<emphasis id="B9409C0C000E2424F6A347197331F9CE" box="[1013,1100,1607,1628]" italics="true" pageId="2" pageNumber="37">Sesarma</emphasis>
(
<emphasis id="B9409C0C000E2424F108471673ABF9CF" box="[1118,1238,1608,1629]" italics="true" pageId="2" pageNumber="37">Episesarma</emphasis>
) 
<emphasis id="B9409C0C000E2424F1BE471972EEF9CF" box="[1256,1427,1607,1629]" italics="true" pageId="2" pageNumber="37">rotundata papuo</emphasis>
</taxonomicName>
- malesiaca 
<bibRefCitation id="EFA53DEF000E2424F462473B76CBF9E9" author="Nobili" box="[308,438,1637,1659]" pageId="2" pageNumber="37" refString="Nobili, G. (1899) Contribuzioni alla conoscienza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali dei Museo Civico di Storia Naturale di Genova, (2) 20, 230 - 282." type="journal article" year="1899">Nobili, 1899</bibRefCitation>
]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B401E000E2424F5C147DA7598F908" blockId="2.[151,1436,1545,1752]" box="[151,741,1667,1690]" pageId="2" pageNumber="37">
<treatmentCitationGroup id="AB246730000E2424F5C147DA7598F908" box="[151,741,1667,1690]" pageId="2" pageNumber="37">
<taxonomicName id="4C343B9D000E2424F5C147DA76BBF908" ID-CoL="7Z28P" box="[151,454,1668,1690]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="rotundata" subGenus="Sesarma">
<emphasis id="B9409C0C000E2424F5C147DA7793F90B" box="[151,238,1668,1689]" italics="true" pageId="2" pageNumber="37">Sesarma</emphasis>
(
<emphasis id="B9409C0C000E2424F5AB47DA7629F90B" box="[253,340,1668,1689]" italics="true" pageId="2" pageNumber="37">Sesarma</emphasis>
) 
<emphasis id="B9409C0C000E2424F43547DA76BBF908" box="[355,454,1668,1690]" italics="true" pageId="2" pageNumber="37">rotundata</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0A95660F000E2424F48947DD75FBF90B" author="Tesch" box="[479,646,1667,1689]" page="193" pageId="2" pageNumber="37" year="1917">
<bibRefCitation id="EFA53DEF000E2424F48947DD752BF90B" author="Tesch" box="[479,598,1667,1689]" pageId="2" pageNumber="37" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, 3, 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch 1917</bibRefCitation>
: 193
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B401E000E2424F5C147FC754DF945" blockId="2.[151,1436,1545,1752]" pageId="2" pageNumber="37">
<treatmentCitationGroup id="AB246730000E2424F5C147FC754DF945" pageId="2" pageNumber="37">
<taxonomicName id="4C343B9D000E2424F5C147FC76FBF92A" ID-CoL="7VK9K" box="[151,390,1698,1720]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B9409C0C000E2424F5C147FC76FBF92A" box="[151,390,1698,1720]" italics="true" pageId="2" pageNumber="37">Labuanium rotundatum</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0A95660F000E2424F4F647FC75E6F92A" author="Serene" box="[416,667,1698,1720]" page="401" pageId="2" pageNumber="37" year="1970">
<bibRefCitation id="EFA53DEF000E2424F4F647FC7517F92A" author="Serene" box="[416,618,1698,1720]" pageId="2" pageNumber="37" refString="Serene, R. &amp; Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say, 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène &amp; Soh 1970</bibRefCitation>
: 401
</treatmentCitation>
(part); 
<treatmentCitation id="0A95660F000E2424F7BE47FC74BCF92A" author="Hicks" box="[744,961,1698,1720]" page="64" pageId="2" pageNumber="37" year="1984">
<bibRefCitation id="EFA53DEF000E2424F7BE47FC74E6F92A" author="Hicks" box="[744,923,1698,1720]" pageId="2" pageNumber="37" refString="Hicks, J., Rumpff, H. &amp; Yorkston, H. (1984) Christmas Crabs. Christmas Island Natural History Association, 1990 second edition, 81 pp." type="book" year="1984">
Hicks 
<emphasis id="B9409C0C000E2424F67D47FD7423F92A" box="[811,862,1698,1720]" italics="true" pageId="2" pageNumber="37">et al.</emphasis>
1984
</bibRefCitation>
: 64
</treatmentCitation>
; 
<treatmentCitation id="0A95660F000E2424F69B47FC7300F92A" author="Davie" box="[973,1149,1698,1720]" page="221" pageId="2" pageNumber="37" year="2002">
<bibRefCitation id="EFA53DEF000E2424F69B47FC7330F92A" author="Davie" box="[973,1101,1698,1720]" pageId="2" pageNumber="37" refString="Davie, P. J. F. (2002) C rustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia. 19.3 B. Pp. 1 - 641. CSIRO Publications." type="book chapter" year="2002">Davie, 2002</bibRefCitation>
: 221
</treatmentCitation>
(part); 
<treatmentCitation id="0A95660F000E2424F19D47FD72E1F92A" author="Ng" box="[1227,1436,1698,1720]" page="614" pageId="2" pageNumber="37" year="2003">
<bibRefCitation id="EFA53DEF000E2424F19D47FD7211F92A" author="Ng" box="[1227,1388,1698,1720]" pageId="2" pageNumber="37" refString="Jeng, M. - S., Liu, H. - C., Tzeng, C. - S. &amp; Ng, P. K. L. (2003) On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana, 76 (2), 227 - 240." type="journal article" year="2003">Ng &amp; Liu 2003</bibRefCitation>
: 614
</treatmentCitation>
(part); Ng 
<emphasis id="B9409C0C000E2424F464479C7618F945" box="[306,357,1729,1751]" italics="true" pageId="2" pageNumber="37">et al.</emphasis>
2008: 221 (in part).
</treatmentCitationGroup>
</paragraph>
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<emphasis id="B9409C0C000E2424F5C1465D76FDF88E" bold="true" box="[151,384,1795,1820]" pageId="2" pageNumber="37">Material examined.</emphasis>
<typeStatus id="548FFEBC000E2424F4D1465A7692F88E" box="[391,495,1796,1820]" pageId="2" pageNumber="37" type="holotype">Holotype</typeStatus>
male (34.8 × 
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) (
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), 
<quantity id="4CCCEDFB000E2424F69D465A7379F88E" box="[971,1028,1796,1820]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="37" unit="m" value="60.0">60 m</quantity>
above beach, on tree trunk, on track to West White Beach, Christmas I., coll. D. Merton, 
<date id="FF8A66DE000E2424F7B4467974D7F8AD" box="[738,938,1831,1855]" pageId="2" pageNumber="37" value="1978-02-11">11 February 1978</date>
. 
<typeStatus id="548FFEBC000E2424F6E34676735BF8D2" box="[949,1062,1832,1856]" pageId="2" pageNumber="37" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D328B97000E2424F16746797306F8AD" box="[1073,1147,1831,1855]" pageId="2" pageNumber="37" type="male">1 male</specimenCount>
(39.5 × 
<quantity id="4CCCEDFB000E2424F18C4679723DF8AD" box="[1242,1344,1831,1855]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="2" pageNumber="37" unit="mm" value="38.5">38.5 mm</quantity>
) (
<collectionCode id="ED25D8DB000E2424F003467972E1F8AD" LSID="urn:lsid:biocol.org:col:34981" box="[1365,1436,1831,1855]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="37">WAM</collectionCode>
C257-89), The Dales, Christmas I., coll. H. Yorkston, ca. 
<date id="FF8A66DE000E2424F674461374C1F8F6" box="[802,956,1868,1893]" pageId="2" pageNumber="37" value="1989-01">January 1989</date>
; 
<specimenCount id="9D328B97000E2424F69E46127357F8F6" box="[968,1066,1868,1892]" pageId="2" pageNumber="37" type="female">1 female</specimenCount>
(35.2 × 
<quantity id="4CCCEDFB000E2424F1DD4612738CF8F6" box="[1163,1265,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="3.36" pageId="2" pageNumber="37" unit="mm" value="33.6">33.6 mm</quantity>
) (
<collectionCode id="ED25D8DB000E2424F05E46127232F8F6" LSID="urn:lsid:biocol.org:col:34981" box="[1288,1359,1868,1892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="37">WAM</collectionCode>
C258- 89), Greta Beach, Christmas I., coll. H. Yorkston, 
<date id="FF8A66DE000E2424F782463174D2F815" box="[724,943,1903,1927]" pageId="2" pageNumber="37" value="1988-12-23">23 December 1988</date>
. Others: 
<specimenCount id="9D328B97000E2424F1414631731EF815" box="[1047,1123,1903,1927]" pageId="2" pageNumber="37" type="male">1 male</specimenCount>
(22.3 × 
<quantity id="4CCCEDFB000E2424F19E46317253F815" box="[1224,1326,1903,1927]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="2" pageNumber="37" unit="mm" value="21.8">21.8 mm</quantity>
) (
<collectionCode id="ED25D8DB000E2424F010463172E1F815" LSID="urn:lsid:biocol.org:col:34992" box="[1350,1436,1903,1927]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="37">RMNH</collectionCode>
1202), Tjibodas, Java, 
<collectingCountry id="F323008E000E2424F4CE46CA7575F83E" box="[408,520,1940,1964]" name="Indonesia" pageId="2" pageNumber="37">Indonesia</collectingCountry>
, coll. J. Boerlarge, 1888; 
<specimenCount id="9D328B97000E2424F67D46CA740BF83E" box="[811,886,1940,1964]" pageId="2" pageNumber="37" type="male">1 male</specimenCount>
(25.5 × 
<quantity id="4CCCEDFB000E2424F68E46CA7343F83E" box="[984,1086,1940,1964]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="2" pageNumber="37" unit="mm" value="24.9">24.9 mm</quantity>
) (
<collectionCode id="ED25D8DB000E2424F10346CA73D6F83E" LSID="urn:lsid:biocol.org:col:34992" box="[1109,1195,1940,1964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="37">RMNH</collectionCode>
1989), Nias, 
<collectingCountry id="F323008E000E2424F01346CA77C4F842" name="Indonesia" pageId="2" pageNumber="37">Indonesia</collectingCountry>
, coll. E. E. W. G. Schröder, 1908; 
<specimenCount id="9D328B97000E2424F76146E675FFF842" box="[567,642,1976,2000]" pageId="2" pageNumber="37" type="male">1 male</specimenCount>
(34.5 × 
<quantity id="4CCCEDFB000E2424F7B446E67435F842" box="[738,840,1976,2000]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" pageId="2" pageNumber="37" unit="mm" value="33.1">33.1 mm</quantity>
) (
<collectionCode id="ED25D8DB000E2424F60846E674EBF842" box="[862,918,1976,2000]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="37">ZRC</collectionCode>
2009.0099), Cuthbert Bay, Andaman Is., coll. P. Biswas, 2001.
</paragraph>
<caption id="DF4B1096000F2425F5C146C77667F87C" httpUri="https://zenodo.org/record/206069/files/figure.png" pageId="3" pageNumber="38" targetBox="[261,1325,193,1924]" targetPageId="3">
<paragraph id="8B8B401E000F2425F5C146C77667F87C" blockId="3.[151,1436,1945,2030]" pageId="3" pageNumber="38">
<emphasis id="B9409C0C000F2425F5C146C7766FF83D" bold="true" box="[151,274,1945,1968]" pageId="3" pageNumber="38">FIGURE 1.</emphasis>
Colours in life of 
<taxonomicName id="4C343B9D000F2425F49A46C475ECF822" box="[460,657,1946,1968]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000F2425F49A46C475ECF822" box="[460,657,1946,1968]" italics="true" pageId="3" pageNumber="38">Labuanium vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000F2425F7C146C4746EF822" box="[663,787,1945,1968]" pageId="3" pageNumber="38" rank="species">
<emphasis id="B9409C0C000F2425F7C146C4746EF822" bold="true" box="[663,787,1945,1968]" pageId="3" pageNumber="38">new species</emphasis>
</taxonomicNameLabel>
. A, B, female (35.2 × 33.6 mm) (WAM C258-89), Greta Beach, Christmas I (courtesy Hugh Yorkston).; C, male (39.5 × 38.5 mm) (WAM C257-89), The Dales, Christmas I (courtesy Peter Greenaway).
</paragraph>
</caption>
<caption id="DF4B109600082422F5C146C77495F87F" httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="4" pageNumber="39" targetBox="[171,1406,193,1900]" targetPageId="4">
<paragraph id="8B8B401E00082422F5C146C77495F87F" blockId="4.[151,1436,1945,2030]" pageId="4" pageNumber="39">
<emphasis id="B9409C0C00082422F5C146C7766BF83D" bold="true" box="[151,278,1945,1968]" pageId="4" pageNumber="39">FIGURE 2.</emphasis>
Overall views of 
<taxonomicName id="4C343B9D00082422F48C46C47530F822" box="[474,589,1946,1968]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9409C0C00082422F48C46C47530F822" box="[474,589,1946,1968]" italics="true" pageId="4" pageNumber="39">Labuanium</emphasis>
</taxonomicName>
species. A, 
<taxonomicName id="4C343B9D00082422F78546C57440F822" box="[723,829,1947,1968]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C00082422F78546C57440F822" box="[723,829,1947,1968]" italics="true" pageId="4" pageNumber="39">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273217700082422F61046C474BBF822" box="[838,966,1945,1968]" pageId="4" pageNumber="39" rank="species">
<emphasis id="B9409C0C00082422F61046C474BBF822" bold="true" box="[838,966,1945,1968]" pageId="4" pageNumber="39">new species</emphasis>
</taxonomicNameLabel>
, holotype male (34.8 × 34.9 mm) (WAM 
<accessionNumber id="9467DDFD00082422F0DD46C477A7F85D" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="4" pageNumber="39">C 13977</accessionNumber>
), Christmas I.; B, 
<taxonomicName id="4C343B9D00082422F4C046E47478F85D" authority="Nobili, 1899" authorityName="Nobili" authorityYear="1899" box="[406,773,1976,1999]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C00082422F4C046E47514F85C" box="[406,617,1976,1999]" italics="true" pageId="4" pageNumber="39">L. papuomalesiacum</emphasis>
(Nobili, 1899)
</taxonomicName>
, male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, 
<taxonomicName id="4C343B9D00082422F0D146E476E3F87F" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C00082422F0D146E47789F87F" italics="true" pageId="4" pageNumber="39">L. scandens</emphasis>
Ng &amp; Liu, 2003
</taxonomicName>
, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan.
</paragraph>
</caption>
<caption id="DF4B109600092423F5C146CA733BF87A" httpUri="https://zenodo.org/record/206071/files/figure.png" pageId="5" pageNumber="40" targetBox="[214,1375,193,1899]" targetPageId="5">
<paragraph id="8B8B401E00092423F5C146CA733BF87A" blockId="5.[151,1436,1940,2025]" pageId="5" pageNumber="40">
<emphasis id="B9409C0C00092423F5C146CA7669F838" bold="true" box="[151,276,1940,1963]" pageId="5" pageNumber="40">FIGURE 3.</emphasis>
Frontal views of carapaces of 
<taxonomicName id="4C343B9D00092423F70146CB75B7F839" box="[599,714,1941,1963]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B9409C0C00092423F70146CB75B7F839" box="[599,714,1941,1963]" italics="true" pageId="5" pageNumber="40">Labuanium</emphasis>
</taxonomicName>
species. A, 
<taxonomicName id="4C343B9D00092423F61D46C874C9F839" box="[843,948,1942,1963]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C00092423F61D46C874C9F839" box="[843,948,1942,1963]" italics="true" pageId="5" pageNumber="40">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273217700092423F6EA46CB7347F839" box="[956,1082,1940,1963]" pageId="5" pageNumber="40" rank="species">
<emphasis id="B9409C0C00092423F6EA46CB7347F839" bold="true" box="[956,1082,1940,1963]" pageId="5" pageNumber="40">new species</emphasis>
</taxonomicNameLabel>
, holotype male (34.8 × 34.9 mm) (WAM 
<accessionNumber id="9467DDFD00092423F5B246EA7642F858" box="[228,319,1972,1994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="5" pageNumber="40">C 13977</accessionNumber>
), Christmas I.; B, 
<taxonomicName id="4C343B9D00092423F4AC46EB7415F858" authority="Nobili, 1899" authorityName="Nobili" authorityYear="1899" box="[506,872,1971,1994]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C00092423F4AC46EB75B3F858" box="[506,718,1972,1994]" italics="true" pageId="5" pageNumber="40">L. papuomalesiacum</emphasis>
(Nobili, 1899)
</taxonomicName>
, male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, 
<taxonomicName id="4C343B9D00092423F58C468D7681F87A" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" box="[218,508,2002,2024]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C00092423F58C468D762FF87A" box="[218,338,2002,2024]" italics="true" pageId="5" pageNumber="40">L. scandens</emphasis>
Ng &amp; Liu, 2003
</taxonomicName>
, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan.
</paragraph>
</caption>
<paragraph id="8B8B401E000A2420F5A942D774E3FC4E" blockId="6.[151,1436,904,988]" pageId="6" pageNumber="41">
<emphasis id="B9409C0C000A2420F5A942D7766FFC0C" bold="true" box="[255,274,905,926]" pageId="6" pageNumber="41">4.</emphasis>
Right third maxillipeds of 
<taxonomicName id="4C343B9D000A2420F77242D675EAFC0C" box="[548,663,904,926]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9409C0C000A2420F77242D675EAFC0C" box="[548,663,904,926]" italics="true" pageId="6" pageNumber="41">Labuanium</emphasis>
</taxonomicName>
species. A, 
<taxonomicName id="4C343B9D000A2420F64442D47404FC0D" box="[786,889,906,927]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum">
<emphasis id="B9409C0C000A2420F64442D47404FC0D" box="[786,889,906,927]" italics="true" pageId="6" pageNumber="41">L. vitatum</emphasis>
</taxonomicName>
<emphasis id="B9409C0C000A2420F62942D774D6FC0C" bold="true" box="[895,939,905,926]" pageId="6" pageNumber="41">new</emphasis>
, 
<typeStatus id="548FFEBC000A2420F15E42D6731CFC0C" box="[1032,1121,904,926]" pageId="6" pageNumber="41" type="holotype">holotype</typeStatus>
male (34.8 × 
<quantity id="4CCCEDFB000A2420F1BB42D77234FC0C" box="[1261,1353,905,926]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="6" pageNumber="41" unit="mm" value="34.9">34.9 mm</quantity>
) (
<collectionCode id="ED25D8DB000A2420F00A42D772E1FC0C" LSID="urn:lsid:biocol.org:col:34981" box="[1372,1436,905,926]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="6" pageNumber="41">WAM</collectionCode>
<accessionNumber id="9467DDFD000A2420F5C142F6778CFC2C" box="[151,241,936,958]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="6" pageNumber="41">C 13977</accessionNumber>
), Christmas I.; B, 
<taxonomicName id="4C343B9D000A2420F4FE42F6746EFC2C" authority="Nobili, 1899" authorityName="Nobili" authorityYear="1899" box="[424,787,935,958]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000A2420F4FE42F67507FC2F" box="[424,634,935,957]" italics="true" pageId="6" pageNumber="41">L. papuomalesiacum</emphasis>
(
<bibRefCitation id="EFA53DEF000A2420F7DE42F97471FC2C" author="Nobili" box="[648,780,935,958]" pageId="6" pageNumber="41" refString="Nobili, G. (1899) Contribuzioni alla conoscienza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali dei Museo Civico di Storia Naturale di Genova, (2) 20, 230 - 282." type="journal article" year="1899">Nobili, 1899</bibRefCitation>
)
</taxonomicName>
, male (38.0 × 
<quantity id="4CCCEDFB000A2420F6F242F67482FC2F" box="[932,1023,936,957]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="6" pageNumber="41" unit="mm" value="37.4">37.4 mm</quantity>
) (
<collectionCode id="ED25D8DB000A2420F14542F6733AFC2F" LSID="urn:lsid:biocol.org:col:34838" box="[1043,1095,936,957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="41">SMF</collectionCode>
1971.18.5.53), Caroline Is.; C, 
<taxonomicName id="4C343B9D000A2420F0D142F676E3FC49" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C000A2420F0D142F67789FC4E" italics="true" pageId="6" pageNumber="41">L. scandens</emphasis>
<bibRefCitation id="EFA53DEF000A2420F5AC429976E3FC49" author="Ng" box="[250,414,966,988]" pageId="6" pageNumber="41" refString="Jeng, M. - S., Liu, H. - C., Tzeng, C. - S. &amp; Ng, P. K. L. (2003) On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana, 76 (2), 227 - 240." type="journal article" year="2003">Ng &amp; Liu, 2003</bibRefCitation>
</taxonomicName>
, male (29.6 × 
<quantity id="4CCCEDFB000A2420F779429875F6FC4E" box="[559,651,966,988]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="6" pageNumber="41" unit="mm" value="28.7">28.7 mm</quantity>
) (
<collectionCode id="ED25D8DB000A2420F7C9429875AFFC49" box="[671,722,966,987]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="41">ZRC</collectionCode>
2002.478), 
<collectingCountry id="F323008E000A2420F61A429874E7FC4E" box="[844,922,966,988]" name="Taiwan" pageId="6" pageNumber="41">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E1395000A2420F59145567548FA66" pageId="6" pageNumber="41" type="diagnosis">
<paragraph id="8B8B401E000A2420F59145567548FA66" blockId="6.[151,1436,1032,2029]" pageId="6" pageNumber="41">
<emphasis id="B9409C0C000A2420F5914556763CFBB3" bold="true" box="[199,321,1032,1057]" pageId="6" pageNumber="41">Diagnosis.</emphasis>
Carapace width and length subequal; antero-dorsal surfaces finely granular, posterior surfaces almost smooth (
<figureCitation id="130F5C9B000A2420F405457276E7FBD6" box="[339,410,1068,1092]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Fig. 2</figureCitation>
A); posterolateral regions with fine granular oblique striae (
<figureCitation id="130F5C9B000A2420F101457273E2FBD6" box="[1111,1183,1068,1092]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Fig. 2</figureCitation>
A); postfrontal cristae prominent, sharp, medially separated by deep cleft (
<figureCitation id="130F5C9B000A2420F78A450E7454FBFA" box="[732,809,1104,1128]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Figs. 2</figureCitation>
A, 3A); merus of third maxilliped in adult male longitudinally ovate, ca. twice as long as broad (Fig. 4A); inner margin of cheliped carpus with prominent triangular tooth, margins distinctly serrated in large males (
<figureCitation id="130F5C9B000A2420F72F45C675C1FB22" box="[633,700,1176,1200]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Fig. 5</figureCitation>
A); outer surface of chela with numerous small, closely arranged rounded granules (
<figureCitation id="130F5C9B000A2420F43A45E276C1FB46" box="[364,444,1212,1236]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Figs. 5</figureCitation>
A, B; 7A, B); dorsal margin of small male chela, with a non-pectinated ridge on outer edge of about 44 small granules, with 2 shorter, less distinct oblique rows of granules posterior to it (
<figureCitation id="130F5C9B000A2420F04145BE7226FB6A" box="[1303,1371,1248,1272]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Fig. 5</figureCitation>
B), in large males, ridges indistinct (
<figureCitation id="130F5C9B000A2420F4BC445A7550FA8E" box="[490,557,1284,1308]" captionStart="FIGURE 7" captionStartId="9.[151,250,1977,2000]" captionTargetBox="[235,1352,267,1946]" captionTargetId="figure@9.[235,1353,267,1946]" captionTargetPageId="9" captionText="FIGURE 7. Labuanium vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I .. A, outer view of left chela; B, fourth right ambulatory leg; C, third right ambulatory leg." httpUri="https://zenodo.org/record/206074/files/figure.png" pageId="6" pageNumber="41">Fig. 7</figureCitation>
B) to almost indiscernible (
<figureCitation id="130F5C9B000A2420F636445A74DEFA8E" box="[864,931,1284,1308]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Fig. 5</figureCitation>
A), with granules appearing relatively larger, more scattered; dorsal surface of cheliped dactylus with row of distinct granules, about 
<quantity id="4CCCEDFB000A2420F128447673C4FAD2" box="[1150,1209,1320,1344]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="41" unit="in" value="12.0">12 in</quantity>
smaller males (
<figureCitation id="130F5C9B000A2420F039447677D8FAF6" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Fig. 5</figureCitation>
B), 10 or 
<quantity id="4CCCEDFB000A2420F44644127635FAF6" box="[272,328,1356,1380]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="6" pageNumber="41" unit="in" value="11.0">11 in</quantity>
larger males (
<figureCitation id="130F5C9B000A2420F4BE44127548FAF6" box="[488,565,1356,1380]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Figs. 5</figureCitation>
A, 7B); outer surface of ambulatory meri weakly rugose, dorsal margin finely granulated (
<figureCitation id="130F5C9B000A2420F448442E7613FA1A" box="[286,366,1392,1416]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Figs. 2</figureCitation>
A, 7C, D); male abdominal somite 6 broader than long, distal part of lateral margins convex, proximal part slightly sinuous (
<figureCitation id="130F5C9B000A2420F4A244CA754AFA3E" box="[500,567,1428,1452]" captionStart="FIGURE 6" captionStartId="8.[151,250,622,645]" captionTargetBox="[151,1436,193,600]" captionTargetId="figure@8.[151,1436,193,601]" captionTargetPageId="8" captionText="FIGURE 6. Thoracic sternal views and abdomens of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206073/files/figure.png" pageId="6" pageNumber="41">Fig. 6</figureCitation>
A); abdominal telson as long as somite 6, subtriangular in shape with rounded tip (
<figureCitation id="130F5C9B000A2420F59144E67670FA42" box="[199,269,1464,1488]" captionStart="FIGURE 6" captionStartId="8.[151,250,622,645]" captionTargetBox="[151,1436,193,600]" captionTargetId="figure@8.[151,1436,193,601]" captionTargetPageId="8" captionText="FIGURE 6. Thoracic sternal views and abdomens of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206073/files/figure.png" pageId="6" pageNumber="41">Fig. 6</figureCitation>
A); G1 relatively stout; distal part bent about 45° from vertical, distal chitinous part relatively short, beak-like, subtruncate (
<figureCitation id="130F5C9B000A2420F4CB44827693FA66" box="[413,494,1500,1524]" captionStart="FIGURE 10" captionStartId="12.[151,250,1729,1752]" captionTargetBox="[161,1425,213,1680]" captionTargetId="figure@12.[151,1436,193,1709]" captionTargetPageId="12" captionText="FIGURE 10. Denuded gonopods of Labuanium species. A – E, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; F, G, L. vitatum new species, male (22.3 × 21.8 mm) (RMNH 1202), Java; H – K, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is. A, B, C, D, F, G, left G 1; H, I, J, K, right G 1, E, left G 2. Scales: A, B, E, F, G, H, I = 1.0 mm; C, D, J, K = 0.5 mm." httpUri="https://zenodo.org/record/206077/files/figure.png" pageId="6" pageNumber="41">Fig. 10</figureCitation>
A–D).
</paragraph>
</subSubSection>
<subSubSection id="C32E1395000A2420F591475F76CFF9AE" pageId="6" pageNumber="41" type="etymology">
<paragraph id="8B8B401E000A2420F591475F76CFF9AE" blockId="6.[151,1436,1032,2029]" pageId="6" pageNumber="41">
<emphasis id="B9409C0C000A2420F591475F7633F98B" bold="true" box="[199,334,1537,1561]" pageId="6" pageNumber="41">Etymology.</emphasis>
The name is Latin meaning for “shun” and “frustrating to find.” It alludes to the naturally secretive habits of the species.
</paragraph>
</subSubSection>
<subSubSection id="C32E1395000A2420F591471774D9F87E" pageId="6" pageNumber="41" type="discussion">
<paragraph id="8B8B401E000A2420F591471773A8F93A" blockId="6.[151,1436,1032,2029]" pageId="6" pageNumber="41">
<emphasis id="B9409C0C000A2420F59147177646F9F3" bold="true" box="[199,315,1609,1633]" pageId="6" pageNumber="41">Remarks.</emphasis>
This species was first recorded from Christmas I. by 
<bibRefCitation id="EFA53DEF000A2420F6DA47167352F9F2" author="George" box="[908,1071,1608,1632]" pageId="6" pageNumber="41" refString="George, R. W. (1978) The Land and Freshwater Crabs of Christmas Island. Unpublished report commissioned by the Australian National Parks and Wildlife Service, 22 pp." type="book" year="1978">George (1978)</bibRefCitation>
under the name of 
<taxonomicName id="4C343B9D000A2420F05347177606F916" authority="De Man, 1889" authorityName="De Man" authorityYear="1889" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="oceanica">
<emphasis id="B9409C0C000A2420F053471777A8F917" italics="true" pageId="6" pageNumber="41">Sesarma oceanica</emphasis>
<collectingCountry id="F323008E000A2420F58D47337781F916" box="[219,252,1645,1668]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="41">De</collectingCountry>
<bibRefCitation id="EFA53DEF000A2420F45447337606F916" author="Man" box="[258,379,1644,1668]" pageId="6" pageNumber="41" refString="Man, J. G. De (1889) Ueber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452, pls. 9, 10." type="journal article" year="1889">
<collectingCountry id="F323008E000A2420F45447337645F916" box="[258,312,1645,1668]" name="Isle of Man" pageId="6" pageNumber="41">Man</collectingCountry>
, 1889
</bibRefCitation>
</taxonomicName>
(his specimen is here designated as the 
<typeStatus id="548FFEBC000A2420F663473274EAF916" box="[821,919,1644,1668]" pageId="6" pageNumber="41" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C343B9D000A2420F6EA473373EBF917" box="[956,1174,1645,1669]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F6EA473373EBF917" box="[956,1174,1645,1669]" italics="true" pageId="6" pageNumber="41">Labuanium vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F1CA47337258F917" box="[1180,1317,1644,1669]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F1CA47337258F917" bold="true" box="[1180,1317,1644,1669]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
). 
<taxonomicName id="4C343B9D000A2420F06A47337781F93B" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="oceanica">
<emphasis id="B9409C0C000A2420F06A47337781F93B" italics="true" pageId="6" pageNumber="41">Sesarma oceanica</emphasis>
</taxonomicName>
is still considered a subjective junior synonym of 
<taxonomicName id="4C343B9D000A2420F66647CF73ACF93A" authority="Hess, 1865" authorityName="Hess" authorityYear="1865" box="[816,1233,1680,1705]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B9409C0C000A2420F66647CF7345F93B" box="[816,1080,1681,1705]" italics="true" pageId="6" pageNumber="41">Labuanium rotundatum</emphasis>
(
<bibRefCitation id="EFA53DEF000A2420F11147CE73B4F93A" author="Hess" box="[1095,1225,1680,1704]" pageId="6" pageNumber="41" refString="Hess, W. (1865) Beitrage zur Kenntnis der Decapoden-Krebse Ost-Australiens. Archiv fur Naturgeschicht e, 36, 127 - 173, pls. 1, 2." type="journal article" year="1865">Hess, 1865</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8B401E000A2420F59147EB74D9F87E" blockId="6.[151,1436,1032,2029]" pageId="6" pageNumber="41">
<taxonomicName id="4C343B9D000A2420F59147EB76DEF95F" box="[199,419,1717,1741]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F59147EB76DEF95F" box="[199,419,1717,1741]" italics="true" pageId="6" pageNumber="41">Labuanium vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F4FA47EB7544F95F" box="[428,569,1716,1741]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F4FA47EB7544F95F" bold="true" box="[428,569,1716,1741]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
is superficially similar to 
<taxonomicName id="4C343B9D000A2420F63E47E87329F95F" box="[872,1108,1717,1741]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000A2420F63E47E87329F95F" box="[872,1108,1717,1741]" italics="true" pageId="6" pageNumber="41">L. papuomalesiacum</emphasis>
</taxonomicName>
but they can be easily separated by their G1 morphology. In 
<taxonomicName id="4C343B9D000A2420F74247847582F963" box="[532,767,1753,1777]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000A2420F74247847582F963" box="[532,767,1753,1777]" italics="true" pageId="6" pageNumber="41">L. papuomalesiacum</emphasis>
</taxonomicName>
, the G1 is relatively longer and the pectinated distal part is proportionately longer and almost straight (
<figureCitation id="130F5C9B000A2420F7D247A275ABF886" box="[644,726,1788,1812]" captionStart="FIGURE 10" captionStartId="12.[151,250,1729,1752]" captionTargetBox="[161,1425,213,1680]" captionTargetId="figure@12.[151,1436,193,1709]" captionTargetPageId="12" captionText="FIGURE 10. Denuded gonopods of Labuanium species. A – E, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; F, G, L. vitatum new species, male (22.3 × 21.8 mm) (RMNH 1202), Java; H – K, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is. A, B, C, D, F, G, left G 1; H, I, J, K, right G 1, E, left G 2. Scales: A, B, E, F, G, H, I = 1.0 mm; C, D, J, K = 0.5 mm." httpUri="https://zenodo.org/record/206077/files/figure.png" pageId="6" pageNumber="41">Fig. 10</figureCitation>
H, I) whereas in 
<taxonomicName id="4C343B9D000A2420F6C047A07374F887" box="[918,1033,1790,1813]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F6C047A07374F887" box="[918,1033,1790,1813]" italics="true" pageId="6" pageNumber="41">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F14447A373E0F887" box="[1042,1181,1788,1813]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F14447A373E0F887" bold="true" box="[1042,1181,1788,1813]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
, the G1 is proportionately shorter and the pectinated distal part is shorter and gently sinuous (
<figureCitation id="130F5C9B000A2420F690467E736AF8AA" box="[966,1047,1824,1848]" captionStart="FIGURE 10" captionStartId="12.[151,250,1729,1752]" captionTargetBox="[161,1425,213,1680]" captionTargetId="figure@12.[151,1436,193,1709]" captionTargetPageId="12" captionText="FIGURE 10. Denuded gonopods of Labuanium species. A – E, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; F, G, L. vitatum new species, male (22.3 × 21.8 mm) (RMNH 1202), Java; H – K, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is. A, B, C, D, F, G, left G 1; H, I, J, K, right G 1, E, left G 2. Scales: A, B, E, F, G, H, I = 1.0 mm; C, D, J, K = 0.5 mm." httpUri="https://zenodo.org/record/206077/files/figure.png" pageId="6" pageNumber="41">Fig. 10</figureCitation>
A, B). The lateral carapace margin of larger adult 
<taxonomicName id="4C343B9D000A2420F41746187551F8CF" box="[321,556,1861,1885]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000A2420F41746187551F8CF" box="[321,556,1861,1885]" italics="true" pageId="6" pageNumber="41">L. papuomalesiacum</emphasis>
</taxonomicName>
is also relatively more convex (
<figureCitation id="130F5C9B000A2420F6CD461A7496F8CE" box="[923,1003,1860,1884]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Figs. 2</figureCitation>
B, 9) (relatively straighter in similarly sized 
<taxonomicName id="4C343B9D000A2420F58F46347631F813" box="[217,332,1898,1921]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F58F46347631F813" box="[217,332,1898,1921]" italics="true" pageId="6" pageNumber="41">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F4034637769DF813" box="[341,480,1896,1921]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F4034637769DF813" bold="true" box="[341,480,1896,1921]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
, 
<figureCitation id="130F5C9B000A2420F4A646367543F812" box="[496,574,1896,1920]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Figs. 2</figureCitation>
A, 8), the merus of the third maxilliped is relatively shorter (Fig. 4B) (relatively longer in 
<taxonomicName id="4C343B9D000A2420F41A46D076C3F837" box="[332,446,1934,1957]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F41A46D076C3F837" box="[332,446,1934,1957]" italics="true" pageId="6" pageNumber="41">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F49046D3752DF837" box="[454,592,1932,1957]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F49046D3752DF837" bold="true" box="[454,592,1932,1957]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
, Fig. 4A), and the dorsomarginal ridge of granules on the chela of 
<taxonomicName id="4C343B9D000A2420F01C46D07644F85B" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000A2420F01C46D07644F85B" italics="true" pageId="6" pageNumber="41">L. papuomalesiacum</emphasis>
</taxonomicName>
is distinct in adult males (
<figureCitation id="130F5C9B000A2420F73246EE75DBF85A" box="[612,678,1968,1992]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Fig. 5</figureCitation>
C) (diffuse and poorly demarcated in large 
<taxonomicName id="4C343B9D000A2420F1C646EC727EF85B" box="[1168,1283,1970,1993]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000A2420F1C646EC727EF85B" box="[1168,1283,1970,1993]" italics="true" pageId="6" pageNumber="41">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000A2420F05C46EF72E9F85B" box="[1290,1428,1968,1993]" pageId="6" pageNumber="41" rank="species">
<emphasis id="B9409C0C000A2420F05C46EF72E9F85B" bold="true" box="[1290,1428,1968,1993]" pageId="6" pageNumber="41">new species</emphasis>
</taxonomicNameLabel>
, 
<figureCitation id="130F5C9B000A2420F5C1468A779FF87E" box="[151,226,2004,2028]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="6" pageNumber="41">Figs. 5</figureCitation>
A, 7B). These differences also apply for 
<taxonomicName id="4C343B9D000A2420F7FD46887452F87F" box="[683,815,2005,2029]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C000A2420F7FD46887452F87F" box="[683,815,2005,2029]" italics="true" pageId="6" pageNumber="41">L. scandens</emphasis>
</taxonomicName>
(
<figureCitation id="130F5C9B000A2420F668468A74FFF87E" box="[830,898,2004,2028]" captionStart="FIGURE 2" captionStartId="4.[151,250,1945,1968]" captionTargetBox="[171,1406,193,1900]" captionTargetId="figure@4.[171,1407,193,1900]" captionTargetPageId="4" captionText="FIGURE 2. Overall views of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, L. scandens Ng &amp; Liu, 2003, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan." httpUri="https://zenodo.org/record/206070/files/figure.png" pageId="6" pageNumber="41">Fig. 2</figureCitation>
C).
</paragraph>
</subSubSection>
<caption id="DF4B1096000B2421F5C146257578F87C" httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="7" pageNumber="42" targetBox="[197,1397,193,1895]" targetPageId="7">
<paragraph id="8B8B401E000B2421F5C146257578F87C" blockId="7.[151,1436,1915,2030]" pageId="7" pageNumber="42">
<emphasis id="B9409C0C000B2421F5C14625766EF803" bold="true" box="[151,275,1915,1938]" pageId="7" pageNumber="42">FIGURE 5.</emphasis>
Dorsal views of major male chelae of 
<taxonomicName id="4C343B9D000B2421F7C94622746FF800" box="[671,786,1916,1938]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9409C0C000B2421F7C94622746FF800" box="[671,786,1916,1938]" italics="true" pageId="7" pageNumber="42">Labuanium</emphasis>
</taxonomicName>
species. A, 
<taxonomicName id="4C343B9D000B2421F6D94623748AF800" box="[911,1015,1917,1938]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000B2421F6D94623748AF800" box="[911,1015,1917,1938]" italics="true" pageId="7" pageNumber="42">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000B2421F6AB46227304F800" box="[1021,1145,1915,1938]" pageId="7" pageNumber="42" rank="species">
<emphasis id="B9409C0C000B2421F6AB46227304F800" bold="true" box="[1021,1145,1915,1938]" pageId="7" pageNumber="42">new species</emphasis>
</taxonomicNameLabel>
, holotype male (34.8 × 34.9 mm) (WAM 
<accessionNumber id="9467DDFD000B2421F44F46C5760EF823" box="[281,371,1947,1969]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="7" pageNumber="42">C 13977</accessionNumber>
), Christmas I.; B, 
<taxonomicName id="4C343B9D000B2421F77C46C275EDF823" box="[554,656,1948,1969]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C000B2421F77C46C275EDF823" box="[554,656,1948,1969]" italics="true" pageId="7" pageNumber="42">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732177000B2421F7C046C5746FF823" box="[662,786,1946,1969]" pageId="7" pageNumber="42" rank="species">
<emphasis id="B9409C0C000B2421F7C046C5746FF823" bold="true" box="[662,786,1946,1969]" pageId="7" pageNumber="42">new species</emphasis>
</taxonomicNameLabel>
, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, 
<taxonomicName id="4C343B9D000B2421F05E46C2760AF842" authority="Nobili, 1899" authorityName="Nobili" authorityYear="1899" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C000B2421F05E46C277A2F85D" italics="true" pageId="7" pageNumber="42">L. papuomalesiacum</emphasis>
(Nobili, 1899)
</taxonomicName>
, male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, 
<taxonomicName id="4C343B9D000B2421F6B946E4726CF85D" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" box="[1007,1297,1977,1999]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C000B2421F6B946E4731AF85D" box="[1007,1127,1977,1999]" italics="true" pageId="7" pageNumber="42">L. scandens</emphasis>
Ng &amp; Liu, 2003
</taxonomicName>
, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan.
</paragraph>
</caption>
<caption id="DF4B10960004242EF5C1433073D9FD53" httpUri="https://zenodo.org/record/206073/files/figure.png" pageId="8" pageNumber="43" targetBox="[151,1436,193,600]" targetPageId="8">
<paragraph id="8B8B401E0004242EF5C1433073D9FD53" blockId="8.[151,1436,622,706]" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF5C143307669FD11" bold="true" box="[151,276,622,645]" pageId="8" pageNumber="43">FIGURE 6.</emphasis>
Thoracic sternal views and abdomens of 
<taxonomicName id="4C343B9D0004242EF79043307447FD16" box="[710,826,622,644]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9409C0C0004242EF79043307447FD16" box="[710,826,622,644]" italics="true" pageId="8" pageNumber="43">Labuanium</emphasis>
</taxonomicName>
species. A, 
<taxonomicName id="4C343B9D0004242EF6EC4331735FFD16" box="[954,1058,623,644]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C0004242EF6EC4331735FFD16" box="[954,1058,623,644]" italics="true" pageId="8" pageNumber="43">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27321770004242EF17C433173DAFD17" box="[1066,1191,622,645]" pageId="8" pageNumber="43" rank="species">
<emphasis id="B9409C0C0004242EF17C433173DAFD17" bold="true" box="[1066,1191,622,645]" pageId="8" pageNumber="43">new species</emphasis>
</taxonomicNameLabel>
, holotype male (34.8 × 34.9 mm) (WAM 
<accessionNumber id="9467DDFD0004242EF41A43D376DAFD31" box="[332,423,653,675]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="8" pageNumber="43">C 13977</accessionNumber>
), Christmas I.; B, 
<taxonomicName id="4C343B9D0004242EF70943D074B6FD31" authority="Nobili, 1899" authorityName="Nobili" authorityYear="1899" box="[607,971,652,675]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="papuomalesiacum">
<emphasis id="B9409C0C0004242EF70943D0744FFD31" box="[607,818,653,675]" italics="true" pageId="8" pageNumber="43">L. papuomalesiacum</emphasis>
(Nobili, 1899)
</taxonomicName>
, male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; C, 
<taxonomicName id="4C343B9D0004242EF46E43F27527FD53" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" box="[312,602,683,705]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C0004242EF46E43F276CDFD53" box="[312,432,683,705]" italics="true" pageId="8" pageNumber="43">L. scandens</emphasis>
Ng &amp; Liu, 2003
</taxonomicName>
, holotype male (42.3 × 41.2 mm) (TMCD 3290), Taiwan.
</paragraph>
</caption>
<subSubSection id="C32E13950004242EF59143B3744FF888" pageId="8" pageNumber="43" type="description">
<paragraph id="8B8B401E0004242EF59143B37787FB00" blockId="8.[151,1437,749,2035]" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF59143B37646FC94" bold="true" box="[199,315,749,774]" pageId="8" pageNumber="43">Variation.</emphasis>
The second anterolateral tooth of the 
<typeStatus id="548FFEBC0004242EF7AA43B17421FC95" box="[764,860,751,775]" pageId="8" pageNumber="43" type="paratype">paratype</typeStatus>
female is more prominent than in the males. The inner angle of the cheliped carpus also has a distinct large triangular tooth which is dorso-ventrally flattened and appears almost lobiform; whereas in males, it is less prominent and the margins appear almost serrated (
<figureCitation id="130F5C9B0004242EF0624268720AFCDC" box="[1332,1399,822,846]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="8" pageNumber="43">Fig. 5</figureCitation>
A). The gastric regions of the males are also slightly more swollen. The two relatively small specimens from Nias (RMNH 1989) and Java (RMNH 1202) (
<figureCitation id="130F5C9B0004242EF735422075D9FC04" box="[611,676,894,918]" captionStart="FIGURE 8" captionStartId="10.[151,250,1939,1962]" captionTargetBox="[151,1436,193,1894]" captionTargetId="figure@10.[151,1436,193,1894]" captionTargetPageId="10" captionText="FIGURE 8. Overall views of Labuanium vitatum new species. A, male (22.3 × 21.8 mm) (RMNH 1202), Java; B, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, male (34.5 × 33.1 mm) (ZRC 2009.0099), Andaman Is." httpUri="https://zenodo.org/record/206075/files/figure.png" pageId="8" pageNumber="43">Fig. 8</figureCitation>
A, B) resemble the 
<typeStatus id="548FFEBC0004242EF6D7422174C1FC05" box="[897,956,895,919]" pageId="8" pageNumber="43">types</typeStatus>
from Christmas I. except that the granules lining the outer marginal ridge on the chela are distinct (
<figureCitation id="130F5C9B0004242EF64F42FC7420FC28" box="[793,861,930,954]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1938]" captionTargetBox="[197,1397,193,1895]" captionTargetId="figure@7.[197,1398,193,1895]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal views of major male chelae of Labuanium species. A, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; B, L. vitatum new species, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is.; D, L. scandens Ng &amp; Liu, 2003, male (29.6 × 28.7 mm) (ZRC 2002.478), Taiwan." httpUri="https://zenodo.org/record/206072/files/figure.png" pageId="8" pageNumber="43">Fig. 5</figureCitation>
B); and the lateral margins of the G1 are relatively more sinuous (
<figureCitation id="130F5C9B0004242EF46B429876F1FC4C" box="[317,396,966,990]" captionStart="FIGURE 10" captionStartId="12.[151,250,1729,1752]" captionTargetBox="[161,1425,213,1680]" captionTargetId="figure@12.[151,1436,193,1709]" captionTargetPageId="12" captionText="FIGURE 10. Denuded gonopods of Labuanium species. A – E, L. vitatum new species, holotype male (34.8 × 34.9 mm) (WAM C 13977), Christmas I.; F, G, L. vitatum new species, male (22.3 × 21.8 mm) (RMNH 1202), Java; H – K, L. papuomalesiacum (Nobili, 1899), male (38.0 × 37.4 mm) (SMF 1971.18.5.53), Caroline Is. A, B, C, D, F, G, left G 1; H, I, J, K, right G 1, E, left G 2. Scales: A, B, E, F, G, H, I = 1.0 mm; C, D, J, K = 0.5 mm." httpUri="https://zenodo.org/record/206077/files/figure.png" pageId="8" pageNumber="43">Fig. 10</figureCitation>
F, G). We also tentatively refer one male from Andaman Is. (ZRC 2009.0099) to this species although it is peculiar in having a relatively smoother dorsal carapace surface, lower anterolateral teeth and proportionately longer ambulatory propodi (
<figureCitation id="130F5C9B0004242EF768455075FDFBB4" box="[574,640,1038,1062]" captionStart="FIGURE 8" captionStartId="10.[151,250,1939,1962]" captionTargetBox="[151,1436,193,1894]" captionTargetId="figure@10.[151,1436,193,1894]" captionTargetPageId="10" captionText="FIGURE 8. Overall views of Labuanium vitatum new species. A, male (22.3 × 21.8 mm) (RMNH 1202), Java; B, male (25.5 × 24.9 mm) (RMNH 1989), Nias; C, male (34.5 × 33.1 mm) (ZRC 2009.0099), Andaman Is." httpUri="https://zenodo.org/record/206075/files/figure.png" pageId="8" pageNumber="43">Fig. 8</figureCitation>
C). Its G1 structure, however, agrees very well with the 
<typeStatus id="548FFEBC0004242EF1AA45507223FBB4" box="[1276,1374,1038,1062]" pageId="8" pageNumber="43" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="4C343B9D0004242EF5E0456D7655FBD8" box="[182,296,1075,1098]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C0004242EF5E0456D7655FBD8" box="[182,296,1075,1098]" italics="true" pageId="8" pageNumber="43">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27321770004242EF466456D76C6FBD9" box="[304,443,1074,1099]" pageId="8" pageNumber="43" rank="species">
<emphasis id="B9409C0C0004242EF466456D76C6FBD9" bold="true" box="[304,443,1074,1099]" pageId="8" pageNumber="43">new species</emphasis>
</taxonomicNameLabel>
. Because this male specimen is not fully hardened (it had moulted shortly before it was preserved) and both its chelae are missing, it nevertheless seems best to refer it to 
<taxonomicName id="4C343B9D0004242EF16B450973CDFBFC" box="[1085,1200,1111,1134]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C0004242EF16B450973CDFBFC" box="[1085,1200,1111,1134]" italics="true" pageId="8" pageNumber="43">L. vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27321770004242EF1E14508723FFBFD" box="[1207,1346,1110,1135]" pageId="8" pageNumber="43" rank="species">
<emphasis id="B9409C0C0004242EF1E14508723FFBFD" bold="true" box="[1207,1346,1110,1135]" pageId="8" pageNumber="43">new species</emphasis>
</taxonomicNameLabel>
, for the moment.
</paragraph>
<paragraph id="8B8B401E0004242EF59145C07454FA20" blockId="8.[151,1437,749,2035]" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF59145C0765DFB25" bold="true" box="[199,288,1182,1207]" pageId="8" pageNumber="43">Colour.</emphasis>
The colour appears to be somewhat variable. An adult male (
<typeStatus id="548FFEBC0004242EF68045C07346FB24" box="[982,1083,1182,1206]" pageId="8" pageNumber="43" type="paratype">paratype</typeStatus>
, 39.5 × 
<quantity id="4CCCEDFB0004242EF1C145C07383FB24" box="[1175,1278,1182,1206]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="8" pageNumber="43" unit="mm" value="38.5">38.5 mm</quantity>
, WAM C257- 89) was a uniform yellowish brown overall (
<figureCitation id="130F5C9B0004242EF7CD459C75A6FB48" box="[667,731,1218,1242]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1968]" captionTargetBox="[261,1325,193,1924]" captionTargetId="figure@3.[261,1325,193,1925]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life of Labuanium vitatum new species. A, B, female (35.2 × 33.6 mm) (WAM C 258 - 89), Greta Beach, Christmas I (courtesy Hugh Yorkston).; C, male (39.5 × 38.5 mm) (WAM C 257 - 89), The Dales, Christmas I (courtesy Peter Greenaway)." httpUri="https://zenodo.org/record/206069/files/figure.png" pageId="8" pageNumber="43">Fig. 1</figureCitation>
C) while a smaller female (
<typeStatus id="548FFEBC0004242EF14D459C73FDFB48" box="[1051,1152,1218,1242]" pageId="8" pageNumber="43" type="paratype">paratype</typeStatus>
, 35.2 × 
<quantity id="4CCCEDFB0004242EF1B7459C7237FB48" box="[1249,1354,1218,1242]" metricMagnitude="-2" metricUnit="m" metricValue="3.36" pageId="8" pageNumber="43" unit="mm" value="33.6">33.6 mm</quantity>
, WAM C258-89) was more brightly coloured, with a purplish carapace with beige lateral margins, with the legs pale purple, and the palm bright purple with red fingers (
<figureCitation id="130F5C9B0004242EF79E44547474FAB0" box="[712,777,1290,1314]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1968]" captionTargetBox="[261,1325,193,1924]" captionTargetId="figure@3.[261,1325,193,1925]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life of Labuanium vitatum new species. A, B, female (35.2 × 33.6 mm) (WAM C 258 - 89), Greta Beach, Christmas I (courtesy Hugh Yorkston).; C, male (39.5 × 38.5 mm) (WAM C 257 - 89), The Dales, Christmas I (courtesy Peter Greenaway)." httpUri="https://zenodo.org/record/206069/files/figure.png" pageId="8" pageNumber="43">Fig. 1</figureCitation>
A, B). The eyes are bright greenish-yellow in both specimens. The 
<typeStatus id="548FFEBC0004242EF4474470760EFAD4" box="[273,371,1326,1350]" pageId="8" pageNumber="43" type="holotype">holotype</typeStatus>
male (34.8 × 
<quantity id="4CCCEDFB0004242EF75B4470750EFAD4" box="[525,627,1326,1350]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="8" pageNumber="43" unit="mm" value="34.9">34.9 mm</quantity>
, WAM 
<accessionNumber id="9467DDFD0004242EF79F44707454FAD4" box="[713,809,1326,1350]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/C13977" pageId="8" pageNumber="43">C 13977</accessionNumber>
) was described on the label as “Deep brownish-purplish on carapace and legs. Hands purplish with vertical (narrow) white stripe between bases of pollex and dactylus on both inner and outer surfaces of hand”. 
<bibRefCitation id="EFA53DEF0004242EF7014428745BFA1C" author="Hicks" box="[599,806,1398,1422]" pageId="8" pageNumber="43" refString="Hicks, J., Rumpff, H. &amp; Yorkston, H. (1984) Christmas Crabs. Christmas Island Natural History Association, 1990 second edition, 81 pp." type="book" year="1984">
Hicks 
<emphasis id="B9409C0C0004242EF7F6442975A4FA1C" box="[672,729,1398,1422]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
(1984
</bibRefCitation>
: 64) notes that it is known as the “White-stripe crab” in Christmas I. because of the beige lateral carapace margins.
</paragraph>
<paragraph id="8B8B401E0004242EF59144E0744FF888" blockId="8.[151,1437,749,2035]" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF59144E076F5FA45" bold="true" box="[199,392,1470,1495]" pageId="8" pageNumber="43">Notes on habits.</emphasis>
Like most of its congeners, 
<taxonomicName id="4C343B9D0004242EF79B44E074D4FA44" box="[717,937,1470,1494]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="vitatum" status="sp. nov.">
<emphasis id="B9409C0C0004242EF79B44E074D4FA44" box="[717,937,1470,1494]" italics="true" pageId="8" pageNumber="43">Labuanium vitatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27321770004242EF6E744E07340FA45" box="[945,1085,1470,1495]" pageId="8" pageNumber="43" rank="species">
<emphasis id="B9409C0C0004242EF6E744E07340FA45" bold="true" box="[945,1085,1470,1495]" pageId="8" pageNumber="43">new species</emphasis>
</taxonomicNameLabel>
is an arboreal species, and has been collected usually at the base of trees in relatively close proximity to the sea (although one specimen was obtained on a vegetated limestone platform at 
<quantity id="4CCCEDFB0004242EF7CA475875A8F98C" box="[668,725,1542,1566]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="43" unit="m" value="60.0">60 m</quantity>
altitude). 
<bibRefCitation id="EFA53DEF0004242EF6104758736EF98C" author="Hicks" box="[838,1043,1542,1566]" pageId="8" pageNumber="43" refString="Hicks, J., Rumpff, H. &amp; Yorkston, H. (1984) Christmas Crabs. Christmas Island Natural History Association, 1990 second edition, 81 pp." type="book" year="1984">
Hicks 
<emphasis id="B9409C0C0004242EF6D9475974BAF98C" box="[911,967,1542,1566]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
(1984
</bibRefCitation>
: 64) notes that individuals grow up to 
<quantity id="4CCCEDFB0004242EF5E34774767BF9D0" box="[181,262,1578,1602]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="43" unit="mm" value="40.0">40 mm</quantity>
in carapace width and spawn about the time of the full moon between November and April. The chief warden on Christmas I., Max Orchard, has spent many years on the island and despite a strong personal interest in crabs, has seen this species only once on the night of 
<date id="FF8A66DE0004242EF655472C74D9F918" box="[771,932,1650,1674]" pageId="8" pageNumber="43" value="2010-04-15">15 April 2010</date>
at Andersons Dale, and he noted “… I was walking back up along the stream in pouring rain and edging around a tree when I caught a glimpse of a crab with bright yellow eyes and brownish carapace on a tree about 1.5 metres above the ground” (M. Orchard, personal communication). 
<collectingCountry id="F323008E0004242EF40947817603F965" box="[351,382,1759,1783]" name="American Samoa" pageId="8" pageNumber="43">As</collectingCountry>
with 
<taxonomicName id="4C343B9D0004242EF4934781751AF964" box="[453,615,1758,1782]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B9409C0C0004242EF4934781751AF964" box="[453,615,1758,1782]" italics="true" pageId="8" pageNumber="43">L. rotundatum</emphasis>
</taxonomicName>
from 
<collectingCountry id="F323008E0004242EF7E44780758BF964" box="[690,758,1758,1782]" name="Guam" pageId="8" pageNumber="43">Guam</collectingCountry>
and 
<taxonomicName id="4C343B9D0004242EF665478174C6F964" box="[819,955,1758,1782]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C0004242EF665478174C6F964" box="[819,955,1758,1782]" italics="true" pageId="8" pageNumber="43">L. scandens</emphasis>
</taxonomicName>
from 
<collectingCountry id="F323008E0004242EF15047807326F964" box="[1030,1115,1758,1782]" name="Taiwan" pageId="8" pageNumber="43">Taiwan</collectingCountry>
, they are apparently mostly active mainly in wet weather and at night (
<bibRefCitation id="EFA53DEF0004242EF722465C745AF888" author="Ng" box="[628,807,1794,1818]" pageId="8" pageNumber="43" refString="Jeng, M. - S., Liu, H. - C., Tzeng, C. - S. &amp; Ng, P. K. L. (2003) On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana, 76 (2), 227 - 240." type="journal article" year="2003">Ng &amp; Liu 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E13950004242EF5914678764CF8F0" pageId="8" pageNumber="43" type="distribution">
<paragraph id="8B8B401E0004242EF5914678764CF8F0" blockId="8.[151,1437,749,2035]" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF5914678761DF8AD" bold="true" box="[199,352,1830,1855]" pageId="8" pageNumber="43">Distribution.</emphasis>
Christmas I. (
<bibRefCitation id="EFA53DEF0004242EF4A8467875BFF8AC" author="Hicks" box="[510,706,1830,1854]" pageId="8" pageNumber="43" refString="Hicks, J., Rumpff, H. &amp; Yorkston, H. (1984) Christmas Crabs. Christmas Island Natural History Association, 1990 second edition, 81 pp." type="book" year="1984">
Hicks 
<emphasis id="B9409C0C0004242EF71E467975FDF8AC" box="[584,640,1830,1854]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
1984
</bibRefCitation>
); 
<collectingCountry id="F323008E0004242EF7804678744DF8AC" box="[726,816,1830,1854]" name="India" pageId="8" pageNumber="43">Nicobar</collectingCountry>
Is. (
<bibRefCitation id="EFA53DEF0004242EF6364678748BF8AC" author="Alcock" box="[864,1014,1830,1854]" pageId="8" pageNumber="43" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (3), 279 - 486." type="journal article" year="1900">Alcock 1900</bibRefCitation>
), Nias (
<bibRefCitation id="EFA53DEF0004242EF11D467873ABF8AC" author="Nobili" box="[1099,1238,1830,1854]" pageId="8" pageNumber="43" refString="Nobili, G. (1899) Contribuzioni alla conoscienza della fauna carcinologica della Papuasia, delle Molucche e dell'Australia. Annali dei Museo Civico di Storia Naturale di Genova, (2) 20, 230 - 282." type="journal article" year="1899">Nobili 1899</bibRefCitation>
, 
<bibRefCitation id="EFA53DEF0004242EF1B646787261F8AC" author="Nobili" box="[1248,1308,1830,1854]" pageId="8" pageNumber="43" refString="Nobili, G. (1900) Decapodi e Stomatopodi Indo-Malesi. Annali del Museo Civico di Storia Naturale &quot; G. Doria &quot;, 40, 473 - 523." type="journal article" year="1900">1900</bibRefCitation>
), west Java (
<bibRefCitation id="EFA53DEF0004242EF5C846147659F8F0" author="Tesch" box="[158,292,1866,1890]" pageId="8" pageNumber="43" refString="Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, 3, 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch 1917</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E13950004242FF591463074DFFF46" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43" type="materials_examined">
<paragraph id="8B8B401E0004242FF591463074DFFF46" blockId="8.[151,1437,749,2035]" lastBlockId="9.[151,1435,151,212]" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
<emphasis id="B9409C0C0004242EF591463076A5F815" bold="true" box="[199,472,1902,1927]" pageId="8" pageNumber="43">Comparative material.</emphasis>
In addition to the specimens listed in 
<bibRefCitation id="EFA53DEF0004242EF6D84631732CF814" author="Ng" box="[910,1105,1902,1927]" pageId="8" pageNumber="43" refString="Jeng, M. - S., Liu, H. - C., Tzeng, C. - S. &amp; Ng, P. K. L. (2003) On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana, 76 (2), 227 - 240." type="journal article" year="2003">Ng &amp; Liu (2003)</bibRefCitation>
for 
<taxonomicName id="4C343B9D0004242EF1D44631725EF814" box="[1154,1315,1902,1926]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" sensu="lato" species="rotundatum">
<emphasis id="B9409C0C0004242EF1D44631725EF814" box="[1154,1315,1902,1926]" italics="true" pageId="8" pageNumber="43">L. rotundatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27321770004242EF07D463172E1F814" box="[1323,1436,1902,1926]" pageId="8" pageNumber="43" sensu="lato">sensu lato</taxonomicNameLabel>
and 
<taxonomicName id="4C343B9D0004242EF59046CD7637F838" box="[198,330,1938,1962]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C0004242EF59046CD7637F838" box="[198,330,1938,1962]" italics="true" pageId="8" pageNumber="43">L. scandens</emphasis>
</taxonomicName>
, the following new material was examined: 
<taxonomicName id="4C343B9D0004242EF66E46CC739CF838" authority="Ng &amp; Liu, 2003" authorityName="Ng &amp; Liu" authorityYear="2003" box="[824,1249,1938,1963]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="scandens">
<emphasis id="B9409C0C0004242EF66E46CC735BF838" box="[824,1062,1938,1962]" italics="true" pageId="8" pageNumber="43">Labuanium scandens</emphasis>
<bibRefCitation id="EFA53DEF0004242EF17A46CD739CF838" author="Ng" box="[1068,1249,1938,1963]" pageId="8" pageNumber="43" refString="Jeng, M. - S., Liu, H. - C., Tzeng, C. - S. &amp; Ng, P. K. L. (2003) On the taxonomy and ecology of Labuanium trapezoideum (Decapoda, Brachyura, Sesarmidae), a crab living on riverine cliffs in Taiwan. Crustaceana, 76 (2), 227 - 240." type="journal article" year="2003">Ng &amp; Liu, 2003</bibRefCitation>
</taxonomicName>
— 
<specimenCount id="9D328B970004242EF1A946CC7239F838" box="[1279,1348,1938,1962]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
(29.6 × 
<quantity id="4CCCEDFB0004242EF5C146E87782F85C" box="[151,255,1974,1998]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="8" pageNumber="43" unit="mm" value="28.7">28.7 mm</quantity>
), 
<specimenCount id="9D328B970004242EF44446E8760EF85C" box="[274,371,1974,1998]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="ED25D8DB0004242EF4D746E876C4F85C" box="[385,441,1974,1998]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="43">ZRC</collectionCode>
2002.478), Hsiang Chiao Wan, Kentin National Park, southern 
<collectingCountry id="F323008E0004242EF1D046E873A6F85C" box="[1158,1243,1974,1998]" name="Taiwan" pageId="8" pageNumber="43">Taiwan</collectingCountry>
, coll. P. K. L. Ng &amp; C. H. Wang, 
<date id="FF8A66DE0004242EF41B46847684F860" box="[333,505,2010,2035]" pageId="8" pageNumber="43" value="2002-08-05">5 August 2002</date>
. 
<taxonomicName id="4C343B9D0004242EF75346847473F860" box="[517,782,2010,2034]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="rotundatum">
<emphasis id="B9409C0C0004242EF75346847473F860" box="[517,782,2010,2034]" italics="true" pageId="8" pageNumber="43">Labuanium rotundatum</emphasis>
</taxonomicName>
— 
<specimenCount id="9D328B970004242EF67A468474FEF860" box="[812,899,2010,2034]" pageId="8" pageNumber="43" type="male">2 males</specimenCount>
(24.0 × 
<quantity id="4CCCEDFB0004242EF6BE4684732DF860" box="[1000,1104,2010,2034]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="8" pageNumber="43" unit="mm" value="24.2">24.2 mm</quantity>
, 26.5 × 
<quantity id="4CCCEDFB0004242EF1F946847264F860" box="[1199,1305,2010,2034]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="8" pageNumber="43" unit="mm" value="27.5">27.5 mm</quantity>
), 
<specimenCount id="9D328B970004242EF078468472E1F860" box="[1326,1436,2010,2034]" pageId="8" pageNumber="43" type="female">2 females</specimenCount>
(24.7 × 
<quantity id="4CCCEDFB0005242FF5B941C9762BFF3D" box="[239,342,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="9" pageNumber="44" unit="mm" value="24.7">24.7 mm</quantity>
; 17.9 × 
<quantity id="4CCCEDFB0005242FF4E741C9756BFF3D" box="[433,534,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="44" unit="mm" value="17.8">17.8 mm</quantity>
) (
<typeStatus id="548FFEBC0005242FF77B41C975ECFF3D" box="[557,657,151,175]" pageId="9" pageNumber="44" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C343B9D0005242FF7E141C67331FF3D" authority="Borradaile, 1900" authorityName="Borradaile" authorityYear="1900" box="[695,1100,151,176]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B9409C0C0005242FF7E141C674FBFF22" box="[695,902,152,176]" italics="true" pageId="9" pageNumber="44">Sesarma gardineri</emphasis>
<bibRefCitation id="EFA53DEF0005242FF6D841C97331FF3D" author="Borradaile" box="[910,1100,151,175]" pageId="9" pageNumber="44" refString="Borradaile, L. A. (1900) On some Crustaceans from the South Pacific. Part IV. The Crabs. Proceedings of the Zoological Society of London, 1900, 568 - 596, pls. 40 - 42." type="proceedings paper" year="1900">Borradaile, 1900</bibRefCitation>
</taxonomicName>
) (Cambridge University Zoology Museum), Ellice Is., Funafuti, 
<collectingCountry id="F323008E0005242FF77141E27505FF46" box="[551,632,188,212]" name="Tuvalu" pageId="9" pageNumber="44">Tuvalu</collectingCountry>
, coll. J. S. Gardiner, 1897.
</paragraph>
</subSubSection>
</treatment>
</document>