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<document id="18E56851F3ACF3B4C459B653F07AEE50" ID-DOI="10.11646/zootaxa.3647.1.6" ID-GBIF-Dataset="18569fb0-f346-456c-8685-831e84073e7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="221386" ID-ZooBank="D57B95E8-3991-4B30-B22D-83978924D05E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460603778594" checkinUser="plazi" docAuthor="Stauffer Jr, Jay R., Black, Kristin &amp; Konings, Adrianus F." docDate="2013" docId="039DED3DA2413D4903E7CDA89C80F861" docLanguage="en" docName="zt03647p136.pdf" docOrigin="Zootaxa 3647 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metriaclima tarakiki Jr, Black &amp; Konings, 2013, new species" docType="treatment" docVersion="10" lastPageNumber="119" masterDocId="FFA49545A2513D5B0370CC749F49FFFD" masterDocTitle="Descriptions of five new species of Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa" masterLastPageNumber="136" masterPageNumber="101" pageNumber="117" updateTime="1698327745829" updateUser="plazi">
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<mods:title id="217D09C390FFCBE4C224231BC2DB0ACC">Descriptions of five new species of Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa</mods:title>
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<mods:namePart id="8E0AEF882F22C368F1A153CB450E5CB8">Stauffer Jr, Jay R.</mods:namePart>
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<mods:date id="B7DEF2412A705EA426C644484A1996FB">2013</mods:date>
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<treatment id="039DED3DA2413D4903E7CDA89C80F861" ID-DOI="http://doi.org/10.5281/zenodo.6153915" ID-GBIF-Taxon="119567620" ID-Zenodo-Dep="6153915" LSID="urn:lsid:plazi:treatment:039DED3DA2413D4903E7CDA89C80F861" httpUri="http://treatment.plazi.org/id/039DED3DA2413D4903E7CDA89C80F861" lastPageId="18" lastPageNumber="119" pageId="16" pageNumber="117">
<subSubSection id="C32E0FA0A2413D4B03E7CDA89D02FD87" pageId="16" pageNumber="117" type="nomenclature">
<paragraph id="8B8B5C2BA2413D4B03E7CDA89D66FE08" blockId="16.[151,559,475,534]" box="[151,559,475,502]" pageId="16" pageNumber="117">
<heading id="D0C3EB47A2413D4B03E7CDA89D66FE08" bold="true" box="[151,559,475,502]" fontSize="11" level="1" pageId="16" pageNumber="117" reason="1">
<emphasis id="B9408039A2413D4B03E7CDA89D66FE08" bold="true" box="[151,559,475,502]" pageId="16" pageNumber="117">
<taxonomicName id="4C3427A8A2413D4B03E7CDA89ED9FE0B" ID-CoL="42HWV" box="[151,400,476,502]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="tarakiki" status="sp. nov.">
<emphasis id="B9408039A2413D4B03E7CDA89ED9FE0B" bold="true" box="[151,400,476,502]" italics="true" pageId="16" pageNumber="117">Metriaclima tarakiki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2733D42A2413D4B02EFCDAF9D66FE08" box="[415,559,475,501]" pageId="16" pageNumber="117" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B8B5C2BA2413D4B03E7CD8A9E44FDEB" blockId="16.[151,559,475,534]" box="[151,269,510,534]" pageId="16" pageNumber="117">
<figureCitation id="130F40AEA2413D4B03E7CD8A9F9EFDEB" box="[151,215,510,534]" captionStart="FIGURE 7" captionStartId="16.[151,250,1895,1918]" captionTargetBox="[151,1435,905,1874]" captionTargetId="figure@16.[151,1436,862,1874]" captionTargetPageId="16" captionText="FIGURE 7. Metriaclima tarakiki. A. Holotype, PSU 4914, adult male, 115.2 mm SL, Tanzania: Higga Reef; B. Male in breeding coloration (approx. 110 mm SL) at Higga Reef, Tanzania; C. Female (approx. 90 mm SL) at Higga Reef, Tanzania." httpUri="https://zenodo.org/record/221392/files/figure.png" pageId="16" pageNumber="117">Fig. 7</figureCitation>
AC
</paragraph>
<paragraph id="8B8B5C2BA2413D4B03E7CE339D02FD87" blockId="16.[151,786,582,635]" pageId="16" pageNumber="117">
<taxonomicName id="4C3427A8A2413D4B03E7CE339E70FDA0" authority="Zebra" authorityName="Zebra" box="[151,313,583,605]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="B9408039A2413D4B03E7CE339E70FDA0" box="[151,313,583,605]" italics="true" pageId="16" pageNumber="117">Pseudotropheus</emphasis>
</taxonomicName>
'
<taxonomicName id="4C3427A8A2413D4B0232CE329E36FDA1" box="[322,383,582,604]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="135" phylum="Chordata" rank="species" species="zebra">Zebra</taxonomicName>
Mbamba Bay Kompakt', Spreinat 1994
<taxonomicName id="4C3427A8A2413D4B03E7CE119E5DFD86" box="[151,276,613,635]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="B9408039A2413D4B03E7CE119E5DFD86" box="[151,276,613,635]" italics="true" pageId="16" pageNumber="117">Metriaclima</emphasis>
</taxonomicName>
sp. '
<taxonomicName id="4C3427A8A2413D4B0230CE119E30FD86" box="[320,377,613,635]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="zebra">zebra</taxonomicName>
slim', Konings 2001
</paragraph>
</subSubSection>
<subSubSection id="C32E0FA0A2413D4B03E7CED79A2BFCDB" pageId="16" pageNumber="117" type="materials_examined">
<paragraph id="8B8B5C2BA2413D4B03E7CED79C41FD22" blockId="16.[151,1436,674,807]" pageId="16" pageNumber="117">
<emphasis id="B9408039A2413D4B03E7CED79E42FD46" bold="true" box="[151,267,675,699]" pageId="16" pageNumber="117">
<typeStatus id="548FE289A2413D4B03E7CED79E4FFD46" box="[151,262,675,699]" pageId="16" pageNumber="117" type="holotype">Holotype</typeStatus>
.
</emphasis>
PSU 4914, adult male,
<quantity id="4CCCF1CEA2413D4B0165CED69DCFFD47" box="[533,646,674,698]" metricMagnitude="-1" metricUnit="m" metricValue="1.152" pageId="16" pageNumber="117" unit="mm" value="115.2">115.2 mm</quantity>
SL,
<geoCoordinate id="EE003AECA2413D4B01CBCED69C19FD47" box="[699,848,674,698]" direction="south" orientation="latitude" pageId="16" pageNumber="117" precision="1" value="-11.306334">11°18.380 S</geoCoordinate>
,
<geoCoordinate id="EE003AECA2413D4B002CCED69CBDFD47" box="[860,1012,674,698]" direction="east" orientation="longitude" pageId="16" pageNumber="117" precision="1" value="34.746517">34°44.791 E</geoCoordinate>
, Higga Reef, Lake
<collectingCountry id="F3231CBBA2413D4B07BDCED69A6CFD46" box="[1229,1317,674,699]" name="Malawi" pageId="16" pageNumber="117">Malaŵi</collectingCountry>
,
<collectingCountry id="F3231CBBA2413D4B0640CED69AD1FD47" box="[1328,1432,674,698]" name="Tanzania" pageId="16" pageNumber="117">Tanzania</collectingCountry>
, Africa, A. F. Konings &amp; J. R. Stauffer Jr.,
<date id="FF8A7AEBA2413D4B011ECEB29C48FD23" box="[622,769,710,734]" pageId="16" pageNumber="117" value="2005-02-13">13 Feb. 2005</date>
.
</paragraph>
<paragraph id="8B8B5C2BA2413D4B03B9CE9F9A2BFCDB" blockId="16.[151,1436,674,807]" pageId="16" pageNumber="117">
<emphasis id="B9408039A2413D4B03B9CE9F9E01FCFE" bold="true" box="[201,328,747,771]" pageId="16" pageNumber="117">
<typeStatus id="548FE289A2413D4B03B9CE9F9E0AFCFE" box="[201,323,747,771]" pageId="16" pageNumber="117" type="paratype">Paratypes</typeStatus>
.
</emphasis>
PSU 4915, 11,
<quantity id="4CCCF1CEA2413D4B0177CE9E9DF3FCFF" box="[519,698,746,770]" metricMagnitude="-2" metricUnit="m" metricValue="9.575" metricValueMax="11.93" metricValueMin="7.22" pageId="16" pageNumber="117" unit="mm" value="95.75" valueMax="119.3" valueMin="72.2">72.2119.3 mm</quantity>
SL, same data as
<typeStatus id="548FE289A2413D4B00E7CE9E9CB4FCFF" box="[919,1021,746,770]" pageId="16" pageNumber="117" type="holotype">holotype</typeStatus>
; PSU 4916, 18, AMNH 257796, 2, (
<quantity id="4CCCF1CEA2413D4B03EECF7A9E0AFCDB" box="[158,323,782,806]" metricMagnitude="-2" metricUnit="m" metricValue="8.635" metricValueMax="9.68" metricValueMin="7.59" pageId="16" pageNumber="117" unit="mm" value="86.35" valueMax="96.8" valueMin="75.9">75.996.8 mm</quantity>
SL), Ngkuyo Island, Lake
<collectingCountry id="F3231CBBA2413D4B0104CF7A9D85FCDA" box="[628,716,782,807]" name="Malawi" pageId="16" pageNumber="117">Malaŵi</collectingCountry>
,
<collectingCountry id="F3231CBBA2413D4B01A8CF7A9C09FCDB" box="[728,832,782,806]" name="Tanzania" pageId="16" pageNumber="117">Tanzania</collectingCountry>
, A. F. Konings &amp; J. R. Stauffer Jr.,
<date id="FF8A7AEBA2413D4B07BCCF7A9A17FCDB" box="[1228,1374,782,806]" pageId="16" pageNumber="117" value="2004-01-29">29 Jan. 2004</date>
.
</paragraph>
</subSubSection>
<caption id="DF4B0CA3A2413D4B03E7CB139ACDF861" httpUri="https://zenodo.org/record/221392/files/figure.png" pageId="16" pageNumber="117" targetBox="[151,1435,905,1874]" targetPageId="16">
<paragraph id="8B8B5C2BA2413D4B03E7CB139ACDF861" blockId="16.[151,1436,1895,1949]" pageId="16" pageNumber="117">
<emphasis id="B9408039A2413D4B03E7CB139E59F880" bold="true" box="[151,272,1895,1918]" pageId="16" pageNumber="117">FIGURE 7</emphasis>
.
<taxonomicName id="4C3427A8A2413D4B0250CB139EBEF880" box="[288,503,1895,1917]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2413D4B0250CB139EBEF880" box="[288,503,1895,1917]" italics="true" pageId="16" pageNumber="117">Metriaclima tarakiki</emphasis>
</taxonomicName>
. A. Holotype, PSU 4914, adult male, 115.2 mm SL, Tanzania: Higga Reef; B. Male in breeding coloration (approx. 110 mm SL) at Higga Reef, Tanzania; C. Female (approx. 90 mm SL) at Higga Reef, Tanzania.
</paragraph>
</caption>
<caption id="DF4B0CA3A2403D4A03E7CCE39B1DFF32" ID-Table-UUID="DF4B0CA3A2403D4A03E7CCE39B1DFF32" httpUri="http://table.plazi.org/id/DF4B0CA3A2403D4A03E7CCE39B1DFF32" pageId="17" pageNumber="118" targetBox="[159,1345,234,2023]" targetIsTable="true" targetPageId="17">
<paragraph id="8B8B5C2BA2403D4A03E7CCE39B1DFF32" blockId="17.[151,1436,151,207]" pageId="17" pageNumber="118">
<emphasis id="B9408039A2403D4A03E7CCE39E40FF52" bold="true" box="[151,265,151,175]" pageId="17" pageNumber="118">TABLE 5.</emphasis>
Morphometric and meristic values for
<taxonomicName id="4C3427A8A2403D4A01DACCEC9CCEFF52" box="[682,903,152,175]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2403D4A01DACCEC9CCEFF52" box="[682,903,152,175]" italics="true" pageId="17" pageNumber="118">Metriaclima tarakiki</emphasis>
</taxonomicName>
from Higga Reef, Tanzania (PSU 4914, holotype; PSU 4915, n=11) and Ngkuyo, Tanzania (PSU 4916, n=20). Ranges include the holotype.
</paragraph>
</caption>
<paragraph id="8B8B5C2BA2403D4A03EFCC9E9B99F81A" pageId="17" pageNumber="118">
<table id="F934AE8BA240C2A403EFCC9E9A08F81A" box="[159,1345,234,2023]" gridcols="5" gridrows="12" pageId="17" pageNumber="118">
<tr id="35045E69A240C2A403EFCC9E9A08FE39" box="[159,1345,234,452]" gridrow="0" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCC9E9D53FE39" box="[159,538,234,452]" gridcol="0" gridrow="0" pageId="17" pageNumber="118">
<taxonomicName id="4C3427A8A2403D4A03EFCC9E9FB0FEB5" authority="Standard" authorityName="Standard" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="tarakiki">Metriaclima tarakiki Standard</taxonomicName>
length, mm Head length, mm Percent standard length Head length
</th>
<th id="76D53715A240C2A401CDCC9E9C6AFE39" box="[701,803,234,452]" gridcol="1" gridrow="0" pageId="17" pageNumber="118">Holotype 115.2 34.0 33.9</th>
<th id="76D53715A240C2A4001DCC9E9CE5FE39" box="[877,940,234,452]" gridcol="2" gridrow="0" pageId="17" pageNumber="118">Mean 90.0 26.6 33.8</th>
<th id="76D53715A240C2A40096CC9E9B13FE39" box="[998,1114,234,452]" gridcol="3" gridrow="0" pageId="17" pageNumber="118">Higga Reef Range 72.2119.3 21.935.8 33.8</th>
<th id="76D53715A240C2A407DBCC9E9A08FE39" box="[1195,1345,234,452]" gridcol="4" gridrow="0" pageId="17" pageNumber="118">Ngkuyo Island Range 75.996.8 21.728.1 34.0</th>
</tr>
<tr id="35045E69A240C2A403EFCDAC9A08FD94" box="[159,1345,472,617]" gridrow="1" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCDAC9D53FD94" box="[159,538,472,617]" gridcol="0" gridrow="1" pageId="17" pageNumber="118">Body depth Snout to dorsal Snout to pelvic Dorsal-fin base length</th>
<td id="76D53715A240C2A401CDCDAC9C6AFD94" box="[701,803,472,617]" gridcol="1" gridrow="1" pageId="17" pageNumber="118">32.9 31.3 36.4 61.7</td>
<td id="76D53715A240C2A4001DCDAC9CE5FD94" box="[877,940,472,617]" gridcol="2" gridrow="1" pageId="17" pageNumber="118">28.8 30.9 36.5 59.9</td>
<td id="76D53715A240C2A40096CDAC9B13FD94" box="[998,1114,472,617]" gridcol="3" gridrow="1" pageId="17" pageNumber="118">28.232.9 28.532.7 35.339.3 59.461.8</td>
<td id="76D53715A240C2A407DBCDAC9A08FD94" box="[1195,1345,472,617]" gridcol="4" gridrow="1" pageId="17" pageNumber="118">25.829.1 29.433.0 34.739.0 56.762.4</td>
</tr>
<tr id="35045E69A240C2A403EFCE099A08FD41" box="[159,1345,637,700]" gridrow="2" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCE099D53FD41" box="[159,538,637,700]" gridcol="0" gridrow="2" pageId="17" pageNumber="118">Anterior dorsal to anterior anal Anterior dorsal to posterior anal</th>
<td id="76D53715A240C2A401CDCE099C6AFD41" box="[701,803,637,700]" gridcol="1" gridrow="2" pageId="17" pageNumber="118">51.9 63.5</td>
<td id="76D53715A240C2A4001DCE099CE5FD41" box="[877,940,637,700]" gridcol="2" gridrow="2" pageId="17" pageNumber="118">48.4 61.6</td>
<td id="76D53715A240C2A40096CE099B13FD41" box="[998,1114,637,700]" gridcol="3" gridrow="2" pageId="17" pageNumber="118">47.651.9 60.963.9</td>
<td id="76D53715A240C2A407DBCE099A08FD41" box="[1195,1345,637,700]" gridcol="4" gridrow="2" pageId="17" pageNumber="118">45.850.0 59.563.7</td>
</tr>
<tr id="35045E69A240C2A403EFCEA49A08FC49" box="[159,1345,720,948]" gridrow="3" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCEA49D53FC49" box="[159,538,720,948]" gridcol="0" gridrow="3" pageId="17" pageNumber="118">Posterior dorsal to anterior anal Posterior dorsal to posterior anal Posterior dorsal to ventral caudal Posterior anal to dorsal caudal Anterior dorsal to pelvic-fin origin Posterior dorsal to pelvic-fin origin</th>
<td id="76D53715A240C2A401CDCEA49C6AFC49" box="[701,803,720,948]" gridcol="1" gridrow="3" pageId="17" pageNumber="118">30.4 17.0 20.2 22.8 35.6 61.9</td>
<td id="76D53715A240C2A4001DCEA49CE5FC49" box="[877,940,720,948]" gridcol="2" gridrow="3" pageId="17" pageNumber="118">27.7 15.3 18.8 21.6 31.7 58.5</td>
<td id="76D53715A240C2A40096CEA49B13FC49" box="[998,1114,720,948]" gridcol="3" gridrow="3" pageId="17" pageNumber="118">26.630.8 15.417.0 18.520.2 20.622.8 31.936.0 56.962.0</td>
<td id="76D53715A240C2A407DBCEA49A08FC49" box="[1195,1345,720,948]" gridcol="4" gridrow="3" pageId="17" pageNumber="118">25.329.0 13.215.5 16.420.5 20.322.6 28.133.1 56.059.5</td>
</tr>
<tr id="35045E69A240C2A403EFCFBC9A08FBA4" box="[159,1345,968,1113]" gridrow="4" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCFBC9D53FBA4" box="[159,538,968,1113]" gridcol="0" gridrow="4" pageId="17" pageNumber="118">Caudal-peduncle length Least caudal-peduncle depth Percent head length Snout length</th>
<td id="76D53715A240C2A401CDCFBC9C6AFBA4" box="[701,803,968,1113]" gridcol="1" gridrow="4" pageId="17" pageNumber="118">15.0 12.3 39.8</td>
<td id="76D53715A240C2A4001DCFBC9CE5FBA4" box="[877,940,968,1113]" gridcol="2" gridrow="4" pageId="17" pageNumber="118">15.4 10.8 37.5</td>
<td id="76D53715A240C2A40096CFBC9B13FBA4" box="[998,1114,968,1113]" gridcol="3" gridrow="4" pageId="17" pageNumber="118">13.715.5 10.712.3 37.142.2</td>
<td id="76D53715A240C2A407DBCFBC9A08FBA4" box="[1195,1345,968,1113]" gridcol="4" gridrow="4" pageId="17" pageNumber="118">14.717.5 9.711.5 32.939.9</td>
</tr>
<tr id="35045E69A240C2A403EFC8199A08FB51" box="[159,1345,1133,1196]" gridrow="5" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFC8199D53FB51" box="[159,538,1133,1196]" gridcol="0" gridrow="5" pageId="17" pageNumber="118">Postorbital head length Horizontal eye diameter</th>
<td id="76D53715A240C2A401CDC8199C6AFB51" box="[701,803,1133,1196]" gridcol="1" gridrow="5" pageId="17" pageNumber="118">39.1 27.2</td>
<td id="76D53715A240C2A4001DC8199CE5FB51" box="[877,940,1133,1196]" gridcol="2" gridrow="5" pageId="17" pageNumber="118">38.7 32.5</td>
<td id="76D53715A240C2A40096C8199B13FB51" box="[998,1114,1133,1196]" gridcol="3" gridrow="5" pageId="17" pageNumber="118">37.740.6 27.235.0</td>
<td id="76D53715A240C2A407DBC8199A08FB51" box="[1195,1345,1133,1196]" gridcol="4" gridrow="5" pageId="17" pageNumber="118">36.040.8 30.636.3</td>
</tr>
<tr id="35045E69A240C2A403EFC8B49A08FAAC" box="[159,1345,1216,1361]" gridrow="6" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFC8B49D53FAAC" box="[159,538,1216,1361]" gridcol="0" gridrow="6" pageId="17" pageNumber="118">Vertical eye diameter Head depth Preorbital depth Cheek depth</th>
<td id="76D53715A240C2A401CDC8B49C6AFAAC" box="[701,803,1216,1361]" gridcol="1" gridrow="6" pageId="17" pageNumber="118">28.4 90.6 23.5 29.9</td>
<td id="76D53715A240C2A4001DC8B49CE5FAAC" box="[877,940,1216,1361]" gridcol="2" gridrow="6" pageId="17" pageNumber="118">32.6 81.9 22.5 26.5</td>
<td id="76D53715A240C2A40096C8B49B13FAAC" box="[998,1114,1216,1361]" gridcol="3" gridrow="6" pageId="17" pageNumber="118">28.233.5 77.496.4 20.525.1 23.730.8</td>
<td id="76D53715A240C2A407DBC8B49A08FAAC" box="[1195,1345,1216,1361]" gridcol="4" gridrow="6" pageId="17" pageNumber="118">30.236.5 73.385.1 20.823.2 23.329.2</td>
</tr>
<tr id="35045E69A240C2A403EFC9119A08FA59" box="[159,1345,1381,1444]" gridrow="7" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFC9119D53FA59" box="[159,538,1381,1444]" gridcol="0" gridrow="7" pageId="17" pageNumber="118">Lower-jaw length Meristics</th>
<td id="76D53715A240C2A401CDC9119C6AFA59" box="[701,803,1381,1444]" gridcol="1" gridrow="7" pageId="17" pageNumber="118">35.2 Holotype</td>
<td id="76D53715A240C2A4001DC9119CE5FA59" box="[877,940,1381,1444]" gridcol="2" gridrow="7" pageId="17" pageNumber="118">37.1 Mode</td>
<td id="76D53715A240C2A40096C9119B13FA59" box="[998,1114,1381,1444]" gridcol="3" gridrow="7" pageId="17" pageNumber="118">35.039.7 Higga Reef</td>
<td id="76D53715A240C2A407DBC9119A08FA59" box="[1195,1345,1381,1444]" gridcol="4" gridrow="7" pageId="17" pageNumber="118">34.838.9 Ngkuyo Island</td>
</tr>
<tr id="35045E69A240C2A403EFC9CC9A08F961" box="[159,1345,1464,1692]" gridrow="8" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFC9CC9D53F961" box="[159,538,1464,1692]" gridcol="0" gridrow="8" pageId="17" pageNumber="118">Dorsal-fin spines Dorsal-fin rays Anal-fin spines Anal-fin rays Pectoral-fin rays Pelvic-fin rays</th>
<td id="76D53715A240C2A401CDC9CC9C6AF961" box="[701,803,1464,1692]" gridcol="1" gridrow="8" pageId="17" pageNumber="118">18 8 3 8 15 5</td>
<td id="76D53715A240C2A4001DC9CC9CE5F961" box="[877,940,1464,1692]" gridcol="2" gridrow="8" pageId="17" pageNumber="118">18 9 3 8 14 5</td>
<td id="76D53715A240C2A40096C9CC9B13F961" box="[998,1114,1464,1692]" gridcol="3" gridrow="8" pageId="17" pageNumber="118">18 89 34 8 1415 5</td>
<td id="76D53715A240C2A407DBC9CC9A08F961" box="[1195,1345,1464,1692]" gridcol="4" gridrow="8" pageId="17" pageNumber="118">1719 810 33 89 1415 5</td>
</tr>
<tr id="35045E69A240C2A403EFCAC49A08F8BC" box="[159,1345,1712,1857]" gridrow="9" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCAC49D53F8BC" box="[159,538,1712,1857]" gridcol="0" gridrow="9" pageId="17" pageNumber="118">Lateral-line scales Pored scales post lateral line Cheek-scale rows Gillrakers 1st ceratobranchial</th>
<td id="76D53715A240C2A401CDCAC49C6AF8BC" box="[701,803,1712,1857]" gridcol="1" gridrow="9" pageId="17" pageNumber="118">31 0 5 11</td>
<td id="76D53715A240C2A4001DCAC49CE5F8BC" box="[877,940,1712,1857]" gridcol="2" gridrow="9" pageId="17" pageNumber="118">32 1 4 11</td>
<td id="76D53715A240C2A40096CAC49B13F8BC" box="[998,1114,1712,1857]" gridcol="3" gridrow="9" pageId="17" pageNumber="118">3132 02 45 1112</td>
<td id="76D53715A240C2A407DBCAC49A08F8BC" box="[1195,1345,1712,1857]" gridcol="4" gridrow="9" pageId="17" pageNumber="118">3233 12 45 1012</td>
</tr>
<tr id="35045E69A240C2A403EFCB219A08F869" box="[159,1345,1877,1940]" gridrow="10" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCB219D53F869" box="[159,538,1877,1940]" gridcol="0" gridrow="10" pageId="17" pageNumber="118">Gillrakers 1st epibranchial Teeth in outer series of left lower jaw</th>
<td id="76D53715A240C2A401CDCB219C6AF869" box="[701,803,1877,1940]" gridcol="1" gridrow="10" pageId="17" pageNumber="118">2 27</td>
<td id="76D53715A240C2A4001DCB219CE5F869" box="[877,940,1877,1940]" gridcol="2" gridrow="10" pageId="17" pageNumber="118">3 2425</td>
<td id="76D53715A240C2A40096CB219B13F869" box="[998,1114,1877,1940]" gridcol="3" gridrow="10" pageId="17" pageNumber="118">23 2431</td>
<td id="76D53715A240C2A407DBCB219A08F869" box="[1195,1345,1877,1940]" gridcol="4" gridrow="10" pageId="17" pageNumber="118">23 2028</td>
</tr>
<tr id="35045E69A240C2A403EFCBDC9A08F81A" box="[159,1345,1960,2023]" gridrow="11" pageId="17" pageNumber="118">
<th id="76D53715A240C2A403EFCBDC9D53F81A" box="[159,538,1960,2023]" gridcol="0" gridrow="11" pageId="17" pageNumber="118">Tooth rows in upper jaw Tooth rows in lower jaw</th>
<td id="76D53715A240C2A401CDCBDC9C6AF81A" box="[701,803,1960,2023]" gridcol="1" gridrow="11" pageId="17" pageNumber="118">6 8</td>
<td id="76D53715A240C2A4001DCBDC9CE5F81A" box="[877,940,1960,2023]" gridcol="2" gridrow="11" pageId="17" pageNumber="118">6 7</td>
<td id="76D53715A240C2A40096CBDC9B13F81A" box="[998,1114,1960,2023]" gridcol="3" gridrow="11" pageId="17" pageNumber="118">57 78</td>
<td id="76D53715A240C2A407DBCBDC9A08F81A" box="[1195,1345,1960,2023]" gridcol="4" gridrow="11" pageId="17" pageNumber="118">48 58</td>
</tr>
</table>
</paragraph>
<subSubSection id="C32E0FA0A2433D4903B9CCE39EC3FDEA" pageId="18" pageNumber="119" type="diagnosis">
<paragraph id="8B8B5C2BA2433D4903B9CCE39EC3FDEA" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">
<emphasis id="B9408039A2433D4903B9CCE39E0AFF4D" bold="true" box="[201,323,151,176]" pageId="18" pageNumber="119">Diagnosis.</emphasis>
The presence of bicuspid teeth in the anterior portion of the outer row on both the upper and lower jaws and a moderately sloped ethmo-vomerine block with a swollen rostral tip place this species in
<taxonomicName id="4C3427A8A2433D49067BCCC89ADCFF29" box="[1291,1429,188,212]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B9408039A2433D49067BCCC89ADCFF29" box="[1291,1429,188,212]" italics="true" pageId="18" pageNumber="119">Metriaclima</emphasis>
</taxonomicName>
. Male
<taxonomicName id="4C3427A8A2433D4903A8CC959E1CFF05" box="[216,341,224,248]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D4903A8CC959E1CFF05" box="[216,341,224,248]" italics="true" pageId="18" pageNumber="119">M. tarakiki</emphasis>
</taxonomicName>
are distinguished from those of the other members of
<taxonomicName id="4C3427A8A2433D4900CBCC949B0CFF05" box="[955,1093,224,248]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B9408039A2433D4900CBCC949B0CFF05" box="[955,1093,224,248]" italics="true" pageId="18" pageNumber="119">Metriaclima</emphasis>
</taxonomicName>
by the blue ground color with distinct black bars, the extension of the lateral bars onto the blue dorsal fin, the blue anal fin, and the absence of a dark submarginal band in the dorsal fin or yellow pigment on the body. While some males of
<taxonomicName id="4C3427A8A2433D4907ACCD5D9A08FEBD" box="[1244,1345,296,320]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B9408039A2433D4907ACCD5D9A08FEBD" box="[1244,1345,296,320]" italics="true" pageId="18" pageNumber="119">M. zebra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3427A8A2433D49060DCD5D9E42FE99" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="fainzilberi">
<emphasis id="B9408039A2433D49060DCD5D9E42FE99" italics="true" pageId="18" pageNumber="119">M. fainzilberi</emphasis>
</taxonomicName>
exhibit extension of the lateral bars into the dorsal fin, these extensions fuse into a broad submarginal band, which is not observed in males of
<taxonomicName id="4C3427A8A2433D490129CD059D9CFE75" box="[601,725,368,392]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D490129CD059D9CFE75" box="[601,725,368,392]" italics="true" pageId="18" pageNumber="119">M. tarakiki</emphasis>
</taxonomicName>
. Male and female
<taxonomicName id="4C3427A8A2433D4900D5CD059B68FE75" box="[933,1057,368,392]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D4900D5CD059B68FE75" box="[933,1057,368,392]" italics="true" pageId="18" pageNumber="119">M. tarakiki</emphasis>
</taxonomicName>
are further distinguished from
<taxonomicName id="4C3427A8A2433D49060DCD059F9CFE51" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B9408039A2433D49060DCD059F9CFE51" italics="true" pageId="18" pageNumber="119">M. zebra</emphasis>
</taxonomicName>
by a shallower body as expressed in the distance between the dorsal fin origin and pelvic fin which ranges from 28.136.0% SL in
<taxonomicName id="4C3427A8A2433D4902DCCDCD9D63FE2D" box="[428,554,440,464]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D4902DCCDCD9D63FE2D" box="[428,554,440,464]" italics="true" pageId="18" pageNumber="119">M. tarakiki</emphasis>
</taxonomicName>
and from 34.240.7% SL in
<taxonomicName id="4C3427A8A2433D490008CDCD9C94FE2D" box="[888,989,440,464]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B9408039A2433D490008CDCD9C94FE2D" box="[888,989,440,464]" italics="true" pageId="18" pageNumber="119">M. zebra</emphasis>
</taxonomicName>
populations from the Tanzanian shores and from Chiwindi in
<collectingCountry id="F3231CBBA2433D4902FECDA89D6CFE09" box="[398,549,476,500]" name="Mozambique" pageId="18" pageNumber="119">Mozambique</collectingCountry>
. On average
<taxonomicName id="4C3427A8A2433D4901C6CDA99C7BFE09" box="[694,818,476,500]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D4901C6CDA99C7BFE09" box="[694,818,476,500]" italics="true" pageId="18" pageNumber="119">M. tarakiki</emphasis>
</taxonomicName>
has more tooth rows on the lower jaw (range 58) than
<taxonomicName id="4C3427A8A2433D4903E7CE759FB3FDE5" box="[151,250,512,536]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B9408039A2433D4903E7CE759FB3FDE5" box="[151,250,512,536]" italics="true" pageId="18" pageNumber="119">M. zebra</emphasis>
</taxonomicName>
(range 35).
</paragraph>
</subSubSection>
<subSubSection id="C32E0FA0A2433D4903B9CE579B4EF915" pageId="18" pageNumber="119" type="description">
<paragraph id="8B8B5C2BA2433D4903B9CE579C02FCC7" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">
<emphasis id="B9408039A2433D4903B9CE579E13FDC1" bold="true" box="[201,346,547,572]" pageId="18" pageNumber="119">Description.</emphasis>
Morphometric and meristic data in
<tableCitation id="C6B66990A2433D490186CE509C04FDC1" box="[758,845,548,572]" captionStart="TABLE 5" captionStartId="17.[151,239,151,175]" captionTargetBox="[159,1345,234,2023]" captionTargetPageId="17" captionText="TABLE 5. Morphometric and meristic values for Metriaclima tarakiki from Higga Reef, Tanzania (PSU 4914, holotype; PSU 4915, n = 11) and Ngkuyo, Tanzania (PSU 4916, n = 20). Ranges include the holotype." httpUri="http://table.plazi.org/id/DF4B0CA3A2403D4A03E7CCE39B1DFF32" pageId="18" pageNumber="119" tableUuid="DF4B0CA3A2403D4A03E7CCE39B1DFF32">Table 5</tableCitation>
. Moderately elongate species (mean BD 28.8% SL) with greatest body depth at about base of 6th dorsal spine. Dorsal body profile with slight curve to caudal peduncle with highest point at about sixth dorsal spine; ventral body profile between pelvic and anal fin slightly convex with upward curve to caudal fin. Dorsal head profile straight to slightly concave between snout tip and interorbital, making about 45° angle with body axis, then round above orbit to dorsal-fin origin; eye (mean 32.5% HL) about one and a half times depth of preorbital and present for ¾ in anterior half of head with posterior orbit margin posterior of vertical median of head. Snout short with isognathous to slightly prognathous jaws; teeth on lower jaw in 58 rows with outer row bicuspid and inner rows tricuspid.
</paragraph>
<paragraph id="8B8B5C2BA2433D4903B9CF339ED2FB9D" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">Dorsal fin with XVIIXIX (mode XVIII) spines and 810 (mode 9) rays. Anal fin with III spines and 8 or 9 (mode 8) rays. First 4 or 5 dorsal spines gradually increasing in length posteriorly with first spine about ½ length of fourth spine; last 13 spines slightly longer posteriorly with last spine longest; dorsal-fin rays with subacuminate tip in males, round in females, third or fourth ray longest, about to middle of caudal fin in males and to base of caudal fin in females. Anal spines longer posteriorly; fourth or fifth ray longest, length to about ¼ caudal fin in some males, to about base if caudal fin in females. Caudal fin subtruncate to emarginate. Pelvic fin not to anal pore in females; length in males about to anal fin. Pectoral fin rounded, paddle-shaped, short, to vertical through base of 9th or 10th dorsal-fin spine.</paragraph>
<paragraph id="8B8B5C2BA2433D4903B9C8189E2FFB31" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">Flank scales large, ctenoid; abrupt difference to small scales on breast and belly; cheek with 4 or 5 rows of small scales. Dorsal-fin rays and anal fin with narrow proximal margin of tiny scales; tiny scales over proximal 90% of caudal fin.</paragraph>
<paragraph id="8B8B5C2BA2433D4903B9C8AC9D28FA59" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">Breeding males with blue/white flank and 79 black lateral bars; caudal peduncle blue/gray; belly gray; breast dark gray. Head blue/black with two blue interorbital bars; throat blue/gray; opercle with light blue and green highlights. Dorsal fin blue to white-blue with some lateral bars on proximal portion of dorsal fin; usually with white marginal band. Proximal 2/3 of caudal fin gray/blue and distal 1/3 dark gray. Anal fin blue/gray; posterior margin clear in some individuals; 28 yellow ocelli. Pectoral fin with gray rays and clear membranes. Pelvic fin black/dark gray with white leading edge.</paragraph>
<paragraph id="8B8B5C2BA2433D4903B9C9C49E69F9A5" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">Females light brown or orange blotch (OB). Non-OB females with brown flank; center of flank scales blue; belly and breast white/beige. Head light brown; blue/black opercular spot; light brown/white throat. Dorsal fin dark gray or brown with white/light-blue distal margin. Proximal ½ of caudal fin gray/ brown and distal ½ light brown to clear. Pectoral fin with gray rays and clear membranes. Pelvic fin with white leading edge, gray rays and clear membranes.</paragraph>
<paragraph id="8B8B5C2BA2433D4903B9CA109B4EF915" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">Orange blotch (OB) females with yellow-gray/orange flank with irregular black blotches; belly white with black blotches; breast yellow with black blotches. Head gray-yellow/orange with black blotches; throat white with black blotches; opercle with blue/green highlights. Dorsal, caudal, and anal fins gray-yellow with black blotches. Pectoral fins with yellow/black rays and clear membranes with black blotches.</paragraph>
</subSubSection>
<subSubSection id="C32E0FA0A2433D4903B9CA809A35F8F1" box="[201,1404,1780,1805]" pageId="18" pageNumber="119" type="distribution">
<paragraph id="8B8B5C2BA2433D4903B9CA809A35F8F1" blockId="18.[151,1437,151,1948]" box="[201,1404,1780,1805]" pageId="18" pageNumber="119">
<emphasis id="B9408039A2433D4903B9CA809E2BF8F0" bold="true" box="[201,354,1780,1805]" pageId="18" pageNumber="119">Distribution.</emphasis>
<taxonomicName id="4C3427A8A2433D490219CA809D1BF8F1" box="[361,594,1780,1804]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D490219CA809D1BF8F1" box="[361,594,1780,1804]" italics="true" pageId="18" pageNumber="119">Metriaclima tarakiki</emphasis>
</taxonomicName>
is known from Ngkuyo Island and Higga Reef, Lake
<collectingCountry id="F3231CBBA2433D4907DCCA809A4DF8F0" box="[1196,1284,1780,1805]" name="Malawi" pageId="18" pageNumber="119">Malaŵi</collectingCountry>
,
<collectingCountry id="F3231CBBA2433D49067FCA809A3EF8F1" box="[1295,1399,1780,1804]" name="Tanzania" pageId="18" pageNumber="119">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E0FA0A2433D4903B9CB6D9FA6F8A9" pageId="18" pageNumber="119" type="etymology">
<paragraph id="8B8B5C2BA2433D4903B9CB6D9FA6F8A9" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">
<emphasis id="B9408039A2433D4903B9CB6D9E06F8CC" bold="true" box="[201,335,1817,1841]" pageId="18" pageNumber="119">Etymology.</emphasis>
The name
<taxonomicName id="4C3427A8A2433D4902A2CB6C9D60F8CD" box="[466,553,1816,1840]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D4902A2CB6C9D60F8CD" box="[466,553,1816,1840]" italics="true" pageId="18" pageNumber="119">tarakiki</emphasis>
</taxonomicName>
, from Swahili, means slim or slender to note the elongate body shape of this species.
</paragraph>
</subSubSection>
<subSubSection id="C32E0FA0A2433D4903B9CB159C80F861" pageId="18" pageNumber="119" type="discussion">
<paragraph id="8B8B5C2BA2433D4903B9CB159C80F861" blockId="18.[151,1437,151,1948]" pageId="18" pageNumber="119">
<emphasis id="B9408039A2433D4903B9CB159E74F884" bold="true" box="[201,317,1889,1913]" pageId="18" pageNumber="119">Remarks.</emphasis>
<taxonomicName id="4C3427A8A2433D490235CB149D66F885" box="[325,559,1888,1912]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="tarakiki">
<emphasis id="B9408039A2433D490235CB149D66F885" box="[325,559,1888,1912]" italics="true" pageId="18" pageNumber="119">Metriaclima tarakiki</emphasis>
</taxonomicName>
exhibits polychromatism with OB females and males present in both known populations (see Konings 2001;
<figureCitation id="130F40AEA2433D490171CBF09D24F861" box="[513,621,1924,1948]" captionStart="FIGURE 14" captionStartId="29.[151,250,1501,1524]" captionTargetBox="[161,1425,462,1468]" captionTargetId="figure@29.[151,1436,445,1481]" captionTargetPageId="29" captionText="FIGURE 14. Plot of the first principal component of the meristic data (PC 1) and the second sheared principal component of the morphometric data (PC 2) of Metriaclima nigrodorsalis from Charlies Bay (n = 20), from Chiloelo (n = 8), from Nkolongwe (n = 20), from Nkhungu Point (n = 20), and from Thundu (n = 20) and Metriaclima phaeos from Cobwé (n = 6); all localities in Mozambique." httpUri="https://zenodo.org/record/221399/files/figure.png" pageId="18" pageNumber="119">Figure 14</figureCitation>
for OB female at Higga Reef).
</paragraph>
</subSubSection>
</treatment>
</document>