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<document id="0197C87BEE827B34C419F8B76524C33E" ID-CLB-Dataset="55739" ID-DOI="10.1080/00222933.2016.1271157" ID-GBIF-Dataset="f91a0e7b-c4c5-458a-858a-9de4cf82e13f" ID-ISSN="1464-5262" ID-Zenodo-Dep="5184295" ID-ZooBank="89612083-9CE6-48E8-8975-1CE5334E098B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628736318312" checkinUser="felipe" docAuthor="Sharaf, Mostafa R., Fisher, Brian L., Collingwood, Cedric A. &amp; Aldawood, Abdulrahman S." docDate="2017" docId="039DB6551630FFD0B499E6D26FDDC9CB" docLanguage="en" docName="JNatHist.51.5-6.317-378.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222933.2016.1271157" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Tetramorium lanuginosum Mayr 1870" docType="treatment" docVersion="9" lastPageNumber="357" masterDocId="FFA4CE2D1616FFF9B53EE3276B59CD7C" masterDocTitle="Ant fauna (Hymenoptera: Formicidae) of the Socotra Archipelago (Yemen): zoogeography, distribution and description of a new species" masterLastPageNumber="378" masterPageNumber="317" pageNumber="354" updateTime="1699076503534" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFF944A214AD32C23779A30827FBED3D">Ant fauna (Hymenoptera: Formicidae) of the Socotra Archipelago (Yemen): zoogeography, distribution and description of a new species</mods:title>
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<mods:namePart id="91935584575E8B4D4EEC3854C0F54DDB">Sharaf, Mostafa R.</mods:namePart>
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<subSubSection id="C32E54C81630FFDFB499E6D26825CB72" box="[423,892,1525,1550]" pageId="38" pageNumber="354" type="nomenclature">
<paragraph id="8B8B07431630FFDFB499E6D26825CB72" blockId="38.[423,892,1525,1585]" box="[423,892,1525,1550]" pageId="38" pageNumber="354">
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<taxonomicName id="4C347CC01630FFDFB499E6D26825CB72" ID-CoL="55PPC" authority="Mayr, 1870" authorityName="Mayr" authorityYear="1870" box="[423,892,1525,1550]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="354" phylum="Arthropoda" rank="species" species="lanuginosum">
<emphasis id="B940DB511630FFDFB499E6D269B6CB72" bold="true" box="[423,751,1525,1550]" italics="true" pageId="38" pageNumber="354">Tetramorium lanuginosum</emphasis>
<bibRefCitation id="EFA57AB21630FFDFB7C7E6D26825CB72" author="Mayr G" box="[761,892,1525,1550]" pageId="38" pageNumber="354" pagination="939 - 996" refId="ref19762" refString="Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien. 20: 939 - 996." type="journal article" year="1870">Mayr, 1870</bibRefCitation>
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<subSubSection id="C32E54C81630FFDFB70BE53F69B7CB4D" box="[565,750,1560,1585]" pageId="38" pageNumber="354" type="description">
<paragraph id="8B8B07431630FFDFB70BE53F69B7CB4D" blockId="38.[423,892,1525,1585]" box="[565,750,1560,1585]" pageId="38" pageNumber="354">
(
<figureCitation id="130F1BC61630FFDFB703E53F69F4CB4D" box="[573,685,1560,1585]" captionStart="Figure 21" captionStartId="40.[160,225,1709,1732]" captionTargetBox="[246,1070,142,1668]" captionTargetId="figure-1@40.[246,1070,142,1678]" captionTargetPageId="40" captionText="Figure 21. Tetramorium lanuginosum. (a) Body in profile. (b) Head in full-face view, CASENT0746619, AntWeb.org, (Zach Lieberman). (c) Distribution map, blue triangles from recent records, red circles from Collingwood et al. (2004)." figureDoi="http://doi.org/10.5281/zenodo.5184337" httpUri="https://zenodo.org/record/5184337/files/figure.png" pageId="38" pageNumber="354">Figure 21</figureCitation>
(ac))
</paragraph>
</subSubSection>
<subSubSection id="C32E54C81630FFD0B59EE56A6A4DCDDB" lastPageId="41" lastPageNumber="357" pageId="38" pageNumber="354" type="reference_group">
<paragraph id="8B8B07431630FFD0B59EE56A6A4DCDDB" blockId="38.[160,1031,1613,1637]" lastBlockId="41.[160,1156,141,410]" lastPageId="41" lastPageNumber="357" pageId="38" pageNumber="354">
<treatmentCitationGroup id="AB24206D1630FFDFB59EE56A6F5ECB19" box="[160,1031,1613,1637]" pageId="38" pageNumber="354">
<treatmentCitation id="0A9521521630FFDFB59EE56A69D8CB19" author="Mayr G" box="[160,641,1613,1637]" page="976" pageId="38" pageNumber="354" year="1870">
<taxonomicName id="4C347CC01630FFDFB59EE56A69D8CB19" ID-CoL="55PPC" authority="Mayr, 1870: 976" authorityName="Mayr" authorityPageNumber="976" authorityYear="1870" box="[160,641,1613,1637]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="354" phylum="Arthropoda" rank="species" species="lanuginosum">
<emphasis id="B940DB511630FFDFB59EE56A6A9DCB19" box="[160,452,1613,1637]" italics="true" pageId="38" pageNumber="354">Tetramorium lanuginosum</emphasis>
<bibRefCitation id="EFA57AB21630FFDFB4F3E56A69D8CB19" author="Mayr G" box="[461,641,1613,1637]" pageId="38" pageNumber="354" pagination="939 - 996" refId="ref19762" refString="Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien. 20: 939 - 996." type="journal article" year="1870">Mayr, 1870: 976</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(w.)
<collectingCountry id="F32347D31630FFDFB785E56A6871CB19" box="[699,808,1613,1637]" name="Indonesia" pageId="38" pageNumber="354">Indonesia</collectingCountry>
(
<collectingRegion id="49F0C9A11630FFDFB608E56A6834CB19" box="[822,877,1613,1637]" country="Indonesia" name="Jawa Timur" pageId="38" pageNumber="354">Java</collectingRegion>
). Indomalaya.
</treatmentCitationGroup>
<emphasis id="B940DB51163FFFD0B59EE3AA6A4DCDDB" bold="true" box="[160,276,141,167]" italics="true" pageId="41" pageNumber="357">Diagnosis</emphasis>
</paragraph>
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<caption id="DF4B57CB1631FFDEB59EE58A6F5ECB9D" ID-DOI="http://doi.org/10.5281/zenodo.5184335" ID-Zenodo-Dep="5184335" httpUri="https://zenodo.org/record/5184335/files/figure.png" pageId="39" pageNumber="355" startId="39.[160,225,1709,1732]" targetBox="[271,1043,142,1678]" targetPageId="39">
<paragraph id="8B8B07431631FFDEB59EE58A6F5ECB9D" blockId="39.[160,1157,1709,1761]" pageId="39" pageNumber="355">
<emphasis id="B940DB511631FFDEB59EE58A6A53CBB8" bold="true" box="[160,266,1709,1732]" pageId="39" pageNumber="355">Figure 20.</emphasis>
<taxonomicName id="4C347CC01631FFDEB42CE58A6AAECBB8" baseAuthorityName="Roger" baseAuthorityYear="1857" box="[274,503,1709,1732]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="355" phylum="Arthropoda" rank="species" species="caldarium">
<emphasis id="B940DB511631FFDEB42CE58A6AAECBB8" box="[274,503,1709,1732]" italics="true" pageId="39" pageNumber="355">Tetramorium caldarium</emphasis>
</taxonomicName>
. (a) Body in profile. (b) Head in full-face view, CASENT0264195, AntWeb.org, (Estella Ortega), KSA. (c) Distribution map from
<bibRefCitation id="EFA57AB21631FFDEB63BE5ED6F5ACB9D" author="Collingwood CA &amp; Pohl H &amp; Guesten R &amp; Wranik W &amp; Van Harten A" box="[773,1027,1738,1761]" pageId="39" pageNumber="355" pagination="473 - 495" refId="ref18239" refString="Collingwood CA, Pohl H, Guesten R, Wranik W, Van Harten A. 2004. The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago. Fauna of Arabia. 20: 473 - 495." type="journal article" year="2004">Collingwood et al. (2004)</bibRefCitation>
.
</paragraph>
</caption>
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<paragraph id="8B8B0743163EFFD1B59EE58A6A80CB83" blockId="40.[160,1157,1709,1791]" pageId="40" pageNumber="356">
<emphasis id="B940DB51163EFFD1B59EE58A6A5ECBB8" bold="true" box="[160,263,1709,1732]" pageId="40" pageNumber="356">Figure 21.</emphasis>
<taxonomicName id="4C347CC0163EFFD1B42EE58A6957CBB8" authorityName="Mayr" authorityYear="1870" box="[272,526,1709,1732]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="356" phylum="Arthropoda" rank="species" species="lanuginosum">
<emphasis id="B940DB51163EFFD1B42EE58A6957CBB8" box="[272,526,1709,1732]" italics="true" pageId="40" pageNumber="356">Tetramorium lanuginosum</emphasis>
</taxonomicName>
. (a) Body in profile. (b) Head in full-face view, CASENT0746619, AntWeb.org, (Zach Lieberman). (c) Distribution map, blue triangles from recent records, red circles from
<bibRefCitation id="EFA57AB2163EFFD1B5E9E5CF6A8CCB83" author="Collingwood CA &amp; Pohl H &amp; Guesten R &amp; Wranik W &amp; Van Harten A" box="[215,469,1768,1791]" pageId="40" pageNumber="356" pagination="473 - 495" refId="ref18239" refString="Collingwood CA, Pohl H, Guesten R, Wranik W, Van Harten A. 2004. The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago. Fauna of Arabia. 20: 473 - 495." type="journal article" year="2004">Collingwood et al. (2004)</bibRefCitation>
.
</paragraph>
</caption>
<subSubSection id="C32E54C8163FFFD0B59EE39569E4CCE6" pageId="41" pageNumber="357" type="description">
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<emphasis id="B940DB51163FFFD0B59EE3956BB4CDB6" box="[160,237,178,202]" italics="true" pageId="41" pageNumber="357">Worker</emphasis>
. Head longer than broad with reticulate-rugose sculpture; anterior clypeal margin with small median impression; frontal carinae strongly developed; antennal scrobes well-developed with distinct margins; eyes of moderate size with 810 ommatidia in longest row; mesosoma convex in profile; metanotal groove absent; propodeal spines long and sharp; petiolar node rounded in profile; gaster smooth and shiny. Body pilosity profuse and relatively long, bifid and simple hairs. Body colour variable from pale brown to dark brown, gaster darker than body.
</paragraph>
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<emphasis id="B940DB51163FFFD0B59EE2E26AD9CCA3" bold="true" box="[160,384,453,479]" italics="true" pageId="41" pageNumber="357">Material examined</emphasis>
</paragraph>
<paragraph id="8B8B0743163FFFD0B59EE2CD68BFCFAE" blockId="41.[160,1156,453,722]" pageId="41" pageNumber="357">
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,
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,
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,
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<collectingDate id="EFCED86B163FFFD0B710E2CD6997CF7E" box="[558,718,490,514]" pageId="41" pageNumber="357" value="2014-04-25">25 April 2014</collectingDate>
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, 26 m, (M.
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.
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leg.),
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(5 w,
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);
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,
<collectingCountry id="F32347D3163FFFD0B721E12B699ACF58" box="[543,707,524,548]" name="Socotra Island" pageId="41" pageNumber="357">Socotra Island</collectingCountry>
, W.
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,
<date id="FF8A2183163FFFD0B676E12B68BFCF58" box="[840,998,524,548]" pageId="41" pageNumber="357" value="2014-04-20">
<collectingDate id="EFCED86B163FFFD0B676E12B68BFCF58" box="[840,998,524,548]" pageId="41" pageNumber="357" value="2014-04-20">
20 April
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, 214 m, (M.
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.
<collectorName id="26C16295163FFFD0B59EE1086BBFCF3B" box="[160,230,559,583]" pageId="41" pageNumber="357">Sharaf</collectorName>
leg.),
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,
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(6 w,
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,
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23 April
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, 372 m, (M.
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.
<collectorName id="26C16295163FFFD0B7A8E1756985CF16" box="[662,732,594,618]" pageId="41" pageNumber="357">Sharaf</collectorName>
leg.),
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,
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(9 w,
<collectionCode id="ED259F86163FFFD0B59EE1536BBFCFF0" box="[160,230,628,652]" pageId="41" pageNumber="357">KSMA</collectionCode>
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<materialsCitation id="3B5C0D1E163FFFD0B5C1E1536903CFD3" ID-GBIF-Occurrence="3762058367" collectingDate="2014-04-21" collectionCode="R, KSMA" collectorName="Sharaf" country="Yemen" elevation="2014372" latitude="12.68401" location="Jo'" longLatPrecision="1" longitude="53.49052" pageId="41" pageNumber="357" specimenCount="1">
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,
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, W.
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o,
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<collectingDate id="EFCED86B163FFFD0B7B0E1536861CFF0" box="[654,824,628,652]" pageId="41" pageNumber="357" value="2014-04-21">
21 April
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, 196 m, (M.
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.
<collectorName id="26C16295163FFFD0B6C9E1536F64CFF0" box="[1015,1085,628,652]" pageId="41" pageNumber="357">Sharaf</collectorName>
leg.),
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,
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, (6 w,
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,
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, W.
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(daitab),
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, 93 m, (M.
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.
<collectorName id="26C16295163FFFD0B494E19D6AA9CFAE" box="[426,496,698,722]" pageId="41" pageNumber="357">Sharaf</collectorName>
leg.),
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,
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(13 w,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C32E54C8163FFFD0B59EE1DA69B2CE03" pageId="41" pageNumber="357" type="distribution">
<paragraph id="8B8B0743163FFFD0B59EE1DA6A2ECE6B" blockId="41.[160,1156,765,895]" box="[160,375,765,791]" pageId="41" pageNumber="357">
<emphasis id="B940DB51163FFFD0B59EE1DA6A2ECE6B" bold="true" box="[160,375,765,791]" italics="true" pageId="41" pageNumber="357">Geographic range</emphasis>
</paragraph>
<paragraph id="8B8B0743163FFFD0B59EE00569B2CE03" blockId="41.[160,1156,765,895]" pageId="41" pageNumber="357">
A cosmopolitan species widely distributed in tropical and subtropical regions of Asia,
<collectingCountry id="F32347D3163FFFD0B59EE0636A5DCE20" box="[160,260,836,860]" name="Australia" pageId="41" pageNumber="357">Australia</collectingCountry>
, Oceania (
<bibRefCitation id="EFA57AB2163FFFD0B445E0636946CE20" author="Wetterer JK" box="[379,543,836,860]" pageId="41" pageNumber="357" pagination="81 - 88" refId="ref21243" refString="Wetterer JK. 2010. Worldwide spread of the wooly ant, Tetramorium lanuginosum (Hymenoptera: Formicidae). Myrmecological News. 13: 81 - 88." type="journal article" year="2010">Wetterer 2010</bibRefCitation>
;
<bibRefCitation id="EFA57AB2163FFFD0B710E0636835CE20" author="Hita Garcia F &amp; Fisher BL" box="[558,876,836,860]" pageId="41" pageNumber="357" pagination="1 - 72" refId="ref19323" refString="Hita Garcia F, Fisher BL. 2011. The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - introduction, definition of species groups, and revision of the T. bicarinatum, T. obesum, T. sericeiventre and T. tosii species groups. Zootaxa. 3039: 1 - 72." type="journal article" year="2011">Hita Garcia and Fisher 2011</bibRefCitation>
),
<collectingCountry id="F32347D3163FFFD0B6BDE06368F6CE20" box="[899,943,836,860]" name="Saudi Arabia" pageId="41" pageNumber="357">KSA</collectingCountry>
(
<bibRefCitation id="EFA57AB2163FFFD0B681E0636A7ECE03" author="Collingwood CA &amp; Agosti D" pageId="41" pageNumber="357" pagination="300 - 385" refId="ref18173" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna of Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
) and
<collectingCountry id="F32347D3163FFFD0B455E0406A99CE03" box="[363,448,871,895]" name="Yemen" pageId="41" pageNumber="357">Socotra</collectingCountry>
(
<bibRefCitation id="EFA57AB2163FFFD0B4F0E0406986CE03" author="Collingwood CA &amp; Pohl H &amp; Guesten R &amp; Wranik W &amp; Van Harten A" box="[462,735,871,895]" pageId="41" pageNumber="357" pagination="473 - 495" refId="ref18239" refString="Collingwood CA, Pohl H, Guesten R, Wranik W, Van Harten A. 2004. The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago. Fauna of Arabia. 20: 473 - 495." type="journal article" year="2004">Collingwood et al. 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E54C8163FFFD0B59EE08D6FDDC9CB" pageId="41" pageNumber="357" type="biology_ecology">
<paragraph id="8B8B0743163FFFD0B59EE08D694CCEB8" blockId="41.[160,1156,938,1207]" box="[160,533,938,964]" pageId="41" pageNumber="357">
<emphasis id="B940DB51163FFFD0B59EE08D694CCEB8" bold="true" box="[160,533,938,964]" italics="true" pageId="41" pageNumber="357">Ecological and biological notes</emphasis>
</paragraph>
<paragraph id="8B8B0743163FFFD0B59EE0E86FDDC9CB" blockId="41.[160,1156,938,1207]" pageId="41" pageNumber="357">
<taxonomicName id="4C347CC0163FFFD0B59EE0E86A9ECE9B" authorityName="Mayr" authorityYear="1870" box="[160,455,975,999]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="357" phylum="Arthropoda" rank="species" species="lanuginosum">
<emphasis id="B940DB51163FFFD0B59EE0E86A9ECE9B" box="[160,455,975,999]" italics="true" pageId="41" pageNumber="357">Tetramorium lanuginosum</emphasis>
</taxonomicName>
is one of the most successful ants occurring on the main island and has varied nesting habitats. A nest was found under the trunk of a fallen, dead palm tree where the soil was moist and rich in decaying organic material. Individuals were collected from dry soil next to a date palm tree. Several specimens were observed foraging on the ground where the soil was moist. Other workers were found foraging on the ground near a small flowing stream where the soil was sandy and moist. This species seems likely to have been introduced to
<collectingCountry id="F32347D3163FFFD0B679E7B868C5C9CB" box="[839,924,1183,1207]" name="Yemen" pageId="41" pageNumber="357">Socotra</collectingCountry>
through commerce.
</paragraph>
</subSubSection>
</treatment>
</document>