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<document id="877875FD7A1CEBC27047CCF2C69A2B12" ID-DOI="10.11646/zootaxa.4991.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042522" ID-ZooBank="842EA450-A680-4D5D-9723-C1D7CCE13E5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624987574070" checkinUser="plazi" docAuthor="Vasanth, M., Selvakumar, C., Subramanian, K. A., Sivaramakrishnan, K. G. &amp; Sinha, Bikramjit" docDate="2021" docId="039D87CCFFA4FF84FF56FBA8FD166EFF" docLanguage="en" docName="zootaxa.4991.3.4.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epeorus (Epeorus) aculeatus Braasch 1990" docType="treatment" docVersion="3" lastPageNumber="503" masterDocId="FFA4FFB4FFA5FF80FFC1FFF6FFD26F53" masterDocTitle="Contribution to the study of Epeorus Eaton, 1881 (Ephemeroptera: Heptageniidae) from India" masterLastPageNumber="522" masterPageNumber="499" pageNumber="500" updateTime="1698978689892" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="52E019C347CE0A0A930B68EC1974A92D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F59D5A2E6932C4906FCFDC1F236D9423">
<mods:title id="1ACCEC7EA9B26576DA5DDE294F6C5518">Contribution to the study of Epeorus Eaton, 1881 (Ephemeroptera: Heptageniidae) from India</mods:title>
</mods:titleInfo>
<mods:name id="39AEAA70B2EA94F1CFFCAEE91E0D253D" type="personal">
<mods:role id="9FC1314853F21FA100F082274DABC3AA">
<mods:roleTerm id="E1476CA8B821716C4FD53D38AE2F0717">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="29F76435652388B6534A0BD1DC133A71">Vasanth, M.</mods:namePart>
<mods:nameIdentifier id="96E6C149397C0E05498D1360535C5F7F" type="ORCID">0000-0003-4446-4889</mods:nameIdentifier>
<mods:affiliation id="19F9283BEDD2EC5E78FF5693E87EE6FE">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India. &amp; University of Madras, Chepauk, Chennai- 600005. &amp; vasan 071994 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4446 - 4889</mods:affiliation>
<mods:nameIdentifier id="665BAEFFCC9B3EF0ABA7ACE8BA6429F5" type="email">vasan071994@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="0CD171126D00A8E84F338C721E1DF427" type="personal">
<mods:role id="3DEC7F0A3F1C3472C60F462DD9DEBBD6">
<mods:roleTerm id="9F642E3AA0EAA63DA18A3F83425EB753">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="412C1B350476CB1CD028A5EE72BDDD01">Selvakumar, C.</mods:namePart>
<mods:nameIdentifier id="0604082E6936A58D469113C7E470FEC3" type="ORCID">0000-0002-4296-5682</mods:nameIdentifier>
<mods:affiliation id="C28C1E770BA289394CCB9FBC21FA9E07">Department of Zoology, The Madura College (Autonomous), Madurai- 625011, Tamil Nadu. &amp; selvaaa 06 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4296 - 5682</mods:affiliation>
<mods:nameIdentifier id="ED3800ACB75F22C6649ADA702C53C9E1" type="email">selvaaa06@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F8958933999F2287B142504D705354DE" type="personal">
<mods:role id="58B80730CA88C0D4E40A3F82D683F495">
<mods:roleTerm id="C3E24A1D05C59B37F42C54B28D1D6913">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7597D2883396799A541D6ADCA9329E9D">Subramanian, K. A.</mods:namePart>
<mods:affiliation id="1AC30CC674F2632BE9597D72A128D885">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="C20E3D129ED258046B09D532CBDB5AF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6FC7CA47616303FB5AC5D55F39E6032">Sivaramakrishnan, K. G.</mods:namePart>
<mods:nameIdentifier id="AF17B0D1F94BBCF6E3C21C0861FD5E2A" type="ORCID">0000-0002-6332-0349</mods:nameIdentifier>
<mods:affiliation id="7250D71EDD5B6C979A9A9D43E2411986">Flat No 3, Door No 7, Gokulam Apartment, Gokulam Colony, West Mambalam, Chennai- 600 033, India. &amp; kgskrishnan @ gmail. com; https: // orcid. org / 0000 - 0002 - 6332 - 0349</mods:affiliation>
<mods:nameIdentifier id="E329B9A52CCB8592AB8EB2AA3A71C562" type="email">kgskrishnan@gmail.com</mods:nameIdentifier>
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<mods:name id="246DA26FB05977026FC46C9846EE9559" type="personal">
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</mods:role>
<mods:namePart id="6CF8246AB44A19A01A3CFE29D6930251">Sinha, Bikramjit</mods:namePart>
<mods:nameIdentifier id="E5F493B31C84049103DC1BB020A6B398" type="ORCID">0000-0002-9904-4964</mods:nameIdentifier>
<mods:affiliation id="81E258FA38DCD263C94E0A68224E7347">Zoological Survey of India, Arunachal Pradesh Regional Centre, Senki Valley, Itanagar- 791113, Arunachal Pradesh, India. &amp; sinhabj @ gmail. com; https: // orcid. org / 0000 - 0002 - 9904 - 4964</mods:affiliation>
<mods:nameIdentifier id="45FC16192DCA2C0A60DF12672B8C93A7" type="email">sinhabj@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="8099C825FCC2E1EB9629DA260A9A91A3">text</mods:typeOfResource>
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<mods:title id="53152C79A89AA4EEF2F715E21BE82813">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="62C472EEE0E97A290E25EF873BCD0D76">
<mods:date id="4CDEDF874753C1C8F9F4D7FA610DE14E">2021</mods:date>
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<mods:number id="0AF63BF12BBCF76822F8BA9C124F9610">2021-06-25</mods:number>
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<mods:detail id="1F5313FAC653949292DAE58AE1A229D3" type="volume">
<mods:number id="6A813FE019AF9D5E11A2874D39C4794F">4991</mods:number>
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<mods:number id="8FE7BBA943F9485FCEC0A4775AC5C7AC">3</mods:number>
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<mods:start id="7F4E484BDC17FA3F468177F94AEDF463">499</mods:start>
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<mods:classification id="15D0229E53D0812AFEA0D49D862935C1">journal article</mods:classification>
<mods:identifier id="6AF50CB2023A93DA26708655D2345B0C" type="DOI">10.11646/zootaxa.4991.3.4</mods:identifier>
<mods:identifier id="F8655C116D965F28E980F377EADF5A22" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="ACB4EAA35000538AAAF68964C4C22E3F" type="Zenodo-Dep">5042522</mods:identifier>
<mods:identifier id="4D7D122578CCD2DF7B40BA1F0B290E66" type="ZooBank">842EA450-A680-4D5D-9723-C1D7CCE13E5D</mods:identifier>
</mods:mods>
<treatment id="039D87CCFFA4FF84FF56FBA8FD166EFF" ID-DOI="http://doi.org/10.5281/zenodo.5042542" ID-Zenodo-Dep="5042542" LSID="urn:lsid:plazi:treatment:039D87CCFFA4FF84FF56FBA8FD166EFF" httpUri="http://treatment.plazi.org/id/039D87CCFFA4FF84FF56FBA8FD166EFF" lastPageId="4" lastPageNumber="503" pageId="1" pageNumber="500">
<subSubSection id="C32E6551FFA4FF81FF56FBA8FD7D6B24" box="[151,687,1117,1144]" pageId="1" pageNumber="500" type="nomenclature">
<paragraph id="8B8B36DAFFA4FF81FF56FBA8FD7D6B24" blockId="1.[151,687,1117,1179]" box="[151,687,1117,1144]" pageId="1" pageNumber="500">
<heading id="D0C381B6FFA4FF81FF56FBA8FD7D6B24" bold="true" box="[151,687,1117,1144]" fontSize="11" level="1" pageId="1" pageNumber="500" reason="1">
<taxonomicName id="4C344D59FFA4FF81FF56FBA8FD7D6B24" ID-CoL="6FNW4" authority="Braasch, 1990" authorityName="Braasch" authorityYear="1990" box="[151,687,1117,1144]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="500" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Epeorus">
<emphasis id="B940EAC8FFA4FF81FF56FBA8FD7D6B24" bold="true" box="[151,687,1117,1144]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF56FBA8FF2E6B2B" bold="true" box="[151,252,1118,1144]" italics="true" pageId="1" pageNumber="500">Epeorus</emphasis>
(
<emphasis id="B940EAC8FFA4FF81FECCFBA8FEA06B2B" bold="true" box="[269,370,1118,1144]" italics="true" pageId="1" pageNumber="500">Epeorus</emphasis>
)
<emphasis id="B940EAC8FFA4FF81FE45FBABFE2A6B2B" bold="true" box="[388,504,1117,1144]" italics="true" pageId="1" pageNumber="500">aculeatus</emphasis>
<bibRefCitation id="EFA54B2BFFA4FF81FE3EFBABFD7D6B24" author="Braasch, D." box="[511,687,1117,1143]" pageId="1" pageNumber="500" pagination="7 - 14" refId="ref10040" refString="Braasch, D. (1990) Neue Eintagsfliegen aus Thailand, nebsteinigen Bemerkungen zuderen generischem Status (Insecta, Ephemeroptera, Heptageniidae). Reichenbachia, 28 (2), 7 - 14." type="journal article" year="1990">Braasch, 1990</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E6551FFA4FF81FF56FB77FECB6BC8" box="[151,281,1153,1179]" pageId="1" pageNumber="500" type="description">
<paragraph id="8B8B36DAFFA4FF81FF56FB77FECB6BC8" blockId="1.[151,687,1117,1179]" box="[151,281,1153,1179]" pageId="1" pageNumber="500">
(
<figureCitation id="130F2A5FFFA4FF81FF5EFB77FEC26BC8" box="[159,272,1153,1179]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–17" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1944,1968]" captionTargetBox-0="[191,1400,181,1917]" captionTargetBox-1="[162,1426,181,1919]" captionTargetId-0="figure-19@2.[188,1400,179,1918]" captionTargetId-1="figure-17@3.[161,1426,181,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." captionText-1="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi-0="http://doi.org/10.5281/zenodo.5042524" figureDoi-1="http://doi.org/10.5281/zenodo.5042528" httpUri-0="https://zenodo.org/record/5042524/files/figure.png" httpUri-1="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Figs 1–17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E6551FFA4FF81FF56FB32FDB76A19" pageId="1" pageNumber="500" type="materials_examined">
<paragraph id="8B8B36DAFFA4FF81FF56FB32FDB76A19" blockId="1.[151,1437,1220,2039]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF56FB32FE576B8D" bold="true" box="[151,389,1220,1246]" pageId="1" pageNumber="500">Material examined:</emphasis>
<materialsCitation id="3B5C3C87FFA4FF81FE4FFB32FB676A51" collectingDate="2017-06-16" collectionCode="SRC-I" collectorName="K. A. Subramanian &amp; M. Vasanth" country="India" elevation="1662" latitude="27.62776" location="Duskilo" longLatPrecision="4" longitude="93.8437" municipality="Lower Subansiri district" pageId="1" pageNumber="500" specimenCode="E 289" specimenCount="1" specimenCount-larva="1" stateProvince="Arunachal Pradesh">
<specimenCount id="9D32FD53FFA4FF81FE4FFB32FE326B8D" box="[398,480,1220,1246]" pageId="1" pageNumber="500" type="larva">1 larva</specimenCount>
,
<collectingCountry id="F323764AFFA4FF81FE2CFB32FDED6B8D" box="[493,575,1220,1246]" name="India" pageId="1" pageNumber="500">INDIA</collectingCountry>
,
<collectingRegion id="49F0F838FFA4FF81FD8BFB32FCF66B8D" box="[586,804,1220,1246]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="500">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="6BEFACA0FFA4FF81FCF1FB33FB946B8D" box="[816,1094,1220,1247]" pageId="1" pageNumber="500">Lower Subansiri district</collectingMunicipality>
,
<location id="8EEB6001FFA4FF81FB92FB32FB7E6B8D" LSID="urn:lsid:plazi:treatment:039D87CCFFA4FF84FF56FBA8FD166EFF:8EEB6001FFA4FF81FB92FB32FB7E6B8D" box="[1107,1196,1220,1246]" country="India" latitude="27.62776" longLatPrecision="4" longitude="93.8437" municipality="Lower Subansiri district" name="Duskilo" pageId="1" pageNumber="500" stateProvince="Arunachal Pradesh">Duskilo</location>
stream,
<geoCoordinate id="EE00501DFFA4FF81FACEFB32FA4A6B8C" box="[1295,1432,1220,1247]" degrees="27.62776" direction="north" orientation="latitude" pageId="1" pageNumber="500" precision="1" value="27.62776">27.62776 N</geoCoordinate>
,
<geoCoordinate id="EE00501DFFA4FF81FF56FB1EFEDF6A50" box="[151,269,1256,1283]" degrees="93.8437" direction="east" orientation="longitude" pageId="1" pageNumber="500" precision="5" value="93.8437">93.8437 E</geoCoordinate>
,
<quantity id="4CCC9B3FFFA4FF81FED8FB1EFEA26A50" box="[281,368,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.6619999999999997" pageId="1" pageNumber="500" unit="m" value="1662.0">
<elevation id="0019D1E9FFA4FF81FED8FB1EFEA26A50" box="[281,368,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.6619999999999997" pageId="1" pageNumber="500" unit="m" value="1662.0">1662 m</elevation>
</quantity>
,
<date id="FF8A101AFFA4FF81FEBCFB1EFE256A51" box="[381,503,1256,1282]" pageId="1" pageNumber="500" value="2017-06-16">
<collectingDate id="EFCEE9F2FFA4FF81FEBCFB1EFE256A51" box="[381,503,1256,1282]" pageId="1" pageNumber="500" value="2017-06-16">16.vi.2017</collectingDate>
</date>
, colls.
<collectorName id="26C1530CFFA4FF81FD87FB1FFCCC6A51" box="[582,798,1256,1283]" pageId="1" pageNumber="500">K. A. Subramanian</collectorName>
and
<collectorName id="26C1530CFFA4FF81FC94FB1FFC076A51" box="[853,981,1256,1283]" pageId="1" pageNumber="500">M. Vasanth</collectorName>
[ZSI/
<collectionCode id="ED25AE1FFFA4FF81FBD4FB1EFB8C6A51" box="[1045,1118,1256,1282]" pageId="1" pageNumber="500">SRC-I</collectionCode>
/
<specimenCode id="DB929EA1FFA4FF81FBA7FB1EFB7C6A51" box="[1126,1198,1255,1282]" collectionCode="SRC-I" pageId="1" pageNumber="500">E 289</specimenCode>
]
</materialsCitation>
;
<materialsCitation id="3B5C3C87FFA4FF81FB00FB1EFDB36A19" collectingDate="2018-03-13" collectionCode="SRC-I" collectorName="Bikramjit Sinha" country="India" county="West Kameng district" elevation="1877" latitude="27.32675" location="Showda village" longLatPrecision="1" longitude="92.2318" municipality="Dirang" pageId="1" pageNumber="500" specimenCode="E 298" specimenCount="6" specimenCount-larva="6" stateProvince="Arunachal Pradesh">
<specimenCount id="9D32FD53FFA4FF81FB00FB1EFACF6A51" box="[1217,1309,1256,1282]" pageId="1" pageNumber="500" type="larva">6 larvae</specimenCount>
,
<collectingRegion id="49F0F838FFA4FF81FAE9FB1EFF216A75" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="500">Arunachal Pradesh</collectingRegion>
,
<collectingCounty id="62EA4E56FFA4FF81FF3DFAFBFE3D6A75" box="[252,495,1292,1319]" pageId="1" pageNumber="500">West Kameng district</collectingCounty>
,
<collectingMunicipality id="6BEFACA0FFA4FF81FE39FAFAFD9B6A75" box="[504,585,1292,1318]" pageId="1" pageNumber="500">Dirang</collectingMunicipality>
,
<location id="8EEB6001FFA4FF81FD92FAFAFCD06A75" LSID="urn:lsid:plazi:treatment:039D87CCFFA4FF84FF56FBA8FD166EFF:8EEB6001FFA4FF81FD92FAFAFCD06A75" box="[595,770,1292,1318]" country="India" county="West Kameng district" latitude="27.32675" longLatPrecision="1" longitude="92.2318" municipality="Dirang" name="Showda village" pageId="1" pageNumber="500" stateProvince="Arunachal Pradesh">Showda village</location>
,
<location id="8EEB6001FFA4FF81FCCAFAFAFC446A75" LSID="urn:lsid:plazi:treatment:039D87CCFFA4FF84FF56FBA8FD166EFF:8EEB6001FFA4FF81FCCAFAFAFC446A75" box="[779,918,1292,1318]" country="India" county="West Kameng district" latitude="27.32675" longLatPrecision="1" longitude="92.2318" municipality="Dirang" name="Dirang river" pageId="1" pageNumber="500" stateProvince="Arunachal Pradesh">Dirang river</location>
,
<geoCoordinate id="EE00501DFFA4FF81FC5EFAFAFBF76A74" box="[927,1061,1292,1319]" degrees="27.32675" direction="north" orientation="latitude" pageId="1" pageNumber="500" precision="1" value="27.32675">27.32675 N</geoCoordinate>
,
<geoCoordinate id="EE00501DFFA4FF81FBEEFAFAFB636A74" box="[1071,1201,1292,1319]" degrees="92.23180" direction="east" orientation="longitude" pageId="1" pageNumber="500" precision="1" value="92.2318">92.23180 E</geoCoordinate>
,
<quantity id="4CCC9B3FFFA4FF81FB7DFAFAFAC36A74" box="[1212,1297,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="1.877" pageId="1" pageNumber="500" unit="m" value="1877.0">
<elevation id="0019D1E9FFA4FF81FB7DFAFAFAC36A74" box="[1212,1297,1292,1319]" metricMagnitude="3" metricUnit="m" metricValue="1.877" pageId="1" pageNumber="500" unit="m" value="1877.0">1877 m</elevation>
</quantity>
,
<date id="FF8A101AFFA4FF81FADDFAFAFA456A75" box="[1308,1431,1292,1318]" pageId="1" pageNumber="500" value="2018-03-13">
<collectingDate id="EFCEE9F2FFA4FF81FADDFAFAFA456A75" box="[1308,1431,1292,1318]" pageId="1" pageNumber="500" value="2018-03-13">13.iii.2018</collectingDate>
</date>
, coll.
<collectorName id="26C1530CFFA4FF81FF0EFAC6FE506A19" box="[207,386,1328,1354]" pageId="1" pageNumber="500">Bikramjit Sinha</collectorName>
[ZSI/
<collectionCode id="ED25AE1FFFA4FF81FE00FAC6FDD86A19" box="[449,522,1328,1354]" pageId="1" pageNumber="500">SRC-I</collectionCode>
/
<specimenCode id="DB929EA1FFA4FF81FDD2FAC6FD896A19" box="[531,603,1327,1354]" collectionCode="SRC-I" pageId="1" pageNumber="500">E 298</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E6551FFA4FF81FF06FAA2FD6468A5" pageId="1" pageNumber="500" type="description">
<paragraph id="8B8B36DAFFA4FF81FF06FAA2FA876A3D" blockId="1.[151,1437,1220,2039]" box="[199,1365,1364,1391]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF06FAA2FE896A3D" bold="true" box="[199,347,1364,1390]" pageId="1" pageNumber="500">Description:</emphasis>
Larva: Body length
<quantity id="4CCC9B3FFFA4FF81FD82FAA2FD4E6A3C" box="[579,668,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="500" unit="mm" value="9.5">9.5 mm</quantity>
; length of cerci
<quantity id="4CCC9B3FFFA4FF81FC91FAA2FC646A3C" box="[848,950,1364,1391]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="500" unit="mm" value="11.5">11.5 mm</quantity>
. Body generally dark brown (
<figureCitation id="130F2A5FFFA4FF81FAC2FAA2FA966A3D" box="[1283,1348,1364,1390]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B8B36DAFFA4FF81FF06FA8EFC8869E1" blockId="1.[151,1437,1220,2039]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF06FA8EFEC26AC1" bold="true" box="[199,272,1400,1426]" pageId="1" pageNumber="500">Head:</emphasis>
Length
<quantity id="4CCC9B3FFFA4FF81FEAEFA8EFE1A6AC0" box="[367,456,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="500" unit="mm" value="1.7">1.7 mm</quantity>
, width
<quantity id="4CCC9B3FFFA4FF81FDDDFA8EFDA76AC0" box="[540,629,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="500" unit="mm" value="2.4">2.4 mm</quantity>
; brown, (
<figureCitation id="130F2A5FFFA4FF81FD21FA8EFCF06AC1" box="[736,802,1400,1426]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 1</figureCitation>
); subquadrangular; anterior and lateral margins convex, with rows of dense hair-like setae directed anteriorly; posterior margin slightly concave. Compound eyes black. Antennal scape and pedicel brown, flagellum pale. Labrum dark brown, deeply cleft with median notch on anterior margin; long hair-like setae laterally (
<figureCitation id="130F2A5FFFA4FF81FD81FA12FD566AAC" box="[576,644,1508,1535]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 5</figureCitation>
). Hypopharynx: Lingua with anterolateral lobes; superlinguae slightly expanded, laterally with rows of dense hair-like setae (
<figureCitation id="130F2A5FFFA4FF81FCC6F9FEFC9F6970" box="[775,845,1544,1571]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 6</figureCitation>
). Mandible without scattered setae along the molar; incisors serrated, outer incisor longer than inner incisor (
<figureCitation id="130F2A5FFFA4FF81FCE1F9DAFC5A6915" box="[800,904,1580,1606]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Figs 7–8</figureCitation>
). Maxilla with row of hair-like setae medially, apical segment of palp with dense row of hair-like setae, basal segment of palp broad (
<figureCitation id="130F2A5FFFA4FF81FB9EF9A6FB716939" box="[1119,1187,1616,1642]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Fig. 9</figureCitation>
). Labium broad, glossae with U-shaped separation, subtriangular; paraglossae slightly expanded laterally; apical segment of each palp pointed, dorsal surface with dense row of comb setae (
<figureCitation id="130F2A5FFFA4FF81FD38F96EFC9F69E1" box="[761,845,1688,1714]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8B8B36DAFFA4FF81FF06F94AFAFB6811" blockId="1.[151,1437,1220,2039]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF06F94AFEFB6985" bold="true" box="[199,297,1724,1750]" pageId="1" pageNumber="500">Thorax:</emphasis>
Pronotum brown without clear marks; mesonotum somewhat darker brown with paramedian pair of darker spots anteriorly (
<figureCitation id="130F2A5FFFA4FF81FE68F916FE3F69A9" box="[425,493,1760,1786]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 1</figureCitation>
). Legs: Femur brown; hypodermal spot on dorsal surface, outer margin with long setae; distal end with blunt spine (
<figureCitation id="130F2A5FFFA4FF81FDDDF8F2FD4D684D" box="[540,671,1796,1822]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Figs 11–13</figureCitation>
); short stout blunt setae scattered on dorsal surface (
<figureCitation id="130F2A5FFFA4FF81FB34F8F2FA86684D" box="[1269,1364,1796,1822]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Fig. 11a</figureCitation>
), tibia brown with long hair-like setae on outer margin; tarsi dark brown. Claw small with 3 denticles (
<figureCitation id="130F2A5FFFA4FF81FB08F8DEFACE6811" box="[1225,1308,1832,1858]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8B8B36DAFFA4FF81FF06F8BAFD6468A5" blockId="1.[151,1437,1220,2039]" pageId="1" pageNumber="500">
<emphasis id="B940EAC8FFA4FF81FF06F8BAFE976835" bold="true" box="[199,325,1868,1894]" pageId="1" pageNumber="500">Abdomen:</emphasis>
Abdominal terga dark brown, terga II–IX with pair of long, acute, submedian spines (
<figureCitation id="130F2A5FFFA4FF81FAD0F8BAFA876835" box="[1297,1365,1868,1894]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 2</figureCitation>
); posterior margin of tergum with hair-like setae medially and with numerous minute spines on segments II–IX (
<figureCitation id="130F2A5FFFA4FF81FA88F886FA5C68D9" box="[1353,1422,1904,1930]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 4</figureCitation>
); segments II–VII with long posterolateral extensions (
<figureCitation id="130F2A5FFFA4FF81FD31F862FCE668FD" box="[752,820,1940,1966]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="1" pageNumber="500">Fig. 3</figureCitation>
); sterna pale yellowish without markings. Lamellae of gill I somewhat extended beneath of abdomen (
<figureCitation id="130F2A5FFFA4FF81FD74F84EFCD86880" box="[693,778,1976,2003]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Fig. 15</figureCitation>
). Gills II–VII with costal ribs arched and anal rib straight (
<figureCitation id="130F2A5FFFA4FF81FF5EF82AFEF368A4" box="[159,289,2012,2039]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="1" pageNumber="500">Figs 16–17</figureCitation>
). Cerci with hair-like setae dorsally.
</paragraph>
</subSubSection>
<caption id="DF4B6652FFA7FF82FF56F861FB176887" ID-DOI="http://doi.org/10.5281/zenodo.5042524" ID-Zenodo-Dep="5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="2" pageNumber="501" startId="2.[151,264,1943,1967]" targetBox="[191,1400,181,1917]" targetPageId="2">
<paragraph id="8B8B36DAFFA7FF82FF56F861FB176887" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="501">
<emphasis id="B940EAC8FFA7FF82FF56F861FEE868FC" bold="true" box="[151,314,1943,1967]" pageId="2" pageNumber="501">FIGURES 1–8.</emphasis>
Larva of
<taxonomicName id="4C344D59FFA7FF82FE5DF86EFD1168E3" authorityName="Braasch" authorityYear="1990" box="[412,707,1944,1968]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="501" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Epeorus">
<emphasis id="B940EAC8FFA7FF82FE5DF86EFE2268FC" box="[412,496,1944,1967]" italics="true" pageId="2" pageNumber="501">Epeorus</emphasis>
(
<emphasis id="B940EAC8FFA7FF82FE3EF86EFD8168FC" box="[511,595,1944,1967]" italics="true" pageId="2" pageNumber="501">Epeorus</emphasis>
)
<emphasis id="B940EAC8FFA7FF82FDA3F86EFD1168E3" box="[610,707,1944,1968]" italics="true" pageId="2" pageNumber="501">aculeatus</emphasis>
</taxonomicName>
<emphasis id="B940EAC8FFA7FF82FD08F861FD0468FC" bold="true" box="[713,726,1943,1967]" pageId="2" pageNumber="501">1</emphasis>
. Dorsal view;
<emphasis id="B940EAC8FFA7FF82FCABF861FCA568FC" bold="true" box="[874,887,1943,1967]" pageId="2" pageNumber="501">2</emphasis>
. Abdomen dorsal view;
<emphasis id="B940EAC8FFA7FF82FBAFF861FBA968FC" bold="true" box="[1134,1147,1943,1967]" pageId="2" pageNumber="501">3</emphasis>
. Abdominal ventral view;
<emphasis id="B940EAC8FFA7FF82FA48F861FA4468FC" bold="true" box="[1417,1430,1943,1967]" pageId="2" pageNumber="501">4</emphasis>
. Posterior margin of abdominal terga V;
<emphasis id="B940EAC8FFA7FF82FDE8F84AFDE46887" bold="true" box="[553,566,1980,2004]" pageId="2" pageNumber="501">5</emphasis>
. Labrum;
<emphasis id="B940EAC8FFA7FF82FD61F84DFD7F6880" bold="true" box="[672,685,1979,2003]" pageId="2" pageNumber="501">6</emphasis>
. Hypopharynx;
<emphasis id="B940EAC8FFA7FF82FC93F84AFC8D6887" bold="true" box="[850,863,1980,2004]" pageId="2" pageNumber="501">7</emphasis>
. Left mandible;
<emphasis id="B940EAC8FFA7FF82FBC9F84DFBC76880" bold="true" box="[1032,1045,1979,2003]" pageId="2" pageNumber="501">8</emphasis>
. Right mandible.
</paragraph>
</caption>
<caption id="DF4B6652FFA6FF83FF56F86EFCEF6887" ID-DOI="http://doi.org/10.5281/zenodo.5042528" ID-Zenodo-Dep="5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="3" pageNumber="502" startId="3.[151,264,1944,1968]" targetBox="[162,1426,181,1919]" targetPageId="3">
<paragraph id="8B8B36DAFFA6FF83FF56F86EFCEF6887" blockId="3.[151,1436,1944,2004]" pageId="3" pageNumber="502">
<emphasis id="B940EAC8FFA6FF83FF56F86EFE9568E3" bold="true" box="[151,327,1944,1968]" pageId="3" pageNumber="502">FIGURES 9–17.</emphasis>
Larva of
<taxonomicName id="4C344D59FFA6FF83FE69F86FFD1D68E3" authorityName="Braasch" authorityYear="1990" box="[424,719,1944,1969]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="502" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Epeorus">
<emphasis id="B940EAC8FFA6FF83FE69F86FFE2E68E3" box="[424,508,1945,1968]" italics="true" pageId="3" pageNumber="502">Epeorus</emphasis>
(
<emphasis id="B940EAC8FFA6FF83FDCAF86FFD8D68E3" box="[523,607,1945,1968]" italics="true" pageId="3" pageNumber="502">Epeorus</emphasis>
)
<emphasis id="B940EAC8FFA6FF83FDAFF86EFD1D68E3" box="[622,719,1944,1968]" italics="true" pageId="3" pageNumber="502">aculeatus</emphasis>
</taxonomicName>
<emphasis id="B940EAC8FFA6FF83FD14F86EFD3068E3" bold="true" box="[725,738,1944,1968]" pageId="3" pageNumber="502">9</emphasis>
. Maxilla;
<emphasis id="B940EAC8FFA6FF83FC8BF86EFCB168E3" bold="true" box="[842,867,1944,1968]" pageId="3" pageNumber="502">10</emphasis>
. Labium;
<emphasis id="B940EAC8FFA6FF83FC0AF86EFC3168E3" bold="true" box="[971,995,1944,1968]" pageId="3" pageNumber="502">11</emphasis>
. Foreleg;
<emphasis id="B940EAC8FFA6FF83FB88F86EFBA168E3" bold="true" box="[1097,1139,1944,1968]" pageId="3" pageNumber="502">11A</emphasis>
. Dorsal surface of femur;
<emphasis id="B940EAC8FFA6FF83FABCF86EFA4468E3" bold="true" box="[1405,1430,1944,1968]" pageId="3" pageNumber="502">12</emphasis>
. Midleg;
<emphasis id="B940EAC8FFA6FF83FF2FF84AFED56887" bold="true" box="[238,263,1980,2004]" pageId="3" pageNumber="502">13</emphasis>
. Hindleg;
<emphasis id="B940EAC8FFA6FF83FEB2F84AFE5E6887" bold="true" box="[371,396,1980,2004]" pageId="3" pageNumber="502">14</emphasis>
. Claw;
<emphasis id="B940EAC8FFA6FF83FE1DF84AFE276887" bold="true" box="[476,501,1980,2004]" pageId="3" pageNumber="502">15</emphasis>
. Gill I;
<emphasis id="B940EAC8FFA6FF83FD84F84AFD8C6887" bold="true" box="[581,606,1980,2004]" pageId="3" pageNumber="502">16</emphasis>
. Gill III;
<emphasis id="B940EAC8FFA6FF83FD01F84AFD0B6887" bold="true" box="[704,729,1980,2004]" pageId="3" pageNumber="502">17</emphasis>
. Gill VII.
</paragraph>
</caption>
<subSubSection id="C32E6551FFA1FF84FF06FF60FD2B6E4F" pageId="4" pageNumber="503" type="diagnosis">
<paragraph id="8B8B36DAFFA1FF84FF06FF60FD2B6E4F" blockId="4.[151,1436,150,429]" pageId="4" pageNumber="503">
<emphasis id="B940EAC8FFA1FF84FF06FF60FE966FE3" bold="true" box="[199,324,150,176]" pageId="4" pageNumber="503">Diagnosis:</emphasis>
The larva of
<taxonomicName id="4C344D59FFA1FF84FE20FF61FD346FE3" authorityName="Braasch" authorityYear="1990" box="[481,742,150,177]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Epeorus">
<emphasis id="B940EAC8FFA1FF84FE20FF61FDEC6FE3" box="[481,574,151,176]" italics="true" pageId="4" pageNumber="503">Epeorus</emphasis>
(
<emphasis id="B940EAC8FFA1FF84FD90FF61FDBB6FE3" box="[593,617,151,176]" italics="true" pageId="4" pageNumber="503">E.</emphasis>
)
<emphasis id="B940EAC8FFA1FF84FDBAFF60FD346FE3" box="[635,742,150,176]" italics="true" pageId="4" pageNumber="503">aculeatus</emphasis>
</taxonomicName>
can be distinguished from other species of this genus by the following combination of characters: (i) posterior margin of abdominal terga II–IX each with pair of long, acute, submedian spines (
<figureCitation id="130F2A5FFFA1FF84FEACFF28FE006FAB" box="[365,466,222,248]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="4" pageNumber="503">Figs 1–2</figureCitation>
); (ii) anterior margin of labrum with deep median notch (
<figureCitation id="130F2A5FFFA1FF84FB97FF28FB486FAA" box="[1110,1178,222,249]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[191,1400,181,1917]" captionTargetId="figure-19@2.[188,1400,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–8. Larva of Epeorus (Epeorus) aculeatus 1. Dorsal view; 2. Abdomen dorsal view; 3. Abdominal ventral view; 4. Posterior margin of abdominal terga V; 5. Labrum; 6. Hypopharynx; 7. Left mandible; 8. Right mandible." figureDoi="http://doi.org/10.5281/zenodo.5042524" httpUri="https://zenodo.org/record/5042524/files/figure.png" pageId="4" pageNumber="503">Fig. 5</figureCitation>
); (iii) femur with short stout blunt setae scattered on dorsal surface (
<figureCitation id="130F2A5FFFA1FF84FD4CFEF4FD3F6E4F" box="[653,749,258,284]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[162,1426,181,1919]" captionTargetId="figure-17@3.[161,1426,181,1920]" captionTargetPageId="3" captionText="FIGURES 9–17. Larva of Epeorus (Epeorus) aculeatus 9. Maxilla; 10. Labium; 11. Foreleg; 11A. Dorsal surface of femur; 12. Midleg; 13. Hindleg; 14. Claw; 15. Gill I; 16. Gill III; 17. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5042528" httpUri="https://zenodo.org/record/5042528/files/figure.png" pageId="4" pageNumber="503">Fig. 11a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6551FFA1FF84FF06FED0FCE16E12" box="[199,819,294,321]" pageId="4" pageNumber="503" type="distribution">
<paragraph id="8B8B36DAFFA1FF84FF06FED0FCE16E12" blockId="4.[151,1436,150,429]" box="[199,819,294,321]" pageId="4" pageNumber="503">
<emphasis id="B940EAC8FFA1FF84FF06FED0FEB16E13" bold="true" box="[199,355,294,320]" pageId="4" pageNumber="503">Distribution:</emphasis>
<collectingCountry id="F323764AFFA1FF84FEABFED0FE1C6E13" box="[362,462,294,320]" name="Thailand" pageId="4" pageNumber="503">Thailand</collectingCountry>
and
<collectingCountry id="F323764AFFA1FF84FDC5FED0FDEC6E13" box="[516,574,294,320]" name="India" pageId="4" pageNumber="503">India</collectingCountry>
(
<collectingRegion id="49F0F838FFA1FF84FD8DFED0FCF56E12" box="[588,807,294,321]" country="India" name="Arunachal Pradesh" pageId="4" pageNumber="503">Arunachal Pradesh</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6551FFA1FF84FF06FEBCFD166EFF" pageId="4" pageNumber="503" type="discussion">
<paragraph id="8B8B36DAFFA1FF84FF06FEBCFD166EFF" blockId="4.[151,1436,150,429]" pageId="4" pageNumber="503">
<emphasis id="B940EAC8FFA1FF84FF06FEBCFEEF6E37" bold="true" box="[199,317,330,356]" pageId="4" pageNumber="503">Remarks:</emphasis>
The larva of this species was described from
<collectingCountry id="F323764AFFA1FF84FC8DFEBCFC626E37" box="[844,944,330,356]" name="Thailand" pageId="4" pageNumber="503">Thailand</collectingCountry>
by
<bibRefCitation id="EFA54B2BFFA1FF84FC1CFEBCFB596E37" author="Braasch, D." box="[989,1163,330,356]" pageId="4" pageNumber="503" pagination="7 - 14" refId="ref10040" refString="Braasch, D. (1990) Neue Eintagsfliegen aus Thailand, nebsteinigen Bemerkungen zuderen generischem Status (Insecta, Ephemeroptera, Heptageniidae). Reichenbachia, 28 (2), 7 - 14." type="journal article" year="1990">Braasch (1990)</bibRefCitation>
and male adult was described by
<bibRefCitation id="EFA54B2BFFA1FF84FED1FE98FD9A6EDA" author="Webb, J. M. &amp; McCafferty, W. P." box="[272,584,366,393]" pageId="4" pageNumber="503" pagination="65 - 68" refId="ref11214" refString="Webb, J. M. &amp; McCafferty, W. P. (2006) First description of the adult male of Epeorus aculeatus Braasch (Ephemeroptera: Heptageniidae). Zootaxa, 1277, 65 - 68. https: // doi. org / 10.11646 / zootaxa. 1277.1.5" type="journal article" year="2006">Webb &amp; McCafferty (2006)</bibRefCitation>
; we provide a supplementary larval description based on our material, which represents the first record of the species for
<collectingCountry id="F323764AFFA1FF84FD42FE64FD126EFF" box="[643,704,402,428]" name="India" pageId="4" pageNumber="503">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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