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<document id="EE0B9DFAAA65894238BB6058A3BDB2C2" ID-DOI="10.11646/zootaxa.3884.1.6" ID-GBIF-Dataset="843aa177-5b7c-4fbf-a56d-e83360cfde05" ID-ISSN="1175-5326" ID-Zenodo-Dep="287455" ID-ZooBank="C08CC21C-4EC5-4C43-9203-73E4E9CE0A5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460964877558" checkinUser="plazi" docAuthor="Sohn, Jae-Cheon" docDate="2014" docId="039D87C67A4AFFC3FF74EC68FCB282A6" docLanguage="en" docName="zt03884p080.pdf" docOrigin="Zootaxa 3884 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spiladarcha tuberculata Sohn, 2014, n. sp." docType="treatment" docVersion="6" lastPageNumber="79" masterDocId="FFA4FFBE7A4EFFC5FFE3EB3FFFF48126" masterDocTitle="Three new species of Spiladarcha Meyrick, 1913 (Lepidoptera: Urodidae) from Costa Rica" masterLastPageNumber="80" masterPageNumber="73" pageNumber="77" updateTime="1698557684868" updateUser="plazi">
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<mods:titleInfo id="E869A432A18776B3BE15F3A40F5F194C">
<mods:title id="34D9A45FB94F42E4AED89ADB69A8CE8A">Three new species of Spiladarcha Meyrick, 1913 (Lepidoptera: Urodidae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="FB7B494B8B8158C12FAECD28414CD3A5" type="personal">
<mods:role id="F68E01918877E0088A65746A22D4B0D0">
<mods:roleTerm id="131DF09AE2E34127760CAABEBD114039">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D67491449AB932BE357A334765975B8B">Sohn, Jae-Cheon</mods:namePart>
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<mods:title id="CA30ECA2A9DBF38E5A5684F77AA78BB7">Zootaxa</mods:title>
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<mods:part id="D9274F7729F02939B998DB72E14D054C">
<mods:date id="0FFE7685E605606ED120A5A2DCC33EF4">2014</mods:date>
<mods:detail id="B3F9E7044C3CDE5E5F88DA562A90FFE3" type="volume">
<mods:number id="86F7DC57431BB8469ED415CA57AE2066">3884</mods:number>
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<mods:classification id="4BD495571510A3543FC3CDC79A728FC6">journal article</mods:classification>
<mods:identifier id="1050916399B353A176480DD2CE50C930" type="DOI">10.11646/zootaxa.3884.1.6</mods:identifier>
<mods:identifier id="36E501A776CE0874BF89BE876205996A" type="GBIF-Dataset">843aa177-5b7c-4fbf-a56d-e83360cfde05</mods:identifier>
<mods:identifier id="EB8499FEEFB253BD5F8504EF665467EE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="477056D6856ACF87ADCB36F4F458481B" type="Zenodo-Dep">287455</mods:identifier>
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<treatment id="039D87C67A4AFFC3FF74EC68FCB282A6" ID-DOI="http://doi.org/10.5281/zenodo.6136466" ID-GBIF-Taxon="127670783" ID-Zenodo-Dep="6136466" LSID="urn:lsid:plazi:treatment:039D87C67A4AFFC3FF74EC68FCB282A6" httpUri="http://treatment.plazi.org/id/039D87C67A4AFFC3FF74EC68FCB282A6" lastPageId="6" lastPageNumber="79" pageId="4" pageNumber="77">
<subSubSection id="C32E655B7A4AFFC1FF74EC68FD2A8654" box="[151,734,1879,1906]" pageId="4" pageNumber="77" type="nomenclature">
<paragraph id="8B8B36D07A4AFFC1FF74EC68FD2A8654" blockId="4.[151,734,1879,1906]" box="[151,734,1879,1906]" pageId="4" pageNumber="77">
<heading id="D0C381BC7A4AFFC1FF74EC68FD2A8654" bold="true" box="[151,734,1879,1906]" fontSize="11" level="1" pageId="4" pageNumber="77" reason="1">
<emphasis id="B940EAC27A4AFFC1FF74EC68FD2A8654" bold="true" box="[151,734,1879,1906]" pageId="4" pageNumber="77">
<taxonomicName id="4C344D537A4AFFC1FF74EC68FE4F8657" box="[151,443,1879,1905]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="B940EAC27A4AFFC1FF74EC68FE4F8657" bold="true" box="[151,443,1879,1905]" italics="true" pageId="4" pageNumber="77">Spiladarcha tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357B97A4AFFC1FE21EC67FDF08654" box="[450,516,1880,1906]" pageId="4" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="130F2A557A4AFFC1FDF0EC67FD898654" box="[531,637,1880,1906]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1467,1489]" captionTargetBox="[211,1371,555,1442]" captionTargetId="figure@2.[203,1384,555,1442]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Adults of Spiladarcha. 1. S. puravida n. sp., male, holotype (inset: labels); 2. S. puravida n. sp., female, paratype; 3. S. septifera n. sp., male, holotype (inset: labels); 4. S. adamskii Sohn, male, paratype; 5. S. tuberculata n. sp., male, holotype (inset: labels); 6. S. tuberculata n. sp., female, paratype." httpUri="https://zenodo.org/record/287456/files/figure.png" pageId="4" pageNumber="77">Figs 5, 6</figureCitation>
,
<figureCitation id="130F2A557A4AFFC1FD68EC67FD5E8654" box="[651,682,1880,1906]" captionStart="FIGURES 7 – 10" captionStartId="3.[151,264,1811,1833]" captionTargetBox="[227,1384,645,1775]" captionTargetId="figure@3.[188,1400,645,1784]" captionTargetPageId="3" captionText="FIGURES 7 – 10. Male genitalia of Spiladarcha (upper—genital capsule; lower—phallus). 7. S. adamskii Sohn, holotype; 8. S. puravida n. sp., holotype; 9. S. septifera n. sp., holotype; 10. S. tuberculata n. sp., paratype." httpUri="https://zenodo.org/record/287457/files/figure.png" pageId="4" pageNumber="77">10</figureCitation>
,
<figureCitation id="130F2A557A4AFFC1FD54EC67FD218654" box="[695,725,1880,1906]" captionStart="FIGURES 11 – 13" captionStartId="5.[151,264,1263,1285]" captionTargetBox="[306,1278,318,1214]" captionTargetId="figure@5.[292,1295,303,1239]" captionTargetPageId="5" captionText="FIGURES 11 – 13. Female genitalia of Spiladarcha. 11. S. adamskii Sohn, paratype; 12. S. tuberculata n. sp., paratype; 13. S. puravida n. sp., paratype." httpUri="https://zenodo.org/record/287458/files/figure.png" pageId="4" pageNumber="77">12</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E655B7A4AFFC0FF74ECA1FB1B81DE" lastPageId="5" lastPageNumber="78" pageId="4" pageNumber="77" type="diagnosis">
<paragraph id="8B8B36D07A4AFFC0FF74ECA1FB1B81DE" blockId="4.[151,1436,1950,2011]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="78" pageId="4" pageNumber="77">
<emphasis id="B940EAC27A4AFFC1FF74ECA1FEE78691" bold="true" box="[151,275,1950,1975]" pageId="4" pageNumber="77">Diagnosis:</emphasis>
<taxonomicName id="4C344D537A4AFFC1FEF9ECA1FDDC8690" box="[282,552,1950,1974]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B940EAC27A4AFFC1FEF9ECA1FDDC8690" box="[282,552,1950,1974]" italics="true" pageId="4" pageNumber="77">Spiladarcha tuberculata</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C344D537A4AFFC1FD5BECA0FCC18691" box="[696,821,1951,1975]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="septifera">
<emphasis id="B940EAC27A4AFFC1FD5BECA0FCC18691" box="[696,821,1951,1975]" italics="true" pageId="4" pageNumber="77">S. septifera</emphasis>
</taxonomicName>
in the forewing pattern and male genital structures, but differs from the latter in having the basal 1/3 of forewing dark brown (pale brown in
<taxonomicName id="4C344D537A4AFFC1FB73ECFCFAE686FD" box="[1168,1298,1987,2011]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="septifera">
<emphasis id="B940EAC27A4AFFC1FB73ECFCFAE686FD" box="[1168,1298,1987,2011]" italics="true" pageId="4" pageNumber="77">S. septifera</emphasis>
</taxonomicName>
), the uncus triangular (trapezoidal in S.
<taxonomicName id="4C344D537A4BFFC0FE2CEBA7FDC58196" box="[463,561,152,176]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="septifera">
<emphasis id="B940EAC27A4BFFC0FE2CEBA7FDC58196" box="[463,561,152,176]" italics="true" pageId="5" pageNumber="78">septifera</emphasis>
</taxonomicName>
), the saccus broader, and the gnathos longer in the male genitalia.
<taxonomicName id="4C344D537A4BFFC0FAF7EBA7FEE381F2" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B940EAC27A4BFFC0FAF7EBA7FEE381F2" italics="true" pageId="5" pageNumber="78">Spiladarcha tuberculata</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C344D537A4BFFC0FE01EB83FD9781F2" box="[482,611,188,212]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="puravida">
<emphasis id="B940EAC27A4BFFC0FE01EB83FD9781F2" box="[482,611,188,212]" italics="true" pageId="5" pageNumber="78">S. puravida</emphasis>
</taxonomicName>
but can be distinguished from the latter in having a costal process of the valva in the male genitalia and only one signum (two signa in
<taxonomicName id="4C344D537A4BFFC0FCACEBDFFC2981DE" box="[847,989,224,248]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="puravidae">
<emphasis id="B940EAC27A4BFFC0FCACEBDFFC2981DE" box="[847,989,224,248]" italics="true" pageId="5" pageNumber="78">S. puravidae</emphasis>
</taxonomicName>
) in the female genitalia.
</paragraph>
</subSubSection>
<caption id="DF4B66587A4BFFC0FF74EFD0FE6E8402" httpUri="https://zenodo.org/record/287458/files/figure.png" pageId="5" pageNumber="78" targetBox="[306,1278,318,1214]" targetPageId="5">
<paragraph id="8B8B36D07A4BFFC0FF74EFD0FE6E8402" blockId="5.[151,1436,1263,1316]" pageId="5" pageNumber="78">
<emphasis id="B940EAC27A4BFFC0FF74EFD0FEA78423" bold="true" box="[151,339,1263,1285]" pageId="5" pageNumber="78">FIGURES 11–13.</emphasis>
Female genitalia of
<taxonomicName id="4C344D537A4BFFC0FDC4EFD0FD568423" box="[551,674,1263,1285]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAC27A4BFFC0FDC4EFD0FD568423" box="[551,674,1263,1285]" italics="true" pageId="5" pageNumber="78">Spiladarcha</emphasis>
</taxonomicName>
. 11.
<taxonomicName id="4C344D537A4BFFC0FD35EFCFFC7E8423" authority="Sohn" authorityName="Sohn" box="[726,906,1263,1285]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="adamskii">
<emphasis id="B940EAC27A4BFFC0FD35EFCFFCB88423" box="[726,844,1263,1285]" italics="true" pageId="5" pageNumber="78">S. adamskii</emphasis>
Sohn
</taxonomicName>
, paratype; 12.
<taxonomicName id="4C344D537A4BFFC0FBC1EFCFFB448423" box="[1058,1200,1263,1285]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="B940EAC27A4BFFC0FBC1EFCFFB448423" box="[1058,1200,1263,1285]" italics="true" pageId="5" pageNumber="78">S. tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357B97A4BFFC0FB54EFCFFB048423" box="[1207,1264,1264,1285]" pageId="5" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
, paratype; 13.
<taxonomicName id="4C344D537A4BFFC0FA6AEFCFFF078402" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="puravida" status="sp. nov.">
<emphasis id="B940EAC27A4BFFC0FA6AEFCFFF078402" italics="true" pageId="5" pageNumber="78">S. puravida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357B97A4BFFC0FF1AEE30FEC58402" box="[249,305,1295,1316]" pageId="5" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
, paratype.
</paragraph>
</caption>
<subSubSection id="C32E655B7A4BFFC3FF24EE6EFB478232" lastPageId="6" lastPageNumber="79" pageId="5" pageNumber="78" type="description">
<paragraph id="8B8B36D07A4BFFC0FF24EE6EFB5A8494" blockId="5.[151,1437,1361,2034]" pageId="5" pageNumber="78">
<emphasis id="B940EAC27A4BFFC0FF24EE6EFEAF844C" bold="true" box="[199,347,1361,1386]" pageId="5" pageNumber="78">Description:</emphasis>
Head—Scales on vertex and frontoclypeus brownish gray with white tips. Scales of antennal scape grayish brown with white tips dorsally, dark brown ventrally; flagellomere gray. Labial palpus upcurved beyond middle of frons, brownish gray, intermixed with white dorsally, dark brown ventrally.
</paragraph>
<paragraph id="8B8B36D07A4BFFC0FF24EE83FD81863C" blockId="5.[151,1437,1361,2034]" pageId="5" pageNumber="78">
Thorax—Patagium, tegulae and mesonotum brownish gray, intermixed with white. Fore- and midleg with coxa dark grayish brown, intermixed with brownish white; femur dark brown on outer surface, pale gray on inner surface; tibia brown on outer surface, pale gray on inner surface; tarsomere dark brown, with narrow, white ring distally; mid-femur intermixed with gray scales on outer surface. Hindleg pale gray. Forewing length
<quantity id="4CCC9B357A4BFFC0FAD1ED16FA688764" box="[1330,1436,1577,1602]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.0" metricValueMin="4.8" pageId="5" pageNumber="78" unit="mm" value="5.4" valueMax="6.0" valueMin="4.8">4.8–6mm</quantity>
(n=8), pale brownish gray, suffused with dark brownish gray on distal 2/3 and with dark orange along posterior margin, speckled with pale brownish gray on distal 1/2 of costal area; a small pale-orange patch present near lower distal margin of discal cell; two large patches of raised black scales present at about basal 1/3, juxtaposed with orange suffusion basally; two pale-orange spots present in tornal area; small black spots of raised scales scattered along veins on upper surface; fringe dark brown, longer on tornus. Hindwing and fringe dark brownish gray; terminal line very narrow, brownish white.
</paragraph>
<paragraph id="8B8B36D07A4BFFC0FF24EC1AFBEC8618" blockId="5.[151,1437,1361,2034]" box="[199,1048,1829,1854]" pageId="5" pageNumber="78">Abdomen—Dorsal area dark brownish gray; ventral area lustrous pale gray.</paragraph>
<paragraph id="8B8B36D07A4BFFC3FF24EC76FE598066" blockId="5.[151,1437,1361,2034]" lastBlockId="6.[151,1437,151,896]" lastPageId="6" lastPageNumber="79" pageId="5" pageNumber="78">
Male genitalia (
<figureCitation id="130F2A557A4BFFC0FE61EC76FE228647" box="[386,470,1865,1890]" captionStart="FIGURES 7 – 10" captionStartId="3.[151,264,1811,1833]" captionTargetBox="[227,1384,645,1775]" captionTargetId="figure@3.[188,1400,645,1784]" captionTargetPageId="3" captionText="FIGURES 7 – 10. Male genitalia of Spiladarcha (upper—genital capsule; lower—phallus). 7. S. adamskii Sohn, holotype; 8. S. puravida n. sp., holotype; 9. S. septifera n. sp., holotype; 10. S. tuberculata n. sp., paratype." httpUri="https://zenodo.org/record/287457/files/figure.png" pageId="5" pageNumber="78">Fig. 10</figureCitation>
)—Uncus triangular, obtuse apically, with spinose bulge anterolaterally, extending toward an oblique ridge on tegumen. Tegumen trapezoidal in posterior 1/3, elongate, rectangular in anterior 2/3, with a large, semicircular emargination medially on anterior margin of dorsal area. Gnathos arms perpendicular to tegumen, upcurved at basal 1/3, apices tapered and medially convergent. Valva 3.2× as long as uncus, narrow, gradually narrower apically, with long hairs on dorsoapical 1/3; costa slanted diagonally; apex narrowly round; sacculus 7/8 as long as valva, with setal zone along distal half; setae wider and longer on apical part; costal process slender, curved at distal 1/3, dilated in basal 1/3, with setae in basal 2/3 and a spiral line of spines along distal 1/3. Juxta slender, broadened basally, with two divergent apical. Vinculum broadly rounded; saccus linguiform, 1/2 as long as uncus. Phallus slender, slightly broadened to coecum; ductus ejaculatorius arising from anterior end of phallus; cornutus absent.
</paragraph>
<paragraph id="8B8B36D07A48FFC3FF24EA73FD11833E" blockId="6.[151,1437,151,896]" pageId="6" pageNumber="79">
Female A8 and genitalia (
<figureCitation id="130F2A557A48FFC3FE12EA73FDB48043" box="[497,576,332,357]" captionStart="FIGURES 11 – 13" captionStartId="5.[151,264,1263,1285]" captionTargetBox="[306,1278,318,1214]" captionTargetId="figure@5.[292,1295,303,1239]" captionTargetPageId="5" captionText="FIGURES 11 – 13. Female genitalia of Spiladarcha. 11. S. adamskii Sohn, paratype; 12. S. tuberculata n. sp., paratype; 13. S. puravida n. sp., paratype." httpUri="https://zenodo.org/record/287458/files/figure.png" pageId="6" pageNumber="79">Fig. 12</figureCitation>
)—Sternite VIII semicircularly emarginated posteromedially, with a pair of digitate, setose projections posteriorly. Papillae anales small, semi-globular. Ovipositor 2× longer than ductus bursae; A9 with semicircular sclerotized area laterally. Apophyses posteriores 1.4× longer than apophyses anteriores. Ductus bursae narrow, with a weakly sclerotized digitate antrum in posterior 1/3; inception of ductus seminalis present near anterior 1/3 of ductus bursae. Corpus bursae elliptical with a signum at middle; signum spiniform, with granulate, semicircular plate basally.
</paragraph>
<paragraph id="8B8B36D07A48FFC3FF24E91BFB478232" blockId="6.[151,1437,151,896]" pageId="6" pageNumber="79">
<emphasis id="B940EAC27A48FFC3FF24E91BFEE1831B" bold="true" box="[199,277,548,573]" pageId="6" pageNumber="79">
<typeStatus id="548F88727A48FFC3FF24E91BFEE5831B" box="[199,273,548,573]" pageId="6" pageNumber="79">Types</typeStatus>
:
</emphasis>
Holotype—male, “
<typeStatus id="548F88727A48FFC3FE16E91BFD78831A" box="[501,652,548,572]" pageId="6" pageNumber="79" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C344D537A48FFC3FD46E91BFCD9831A" box="[677,813,548,572]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAC27A48FFC3FD46E91BFCD9831A" box="[677,813,548,572]" italics="true" pageId="6" pageNumber="79">Spiladarcha</emphasis>
</taxonomicName>
|
<taxonomicName id="4C344D537A48FFC3FCA4E91BFC33831A" box="[839,967,548,572]" class="Insecta" family="Urodidae" genus="Spiladarcha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="79" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B940EAC27A48FFC3FCA4E91BFC33831A" box="[839,967,548,572]" italics="true" pageId="6" pageNumber="79">tuberculata</emphasis>
</taxonomicName>
|
<bibRefCitation id="EFA54B217A48FFC3FC02E91BFB86831A" author="SOHN" box="[993,1138,548,572]" pageId="6" pageNumber="79" refString="Sohn, J. - C. (2014) Two new species of Incawockia Heppner (Lepidoptera: Urodidae) from Costa Rica and Brazil. Proceedings of the Entomological Society of Washington, 116 (1), 126 - 132. http: // dx. doi. org / 10.4289 / 0013 - 8797.116.1.126" type="journal article" year="2014">SOHN 2014</bibRefCitation>
” [red label enclosed with black lines], “
<collectingCountry id="F32376407A48FFC3FED5E977FE158346" box="[310,481,584,608]" name="Costa Rica" pageId="6" pageNumber="79">COSTA RICA</collectingCountry>
, Heredia: | Est. Biol. La Selva, 50- |
<quantity id="4CCC9B357A48FFC3FC77E978FC238346" box="[916,983,583,608]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="79" unit="m" value="150.0">150m</quantity>
,
<geoCoordinate id="EE0050177A48FFC3FC06E978FBB58346" box="[997,1089,583,608]" direction="north" orientation="latitude" pageId="6" pageNumber="79" precision="925" value="10.433333">10°26'N</geoCoordinate>
<geoCoordinate id="EE0050177A48FFC3FBA8E977FB5A8346" box="[1099,1198,584,608]" direction="west" orientation="longitude" pageId="6" pageNumber="79" precision="925" value="-84.01667">84°01'W</geoCoordinate>
|
<date id="FF8A10107A48FFC3FB24E977FACB8346" box="[1223,1343,584,608]" pageId="6" pageNumber="79" value="1994-06">June 1994</date>
, INBio- OET”, “
<date id="FF8A10107A48FFC3FF11E953FE6683A2" box="[242,402,620,645]" pageId="6" pageNumber="79" value="1994-06-29">29 Junio 1994</date>
| Lab. Area | L/04/108”, the dissected abdomen in a glycerin-filled plastic envelope attached with the specimen, INBIO.
<typeStatus id="548F88727A48FFC3FE2CE9AFFDC8838E" box="[463,572,656,680]" pageId="6" pageNumber="79" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D32FD597A48FFC3FDAFE9AFFD86838E" box="[588,626,656,680]" pageId="6" pageNumber="79" type="generic">4♂</specimenCount>
, 3♀)—1♀, same locality as
<typeStatus id="548F88727A48FFC3FC57E9B0FBED838E" box="[948,1049,655,680]" pageId="6" pageNumber="79" type="holotype">holotype</typeStatus>
,
<date id="FF8A10107A48FFC3FBC7E9AFFB37838E" box="[1060,1219,655,680]" pageId="6" pageNumber="79" value="1993-04-19">19 April 1993</date>
(INBio-OET);
<specimenCount id="9D32FD597A48FFC3FA91E9AFFA6C838E" box="[1394,1432,656,680]" pageId="6" pageNumber="79" type="generic">1♂</specimenCount>
. ditto,
<date id="FF8A10107A48FFC3FF3BE98BFE6B83EA" box="[216,415,692,717]" pageId="6" pageNumber="79" value="1994-02-14">14 February 1994</date>
(INBio-OET);
<specimenCount id="9D32FD597A48FFC3FDADE98BFD8783EA" box="[590,627,692,716]" pageId="6" pageNumber="79" type="generic">2♂</specimenCount>
, 1♀, ditto,
<date id="FF8A10107A48FFC3FD12E98BFC2183EB" box="[753,981,692,717]" pageId="6" pageNumber="79" value="1996-02-05" valueMax="1996-02-14" valueMin="1996-02-05">5–14 February 1996</date>
(INBio-OET), [GSN] SJC-767 (♀); 1♀, ditto,
<date id="FF8A10107A48FFC3FF3DE9E7FE6183D6" box="[222,405,727,752]" pageId="6" pageNumber="79" value="1996-03-13">13 March 1996</date>
(INBio-OET), [GSN] SJC-760;
<specimenCount id="9D32FD597A48FFC3FCFFE9E7FCB483D6" box="[796,832,728,752]" pageId="6" pageNumber="79" type="generic">1♂</specimenCount>
, Puntarenas Prov., Manuel Antonio National Park, Quepos, alt.
<quantity id="4CCC9B357A48FFC3FEC7E9C3FEAD8232" box="[292,345,764,788]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="79" unit="m" value="80.0">80m</quantity>
,
<date id="FF8A10107A48FFC3FE87E9C3FE188232" box="[356,492,764,789]" pageId="6" pageNumber="79" value="1991-03">March 1991</date>
(R Zuniga), [GSN] SJC-529, [WSN] SJC-W022, all in INBIO.
</paragraph>
</subSubSection>
<subSubSection id="C32E655B7A48FFC3FF24E820FD12821E" box="[199,742,799,824]" pageId="6" pageNumber="79" type="distribution">
<paragraph id="8B8B36D07A48FFC3FF24E820FD12821E" blockId="6.[151,1437,151,896]" box="[199,742,799,824]" pageId="6" pageNumber="79">
<emphasis id="B940EAC27A48FFC3FF24E820FE96821E" bold="true" box="[199,354,799,824]" pageId="6" pageNumber="79">Distribution:</emphasis>
<collectingCountry id="F32376407A48FFC3FE8AE81FFE10821E" box="[361,484,799,824]" name="Costa Rica" pageId="6" pageNumber="79">Costa Rica</collectingCountry>
(Heredia, Puntarenas).
</paragraph>
</subSubSection>
<subSubSection id="C32E655B7A48FFC3FF24E87BFCB282A6" pageId="6" pageNumber="79" type="etymology">
<paragraph id="8B8B36D07A48FFC3FF24E87BFCB282A6" blockId="6.[151,1437,151,896]" pageId="6" pageNumber="79">
<emphasis id="B940EAC27A48FFC3FF24E87BFEA5827B" bold="true" box="[199,337,836,861]" pageId="6" pageNumber="79">Etymology:</emphasis>
The species epithet is derived from the Latin word ‘tuber’, meaning “swelling” and refers to the patch of raised scales on the antemedian line of the forewing.
</paragraph>
</subSubSection>
</treatment>
</document>
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