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<document id="9A79B91465A0C453B41CCB6B68288EB8" ID-CLB-Dataset="3143" ID-DOI="10.11646/zootaxa.5072.5.1" ID-GBIF-Dataset="5758176c-96f6-43b9-b9a9-b2fac894dca8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748979" ID-ZooBank="B23A6B99-6D47-469F-8A4F-8AD41D77151A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638425414087" checkinUser="plazi" docAuthor="Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee &amp; Wang, Ying-Yong" docDate="2021" docId="039D87C3FFC90928FF226EC3FD29FF47" docLanguage="en" docName="zootaxa.5072.5.1.pdf" docOrigin="Zootaxa 5072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boulenophrys yingdeensis Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="420" masterDocId="FFA4FFBBFFC6093CFFB56B09FFADFF92" masterDocTitle="Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China" masterLastPageNumber="438" masterPageNumber="401" pageNumber="416" updateTime="1699471732113" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3616EF009571A41A2347DCBE6F9187A5">Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China</mods:title>
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<mods:namePart id="92286A3154CCA89984F104BFBB170813">Qi, Shuo</mods:namePart>
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<mods:namePart id="E454D7B1FC5C55030F48098942F0AA8A">Lyu, Zhi-Tong</mods:namePart>
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<mods:affiliation id="7FC8CE39BE242E76F0EEC7D580598D93">State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China &amp; lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859 &amp; These authors contributed equally to this work</mods:affiliation>
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<mods:namePart id="DB0C4F7D5840770AD81ECC79A3B20977">Wang, Jian</mods:namePart>
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<mods:affiliation id="39D32676D00DEB6956578780AE97AB16">State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China &amp; wangj- 1994 @ outlook. com; https: // orcid. org / 0000 - 0003 - 4249 - 7767</mods:affiliation>
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<mods:namePart id="3820177415E1F15AB0AD20051455E868">Dai, Ke-Yuan</mods:namePart>
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<mods:affiliation id="9B687E4BA78F2558CA123AA8B90D5ED3">Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California 92515, USA &amp; Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA &amp; lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698</mods:affiliation>
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<taxonomicName id="4C344D56FFC90933FF226EC3FE79FA77" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[151,468,1482,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFC90933FF226EC3FE79FA77" bold="true" box="[151,468,1482,1509]" italics="true" pageId="15" pageNumber="416">Boulenophrys yingdeensis</emphasis>
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<taxonomicNameLabel id="A27357BCFFC90933FE696EC2FD94FA77" box="[476,569,1483,1509]" pageId="15" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
Qi, Lyu, Wang &amp; Wang
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<paragraph id="8B8B36D5FFC90933FF226EE6FD4DF99B" blockId="15.[151,927,1482,1611]" box="[151,736,1519,1545]" pageId="15" pageNumber="416">Yingde Horned Toad / ying de jiao chan (Ȓ德fflDz)</paragraph>
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<paragraph id="8B8B36D5FFC90933FF226D1AFEE6F9BF" blockId="15.[151,927,1482,1611]" box="[151,331,1554,1581]" pageId="15" pageNumber="416">
<figureCitation id="130F2A50FFC90933FF226D1AFEAFF9BF" box="[151,258,1555,1581]" captionStart="FIGURE 4" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[151,1435,183,1580]" captionTargetId="figure-18@17.[151,1436,181,1581]" captionTargetPageId="17" captionText="FIGURE 4. Boulenophrys yingdeensis sp. nov. in life. A: holotype (SYS a002099), male; B: paratype (SYS a004721), female; 1: dorsolateral view; 2: ventral view; 3: hand; 4: foot." figureDoi="http://doi.org/10.5281/zenodo.5748987" httpUri="https://zenodo.org/record/5748987/files/figure.png" pageId="15" pageNumber="416">Figures 4</figureCitation>
,
<figureCitation id="130F2A50FFC90933FEBB6D1BFEB3F9BF" box="[270,286,1554,1581]" captionStart="FIGURE 5" captionStartId="19.[151,250,1050,1074]" captionTargetBox="[158,1430,182,1026]" captionTargetId="figure-18@19.[157,1430,181,1026]" captionTargetPageId="19" captionText="FIGURE 5. Color variation of Boulenophrys yingdeensis sp. nov. from the type locality. These individuals were captured, photographed, and finally released." figureDoi="http://doi.org/10.5281/zenodo.5748989" httpUri="https://zenodo.org/record/5748989/files/figure.png" pageId="15" pageNumber="416">5</figureCitation>
,
<figureCitation id="130F2A50FFC90933FE9F6D1AFEE6F9BF" box="[298,331,1555,1581]" captionStart="FIGURE 7" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[313,1274,181,1919]" captionTargetId="figure-21@24.[313,1274,181,1920]" captionTargetPageId="24" captionText="FIGURE 7. Comparisons of lateral position of the head with three new Boulenophrys species in preservative. The white arrows marking tubercles bearing tiny spine in Boulenophrys yunkaiensis sp. nov.. A: Boulenophrys yaoshanensis sp. nov. (SYS a002189); B: Boulenophrys yingdeensis sp. nov. (SYS a002099); C: Boulenophrys yunkaiensis sp. nov. (SYS a004637)." figureDoi="http://doi.org/10.5281/zenodo.5748993" httpUri="https://zenodo.org/record/5748993/files/figure.png" pageId="15" pageNumber="416">7B</figureCitation>
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<emphasis id="B940EAC7FFC90933FF226D3AFE88F9D9" bold="true" box="[151,293,1587,1611]" inLineHeading="true" pageId="15" pageNumber="416" reason="1">Chresonymy.</emphasis>
<taxonomicName id="4C344D56FFC90933FE996D3AFE63F9D9" box="[300,462,1587,1611]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="undefined-4">
<emphasis id="B940EAC7FFC90933FE996D3AFE33F9D9" box="[300,414,1587,1611]" italics="true" pageId="15" pageNumber="416">Megophrys</emphasis>
sp 4
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(SYS a002100, 4721, 5447)—
<bibRefCitation id="EFA54B24FFC90933FCBD6D3AFC32F9D9" author="Liu, Z. Y. &amp; Chen, G. L. &amp; Zhu, T. Q. &amp; Zeng, Z. C. &amp; Lyu, Z. T. &amp; Wang, J. &amp; Messenger, K. &amp; Greenberg, A. J. &amp; Guo, Z. X. &amp; Yang, Z. H. &amp; Shi, S. H. &amp; Wang, Y. Y." box="[776,927,1587,1611]" pageId="15" pageNumber="416" pagination="723 - 731" refId="ref16886" refString="Liu, Z. Y., Chen, G. L., Zhu, T. Q., Zeng, Z. C., Lyu, Z. T., Wang, J., Messenger, K., Greenberg, A. J., Guo, Z. X., Yang, Z. H., Shi, S. H. &amp; Wang, Y. Y. (2018) Prevalence of cryptic species in morphologically uniform taxa - Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution, 127, 723 - 731. https: // doi. org / 10.1016 / j. ympev. 2018.06.020" type="journal article" year="2018">
Liu
<emphasis id="B940EAC7FFC90933FC866D3DFCF2F9D9" box="[819,863,1587,1611]" italics="true" pageId="15" pageNumber="416">et al</emphasis>
. 2018
</bibRefCitation>
</paragraph>
</subSubSection>
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<materialsCitation id="3B5C3C88FFC90933FF226D70FAAFF925" ID-GBIF-Occurrence="3413047303" collectingDate="2013-04-26" country="China" county="Yingde City" elevation="357" latitude="24.4435" location="Run-Lin Li" longLatPrecision="7" longitude="113.3034" municipality="Shimentai Nature Reserve" pageId="15" pageNumber="416" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
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<typeStatus id="548F8877FFC90933FF226D70FEABF901" box="[151,262,1657,1683]" pageId="15" pageNumber="416" type="holotype">Holotype</typeStatus>
.
</emphasis>
SYS a002099 (
<figureCitation id="130F2A50FFC90933FE0B6D70FDEDF901" box="[446,576,1657,1683]" captionStart="FIGURE 4" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[151,1435,183,1580]" captionTargetId="figure-18@17.[151,1436,181,1581]" captionTargetPageId="17" captionText="FIGURE 4. Boulenophrys yingdeensis sp. nov. in life. A: holotype (SYS a002099), male; B: paratype (SYS a004721), female; 1: dorsolateral view; 2: ventral view; 3: hand; 4: foot." figureDoi="http://doi.org/10.5281/zenodo.5748987" httpUri="https://zenodo.org/record/5748987/files/figure.png" pageId="15" pageNumber="416">Figures 4A</figureCitation>
,
<figureCitation id="130F2A50FFC90933FDF86D73FDDDF906" box="[589,624,1658,1684]" captionStart="FIGURE 7" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[313,1274,181,1919]" captionTargetId="figure-21@24.[313,1274,181,1920]" captionTargetPageId="24" captionText="FIGURE 7. Comparisons of lateral position of the head with three new Boulenophrys species in preservative. The white arrows marking tubercles bearing tiny spine in Boulenophrys yunkaiensis sp. nov.. A: Boulenophrys yaoshanensis sp. nov. (SYS a002189); B: Boulenophrys yingdeensis sp. nov. (SYS a002099); C: Boulenophrys yunkaiensis sp. nov. (SYS a004637)." figureDoi="http://doi.org/10.5281/zenodo.5748993" httpUri="https://zenodo.org/record/5748993/files/figure.png" pageId="15" pageNumber="416">7B</figureCitation>
), adult male, collected on
<date id="FF8A1015FFC90933FC296D70FB94F901" box="[924,1081,1657,1683]" pageId="15" pageNumber="416" value="2013-04-26">
<collectingDate id="EFCEE9FDFFC90933FC296D70FB94F901" box="[924,1081,1657,1683]" pageId="15" pageNumber="416" value="2013-04-26">26 April 2013</collectingDate>
</date>
by
<location id="8EEB600EFFC90933FBD06D70FB48F901" LSID="urn:lsid:plazi:treatment:039D87C3FFC90928FF226EC3FD29FF47:8EEB600EFFC90933FBD06D70FB48F901" box="[1125,1253,1657,1683]" country="China" county="Yingde City" latitude="24.4435" longLatPrecision="7" longitude="113.3034" municipality="Shimentai Nature Reserve" name="Run-Lin Li" pageId="15" pageNumber="416" stateProvince="Guangdong">Run-Lin Li</location>
from
<collectingMunicipality id="6BEFACAFFFC90933FA9E6D70FEE9F92A" pageId="15" pageNumber="416">Shimentai Nature Reserve</collectingMunicipality>
(
<geoCoordinate id="EE005012FFC90933FEE66D94FE7FF92A" box="[339,466,1693,1720]" degrees="24.4435" direction="north" orientation="latitude" pageId="15" pageNumber="416" precision="5" value="24.4435">24.4435°N</geoCoordinate>
,
<geoCoordinate id="EE005012FFC90933FE686D94FDC8F925" box="[477,613,1693,1719]" degrees="113.3034" direction="east" orientation="longitude" pageId="15" pageNumber="416" precision="5" value="113.3034">113.3034°E</geoCoordinate>
; ca
<elevation id="0019D1E6FFC90933FD256D94FCBEF92A" box="[656,787,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="3.57" pageId="15" pageNumber="416" unit="m" value="357.0">
<quantity id="4CCC9B30FFC90933FD256D94FD7AF92A" box="[656,727,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="3.57" pageId="15" pageNumber="416" unit="m" value="357.0">357 m</quantity>
a.s.l.
</elevation>
),
<collectingCounty id="62EA4E59FFC90933FC9D6D94FC1CF925" box="[808,945,1693,1719]" pageId="15" pageNumber="416">Yingde City</collectingCounty>
,
<collectingRegion id="49F0F837FFC90933FC086D94FB01F925" box="[957,1196,1693,1719]" country="China" name="Guangdong" pageId="15" pageNumber="416">Guangdong Province</collectingRegion>
,
<collectingCountry id="F3237645FFC90933FB0D6D94FB52F925" box="[1208,1279,1693,1719]" name="China" pageId="15" pageNumber="416">China</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8B36D5FFC90933FF726DC8FABBF8FE" blockId="15.[151,1437,1657,2008]" pageId="15" pageNumber="416">
<materialsCitation id="3B5C3C88FFC90933FF726DC8FABFF8FE" ID-GBIF-Occurrence="3413047313" collectingDate="2012-04-23" collectingDateMax="2018-08-20" collectingDateMin="2012-04-23" collectionCode="CIB" country="China" county="Yingde City" elevation="357" latitude="24.4435" location="Run-Lin Li" longLatPrecision="7" longitude="113.3034" municipality="Shimentai Nature Reserve" pageId="15" pageNumber="416" specimenCode="CIB 116084" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="B940EAC7FFC90933FF726DC8FEEBF949" bold="true" box="[199,326,1729,1755]" pageId="15" pageNumber="416">
<typeStatus id="548F8877FFC90933FF726DC8FEECF949" box="[199,321,1729,1755]" pageId="15" pageNumber="416" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Eight adult specimens from the same locality as the holotype: female SYS a001563 collected on
<date id="FF8A1015FFC90933FF226DECFE9AF96D" box="[151,311,1765,1791]" pageId="15" pageNumber="416" value="2012-04-23">
<collectingDate id="EFCEE9FDFFC90933FF226DECFE9AF96D" box="[151,311,1765,1791]" pageId="15" pageNumber="416" value="2012-04-23">23 April 2012</collectingDate>
</date>
by Run-Lin Li; female SYS a004721 collected on
<date id="FF8A1015FFC90933FC306DECFB89F96D" box="[901,1060,1765,1791]" pageId="15" pageNumber="416" value="2016-04-29">
<collectingDate id="EFCEE9FDFFC90933FC306DECFB89F96D" box="[901,1060,1765,1791]" pageId="15" pageNumber="416" value="2016-04-29">29 April 2016</collectingDate>
</date>
by Ying-Yong Wang, Jian Wang and Zhi-Tong Lyu; female SYS a005447 and male SYS a005449 collected on
<date id="FF8A1015FFC90933FBA36C00FB67F8B1" box="[1046,1226,1801,1828]" pageId="15" pageNumber="416" value="2016-08-19">
<collectingDate id="EFCEE9FDFFC90933FBA36C00FB67F8B1" box="[1046,1226,1801,1828]" pageId="15" pageNumber="416" value="2016-08-19">19 August 2016</collectingDate>
</date>
by Zhi-Tong Lyu; male SYS a007114/
<specimenCode id="DB929EAEFFC90933FEC26C24FE50F8D5" box="[375,509,1837,1863]" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" pageId="15" pageNumber="416">CIB 116084</specimenCode>
collected on
<date id="FF8A1015FFC90933FD246C24FC88F8D5" box="[657,805,1837,1864]" pageId="15" pageNumber="416" value="2018-06-20">
<collectingDate id="EFCEE9FDFFC90933FD246C24FC88F8D5" box="[657,805,1837,1864]" pageId="15" pageNumber="416" value="2018-06-20">20 June 2018</collectingDate>
</date>
by Hong-Hui Chen, Jia-He Li and Jian Wang; male SYS a007405 and
<specimenCount id="9D32FD5CFFC90933FE9B6C5BFE1BF8F9" box="[302,438,1873,1900]" pageId="15" pageNumber="416" type="female">two females</specimenCount>
SYS a007406 and SYS a007407 collected on
<date id="FF8A1015FFC90933FC0A6C58FBDCF8F9" box="[959,1137,1873,1900]" pageId="15" pageNumber="416" value="2018-08-20">
<collectingDate id="EFCEE9FDFFC90933FC0A6C58FBDCF8F9" box="[959,1137,1873,1900]" pageId="15" pageNumber="416" value="2018-08-20">20 August 2018</collectingDate>
</date>
by Jian Wang
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E655EFFC90933FF726C7CFAD7F81D" box="[199,1402,1909,1936]" pageId="15" pageNumber="416" type="etymology">
<paragraph id="8B8B36D5FFC90933FF726C7CFAD7F81D" blockId="15.[151,1437,1657,2008]" box="[199,1402,1909,1936]" pageId="15" pageNumber="416">
<emphasis id="B940EAC7FFC90933FF726C7CFEE3F81D" bold="true" box="[199,334,1909,1935]" pageId="15" pageNumber="416">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C344D56FFC90933FD8C6C7CFD11F81D" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[569,700,1909,1935]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="416" phylum="Chordata" rank="species" species="yingdeensis">
<emphasis id="B940EAC7FFC90933FD8C6C7CFD11F81D" box="[569,700,1909,1935]" italics="true" pageId="15" pageNumber="416">yingdeensis</emphasis>
</taxonomicName>
refers to its
<typeStatus id="548F8877FFC90933FCF26C7FFCDAF802" box="[839,887,1910,1936]" pageId="15" pageNumber="416">type</typeStatus>
locality, Yingde City in northern
<collectingRegion id="49F0F837FFC90933FB5A6C7CFAD8F81D" box="[1263,1397,1909,1935]" country="China" name="Guangdong" pageId="15" pageNumber="416">Guangdong</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E655EFFC9092CFF726C90FCFDF92C" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="diagnosis">
<paragraph id="8B8B36D5FFC9092CFF726C90FF4DFE43" blockId="15.[151,1437,1657,2008]" lastBlockId="16.[151,1437,150,2013]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
<emphasis id="B940EAC7FFC90933FF726C90FEEFF821" bold="true" box="[199,322,1945,1971]" pageId="15" pageNumber="416">Diagnosis.</emphasis>
(1) Small body size, SVL
<quantity id="4CCC9B30FFC90933FDDE6C90FCA3F826" box="[619,782,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="3.425" metricValueMax="3.53" metricValueMin="3.32" pageId="15" pageNumber="416" unit="mm" value="34.25" valueMax="35.3" valueMin="33.2">33.235.3 mm</quantity>
(34.2 ± 1.0, N = 4) in adult males and SVL
<quantity id="4CCC9B30FFC90933FB4F6C90FA30F826" box="[1274,1437,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="4.1049999999999995" metricValueMax="4.58" metricValueMin="3.63" pageId="15" pageNumber="416" unit="mm" value="41.05" valueMax="45.8" valueMin="36.3">36.345.8 mm</quantity>
(40.7 ± 4.2, N = 5) in adult females; (2) snout rounded in dorsal view; (3) tympanum boundary clear, ED/TD
<quantity id="4CCC9B30FFC9092CFAE86CB4FF4AFF23" lastPageId="16" lastPageNumber="417" metricMagnitude="-2" metricUnit="m" metricValue="4.571999999999999" metricValueMax="4.953" metricValueMin="4.191" pageId="15" pageNumber="416" unit="in" value="1.7999999999999998" valueMax="1.95" valueMin="1.65">1.65 1.95 in</quantity>
males,
<quantity id="4CCC9B30FFD6092CFEF56B9EFE62FF23" box="[320,463,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.5339" metricValueMax="5.308599999999999" metricValueMin="3.7592" pageId="16" pageNumber="417" unit="in" value="1.785" valueMax="2.09" valueMin="1.48">1.482.09 in</quantity>
females; (4) vomerine ridge prominent bearing vomerine teeth; (5) margin of tongue rounded, not notched behind; (6) hindlimbs slender, heels overlapping or just meeting and tibio-tarsal articulation reaching forward between tympanum to posterior corner of eye; (7) tibia 0.460.48 of SVL and foot 0.610.67 of SVL in males, while tibia 0.440.46 of SVL and foot 0.610.66 of SVL in females; (8) toes without lateral fringes and with only rudimentary webbing; (9) presence of small, horn-like tubercle at the edge of upper eyelid; (10) dorsal skin smooth, a discontinuous “Y” or “X”-shaped ridge on the mid-dorsum, two discontinuous dorsolateral ridges on two side on the dorsum; (11) skin of flanks smooth with small conical tubercles; (12) single subgular vocal sac in males; (13) presence of villiform black nuptial spines on the dorsal surface of the first and second fingers in adult males.
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF726AD3FC70FD8B" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
<emphasis id="B940EAC7FFD6092CFF726AD3FEC0FE67" bold="true" box="[199,365,474,501]" pageId="16" pageNumber="417">Comparisons.</emphasis>
Comparative data of
<taxonomicName id="4C344D56FFD6092CFDD46AD2FC2AFE67" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[609,903,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFDD46AD2FC2AFE67" box="[609,903,475,501]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFC3B6AD2FC45FE67" bold="true" box="[910,1000,475,501]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFC3B6AD2FC45FE67" box="[910,1000,475,501]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C344D56FFD6092CFB986AD2FB4CFE67" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[1069,1249,475,501]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yaoshanensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFB986AD2FB4CFE67" box="[1069,1249,475,501]" italics="true" pageId="16" pageNumber="417">B. yaoshanensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFB5C6AD2FAEEFE67" bold="true" box="[1257,1347,475,501]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFB5C6AD2FAEEFE67" box="[1257,1347,475,501]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the other recognized members of the genus
<taxonomicName id="4C344D56FFD6092CFDE26AF6FD5EFD8B" authorityName="Fei, Ye &amp; Jiang" authorityYear="2016" box="[599,755,511,537]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B940EAC7FFD6092CFDE26AF6FD5EFD8B" box="[599,755,511,537]" italics="true" pageId="16" pageNumber="417">Boulenophrys</emphasis>
</taxonomicName>
are listed in
<tableCitation id="C6B6036EFFD6092CFC366AF6FC7AFD8B" box="[899,983,511,537]" captionStart="TABLE 3" captionStartId="8.[150,243,507,534]" captionTargetPageId="8" captionText="TABLE 3. Diagnostic characters separating all 61 species of the Boulenophrys." httpUri="http://table.plazi.org/id/DF4B665DFFCE0934FF236AF2FBBDFD84" pageId="16" pageNumber="417" tableUuid="DF4B665DFFCE0934FF236AF2FBBDFD84">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF72692AFE79FD5C" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Having a smaller body size with SVL
<quantity id="4CCC9B30FFD6092CFD25692BFC9AFDAF" box="[656,823,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.425" metricValueMax="3.53" metricValueMin="3.32" pageId="16" pageNumber="417" unit="mm" value="34.25" valueMax="35.3" valueMin="33.2">33.235.3 mm</quantity>
in males,
<taxonomicName id="4C344D56FFD6092CFC03692AFB4DFDAF" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[950,1248,547,573]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFC03692AFB4DFDAF" box="[950,1248,547,573]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFB5E692AFAE4FDAF" bold="true" box="[1259,1353,547,573]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFB5E692AFAE4FDAF" box="[1259,1353,547,573]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from seven congeners whose SVL&gt;
<quantity id="4CCC9B30FFD6092CFDFF694FFD30FDF3" box="[586,669,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="417" unit="mm" value="50.0">50 mm</quantity>
in males, including
<taxonomicName id="4C344D56FFD6092CFC3A694EFB92FDF3" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Shen" baseAuthorityYear="1994" box="[911,1087,583,609]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="caudoprocta">
<emphasis id="B940EAC7FFD6092CFC3A694EFB92FDF3" box="[911,1087,583,609]" italics="true" pageId="16" pageNumber="417">B. caudoprocta</emphasis>
</taxonomicName>
(
<quantity id="4CCC9B30FFD6092CFBE4694EFB17FDF3" box="[1105,1210,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="8.129999999999999" pageId="16" pageNumber="417" unit="mm" value="81.3">81.3 mm</quantity>
in single male),
<taxonomicName id="4C344D56FFD6092CFA31694EFE9EFD17" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B940EAC7FFD6092CFA31694EFE9EFD17" italics="true" pageId="16" pageNumber="417">B. jingdongensis</emphasis>
</taxonomicName>
(53.0
<quantity id="4CCC9B30FFD6092CFE336963FE40FD17" box="[390,493,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" pageId="16" pageNumber="417" unit="mm" value="56.5">56.5 mm</quantity>
in males),
<taxonomicName id="4C344D56FFD6092CFDC66962FD57FD17" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang &amp; Zhou" baseAuthorityYear="2017" box="[627,762,619,645]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="B940EAC7FFD6092CFDC66962FD57FD17" box="[627,762,619,645]" italics="true" pageId="16" pageNumber="417">B. liboensis</emphasis>
</taxonomicName>
(
<quantity id="4CCC9B30FFD6092CFCB96963FC19FD17" box="[780,948,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" metricValueMax="6.77" metricValueMin="6.05" pageId="16" pageNumber="417" unit="mm" value="64.1" valueMax="67.7" valueMin="60.5">60.567.7 mm</quantity>
in males),
<taxonomicName id="4C344D56FFD6092CFB8F6962FB6FFD17" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang &amp; Zhao" baseAuthorityYear="2020" box="[1082,1218,619,645]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B940EAC7FFD6092CFB8F6962FB6FFD17" box="[1082,1218,619,645]" italics="true" pageId="16" pageNumber="417">B. mirabilis</emphasis>
</taxonomicName>
(
<quantity id="4CCC9B30FFD6092CFB616963FAD1FD17" box="[1236,1404,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.859999999999999" metricValueMax="6.14" metricValueMin="5.58" pageId="16" pageNumber="417" unit="mm" value="58.599999999999994" valueMax="61.4" valueMin="55.8">55.861.4 mm</quantity>
in males),
<taxonomicName id="4C344D56FFD6092CFF456986FE3CFD3A" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[240,401,655,680]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="B940EAC7FFD6092CFF456986FE3CFD3A" box="[240,401,655,680]" italics="true" pageId="16" pageNumber="417">B. omeimontis</emphasis>
</taxonomicName>
(56.0
<quantity id="4CCC9B30FFD6092CFE546987FDEBFD3B" box="[481,582,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" pageId="16" pageNumber="417" unit="mm" value="59.5">59.5 mm</quantity>
in males),
<taxonomicName id="4C344D56FFD6092CFD766986FCDCFD3B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Jiang, Ye &amp; Fei" baseAuthorityYear="2008" box="[707,881,655,681]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B940EAC7FFD6092CFD766986FCDCFD3B" box="[707,881,655,681]" italics="true" pageId="16" pageNumber="417">B. sangzhiensis</emphasis>
</taxonomicName>
(
<quantity id="4CCC9B30FFD6092CFC356987FC4BFD3B" box="[896,998,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="16" pageNumber="417" unit="mm" value="54.7">54.7 mm</quantity>
in single male), and
<taxonomicName id="4C344D56FFD6092CFB666986FA30FD3B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" box="[1235,1437,655,681]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="B940EAC7FFD6092CFB666986FA30FD3B" box="[1235,1437,655,681]" italics="true" pageId="16" pageNumber="417">B. shuichengensis</emphasis>
</taxonomicName>
(102.0
<quantity id="4CCC9B30FFD6092CFF5869BAFEF3FD5F" box="[237,350,691,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.183" pageId="16" pageNumber="417" unit="mm" value="118.3">118.3 mm</quantity>
in males).
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF7269DEFAFAFCCC" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Having relatively longer shanks with the heels overlapping or meeting when the flexed hindlimbs are held at right angles to the body axis,
<taxonomicName id="4C344D56FFD6092CFE4569F2FCB5FC87" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[496,792,763,789]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFE4569F2FCB5FC87" box="[496,792,763,789]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFC9469F2FCD0FC87" bold="true" box="[801,893,763,789]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFC9469F2FCD0FC87" box="[801,893,763,789]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from the following nine congeners,
<taxonomicName id="4C344D56FFD6092CFEE76816FE1FFCAA" baseAuthorityName="Wang, Li &amp; Jin" baseAuthorityYear="2014" box="[338,434,799,824]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="acuta">
<emphasis id="B940EAC7FFD6092CFEE76816FE1FFCAA" box="[338,434,799,824]" italics="true" pageId="16" pageNumber="417">B. acuta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE776816FDC0FCAB" baseAuthorityName="Inger &amp; Romer" baseAuthorityYear="1961" box="[450,621,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="brachykolos">
<emphasis id="B940EAC7FFD6092CFE776816FDC0FCAB" box="[450,621,799,825]" italics="true" pageId="16" pageNumber="417">B. brachykolos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDC86816FD75FCAB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng, Wang &amp; Wang" baseAuthorityYear="2021" box="[637,728,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="daoji">
<emphasis id="B940EAC7FFD6092CFDC86816FD75FCAB" box="[637,728,799,825]" italics="true" pageId="16" pageNumber="417">B. daoji</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD5D6816FC1CFCAB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" baseAuthorityYear="2021" box="[744,945,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="dongguanensis">
<emphasis id="B940EAC7FFD6092CFD5D6816FC1CFCAB" box="[744,945,799,825]" italics="true" pageId="16" pageNumber="417">B. dongguanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC746816FBE8FCAB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Liu, Lyu, Zeng &amp; Wang" baseAuthorityYear="2017" box="[961,1093,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="insularis">
<emphasis id="B940EAC7FFD6092CFC746816FBE8FCAB" box="[961,1093,799,825]" italics="true" pageId="16" pageNumber="417">B. insularis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBE06816FAACFCAB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Zeng &amp; Wang" baseAuthorityYear="2019" box="[1109,1281,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="nankunensis">
<emphasis id="B940EAC7FFD6092CFBE06816FAACFCAB" box="[1109,1281,799,825]" italics="true" pageId="16" pageNumber="417">B. nankunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFAA46816FAD9FCAB" baseAuthorityName="Wang, Li &amp; Zhao" baseAuthorityYear="2014" box="[1297,1396,799,825]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="obesa">
<emphasis id="B940EAC7FFD6092CFAA46816FAD9FCAB" box="[1297,1396,799,825]" italics="true" pageId="16" pageNumber="417">B. obesa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316816FEB4FCCF" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Messenger &amp; Dahn" baseAuthorityYear="2019" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="ombrophila">
<emphasis id="B940EAC7FFD6092CFA316816FEB4FCCF" italics="true" pageId="16" pageNumber="417">B. ombrophila</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C344D56FFD6092CFEE3684AFDABFCCE" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" box="[342,518,835,860]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wugongensis">
<emphasis id="B940EAC7FFD6092CFEE3684AFDABFCCE" box="[342,518,835,860]" italics="true" pageId="16" pageNumber="417">B. wugongensis</emphasis>
</taxonomicName>
(vs. all of which have relatively shorter shanks with the heels not meeting).
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF72686EFDA9FBC8" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Having vomerine teeth,
<taxonomicName id="4C344D56FFD6092CFE62686EFD50FC13" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[471,765,871,897]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFE62686EFD50FC13" box="[471,765,871,897]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFCB1686EFCF0FC13" bold="true" box="[772,861,871,897]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFCB1686EFCF0FC13" box="[772,861,871,897]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C344D56FFD6092CFC47686EFBFDFC12" baseAuthorityName="Wang, Li &amp; Jin" baseAuthorityYear="2014" box="[1010,1104,871,896]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="acuta">
<emphasis id="B940EAC7FFD6092CFC47686EFBFDFC12" box="[1010,1104,871,896]" italics="true" pageId="16" pageNumber="417">B. acuta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBEB686EFB6FFC13" box="[1118,1218,871,897]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="angka">
<emphasis id="B940EAC7FFD6092CFBEB686EFB6FFC13" box="[1118,1218,871,897]" italics="true" pageId="16" pageNumber="417">B. angka</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB64686EFADBFC13" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Li, Lu, Liu &amp; Wang" baseAuthorityYear="2020" box="[1233,1398,871,897]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="anlongensis">
<emphasis id="B940EAC7FFD6092CFB64686EFADBFC13" box="[1233,1398,871,897]" italics="true" pageId="16" pageNumber="417">B. anlongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA31686EFEECFC37" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wu, Li, Liu, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baishanzuensis">
<emphasis id="B940EAC7FFD6092CFA31686EFEECFC37" italics="true" pageId="16" pageNumber="417">B. baishanzuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEF86882FE53FC37" baseAuthorityName="Ye, Fei &amp; Xie" baseAuthorityYear="2007" box="[333,510,907,933]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="B940EAC7FFD6092CFEF86882FE53FC37" box="[333,510,907,933]" italics="true" pageId="16" pageNumber="417">B. baolongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDBC6882FD6BFC37" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" box="[521,710,907,933]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="B940EAC7FFD6092CFDBC6882FD6BFC37" box="[521,710,907,933]" italics="true" pageId="16" pageNumber="417">B. binchuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD666882FCDAFC37" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Jiang, Fei &amp; Ye" baseAuthorityYear="2009" box="[723,887,907,933]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="binlingensis">
<emphasis id="B940EAC7FFD6092CFD666882FCDAFC37" box="[723,887,907,933]" italics="true" pageId="16" pageNumber="417">B. binlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC366882FBA8FC37" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[899,1029,907,933]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="B940EAC7FFD6092CFC366882FBA8FC37" box="[899,1029,907,933]" italics="true" pageId="16" pageNumber="417">B. boettgeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBA46882FB1AFC37" baseAuthorityName="Inger &amp; Romer" baseAuthorityYear="1961" box="[1041,1207,907,933]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="brachykolos">
<emphasis id="B940EAC7FFD6092CFBA46882FB1AFC37" box="[1041,1207,907,933]" italics="true" pageId="16" pageNumber="417">B. brachykolos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB766882FAD5FC37" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Nguyen, Pham, Nguyen, Luong &amp; Ziegler" baseAuthorityYear="2020" box="[1219,1400,907,933]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="caobangensis">
<emphasis id="B940EAC7FFD6092CFB766882FAD5FC37" box="[1219,1400,907,933]" italics="true" pageId="16" pageNumber="417">B. caobangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316882FE92FC5A" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng &amp; Zhou" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="congjiangensis">
<emphasis id="B940EAC7FFD6092CFA316882FE92FC5A" italics="true" pageId="16" pageNumber="417">B. congjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEF868A6FE04FC5B" baseAuthorityName="Wang &amp; Liu" baseAuthorityYear="2014" box="[333,425,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="cheni">
<emphasis id="B940EAC7FFD6092CFEF868A6FE04FC5B" box="[333,425,943,969]" italics="true" pageId="16" pageNumber="417">B. cheni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE0368A6FDF3FC5B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Xu, Li, Liu, Wei &amp; Wang" baseAuthorityYear="2020" box="[438,606,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="chishuiensis">
<emphasis id="B940EAC7FFD6092CFE0368A6FDF3FC5B" box="[438,606,943,969]" italics="true" pageId="16" pageNumber="417">B. chishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDDE68A6FD69FC5B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng, Wang &amp; Wang" baseAuthorityYear="2021" box="[619,708,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="daoji">
<emphasis id="B940EAC7FFD6092CFDDE68A6FD69FC5B" box="[619,708,943,969]" italics="true" pageId="16" pageNumber="417">B. daoji</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD6468A6FC0EFC5B" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="2005" box="[721,931,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="huangshanensis">
<emphasis id="B940EAC7FFD6092CFD6468A6FC0EFC5B" box="[721,931,943,969]" italics="true" pageId="16" pageNumber="417">B. huangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC0568A6FBBDFC5A" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang &amp; Wu" baseAuthorityYear="2020" box="[944,1040,943,968]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jiangi">
<emphasis id="B940EAC7FFD6092CFC0568A6FBBDFC5A" box="[944,1040,943,968]" italics="true" pageId="16" pageNumber="417">B. jiangi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBAB68A6FB6CFC5B" baseAuthorityName="Pope" baseAuthorityYear="1929" box="[1054,1217,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="B940EAC7FFD6092CFBAB68A6FB6CFC5B" box="[1054,1217,943,969]" italics="true" pageId="16" pageNumber="417">B. kuatunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB7A68A6FADAFC5B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Li, Xu, Liu, Jiang, Wei &amp; Wang" baseAuthorityYear="2018" box="[1231,1399,943,969]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="leishanensis">
<emphasis id="B940EAC7FFD6092CFB7A68A6FADAFC5B" box="[1231,1399,943,969]" italics="true" pageId="16" pageNumber="417">B. leishanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA3168A6FF11FC7F" baseAuthorityName="Wang &amp; Yang" baseAuthorityYear="2014" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lini">
<emphasis id="B940EAC7FFD6092CFA3168A6FF11FC7F" italics="true" pageId="16" pageNumber="417">B. lini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF7C68DAFEF1FC7F" baseAuthorityName="Wang, Liu &amp; Jiang" baseAuthorityYear="2017" box="[201,348,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lishuiensis">
<emphasis id="B940EAC7FFD6092CFF7C68DAFEF1FC7F" box="[201,348,979,1005]" italics="true" pageId="16" pageNumber="417">B. lishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEDD68DAFDACFC7F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang &amp; Wang" baseAuthorityYear="2021" box="[360,513,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lushuiensis">
<emphasis id="B940EAC7FFD6092CFEDD68DAFDACFC7F" box="[360,513,979,1005]" italics="true" pageId="16" pageNumber="417">B. lushuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDBB68DAFDC3FC7E" box="[526,622,979,1004]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="minor">
<emphasis id="B940EAC7FFD6092CFDBB68DAFDC3FC7E" box="[526,622,979,1004]" italics="true" pageId="16" pageNumber="417">B. minor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDCF68DAFD51FC7F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang &amp; Zhao" baseAuthorityYear="2020" box="[634,764,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B940EAC7FFD6092CFDCF68DAFD51FC7F" box="[634,764,979,1005]" italics="true" pageId="16" pageNumber="417">B. mirabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCBD68DAFC6DFC7F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" box="[776,960,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="mufumontana">
<emphasis id="B940EAC7FFD6092CFCBD68DAFC6DFC7F" box="[776,960,979,1005]" italics="true" pageId="16" pageNumber="417">B. mufumontana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC7968DAFB87FC7F" baseAuthorityName="Wang, Li &amp; Zhao" baseAuthorityYear="2014" box="[972,1066,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="obesa">
<emphasis id="B940EAC7FFD6092CFC7968DAFB87FC7F" box="[972,1066,979,1005]" italics="true" pageId="16" pageNumber="417">B. obesa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB8368DAFB78FC7F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Messenger &amp; Dahn" baseAuthorityYear="2019" box="[1078,1237,979,1005]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="ombrophila">
<emphasis id="B940EAC7FFD6092CFB8368DAFB78FC7F" box="[1078,1237,979,1005]" italics="true" pageId="16" pageNumber="417">B. ombrophila</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB5468DAFA38FC7E" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" box="[1249,1429,979,1004]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="B940EAC7FFD6092CFB5468DAFA38FC7E" box="[1249,1429,979,1004]" italics="true" pageId="16" pageNumber="417">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF2268FEFECFFB83" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" box="[151,354,1015,1041]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="B940EAC7FFD6092CFF2268FEFECFFB83" box="[151,354,1015,1041]" italics="true" pageId="16" pageNumber="417">B. shuichengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEC768FEFDE8FB83" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Deng, Liu, Wu &amp; Liu" baseAuthorityYear="2019" box="[370,581,1015,1041]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shunhuangensis">
<emphasis id="B940EAC7FFD6092CFEC768FEFDE8FB83" box="[370,581,1015,1041]" italics="true" pageId="16" pageNumber="417">B. shunhuangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDE068FEFD65FB82" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" box="[597,712,1015,1040]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="spinata">
<emphasis id="B940EAC7FFD6092CFDE068FEFD65FB82" box="[597,712,1015,1040]" italics="true" pageId="16" pageNumber="417">B. spinata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD6D68FEFC17FB83" baseAuthorityName="Shen, Mo &amp; Li" baseAuthorityYear="2010" box="[728,954,1015,1041]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="tuberogranulatus">
<emphasis id="B940EAC7FFD6092CFD6D68FEFC17FB83" box="[728,954,1015,1041]" italics="true" pageId="16" pageNumber="417">B. tuberogranulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC7E68FEFBD0FB82" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" box="[971,1149,1015,1040]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wugongensis">
<emphasis id="B940EAC7FFD6092CFC7E68FEFBD0FB82" box="[971,1149,1015,1040]" italics="true" pageId="16" pageNumber="417">B. wugongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB3968FEFAD9FB83" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" box="[1164,1396,1015,1041]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wuliangshanensis">
<emphasis id="B940EAC7FFD6092CFB3968FEFAD9FB83" box="[1164,1396,1015,1041]" italics="true" pageId="16" pageNumber="417">B. wuliangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA3168FEFE88FBA7" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wushanensis">
<emphasis id="B940EAC7FFD6092CFA3168FEFE88FBA7" italics="true" pageId="16" pageNumber="417">B. wushanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE836F12FE55FBA6" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" box="[310,504,1051,1076]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="xiangnanensis">
<emphasis id="B940EAC7FFD6092CFE836F12FE55FBA6" box="[310,504,1051,1076]" italics="true" pageId="16" pageNumber="417">B. xiangnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDBC6F12FD04FBA6" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Wu, Peng, Shi, Lu &amp; Wu" baseAuthorityYear="2020" box="[521,681,1051,1076]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="xianjuensis">
<emphasis id="B940EAC7FFD6092CFDBC6F12FD04FBA6" box="[521,681,1051,1076]" italics="true" pageId="16" pageNumber="417">B. xianjuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD0E6F12FC2EFBA6" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng, &amp; Wang" baseAuthorityYear="2020" box="[699,899,1051,1076]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yangmingensis">
<emphasis id="B940EAC7FFD6092CFD0E6F12FC2EFBA6" box="[699,899,1051,1076]" italics="true" pageId="16" pageNumber="417">B. yangmingensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C344D56FFD6092CFC726F12FBD3FBA7" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[967,1150,1051,1077]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yaoshanensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFC726F12FBD3FBA7" box="[967,1150,1051,1077]" italics="true" pageId="16" pageNumber="417">B. yaoshanensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFB3D6F12FB48FBA7" bold="true" box="[1160,1253,1051,1077]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFB3D6F12FB48FBA7" box="[1160,1253,1051,1077]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(vs. absence of vomerine teeth in these species).
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF726F6AFEFBFA9C" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Having an unnotched tongue,
<taxonomicName id="4C344D56FFD6092CFDAB6F6AFCE8FBEF" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[542,837,1123,1149]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFDAB6F6AFCE8FBEF" box="[542,837,1123,1149]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFCFB6F6AFC04FBEF" bold="true" box="[846,937,1123,1149]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFCFB6F6AFC04FBEF" box="[846,937,1123,1149]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C344D56FFD6092CFB546F6AFA38FBEF" baseAuthorityName="Ye, Fei &amp; Xie" baseAuthorityYear="2007" box="[1249,1429,1123,1149]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="B940EAC7FFD6092CFB546F6AFA38FBEF" box="[1249,1429,1123,1149]" italics="true" pageId="16" pageNumber="417">B. baolongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF226F8EFEECFB33" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Jiang, Fei &amp; Ye" baseAuthorityYear="2009" box="[151,321,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="binlingensis">
<emphasis id="B940EAC7FFD6092CFF226F8EFEECFB33" box="[151,321,1159,1185]" italics="true" pageId="16" pageNumber="417">B. binlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEE76F8EFE77FB33" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[338,474,1159,1185]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="B940EAC7FFD6092CFEE76F8EFE77FB33" box="[338,474,1159,1185]" italics="true" pageId="16" pageNumber="417">B. boettgeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE5E6F8EFDE6FB33" baseAuthorityName="Wang &amp; Liu" baseAuthorityYear="2014" box="[491,587,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="cheni">
<emphasis id="B940EAC7FFD6092CFE5E6F8EFDE6FB33" box="[491,587,1159,1185]" italics="true" pageId="16" pageNumber="417">B. cheni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDE96F8EFC8AFB33" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" box="[604,807,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="hoanglienensis">
<emphasis id="B940EAC7FFD6092CFDE96F8EFC8AFB33" box="[604,807,1159,1185]" italics="true" pageId="16" pageNumber="417">B. hoanglienensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC8D6F8EFBA0FB33" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="2005" box="[824,1037,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="huangshanensis">
<emphasis id="B940EAC7FFD6092CFC8D6F8EFBA0FB33" box="[824,1037,1159,1185]" italics="true" pageId="16" pageNumber="417">B. huangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBAA6F8EFB0EFB33" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Liu, Lyu, Zeng &amp; Wang" baseAuthorityYear="2017" box="[1055,1187,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="insularis">
<emphasis id="B940EAC7FFD6092CFBAA6F8EFB0EFB33" box="[1055,1187,1159,1185]" italics="true" pageId="16" pageNumber="417">B. insularis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB006F8EFADEFB33" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" box="[1205,1395,1159,1185]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B940EAC7FFD6092CFB006F8EFADEFB33" box="[1205,1395,1159,1185]" italics="true" pageId="16" pageNumber="417">B. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316F8EFEB5FB57" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" baseAuthorityYear="2021" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jiulianensis">
<emphasis id="B940EAC7FFD6092CFA316F8EFEB5FB57" italics="true" pageId="16" pageNumber="417">B. jiulianensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE936FA2FE64FB57" baseAuthorityName="Pope" baseAuthorityYear="1929" box="[294,457,1195,1221]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="B940EAC7FFD6092CFE936FA2FE64FB57" box="[294,457,1195,1221]" italics="true" pageId="16" pageNumber="417">B. kuatunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE626FA2FDF6FB57" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang &amp; Zhou" baseAuthorityYear="2017" box="[471,603,1195,1221]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="B940EAC7FFD6092CFE626FA2FDF6FB57" box="[471,603,1195,1221]" italics="true" pageId="16" pageNumber="417">B. liboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDDD6FA2FCAEFB57" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang &amp; Wang" baseAuthorityYear="2021" box="[616,771,1195,1221]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lushuiensis">
<emphasis id="B940EAC7FFD6092CFDDD6FA2FCAEFB57" box="[616,771,1195,1221]" italics="true" pageId="16" pageNumber="417">B. lushuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCA76FA2FCD9FB56" box="[786,884,1195,1220]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="minor">
<emphasis id="B940EAC7FFD6092CFCA76FA2FCD9FB56" box="[786,884,1195,1220]" italics="true" pageId="16" pageNumber="417">B. minor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC346FA2FB82FB57" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang, Liu &amp; Wang" baseAuthorityYear="2019" box="[897,1071,1195,1221]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="nanlingensis">
<emphasis id="B940EAC7FFD6092CFC346FA2FB82FB57" box="[897,1071,1195,1221]" italics="true" pageId="16" pageNumber="417">B. nanlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB896FA2FB71FB56" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[1084,1244,1195,1220]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="B940EAC7FFD6092CFB896FA2FB71FB56" box="[1084,1244,1195,1220]" italics="true" pageId="16" pageNumber="417">B. omeimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB5F6FA2FA38FB57" box="[1258,1429,1195,1221]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="qianbeiensis">
<emphasis id="B940EAC7FFD6092CFB5F6FA2FA38FB57" box="[1258,1429,1195,1221]" italics="true" pageId="16" pageNumber="417">B. qianbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF226FC6FEE9FB7B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Jiang, Ye &amp; Fei" baseAuthorityYear="2008" box="[151,324,1231,1257]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B940EAC7FFD6092CFF226FC6FEE9FB7B" box="[151,324,1231,1257]" italics="true" pageId="16" pageNumber="417">B. sangzhiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEE76FC6FDA5FB7A" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" box="[338,520,1231,1256]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="B940EAC7FFD6092CFEE76FC6FDA5FB7A" box="[338,520,1231,1256]" italics="true" pageId="16" pageNumber="417">B. sanmingensis</emphasis>
</taxonomicName>
,
<emphasis id="B940EAC7FFD6092CFDA36FC6FCFBFB7A" box="[534,854,1231,1257]" italics="true" pageId="16" pageNumber="417">
<taxonomicName id="4C344D56FFD6092CFDA36FC6FD72FB7B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" box="[534,735,1231,1257]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shuichengensis">B. shuichengensis</taxonomicName>
<taxonomicName id="4C344D56FFD6092CFD506FC6FCFBFB7A" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" box="[741,854,1231,1256]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="spinata">B. spinata</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C344D56FFD6092CFC216FC6FB94FB7B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang &amp; Lyu" baseAuthorityYear="2021" box="[916,1081,1231,1257]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="tongboensis">
<emphasis id="B940EAC7FFD6092CFC216FC6FB94FB7B" box="[916,1081,1231,1257]" italics="true" pageId="16" pageNumber="417">B. tongboensis</emphasis>
</taxonomicName>
(vs. tongue notched posteriorly in these species).
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF726E1EFED8F99A" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Lacking lateral fringes on toes,
<taxonomicName id="4C344D56FFD6092CFD806E1EFCF0FAA3" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[565,861,1303,1329]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFD806E1EFCF0FAA3" box="[565,861,1303,1329]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFCD36E1EFC6FFAA3" bold="true" box="[870,962,1303,1329]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFCD36E1EFC6FFAA3" box="[870,962,1303,1329]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C344D56FFD6092CFBE86E1EFB11FAA2" baseAuthorityName="Wang, Li &amp; Jin" baseAuthorityYear="2014" box="[1117,1212,1303,1329]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="acuta">
<emphasis id="B940EAC7FFD6092CFBE86E1EFBC3FAA2" box="[1117,1134,1303,1328]" italics="true" pageId="16" pageNumber="417">B</emphasis>
.
<emphasis id="B940EAC7FFD6092CFBCB6E1EFB11FAA2" box="[1150,1212,1303,1328]" italics="true" pageId="16" pageNumber="417">acuta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB786E1EFAD9FAA3" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Li, Lu, Liu &amp; Wang" baseAuthorityYear="2020" box="[1229,1396,1303,1329]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="anlongensis">
<emphasis id="B940EAC7FFD6092CFB786E1EFAD9FAA3" box="[1229,1396,1303,1329]" italics="true" pageId="16" pageNumber="417">B. anlongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316E1EFEECFAC7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wu, Li, Liu, Wang &amp; Wu" baseAuthorityYear="2020" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baishanzuensis">
<emphasis id="B940EAC7FFD6092CFA316E1EFA38FAA2" box="[1412,1429,1303,1328]" italics="true" pageId="16" pageNumber="417">B</emphasis>
.
<emphasis id="B940EAC7FFD6092CFF226E32FEECFAC7" box="[151,321,1339,1365]" italics="true" pageId="16" pageNumber="417">baishanzuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEF96E32FDA5FAC7" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" box="[332,520,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="binchuanensis">
<emphasis id="B940EAC7FFD6092CFEF96E32FDA5FAC7" box="[332,520,1339,1365]" italics="true" pageId="16" pageNumber="417">B. binchuanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDA66E32FD39FAC7" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[531,660,1339,1365]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="B940EAC7FFD6092CFDA66E32FD39FAC7" box="[531,660,1339,1365]" italics="true" pageId="16" pageNumber="417">B. boettgeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD2A6E32FCCEFAC6" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Luo, Wang, Wang, Lu, Wang, Deng &amp; Zhou" baseAuthorityYear="2021" box="[671,867,1339,1364]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="congjiangensis">
<emphasis id="B940EAC7FFD6092CFD2A6E32FCCEFAC6" box="[671,867,1339,1364]" italics="true" pageId="16" pageNumber="417">B. congjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCDB6E32FC6AFAC7" baseAuthorityName="Wang &amp; Liu" baseAuthorityYear="2014" box="[878,967,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="cheni">
<emphasis id="B940EAC7FFD6092CFCDB6E32FC6AFAC7" box="[878,967,1339,1365]" italics="true" pageId="16" pageNumber="417">B. cheni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC676E32FBDFFAC7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang &amp; Wang" baseAuthorityYear="2021" box="[978,1138,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="daiyunensis">
<emphasis id="B940EAC7FFD6092CFC676E32FBDFFAC7" box="[978,1138,1339,1365]" italics="true" pageId="16" pageNumber="417">B. daiyunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBC86E32FB7EFAC7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng, Wang &amp; Wang" baseAuthorityYear="2021" box="[1149,1235,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="daoji">
<emphasis id="B940EAC7FFD6092CFBC86E32FB7EFAC7" box="[1149,1235,1339,1365]" italics="true" pageId="16" pageNumber="417">B. daoji</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB6B6E32FA3BFAC7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" box="[1246,1430,1339,1365]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B940EAC7FFD6092CFB6B6E32FA3BFAC7" box="[1246,1430,1339,1365]" italics="true" pageId="16" pageNumber="417">B. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF226E56FEFFFAEA" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang" baseAuthorityYear="2012" box="[151,338,1375,1400]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="B940EAC7FFD6092CFF226E56FEFFFAEA" box="[151,338,1375,1400]" italics="true" pageId="16" pageNumber="417">B. jinggangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEEB6E56FE4CFAEB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang &amp; Zhou" baseAuthorityYear="2017" box="[350,481,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="liboensis">
<emphasis id="B940EAC7FFD6092CFEEB6E56FE4CFAEB" box="[350,481,1375,1401]" italics="true" pageId="16" pageNumber="417">B. liboensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE586E56FD9CFAEB" baseAuthorityName="Wang &amp; Yang" baseAuthorityYear="2014" box="[493,561,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lini">
<emphasis id="B940EAC7FFD6092CFE586E56FD9CFAEB" box="[493,561,1375,1401]" italics="true" pageId="16" pageNumber="417">B. lini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD8B6E56FD74FAEB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Shi, Li, Zhu, Jiang, Jiang &amp; Wang" baseAuthorityYear="2021" box="[574,729,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lushuiensis">
<emphasis id="B940EAC7FFD6092CFD8B6E56FD74FAEB" box="[574,729,1375,1401]" italics="true" pageId="16" pageNumber="417">B. lushuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD536E56FCC7FAEB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang &amp; Zhao" baseAuthorityYear="2020" box="[742,874,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B940EAC7FFD6092CFD536E56FCC7FAEB" box="[742,874,1375,1401]" italics="true" pageId="16" pageNumber="417">B. mirabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCC26E56FB82FAEB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Lyu &amp; Wang" baseAuthorityYear="2019" box="[887,1071,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="mufumontana">
<emphasis id="B940EAC7FFD6092CFCC26E56FB82FAEB" box="[887,1071,1375,1401]" italics="true" pageId="16" pageNumber="417">B. mufumontana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB886E56FB44FAEB" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Wang, Liu &amp; Wang" baseAuthorityYear="2019" box="[1085,1257,1375,1401]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="nanlingensis">
<emphasis id="B940EAC7FFD6092CFB886E56FB44FAEB" box="[1085,1257,1375,1401]" italics="true" pageId="16" pageNumber="417">B. nanlingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB436E56FA38FAEA" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[1270,1429,1375,1400]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="omeimontis">
<emphasis id="B940EAC7FFD6092CFB436E56FA38FAEA" box="[1270,1429,1375,1400]" italics="true" pageId="16" pageNumber="417">B. omeimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF226E8AFE23FA0F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[151,398,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="palpebralespinosa">
<emphasis id="B940EAC7FFD6092CFF226E8AFE23FA0F" box="[151,398,1411,1437]" italics="true" pageId="16" pageNumber="417">B. palpebralespinosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE116E8AFDFAFA0F" box="[420,599,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="qianbeiensis">
<emphasis id="B940EAC7FFD6092CFE116E8AFDFAFA0F" box="[420,599,1411,1437]" italics="true" pageId="16" pageNumber="417">B. qianbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDD86E8AFCA5FA0F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong &amp; Rowley" baseAuthorityYear="2017" box="[621,776,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="rubrimera">
<emphasis id="B940EAC7FFD6092CFDD86E8AFCA5FA0F" box="[621,776,1411,1437]" italics="true" pageId="16" pageNumber="417">B. rubrimera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCAB6E8AFC7EFA0F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Jiang, Ye &amp; Fei" baseAuthorityYear="2008" box="[798,979,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sangzhiensis">
<emphasis id="B940EAC7FFD6092CFCAB6E8AFC7EFA0F" box="[798,979,1411,1437]" italics="true" pageId="16" pageNumber="417">B. sangzhiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC5C6E8AFB05FA0E" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu &amp; Wang" baseAuthorityYear="2021" box="[1001,1192,1411,1436]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="sanmingensis">
<emphasis id="B940EAC7FFD6092CFC5C6E8AFB05FA0E" box="[1001,1192,1411,1436]" italics="true" pageId="16" pageNumber="417">B. sanmingensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB0B6E8AFAC3FA0F" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Liu, &amp; Wang" baseAuthorityYear="2020" box="[1214,1390,1411,1437]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shimentaina">
<emphasis id="B940EAC7FFD6092CFB0B6E8AFAC3FA0F" box="[1214,1390,1411,1437]" italics="true" pageId="16" pageNumber="417">B. shimentaina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316E8AFEECFA53" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="B940EAC7FFD6092CFA316E8AFEECFA53" italics="true" pageId="16" pageNumber="417">B. shuichengensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEFA6EAEFE6CFA52" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" box="[335,449,1447,1472]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="spinata">
<emphasis id="B940EAC7FFD6092CFEFA6EAEFE6CFA52" box="[335,449,1447,1472]" italics="true" pageId="16" pageNumber="417">B. spinata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE656EAEFD22FA52" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng &amp; Wang" baseAuthorityYear="2020" box="[464,655,1447,1472]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="xiangnanensis">
<emphasis id="B940EAC7FFD6092CFE656EAEFD22FA52" box="[464,655,1447,1472]" italics="true" pageId="16" pageNumber="417">B. xiangnanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD2B6EAEFC91FA52" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang, Wu, Peng, Shi, Lu &amp; Wu" baseAuthorityYear="2020" box="[670,828,1447,1472]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="xianjuensis">
<emphasis id="B940EAC7FFD6092CFD2B6EAEFC91FA52" box="[670,828,1447,1472]" italics="true" pageId="16" pageNumber="417">B. xianjuensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C344D56FFD6092CFCCF6EAEFBEDFA52" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Zeng, &amp; Wang" baseAuthorityYear="2020" box="[890,1088,1447,1472]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yangmingensis">
<emphasis id="B940EAC7FFD6092CFCCF6EAEFBEDFA52" box="[890,1088,1447,1472]" italics="true" pageId="16" pageNumber="417">B. yangmingensis</emphasis>
</taxonomicName>
(vs. presence of lateral fringes on toes in these species); and from
<taxonomicName id="4C344D56FFD6092CFD806EC2FD48FA77" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" box="[565,741,1483,1509]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wushanensis">
<emphasis id="B940EAC7FFD6092CFD806EC2FD48FA77" box="[565,741,1483,1509]" italics="true" pageId="16" pageNumber="417">B. wushanensis</emphasis>
</taxonomicName>
(vs. presence of wide lateral fringes on toes in males while lacking in females).
</paragraph>
<paragraph id="8B8B36D5FFD6092CFF726D1AFCFDF92C" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
Having rudimentary webbing on toes,
<taxonomicName id="4C344D56FFD6092CFD346D1AFC04F9BF" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[641,937,1555,1581]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD6092CFD346D1AFC04F9BF" box="[641,937,1555,1581]" italics="true" pageId="16" pageNumber="417">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD6092CFC076D1AFBA3F9BF" bold="true" box="[946,1038,1555,1581]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A27357BCFFD6092CFC076D1AFBA3F9BF" box="[946,1038,1555,1581]" pageId="16" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C344D56FFD6092CFB1C6D1AFAD9F9BF" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wu, Li, Liu, Wang &amp; Wu" baseAuthorityYear="2020" box="[1193,1396,1555,1581]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baishanzuensis">
<emphasis id="B940EAC7FFD6092CFB1C6D1AFAD9F9BF" box="[1193,1396,1555,1581]" italics="true" pageId="16" pageNumber="417">B. baishanzuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFA316D1AFE86F9C3" baseAuthorityName="Ye, Fei &amp; Xie" baseAuthorityYear="2007" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="baolongensis">
<emphasis id="B940EAC7FFD6092CFA316D1AFE86F9C3" italics="true" pageId="16" pageNumber="417">B. baolongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE8B6D3EFE5DF9C3" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Rao &amp; Yang" baseAuthorityYear="1997" box="[318,496,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="daweimontis">
<emphasis id="B940EAC7FFD6092CFE8B6D3EFE5DF9C3" box="[318,496,1591,1617]" italics="true" pageId="16" pageNumber="417">B. daweimontis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFDB66D3EFD12F9C3" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong &amp; Rowley" baseAuthorityYear="2018" box="[515,703,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="fansipanensis">
<emphasis id="B940EAC7FFD6092CFDB66D3EFD12F9C3" box="[515,703,1591,1617]" italics="true" pageId="16" pageNumber="417">B. fansipanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD676D3EFCEFF9C3" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong &amp; Rowley" baseAuthorityYear="2021" box="[722,834,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="frigida">
<emphasis id="B940EAC7FFD6092CFD676D3EFCEFF9C3" box="[722,834,1591,1617]" italics="true" pageId="16" pageNumber="417">B. frigida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFCE36D3EFB80F9C3" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="2005" box="[854,1069,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="huangshanensis">
<emphasis id="B940EAC7FFD6092CFCE36D3EFB80F9C3" box="[854,1069,1591,1617]" italics="true" pageId="16" pageNumber="417">B. huangshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFBF56D3EFB45F9C3" baseAuthorityName="Pope" baseAuthorityYear="1929" box="[1088,1256,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="B940EAC7FFD6092CFBF56D3EFB45F9C3" box="[1088,1256,1591,1617]" italics="true" pageId="16" pageNumber="417">B. kuatunensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFB4E6D3EFA38F9C3" baseAuthorityName="Wang, Liu &amp; Jiang" baseAuthorityYear="2017" box="[1275,1429,1591,1617]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="lishuiensis">
<emphasis id="B940EAC7FFD6092CFB4E6D3EFA38F9C3" box="[1275,1429,1591,1617]" italics="true" pageId="16" pageNumber="417">B. lishuiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFF226D52FE96F9E7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Messenger &amp; Dahn" baseAuthorityYear="2019" box="[151,315,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="ombrophila">
<emphasis id="B940EAC7FFD6092CFF226D52FE96F9E7" box="[151,315,1627,1653]" italics="true" pageId="16" pageNumber="417">B. ombrophila</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFEFF6D52FE72F9E7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong &amp; Rowley" baseAuthorityYear="2017" box="[330,479,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="rubrimera">
<emphasis id="B940EAC7FFD6092CFEFF6D52FE72F9E7" box="[330,479,1627,1653]" italics="true" pageId="16" pageNumber="417">B. rubrimera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFE5A6D52FD3BF9E7" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Wang &amp; Lyu" baseAuthorityYear="2021" box="[495,662,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="tongboensis">
<emphasis id="B940EAC7FFD6092CFE5A6D52FD3BF9E7" box="[495,662,1627,1653]" italics="true" pageId="16" pageNumber="417">B. tongboensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C344D56FFD6092CFD6D6D52FC6DF9E7" baseAuthorityName="Ye &amp; Fei" baseAuthorityYear="1995" box="[728,960,1627,1653]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="wuliangshanensis">
<emphasis id="B940EAC7FFD6092CFD6D6D52FC6DF9E7" box="[728,960,1627,1653]" italics="true" pageId="16" pageNumber="417">B. wuliangshanensis</emphasis>
</taxonomicName>
(vs. absence of webbing on toes in these species); and from
<taxonomicName id="4C344D56FFD6092CFEDA6D76FD86F90B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Fei &amp; Ye" baseAuthorityYear="1983" box="[367,555,1663,1689]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="jingdongensis">
<emphasis id="B940EAC7FFD6092CFEDA6D76FD86F90B" box="[367,555,1663,1689]" italics="true" pageId="16" pageNumber="417">B. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFD8C6D76FC85F90B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[569,808,1663,1689]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="palpebralespinosa">
<emphasis id="B940EAC7FFD6092CFD8C6D76FC85F90B" box="[569,808,1663,1689]" italics="true" pageId="16" pageNumber="417">B. palpebralespinosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC836D76FC4FF90B" box="[822,994,1663,1689]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="qianbeiensis">
<emphasis id="B940EAC7FFD6092CFC836D76FC4FF90B" box="[822,994,1663,1689]" italics="true" pageId="16" pageNumber="417">B. qianbeiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D56FFD6092CFC446D76FB16F90B" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Tian &amp; Sun" baseAuthorityYear="1995" box="[1009,1211,1663,1689]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="shuichengensis">
<emphasis id="B940EAC7FFD6092CFC446D76FB16F90B" box="[1009,1211,1663,1689]" italics="true" pageId="16" pageNumber="417">B. shuichengensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C344D56FFD6092CFB4C6D76FAC6F90A" baseAuthorityName="Liu &amp; Hu" baseAuthorityYear="1973" box="[1273,1387,1663,1688]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="417" phylum="Chordata" rank="species" species="spinata">
<emphasis id="B940EAC7FFD6092CFB4C6D76FAC6F90A" box="[1273,1387,1663,1688]" italics="true" pageId="16" pageNumber="417">B. spinata</emphasis>
</taxonomicName>
(vs. presence of well-developed webbing on toes in these species).
</paragraph>
</subSubSection>
<subSubSection id="C32E655EFFD6092FFF726DCFFD0EF8FD" lastPageId="19" lastPageNumber="420" pageId="16" pageNumber="417" type="description">
<paragraph id="8B8B36D5FFD6092CFF726DCFFF49F84F" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="417">
<emphasis id="B940EAC7FFD6092CFF726DCFFE47F973" bold="true" box="[199,490,1734,1761]" pageId="16" pageNumber="417">
Description of
<typeStatus id="548F8877FFD6092CFEC96DCEFE48F973" box="[380,485,1735,1761]" pageId="16" pageNumber="417" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male. small body size, SVL
<quantity id="4CCC9B30FFD6092CFC336DCFFC42F973" box="[902,1007,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="16" pageNumber="417" unit="mm" value="35.3">35.3 mm</quantity>
; head width slightly larger than head length, HWD/HDL 1.07; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size, ED 0.30 of HDL, pupil vertical, near diamond-shaped; nostril oblique-ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanum boundary clear, ED/TD 1.68; large ovoid choanae at the base of the maxilla; vomerine ridge prominent, vomerine teeth present, maxillary teeth present; margin of tongue rounded, not notched behind; presence of a single subgular vocal sac, a pair of slit-like openings at posterior of jaw.
</paragraph>
<caption id="DF4B665DFFD7092DFF226D4CFD1BF913" ID-DOI="http://doi.org/10.5281/zenodo.5748987" ID-Zenodo-Dep="5748987" httpUri="https://zenodo.org/record/5748987/files/figure.png" pageId="17" pageNumber="418" startId="17.[151,250,1605,1629]" targetBox="[151,1435,183,1580]" targetPageId="17">
<paragraph id="8B8B36D5FFD7092DFF226D4CFD1BF913" blockId="17.[151,1437,1605,1665]" pageId="17" pageNumber="418">
<emphasis id="B940EAC7FFD7092DFF226D4CFEBEF9CF" bold="true" box="[151,275,1605,1629]" pageId="17" pageNumber="418">FIGURE 4.</emphasis>
<taxonomicName id="4C344D56FFD7092DFEAC6D4CFD8FF9CF" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[281,546,1605,1629]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="17" pageNumber="418" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD7092DFEAC6D4CFD8FF9CF" box="[281,546,1605,1629]" italics="true" pageId="17" pageNumber="418">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD7092DFD9D6D4CFDD4F9CF" bold="true" box="[552,633,1605,1629]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="A27357BCFFD7092DFD9D6D4CFDD4F9CF" box="[552,633,1605,1629]" pageId="17" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. A: holotype (SYS a002099), male; B: paratype (SYS a004721), female; 1: dorsolateral view; 2: ventral view; 3: hand; 4: foot.
</paragraph>
</caption>
<paragraph id="8B8B36D5FFD7092DFF726DA6FF4BF87B" blockId="17.[151,1437,1711,2025]" pageId="17" pageNumber="418">Radio-ulnar length 0.20 of SVL and hand 0.26 of SVL; hand without webbing, fingers without lateral fringes, relative finger length I &lt;II &lt;IV &lt;III; tips of fingers slightly dilated, round; subarticular tubercles present; inner metacarpal tubercle observably enlarged and the outer one smaller; a single nuptial pad on the dorsal surface of the first and second fingers. Hindlimbs slender, tibio-tarsal articulation reaching forward the posterior corner of the eye when hindlimb is stretched along the side of the body; heels overlapping when the flexed hindlimbs are held at right angles to the body axis; tibia length 0.48 of SVL and foot length 0.67 of SVL; relative toe length I &lt;II &lt;V &lt;III &lt;IV; tips of toes round and slightly dilated; toes without lateral fringes and with rudiment of webs; ventromedial callous pad on toe I is round, on others are long strip shape; inner metatarsal tubercle long ovoid and the outer one absent.</paragraph>
<caption id="DF4B665DFFD4092EFF2369A8F98FFD2E" ID-Table-UUID="DF4B665DFFD4092EFF2369A8F98FFD2E" box="[150,1570,673,700]" httpUri="http://table.plazi.org/id/DF4B665DFFD4092EFF2369A8F98FFD2E" pageId="18" pageNumber="419" startId="18.[150,243,673,700]" targetBox="[164,2010,715,1542]" targetIsTable="true" targetPageId="18">
<paragraph id="8B8B36D5FFD4092EFF2369A8F98FFD2E" blockId="18.[150,1570,673,700]" box="[150,1570,673,700]" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF2369A8FEA2FD2E" bold="true" box="[150,271,673,700]" pageId="18" pageNumber="419">TABLE 5.</emphasis>
Measurements (in mm) and ratios of the type series of
<taxonomicName id="4C344D56FFD4092EFCCF69A8FB0DFD2E" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[890,1184,673,700]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="18" pageNumber="419" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD4092EFCCF69A8FB0DFD2E" box="[890,1184,673,700]" italics="true" pageId="18" pageNumber="419">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27357BCFFD4092EFB1369A8FAADFD2E" box="[1190,1280,673,700]" pageId="18" pageNumber="419" rank="species">
<emphasis id="B940EAC7FFD4092EFB1369A8FB56FD2E" bold="true" box="[1190,1275,673,700]" pageId="18" pageNumber="419">sp. nov</emphasis>
.
</taxonomicNameLabel>
, * indicates the holotype.
</paragraph>
</caption>
<paragraph id="8B8B36D5FFD4092EFEEE69C2F8D5F994" pageId="18" pageNumber="419">
<table id="F934C475FFD4F6C3FF1169C2F877F994" box="[164,2010,715,1542]" gridcols="10" gridrows="18" pageId="18" pageNumber="419">
<tr id="35043497FFD4F6C3FF1169C2F877FC9A" box="[164,2010,715,776]" gridrow="0" pageId="18" pageNumber="419" rowspan-0="1">
<th id="76D55DEBFFD4F6C3FEEE69C2FE58FC9A" box="[347,501,715,776]" gridcol="1" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFEEE69C2FE58FD76" bold="true" box="[347,501,715,740]" pageId="18" pageNumber="419">SYS a002099*</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FDA369C2FD09FC9A" box="[534,676,715,776]" gridcol="2" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFDA369C2FD09FD76" bold="true" box="[534,676,715,740]" pageId="18" pageNumber="419">SYS a005449</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FD7869C2FCCDFC9A" box="[717,864,715,776]" gridcol="3" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFD7869C2FCE1FC9A" bold="true" pageId="18" pageNumber="419">SYS a007114/ CIB 116084</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FC0369C2FBE9FC9A" box="[950,1092,715,776]" gridcol="4" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFC0369C2FBE9FD76" bold="true" box="[950,1092,715,740]" pageId="18" pageNumber="419">SYS a007405</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FBDB69C2FB51FC9A" box="[1134,1276,715,776]" gridcol="5" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFBDB69C2FB51FD76" bold="true" box="[1134,1276,715,740]" pageId="18" pageNumber="419">SYS a001563</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FA9069C2FA1EFC9A" box="[1317,1459,715,776]" gridcol="6" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFA9069C2FA1EFD76" bold="true" box="[1317,1459,715,740]" pageId="18" pageNumber="419">SYS a004721</emphasis>
</th>
<th id="76D55DEBFFD4F6C3FA6869C2F9C6FC9A" box="[1501,1643,715,776]" gridcol="7" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFA6869C2F9C6FD76" bold="true" box="[1501,1643,715,740]" pageId="18" pageNumber="419">SYS a005447</emphasis>
</th>
<th id="76D55DEBFFD4F6C3F92169C2F88FFC9A" box="[1684,1826,715,776]" gridcol="8" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EF92169C2F88FFD76" bold="true" box="[1684,1826,715,740]" pageId="18" pageNumber="419">SYS a007406</emphasis>
</th>
<th id="76D55DEBFFD4F6C3F8F969C2F877FC9A" box="[1868,2010,715,776]" gridcol="9" gridrow="0" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EF8F969C2F877FD76" bold="true" box="[1868,2010,715,740]" pageId="18" pageNumber="419">SYS a007407</emphasis>
</th>
</tr>
<tr id="35043497FFD4F6C3FF11681EF877FCA2" box="[164,2010,791,816]" gridrow="1" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF11681EFEA3FCA2" box="[164,270,791,816]" gridcol="0" gridrow="1" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF11681EFF67FCA2" bold="true" box="[164,202,791,816]" pageId="18" pageNumber="419">Sex</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE681EFE58FCA2" box="[347,501,791,816]" gridcol="1" gridrow="1" pageId="18" pageNumber="419">Male</td>
<td id="76D55DEBFFD4F6C3FDA3681EFD09FCA2" box="[534,676,791,816]" gridcol="2" gridrow="1" pageId="18" pageNumber="419">Male</td>
<td id="76D55DEBFFD4F6C3FD78681EFCCDFCA2" box="[717,864,791,816]" gridcol="3" gridrow="1" pageId="18" pageNumber="419">Male</td>
<td id="76D55DEBFFD4F6C3FC03681EFBE9FCA2" box="[950,1092,791,816]" gridcol="4" gridrow="1" pageId="18" pageNumber="419">Male</td>
<td id="76D55DEBFFD4F6C3FBDB681EFB51FCA2" box="[1134,1276,791,816]" gridcol="5" gridrow="1" pageId="18" pageNumber="419">Female</td>
<td id="76D55DEBFFD4F6C3FA90681EFA1EFCA2" box="[1317,1459,791,816]" gridcol="6" gridrow="1" pageId="18" pageNumber="419">Female</td>
<td id="76D55DEBFFD4F6C3FA68681EF9C6FCA2" box="[1501,1643,791,816]" gridcol="7" gridrow="1" pageId="18" pageNumber="419">Female</td>
<td id="76D55DEBFFD4F6C3F921681EF88FFCA2" box="[1684,1826,791,816]" gridcol="8" gridrow="1" pageId="18" pageNumber="419">Female</td>
<td id="76D55DEBFFD4F6C3F8F9681EF877FCA2" box="[1868,2010,791,816]" gridcol="9" gridrow="1" pageId="18" pageNumber="419">Female</td>
</tr>
<tr id="35043497FFD4F6C3FF11684DF877FCCF" box="[164,2010,836,861]" gridrow="2" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF11684DFEA3FCCF" box="[164,270,836,861]" gridcol="0" gridrow="2" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF11684DFF78FCCF" bold="true" box="[164,213,836,861]" pageId="18" pageNumber="419">SVL</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE684DFE58FCCF" box="[347,501,836,861]" gridcol="1" gridrow="2" pageId="18" pageNumber="419">35.3</td>
<td id="76D55DEBFFD4F6C3FDA3684DFD09FCCF" box="[534,676,836,861]" gridcol="2" gridrow="2" pageId="18" pageNumber="419">33.4</td>
<td id="76D55DEBFFD4F6C3FD78684DFCCDFCCF" box="[717,864,836,861]" gridcol="3" gridrow="2" pageId="18" pageNumber="419">33.2</td>
<td id="76D55DEBFFD4F6C3FC03684DFBE9FCCF" box="[950,1092,836,861]" gridcol="4" gridrow="2" pageId="18" pageNumber="419">34.8</td>
<td id="76D55DEBFFD4F6C3FBDB684DFB51FCCF" box="[1134,1276,836,861]" gridcol="5" gridrow="2" pageId="18" pageNumber="419">45.8</td>
<td id="76D55DEBFFD4F6C3FA90684DFA1EFCCF" box="[1317,1459,836,861]" gridcol="6" gridrow="2" pageId="18" pageNumber="419">37.4</td>
<td id="76D55DEBFFD4F6C3FA68684DF9C6FCCF" box="[1501,1643,836,861]" gridcol="7" gridrow="2" pageId="18" pageNumber="419">39.9</td>
<td id="76D55DEBFFD4F6C3F921684DF88FFCCF" box="[1684,1826,836,861]" gridcol="8" gridrow="2" pageId="18" pageNumber="419">36.3</td>
<td id="76D55DEBFFD4F6C3F8F9684DF877FCCF" box="[1868,2010,836,861]" gridcol="9" gridrow="2" pageId="18" pageNumber="419">44.2</td>
</tr>
<tr id="35043497FFD4F6C3FF116878F877FC19" box="[164,2010,881,907]" gridrow="3" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116878FEA3FC19" box="[164,270,881,907]" gridcol="0" gridrow="3" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116878FF76FC18" bold="true" box="[164,219,881,906]" pageId="18" pageNumber="419">HDL</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6878FE58FC19" box="[347,501,881,907]" gridcol="1" gridrow="3" pageId="18" pageNumber="419">12.4</td>
<td id="76D55DEBFFD4F6C3FDA36878FD09FC19" box="[534,676,881,907]" gridcol="2" gridrow="3" pageId="18" pageNumber="419">11.3</td>
<td id="76D55DEBFFD4F6C3FD786878FCCDFC19" box="[717,864,881,907]" gridcol="3" gridrow="3" pageId="18" pageNumber="419">11.1</td>
<td id="76D55DEBFFD4F6C3FC036878FBE9FC19" box="[950,1092,881,907]" gridcol="4" gridrow="3" pageId="18" pageNumber="419">11.9</td>
<td id="76D55DEBFFD4F6C3FBDB6878FB51FC19" box="[1134,1276,881,907]" gridcol="5" gridrow="3" pageId="18" pageNumber="419">15.2</td>
<td id="76D55DEBFFD4F6C3FA906878FA1EFC19" box="[1317,1459,881,907]" gridcol="6" gridrow="3" pageId="18" pageNumber="419">13.3</td>
<td id="76D55DEBFFD4F6C3FA686878F9C6FC19" box="[1501,1643,881,907]" gridcol="7" gridrow="3" pageId="18" pageNumber="419">12.9</td>
<td id="76D55DEBFFD4F6C3F9216878F88FFC19" box="[1684,1826,881,907]" gridcol="8" gridrow="3" pageId="18" pageNumber="419">11.7</td>
<td id="76D55DEBFFD4F6C3F8F96878F877FC19" box="[1868,2010,881,907]" gridcol="9" gridrow="3" pageId="18" pageNumber="419">14.0</td>
</tr>
<tr id="35043497FFD4F6C3FF116896F877FC2A" box="[164,2010,927,952]" gridrow="4" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116896FEA3FC2A" box="[164,270,927,952]" gridcol="0" gridrow="4" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116896FF4EFC2A" bold="true" box="[164,227,927,952]" pageId="18" pageNumber="419">HDW</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6896FE58FC2A" box="[347,501,927,952]" gridcol="1" gridrow="4" pageId="18" pageNumber="419">13.3</td>
<td id="76D55DEBFFD4F6C3FDA36896FD09FC2A" box="[534,676,927,952]" gridcol="2" gridrow="4" pageId="18" pageNumber="419">11.6</td>
<td id="76D55DEBFFD4F6C3FD786896FCCDFC2A" box="[717,864,927,952]" gridcol="3" gridrow="4" pageId="18" pageNumber="419">11.9</td>
<td id="76D55DEBFFD4F6C3FC036896FBE9FC2A" box="[950,1092,927,952]" gridcol="4" gridrow="4" pageId="18" pageNumber="419">11.6</td>
<td id="76D55DEBFFD4F6C3FBDB6896FB51FC2A" box="[1134,1276,927,952]" gridcol="5" gridrow="4" pageId="18" pageNumber="419">15.8</td>
<td id="76D55DEBFFD4F6C3FA906896FA1EFC2A" box="[1317,1459,927,952]" gridcol="6" gridrow="4" pageId="18" pageNumber="419">14.2</td>
<td id="76D55DEBFFD4F6C3FA686896F9C6FC2A" box="[1501,1643,927,952]" gridcol="7" gridrow="4" pageId="18" pageNumber="419">13.1</td>
<td id="76D55DEBFFD4F6C3F9216896F88FFC2A" box="[1684,1826,927,952]" gridcol="8" gridrow="4" pageId="18" pageNumber="419">12.2</td>
<td id="76D55DEBFFD4F6C3F8F96896F877FC2A" box="[1868,2010,927,952]" gridcol="9" gridrow="4" pageId="18" pageNumber="419">14.5</td>
</tr>
<tr id="35043497FFD4F6C3FF1168C5F877FC74" box="[164,2010,972,998]" gridrow="5" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF1168C5FEA3FC74" box="[164,270,972,998]" gridcol="0" gridrow="5" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF1168C5FF78FC77" bold="true" box="[164,213,972,997]" pageId="18" pageNumber="419">SNT</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE68C5FE58FC74" box="[347,501,972,998]" gridcol="1" gridrow="5" pageId="18" pageNumber="419">4.1</td>
<td id="76D55DEBFFD4F6C3FDA368C5FD09FC74" box="[534,676,972,998]" gridcol="2" gridrow="5" pageId="18" pageNumber="419">3.8</td>
<td id="76D55DEBFFD4F6C3FD7868C5FCCDFC74" box="[717,864,972,998]" gridcol="3" gridrow="5" pageId="18" pageNumber="419">4.1</td>
<td id="76D55DEBFFD4F6C3FC0368C5FBE9FC74" box="[950,1092,972,998]" gridcol="4" gridrow="5" pageId="18" pageNumber="419">4.0</td>
<td id="76D55DEBFFD4F6C3FBDB68C5FB51FC74" box="[1134,1276,972,998]" gridcol="5" gridrow="5" pageId="18" pageNumber="419">4.7</td>
<td id="76D55DEBFFD4F6C3FA9068C5FA1EFC74" box="[1317,1459,972,998]" gridcol="6" gridrow="5" pageId="18" pageNumber="419">4.8</td>
<td id="76D55DEBFFD4F6C3FA6868C5F9C6FC74" box="[1501,1643,972,998]" gridcol="7" gridrow="5" pageId="18" pageNumber="419">4.4</td>
<td id="76D55DEBFFD4F6C3F92168C5F88FFC74" box="[1684,1826,972,998]" gridcol="8" gridrow="5" pageId="18" pageNumber="419">4.2</td>
<td id="76D55DEBFFD4F6C3F8F968C5F877FC74" box="[1868,2010,972,998]" gridcol="9" gridrow="5" pageId="18" pageNumber="419">4.8</td>
</tr>
<tr id="35043497FFD4F6C3FF1168F0F877FB81" box="[164,2010,1017,1043]" gridrow="6" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF1168F0FEA3FB81" box="[164,270,1017,1043]" gridcol="0" gridrow="6" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF1168F0FF7FFB80" bold="true" box="[164,210,1017,1042]" pageId="18" pageNumber="419">IND</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE68F0FE58FB81" box="[347,501,1017,1043]" gridcol="1" gridrow="6" pageId="18" pageNumber="419">4.1</td>
<td id="76D55DEBFFD4F6C3FDA368F0FD09FB81" box="[534,676,1017,1043]" gridcol="2" gridrow="6" pageId="18" pageNumber="419">3.7</td>
<td id="76D55DEBFFD4F6C3FD7868F0FCCDFB81" box="[717,864,1017,1043]" gridcol="3" gridrow="6" pageId="18" pageNumber="419">4.0</td>
<td id="76D55DEBFFD4F6C3FC0368F0FBE9FB81" box="[950,1092,1017,1043]" gridcol="4" gridrow="6" pageId="18" pageNumber="419">3.8</td>
<td id="76D55DEBFFD4F6C3FBDB68F0FB51FB81" box="[1134,1276,1017,1043]" gridcol="5" gridrow="6" pageId="18" pageNumber="419">4.3</td>
<td id="76D55DEBFFD4F6C3FA9068F0FA1EFB81" box="[1317,1459,1017,1043]" gridcol="6" gridrow="6" pageId="18" pageNumber="419">4.4</td>
<td id="76D55DEBFFD4F6C3FA6868F0F9C6FB81" box="[1501,1643,1017,1043]" gridcol="7" gridrow="6" pageId="18" pageNumber="419">4.3</td>
<td id="76D55DEBFFD4F6C3F92168F0F88FFB81" box="[1684,1826,1017,1043]" gridcol="8" gridrow="6" pageId="18" pageNumber="419">3.8</td>
<td id="76D55DEBFFD4F6C3F8F968F0F877FB81" box="[1868,2010,1017,1043]" gridcol="9" gridrow="6" pageId="18" pageNumber="419">4.2</td>
</tr>
<tr id="35043497FFD4F6C3FF116F2EF877FBD2" box="[164,2010,1063,1088]" gridrow="7" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116F2EFEA3FBD2" box="[164,270,1063,1088]" gridcol="0" gridrow="7" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116F2EFF79FBD2" bold="true" box="[164,212,1063,1088]" pageId="18" pageNumber="419">IOD</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6F2EFE58FBD2" box="[347,501,1063,1088]" gridcol="1" gridrow="7" pageId="18" pageNumber="419">3.6</td>
<td id="76D55DEBFFD4F6C3FDA36F2EFD09FBD2" box="[534,676,1063,1088]" gridcol="2" gridrow="7" pageId="18" pageNumber="419">3.0</td>
<td id="76D55DEBFFD4F6C3FD786F2EFCCDFBD2" box="[717,864,1063,1088]" gridcol="3" gridrow="7" pageId="18" pageNumber="419">3.3</td>
<td id="76D55DEBFFD4F6C3FC036F2EFBE9FBD2" box="[950,1092,1063,1088]" gridcol="4" gridrow="7" pageId="18" pageNumber="419">3.3</td>
<td id="76D55DEBFFD4F6C3FBDB6F2EFB51FBD2" box="[1134,1276,1063,1088]" gridcol="5" gridrow="7" pageId="18" pageNumber="419">4.0</td>
<td id="76D55DEBFFD4F6C3FA906F2EFA1EFBD2" box="[1317,1459,1063,1088]" gridcol="6" gridrow="7" pageId="18" pageNumber="419">3.5</td>
<td id="76D55DEBFFD4F6C3FA686F2EF9C6FBD2" box="[1501,1643,1063,1088]" gridcol="7" gridrow="7" pageId="18" pageNumber="419">3.8</td>
<td id="76D55DEBFFD4F6C3F9216F2EF88FFBD2" box="[1684,1826,1063,1088]" gridcol="8" gridrow="7" pageId="18" pageNumber="419">3.4</td>
<td id="76D55DEBFFD4F6C3F8F96F2EF877FBD2" box="[1868,2010,1063,1088]" gridcol="9" gridrow="7" pageId="18" pageNumber="419">3.9</td>
</tr>
<tr id="35043497FFD4F6C3FF116F5DF877FBFC" box="[164,2010,1108,1134]" gridrow="8" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116F5DFEA3FBFC" box="[164,270,1108,1134]" gridcol="0" gridrow="8" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116F5DFF6AFBFF" bold="true" box="[164,199,1108,1133]" pageId="18" pageNumber="419">ED</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6F5DFE58FBFC" box="[347,501,1108,1134]" gridcol="1" gridrow="8" pageId="18" pageNumber="419">3.7</td>
<td id="76D55DEBFFD4F6C3FDA36F5DFD09FBFC" box="[534,676,1108,1134]" gridcol="2" gridrow="8" pageId="18" pageNumber="419">3.9</td>
<td id="76D55DEBFFD4F6C3FD786F5DFCCDFBFC" box="[717,864,1108,1134]" gridcol="3" gridrow="8" pageId="18" pageNumber="419">3.9</td>
<td id="76D55DEBFFD4F6C3FC036F5DFBE9FBFC" box="[950,1092,1108,1134]" gridcol="4" gridrow="8" pageId="18" pageNumber="419">3.8</td>
<td id="76D55DEBFFD4F6C3FBDB6F5DFB51FBFC" box="[1134,1276,1108,1134]" gridcol="5" gridrow="8" pageId="18" pageNumber="419">4.9</td>
<td id="76D55DEBFFD4F6C3FA906F5DFA1EFBFC" box="[1317,1459,1108,1134]" gridcol="6" gridrow="8" pageId="18" pageNumber="419">4.6</td>
<td id="76D55DEBFFD4F6C3FA686F5DF9C6FBFC" box="[1501,1643,1108,1134]" gridcol="7" gridrow="8" pageId="18" pageNumber="419">3.7</td>
<td id="76D55DEBFFD4F6C3F9216F5DF88FFBFC" box="[1684,1826,1108,1134]" gridcol="8" gridrow="8" pageId="18" pageNumber="419">3.6</td>
<td id="76D55DEBFFD4F6C3F8F96F5DF877FBFC" box="[1868,2010,1108,1134]" gridcol="9" gridrow="8" pageId="18" pageNumber="419">4.2</td>
</tr>
<tr id="35043497FFD4F6C3FF116F8BF877FB09" box="[164,2010,1154,1179]" gridrow="9" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116F8BFEA3FB09" box="[164,270,1154,1179]" gridcol="0" gridrow="9" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116F8BFF6AFB09" bold="true" box="[164,199,1154,1179]" pageId="18" pageNumber="419">TD</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6F8BFE58FB09" box="[347,501,1154,1179]" gridcol="1" gridrow="9" pageId="18" pageNumber="419">2.2</td>
<td id="76D55DEBFFD4F6C3FDA36F8BFD09FB09" box="[534,676,1154,1179]" gridcol="2" gridrow="9" pageId="18" pageNumber="419">2.1</td>
<td id="76D55DEBFFD4F6C3FD786F8BFCCDFB09" box="[717,864,1154,1179]" gridcol="3" gridrow="9" pageId="18" pageNumber="419">2.0</td>
<td id="76D55DEBFFD4F6C3FC036F8BFBE9FB09" box="[950,1092,1154,1179]" gridcol="4" gridrow="9" pageId="18" pageNumber="419">2.3</td>
<td id="76D55DEBFFD4F6C3FBDB6F8BFB51FB09" box="[1134,1276,1154,1179]" gridcol="5" gridrow="9" pageId="18" pageNumber="419">3.0</td>
<td id="76D55DEBFFD4F6C3FA906F8BFA1EFB09" box="[1317,1459,1154,1179]" gridcol="6" gridrow="9" pageId="18" pageNumber="419">2.2</td>
<td id="76D55DEBFFD4F6C3FA686F8BF9C6FB09" box="[1501,1643,1154,1179]" gridcol="7" gridrow="9" pageId="18" pageNumber="419">2.5</td>
<td id="76D55DEBFFD4F6C3F9216F8BF88FFB09" box="[1684,1826,1154,1179]" gridcol="8" gridrow="9" pageId="18" pageNumber="419">2.0</td>
<td id="76D55DEBFFD4F6C3F8F96F8BF877FB09" box="[1868,2010,1154,1179]" gridcol="9" gridrow="9" pageId="18" pageNumber="419">2.7</td>
</tr>
<tr id="35043497FFD4F6C3FF116FA6F877FB5A" box="[164,2010,1199,1224]" gridrow="10" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116FA6FEA3FB5A" box="[164,270,1199,1224]" gridcol="0" gridrow="10" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116FA6FF75FB5A" bold="true" box="[164,216,1199,1224]" pageId="18" pageNumber="419">TED</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6FA6FE58FB5A" box="[347,501,1199,1224]" gridcol="1" gridrow="10" pageId="18" pageNumber="419">2.3</td>
<td id="76D55DEBFFD4F6C3FDA36FA6FD09FB5A" box="[534,676,1199,1224]" gridcol="2" gridrow="10" pageId="18" pageNumber="419">2.1</td>
<td id="76D55DEBFFD4F6C3FD786FA6FCCDFB5A" box="[717,864,1199,1224]" gridcol="3" gridrow="10" pageId="18" pageNumber="419">1.9</td>
<td id="76D55DEBFFD4F6C3FC036FA6FBE9FB5A" box="[950,1092,1199,1224]" gridcol="4" gridrow="10" pageId="18" pageNumber="419">2.2</td>
<td id="76D55DEBFFD4F6C3FBDB6FA6FB51FB5A" box="[1134,1276,1199,1224]" gridcol="5" gridrow="10" pageId="18" pageNumber="419">2.4</td>
<td id="76D55DEBFFD4F6C3FA906FA6FA1EFB5A" box="[1317,1459,1199,1224]" gridcol="6" gridrow="10" pageId="18" pageNumber="419">1.9</td>
<td id="76D55DEBFFD4F6C3FA686FA6F9C6FB5A" box="[1501,1643,1199,1224]" gridcol="7" gridrow="10" pageId="18" pageNumber="419">2.5</td>
<td id="76D55DEBFFD4F6C3F9216FA6F88FFB5A" box="[1684,1826,1199,1224]" gridcol="8" gridrow="10" pageId="18" pageNumber="419">2.1</td>
<td id="76D55DEBFFD4F6C3F8F96FA6F877FB5A" box="[1868,2010,1199,1224]" gridcol="9" gridrow="10" pageId="18" pageNumber="419">2.6</td>
</tr>
<tr id="35043497FFD4F6C3FF116FD5F877FB64" box="[164,2010,1244,1270]" gridrow="11" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116FD5FEA3FB64" box="[164,270,1244,1270]" gridcol="0" gridrow="11" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116FD5FF71FB67" bold="true" box="[164,220,1244,1269]" pageId="18" pageNumber="419">HND</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6FD5FE58FB64" box="[347,501,1244,1270]" gridcol="1" gridrow="11" pageId="18" pageNumber="419">9.2</td>
<td id="76D55DEBFFD4F6C3FDA36FD5FD09FB64" box="[534,676,1244,1270]" gridcol="2" gridrow="11" pageId="18" pageNumber="419">7.2</td>
<td id="76D55DEBFFD4F6C3FD786FD5FCCDFB64" box="[717,864,1244,1270]" gridcol="3" gridrow="11" pageId="18" pageNumber="419">8.3</td>
<td id="76D55DEBFFD4F6C3FC036FD5FBE9FB64" box="[950,1092,1244,1270]" gridcol="4" gridrow="11" pageId="18" pageNumber="419">8.7</td>
<td id="76D55DEBFFD4F6C3FBDB6FD5FB51FB64" box="[1134,1276,1244,1270]" gridcol="5" gridrow="11" pageId="18" pageNumber="419">11.3</td>
<td id="76D55DEBFFD4F6C3FA906FD5FA1EFB64" box="[1317,1459,1244,1270]" gridcol="6" gridrow="11" pageId="18" pageNumber="419">9.0</td>
<td id="76D55DEBFFD4F6C3FA686FD5F9C6FB64" box="[1501,1643,1244,1270]" gridcol="7" gridrow="11" pageId="18" pageNumber="419">9.6</td>
<td id="76D55DEBFFD4F6C3F9216FD5F88FFB64" box="[1684,1826,1244,1270]" gridcol="8" gridrow="11" pageId="18" pageNumber="419">9.1</td>
<td id="76D55DEBFFD4F6C3F8F96FD5F877FB64" box="[1868,2010,1244,1270]" gridcol="9" gridrow="11" pageId="18" pageNumber="419">10.2</td>
</tr>
<tr id="35043497FFD4F6C3FF116E03F877FAB1" box="[164,2010,1290,1315]" gridrow="12" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116E03FEA3FAB1" box="[164,270,1290,1315]" gridcol="0" gridrow="12" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116E03FF76FAB1" bold="true" box="[164,219,1290,1315]" pageId="18" pageNumber="419">RAD</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6E03FE58FAB1" box="[347,501,1290,1315]" gridcol="1" gridrow="12" pageId="18" pageNumber="419">6.9</td>
<td id="76D55DEBFFD4F6C3FDA36E03FD09FAB1" box="[534,676,1290,1315]" gridcol="2" gridrow="12" pageId="18" pageNumber="419">7.1</td>
<td id="76D55DEBFFD4F6C3FD786E03FCCDFAB1" box="[717,864,1290,1315]" gridcol="3" gridrow="12" pageId="18" pageNumber="419">7.4</td>
<td id="76D55DEBFFD4F6C3FC036E03FBE9FAB1" box="[950,1092,1290,1315]" gridcol="4" gridrow="12" pageId="18" pageNumber="419">6.8</td>
<td id="76D55DEBFFD4F6C3FBDB6E03FB51FAB1" box="[1134,1276,1290,1315]" gridcol="5" gridrow="12" pageId="18" pageNumber="419">9.2</td>
<td id="76D55DEBFFD4F6C3FA906E03FA1EFAB1" box="[1317,1459,1290,1315]" gridcol="6" gridrow="12" pageId="18" pageNumber="419">8.4</td>
<td id="76D55DEBFFD4F6C3FA686E03F9C6FAB1" box="[1501,1643,1290,1315]" gridcol="7" gridrow="12" pageId="18" pageNumber="419">7.7</td>
<td id="76D55DEBFFD4F6C3F9216E03F88FFAB1" box="[1684,1826,1290,1315]" gridcol="8" gridrow="12" pageId="18" pageNumber="419">7.8</td>
<td id="76D55DEBFFD4F6C3F8F96E03F877FAB1" box="[1868,2010,1290,1315]" gridcol="9" gridrow="12" pageId="18" pageNumber="419">8.9</td>
</tr>
<tr id="35043497FFD4F6C3FF116E3EF877FAC2" box="[164,2010,1335,1360]" gridrow="13" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116E3EFEA3FAC2" box="[164,270,1335,1360]" gridcol="0" gridrow="13" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116E3EFF7DFAC2" bold="true" box="[164,208,1335,1360]" pageId="18" pageNumber="419">TIB</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6E3EFE58FAC2" box="[347,501,1335,1360]" gridcol="1" gridrow="13" pageId="18" pageNumber="419">16.8</td>
<td id="76D55DEBFFD4F6C3FDA36E3EFD09FAC2" box="[534,676,1335,1360]" gridcol="2" gridrow="13" pageId="18" pageNumber="419">15.2</td>
<td id="76D55DEBFFD4F6C3FD786E3EFCCDFAC2" box="[717,864,1335,1360]" gridcol="3" gridrow="13" pageId="18" pageNumber="419">15.9</td>
<td id="76D55DEBFFD4F6C3FC036E3EFBE9FAC2" box="[950,1092,1335,1360]" gridcol="4" gridrow="13" pageId="18" pageNumber="419">16.2</td>
<td id="76D55DEBFFD4F6C3FBDB6E3EFB51FAC2" box="[1134,1276,1335,1360]" gridcol="5" gridrow="13" pageId="18" pageNumber="419">20.9</td>
<td id="76D55DEBFFD4F6C3FA906E3EFA1EFAC2" box="[1317,1459,1335,1360]" gridcol="6" gridrow="13" pageId="18" pageNumber="419">16.9</td>
<td id="76D55DEBFFD4F6C3FA686E3EF9C6FAC2" box="[1501,1643,1335,1360]" gridcol="7" gridrow="13" pageId="18" pageNumber="419">17.5</td>
<td id="76D55DEBFFD4F6C3F9216E3EF88FFAC2" box="[1684,1826,1335,1360]" gridcol="8" gridrow="13" pageId="18" pageNumber="419">16.6</td>
<td id="76D55DEBFFD4F6C3F8F96E3EF877FAC2" box="[1868,2010,1335,1360]" gridcol="9" gridrow="13" pageId="18" pageNumber="419">19.6</td>
</tr>
<tr id="35043497FFD4F6C3FF116E6DF877FAEC" box="[164,2010,1380,1406]" gridrow="14" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116E6DFEA3FAEC" box="[164,270,1380,1406]" gridcol="0" gridrow="14" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116E6DFF78FAEF" bold="true" box="[164,213,1380,1405]" pageId="18" pageNumber="419">FTL</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6E6DFE58FAEC" box="[347,501,1380,1406]" gridcol="1" gridrow="14" pageId="18" pageNumber="419">23.8</td>
<td id="76D55DEBFFD4F6C3FDA36E6DFD09FAEC" box="[534,676,1380,1406]" gridcol="2" gridrow="14" pageId="18" pageNumber="419">20.3</td>
<td id="76D55DEBFFD4F6C3FD786E6DFCCDFAEC" box="[717,864,1380,1406]" gridcol="3" gridrow="14" pageId="18" pageNumber="419">21.9</td>
<td id="76D55DEBFFD4F6C3FC036E6DFBE9FAEC" box="[950,1092,1380,1406]" gridcol="4" gridrow="14" pageId="18" pageNumber="419">22.1</td>
<td id="76D55DEBFFD4F6C3FBDB6E6DFB51FAEC" box="[1134,1276,1380,1406]" gridcol="5" gridrow="14" pageId="18" pageNumber="419">30.1</td>
<td id="76D55DEBFFD4F6C3FA906E6DFA1EFAEC" box="[1317,1459,1380,1406]" gridcol="6" gridrow="14" pageId="18" pageNumber="419">22.9</td>
<td id="76D55DEBFFD4F6C3FA686E6DF9C6FAEC" box="[1501,1643,1380,1406]" gridcol="7" gridrow="14" pageId="18" pageNumber="419">25.5</td>
<td id="76D55DEBFFD4F6C3F9216E6DF88FFAEC" box="[1684,1826,1380,1406]" gridcol="8" gridrow="14" pageId="18" pageNumber="419">24.1</td>
<td id="76D55DEBFFD4F6C3F8F96E6DF877FAEC" box="[1868,2010,1380,1406]" gridcol="9" gridrow="14" pageId="18" pageNumber="419">27.6</td>
</tr>
<tr id="35043497FFD4F6C3FF116E9BF877FA39" box="[164,2010,1426,1451]" gridrow="15" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116E9BFEA3FA39" box="[164,270,1426,1451]" gridcol="0" gridrow="15" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116E9BFF5CFA39" bold="true" box="[164,241,1426,1451]" pageId="18" pageNumber="419">ED/TD</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6E9BFE58FA39" box="[347,501,1426,1451]" gridcol="1" gridrow="15" pageId="18" pageNumber="419">1.68</td>
<td id="76D55DEBFFD4F6C3FDA36E9BFD09FA39" box="[534,676,1426,1451]" gridcol="2" gridrow="15" pageId="18" pageNumber="419">1.86</td>
<td id="76D55DEBFFD4F6C3FD786E9BFCCDFA39" box="[717,864,1426,1451]" gridcol="3" gridrow="15" pageId="18" pageNumber="419">1.95</td>
<td id="76D55DEBFFD4F6C3FC036E9BFBE9FA39" box="[950,1092,1426,1451]" gridcol="4" gridrow="15" pageId="18" pageNumber="419">1.65</td>
<td id="76D55DEBFFD4F6C3FBDB6E9BFB51FA39" box="[1134,1276,1426,1451]" gridcol="5" gridrow="15" pageId="18" pageNumber="419">1.63</td>
<td id="76D55DEBFFD4F6C3FA906E9BFA1EFA39" box="[1317,1459,1426,1451]" gridcol="6" gridrow="15" pageId="18" pageNumber="419">2.09</td>
<td id="76D55DEBFFD4F6C3FA686E9BF9C6FA39" box="[1501,1643,1426,1451]" gridcol="7" gridrow="15" pageId="18" pageNumber="419">1.48</td>
<td id="76D55DEBFFD4F6C3F9216E9BF88FFA39" box="[1684,1826,1426,1451]" gridcol="8" gridrow="15" pageId="18" pageNumber="419">1.80</td>
<td id="76D55DEBFFD4F6C3F8F96E9BF877FA39" box="[1868,2010,1426,1451]" gridcol="9" gridrow="15" pageId="18" pageNumber="419">1.56</td>
</tr>
<tr id="35043497FFD4F6C3FF116EB6F877FA4A" box="[164,2010,1471,1496]" gridrow="16" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116EB6FEA3FA4A" box="[164,270,1471,1496]" gridcol="0" gridrow="16" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116EB6FEA5FA4A" bold="true" box="[164,264,1471,1496]" pageId="18" pageNumber="419">TIB/SVL</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6EB6FE58FA4A" box="[347,501,1471,1496]" gridcol="1" gridrow="16" pageId="18" pageNumber="419">0.48</td>
<td id="76D55DEBFFD4F6C3FDA36EB6FD09FA4A" box="[534,676,1471,1496]" gridcol="2" gridrow="16" pageId="18" pageNumber="419">0.46</td>
<td id="76D55DEBFFD4F6C3FD786EB6FCCDFA4A" box="[717,864,1471,1496]" gridcol="3" gridrow="16" pageId="18" pageNumber="419">0.48</td>
<td id="76D55DEBFFD4F6C3FC036EB6FBE9FA4A" box="[950,1092,1471,1496]" gridcol="4" gridrow="16" pageId="18" pageNumber="419">0.47</td>
<td id="76D55DEBFFD4F6C3FBDB6EB6FB51FA4A" box="[1134,1276,1471,1496]" gridcol="5" gridrow="16" pageId="18" pageNumber="419">0.46</td>
<td id="76D55DEBFFD4F6C3FA906EB6FA1EFA4A" box="[1317,1459,1471,1496]" gridcol="6" gridrow="16" pageId="18" pageNumber="419">0.45</td>
<td id="76D55DEBFFD4F6C3FA686EB6F9C6FA4A" box="[1501,1643,1471,1496]" gridcol="7" gridrow="16" pageId="18" pageNumber="419">0.44</td>
<td id="76D55DEBFFD4F6C3F9216EB6F88FFA4A" box="[1684,1826,1471,1496]" gridcol="8" gridrow="16" pageId="18" pageNumber="419">0.46</td>
<td id="76D55DEBFFD4F6C3F8F96EB6F877FA4A" box="[1868,2010,1471,1496]" gridcol="9" gridrow="16" pageId="18" pageNumber="419">0.44</td>
</tr>
<tr id="35043497FFD4F6C3FF116EE5F877F994" box="[164,2010,1516,1542]" gridrow="17" pageId="18" pageNumber="419">
<th id="76D55DEBFFD4F6C3FF116EE5FEA3F994" box="[164,270,1516,1542]" gridcol="0" gridrow="17" pageId="18" pageNumber="419">
<emphasis id="B940EAC7FFD4092EFF116EE5FEA3F997" bold="true" box="[164,270,1516,1541]" pageId="18" pageNumber="419">FTL/SVL</emphasis>
</th>
<td id="76D55DEBFFD4F6C3FEEE6EE5FE58F994" box="[347,501,1516,1542]" gridcol="1" gridrow="17" pageId="18" pageNumber="419">0.67</td>
<td id="76D55DEBFFD4F6C3FDA36EE5FD09F994" box="[534,676,1516,1542]" gridcol="2" gridrow="17" pageId="18" pageNumber="419">0.61</td>
<td id="76D55DEBFFD4F6C3FD786EE5FCCDF994" box="[717,864,1516,1542]" gridcol="3" gridrow="17" pageId="18" pageNumber="419">0.66</td>
<td id="76D55DEBFFD4F6C3FC036EE5FBE9F994" box="[950,1092,1516,1542]" gridcol="4" gridrow="17" pageId="18" pageNumber="419">0.64</td>
<td id="76D55DEBFFD4F6C3FBDB6EE5FB51F994" box="[1134,1276,1516,1542]" gridcol="5" gridrow="17" pageId="18" pageNumber="419">0.66</td>
<td id="76D55DEBFFD4F6C3FA906EE5FA1EF994" box="[1317,1459,1516,1542]" gridcol="6" gridrow="17" pageId="18" pageNumber="419">0.61</td>
<td id="76D55DEBFFD4F6C3FA686EE5F9C6F994" box="[1501,1643,1516,1542]" gridcol="7" gridrow="17" pageId="18" pageNumber="419">0.64</td>
<td id="76D55DEBFFD4F6C3F9216EE5F88FF994" box="[1684,1826,1516,1542]" gridcol="8" gridrow="17" pageId="18" pageNumber="419">0.66</td>
<td id="76D55DEBFFD4F6C3F8F96EE5F877F994" box="[1868,2010,1516,1542]" gridcol="9" gridrow="17" pageId="18" pageNumber="419">0.62</td>
</tr>
</table>
</paragraph>
<caption id="DF4B665DFFD5092FFF226F13FE50FBC4" ID-DOI="http://doi.org/10.5281/zenodo.5748989" ID-Zenodo-Dep="5748989" httpUri="https://zenodo.org/record/5748989/files/figure.png" pageId="19" pageNumber="420" startId="19.[151,250,1050,1074]" targetBox="[158,1430,182,1026]" targetPageId="19">
<paragraph id="8B8B36D5FFD5092FFF226F13FE50FBC4" blockId="19.[151,1437,1050,1110]" pageId="19" pageNumber="420">
<emphasis id="B940EAC7FFD5092FFF226F13FEBBFBA0" bold="true" box="[151,278,1050,1074]" pageId="19" pageNumber="420">FIGURE 5.</emphasis>
Color variation of
<taxonomicName id="4C344D56FFD5092FFE566F13FD42FBA0" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[483,751,1050,1074]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD5092FFE566F13FD42FBA0" box="[483,751,1050,1074]" italics="true" pageId="19" pageNumber="420">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD5092FFD4D6F13FCE6FBA0" bold="true" box="[760,843,1050,1074]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A27357BCFFD5092FFD4D6F13FCE6FBA0" box="[760,843,1050,1074]" pageId="19" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the type locality. These individuals were captured, photographed, and finally released.
</paragraph>
</caption>
<paragraph id="8B8B36D5FFD5092FFF726F8DFB69FAC0" blockId="19.[151,1437,1156,2011]" pageId="19" pageNumber="420">Skin of dorsum bearing small granules; skin of flanks smooth with small conical tubercles; a small, horn-like prominent tubercle on the edge of upper eyelid; clear supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; a discontinuous “X” shaped ridge on the mid-dorsum and two discontinuous dorsolateral ridges on two sides at the mid-back; sparse tubercles on the dorsal shank and thigh; ventral surface smooth; several tubercles on posterior hindlimbs; small pectoral gland closer to axilla; a single femoral positioned subequally distant from knees and cloaca on posterior surface of each thigh.</paragraph>
<paragraph id="8B8B36D5FFD5092FFF726E55FB14F9E0" blockId="19.[151,1437,1156,2011]" pageId="19" pageNumber="420">
<emphasis id="B940EAC7FFD5092FFF726E55FD88FAE4" bold="true" box="[199,549,1372,1398]" pageId="19" pageNumber="420">
Coloration of
<typeStatus id="548F8877FFD5092FFED86E55FE79FAE4" box="[365,468,1372,1398]" pageId="19" pageNumber="420" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsal surface of body olive-brown with an inverted brown triangular marking between eyes; an “X” shaped marking on the mid-dorsum. Forearms and hindlimbs with brown transverse bands. Supratympanic fold with a continuous white line; a dark vertical band below the eye, from the inferior margin of the eye to the upper lip. Ventral surface of throat and chest grayish brown with brown patches, an indistinct longitudinal stripe on surface of throat; a pair of dark brown longitudinal stripes scattered on surface of lateroventral flanks; ventral surface of body white with brown patches; ventral surface of limbs grayish brown with dark brown spots and blotches; ventral surfaces of hands and ventral surfaces of feet brown, tips of digits pale-brown; metacarpal tubercle and metatarsal tubercle reddish. Pectoral gland and femoral gland white. Iris yellowish brown.
</paragraph>
<paragraph id="8B8B36D5FFD5092FFF726D75FB65F94C" blockId="19.[151,1437,1156,2011]" pageId="19" pageNumber="420">
<emphasis id="B940EAC7FFD5092FFF726D75FD32F904" bold="true" box="[199,671,1660,1686]" pageId="19" pageNumber="420">
Coloration of
<typeStatus id="548F8877FFD5092FFEC66D75FE77F904" box="[371,474,1660,1686]" pageId="19" pageNumber="420" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Olive-brown faded to greyish brown dorsally. Triangular marking between eyes, “X” shaped marking on the mid-dorsum, transverse bands on dorsal forearms and hind limbs became indistinct. Color of ventral surface faded to greyish white all bands and spots became indistinct.
</paragraph>
<paragraph id="8B8B36D5FFD5092FFF726DE1FD0EF8FD" blockId="19.[151,1437,1156,2011]" pageId="19" pageNumber="420">
<emphasis id="B940EAC7FFD5092FFF726DE1FDC7F890" bold="true" box="[199,618,1768,1794]" pageId="19" pageNumber="420">Variation and sexual dimorphism.</emphasis>
<materialsCitation id="3B5C3C88FFD5092FFDC16DE0FD32F8FD" ID-GBIF-Occurrence="3413047314" collectionCode="SVL" pageId="19" pageNumber="420" specimenCount="1">
Measurement data of
<typeStatus id="548F8877FFD5092FFCC16DE0FC09F891" box="[884,932,1769,1795]" pageId="19" pageNumber="420">type</typeStatus>
series are listed in
<tableCitation id="C6B6036EFFD5092FFB3B6DE1FB4AF891" box="[1166,1255,1768,1795]" captionStart="TABLE 5" captionStartId="18.[150,243,673,700]" captionTargetPageId="18" captionText="TABLE 5. Measurements (in mm) and ratios of the type series of Boulenophrys yingdeensis sp. nov., * indicates the holotype." httpUri="http://table.plazi.org/id/DF4B665DFFD4092EFF2369A8F98FFD2E" pageId="19" pageNumber="420" tableUuid="DF4B665DFFD4092EFF2369A8F98FFD2E">Table 5</tableCitation>
. Females (
<collectionCode id="ED25AE10FFD5092FFAD26DE1FA30F890" box="[1383,1437,1768,1794]" pageId="19" pageNumber="420">SVL</collectionCode>
<quantity id="4CCC9B30FFD5092FFF226C05FE96F8B5" box="[151,315,1804,1831]" metricMagnitude="-2" metricUnit="m" metricValue="4.1049999999999995" metricValueMax="4.58" metricValueMin="3.63" pageId="19" pageNumber="420" unit="mm" value="41.05" valueMax="45.8" valueMin="36.3">36.345.8 mm</quantity>
) are larger than males (
<collectionCode id="ED25AE10FFD5092FFDF56C05FDDBF8B4" box="[576,630,1804,1830]" pageId="19" pageNumber="420">SVL</collectionCode>
<quantity id="4CCC9B30FFD5092FFDC96C05FC8EF8B5" box="[636,803,1804,1831]" metricMagnitude="-2" metricUnit="m" metricValue="3.425" metricValueMax="3.53" metricValueMin="3.32" pageId="19" pageNumber="420" unit="mm" value="34.25" valueMax="35.3" valueMin="33.2">33.235.3 mm</quantity>
). Presence of villiform black nuptial spines on the dorsal surface of the first and second fingers in adult males. Body coloration is quite variable, light brown, reddish brown, olive brown or dark brown dorsally (
<figureCitation id="130F2A50FFD5092FFD866C5DFD38F8FD" box="[563,661,1876,1903]" captionStart="FIGURE 5" captionStartId="19.[151,250,1050,1074]" captionTargetBox="[158,1430,182,1026]" captionTargetId="figure-18@19.[157,1430,181,1026]" captionTargetPageId="19" captionText="FIGURE 5. Color variation of Boulenophrys yingdeensis sp. nov. from the type locality. These individuals were captured, photographed, and finally released." figureDoi="http://doi.org/10.5281/zenodo.5748989" httpUri="https://zenodo.org/record/5748989/files/figure.png" pageId="19" pageNumber="420">Figure 5</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E655EFFD50928FF726C71FD29FF47" lastPageId="20" lastPageNumber="421" pageId="19" pageNumber="420" type="distribution">
<paragraph id="8B8B36D5FFD50928FF726C71FD29FF47" blockId="19.[151,1437,1156,2011]" lastBlockId="20.[151,1436,151,213]" lastPageId="20" lastPageNumber="421" pageId="19" pageNumber="420">
<emphasis id="B940EAC7FFD5092FFF726C71FE5CF800" bold="true" box="[199,497,1912,1938]" pageId="19" pageNumber="420">Distribution and ecology.</emphasis>
Currently,
<taxonomicName id="4C344D56FFD5092FFDDB6C71FC3EF800" authority="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang, 2021" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" box="[622,915,1912,1938]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="yingdeensis" status="sp. nov.">
<emphasis id="B940EAC7FFD5092FFDDB6C71FC3EF800" box="[622,915,1912,1938]" italics="true" pageId="19" pageNumber="420">Boulenophrys yingdeensis</emphasis>
</taxonomicName>
<emphasis id="B940EAC7FFD5092FFC2C6C71FC5FF800" bold="true" box="[921,1010,1912,1938]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A27357BCFFD5092FFC2C6C71FC5FF800" box="[921,1010,1912,1938]" pageId="19" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from Shimentai Nature Reserve,
<collectingRegion id="49F0F837FFD5092FFF4B6C95FE29F824" box="[254,388,1948,1974]" country="China" name="Guangdong" pageId="19" pageNumber="420">Guangdong</collectingRegion>
,
<collectingCountry id="F3237645FFD5092FFE3B6C95FE79F824" box="[398,468,1948,1974]" name="China" pageId="19" pageNumber="420">China</collectingCountry>
, and sympatric with
<taxonomicName id="4C344D56FFD5092FFD0C6C94FCCDF824" authorityName="Qi &amp; Lyu &amp; Wang &amp; Mo &amp; Zeng &amp; Zeng &amp; Dai &amp; Li &amp; Grismer &amp; Wang" authorityYear="2021" baseAuthorityName="Lyu, Liu, &amp; Wang" baseAuthorityYear="2020" box="[697,864,1948,1974]" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" pageId="19" pageNumber="420" phylum="Chordata" rank="species" species="shimentaina">
<emphasis id="B940EAC7FFD5092FFD0C6C94FCCDF824" box="[697,864,1948,1974]" italics="true" pageId="19" pageNumber="420">B. shimentaina</emphasis>
</taxonomicName>
. All individuals were found in evergreen secondary forest, near lowland streams and nearbyleaf litter at elevations between
<quantity id="4CCC9B30FFD5092FFC746CC9FBEEF849" box="[961,1091,1984,2011]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="19" pageNumber="420" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
300
<specimenCount id="9D32FD5CFFD5092FFC4D6CC9FBEEF849" box="[1016,1091,1984,2011]" pageId="19" pageNumber="420" type="generic">400 m</specimenCount>
</quantity>
. Males perch and call on plant leaves from April to June, suggesting their breeding season corresponds to this period. Females were found on the forest floor and tadpoles were not observed.
</paragraph>
</subSubSection>
</treatment>
</document>