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<mods:title id="9816DEADD8AAB808EBD84141BD31F627">A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
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<heading id="D0C381BA186AFFDCFF01F9E3FD7B4E5C" bold="true" box="[151,759,1575,1599]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="483" reason="2">
<taxonomicName id="4C344D55186AFFDCFF01F9E3FD7B4E5C" ID-CoL="4DJZ9" authority="(Bosenberg &amp; Strand, 1906)" baseAuthorityName="Bosenberg &amp; Strand" baseAuthorityYear="1906" box="[151,759,1575,1599]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="pseudoannulata">
<emphasis id="B940EAC4186AFFDCFF01F9E3FD7B4E5C" bold="true" box="[151,759,1575,1599]" pageId="4" pageNumber="483">
<emphasis id="B940EAC4186AFFDCFF01F9E3FE264E5C" bold="true" box="[151,426,1575,1599]" italics="true" pageId="4" pageNumber="483">Pardosa pseudoannulata</emphasis>
(
<bibRefCitation id="EFA54B27186AFFDCFE2CF9E3FD7D4E5C" author="Bosenberg W &amp; Strand E" box="[442,753,1575,1599]" pageId="4" pageNumber="483" pagination="93 - 422" refId="ref4602" refString="Bosenberg W &amp; Strand E (1906) Japanische Spinnen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 30: 93 - 422." type="journal article" year="1906">Bösenberg &amp; Strand, 1906</bibRefCitation>
)
</emphasis>
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<paragraph id="8B8B36D6186AFFDCFEFCF983FDA94E03" blockId="4.[151,759,1575,1632]" box="[362,549,1607,1632]" pageId="4" pageNumber="483">
(
<figureCitation id="130F2A53186AFFDCFEE7F983FE794E03" box="[369,501,1607,1632]" captionStart="Figs" captionStartId="6.[135,179,2035,2057]" captionTargetBox="[169,1435,1042,2018]" captionTargetId="figure-59@6.[167,1440,1042,2023]" captionTargetPageId="6" captionText="Figs. 3641. Copulatory organs of Passiena torbjoerni (36), Pardosa pseudoannulata (3739); epigyne and habitus of P. pusiola (4041). 36, cymbium with bulb removed, ventral; 37, palp, ventral; 38, macerated epigyne; 39, endogyne, 40, drawing of epigyne (syntype); 41, living specimen from Singapore (photo taken by David Court). Scale = 0.25 mm (unless otherwise specified)." figureDoi="http://doi.org/10.5281/zenodo.5344578" httpUri="https://zenodo.org/record/5344578/files/figure.png" pageId="4" pageNumber="483">Figs. 3739</figureCitation>
,
<figureCitation id="130F2A53186AFFDCFD94F98CFD924E03" box="[514,542,1608,1632]" captionStart="Fig" captionStartId="7.[113,148,1339,1361]" captionTargetBox="[115,1452,189,1324]" captionTargetId="figure-17@7.[101,1462,181,1333]" captionTargetPageId="7" captionText="Fig. 42. Map of collecting localities." figureDoi="http://doi.org/10.5281/zenodo.5344580" httpUri="https://zenodo.org/record/5344580/files/figure.png" pageId="4" pageNumber="483">42</figureCitation>
)
</paragraph>
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<subSubSection id="C32E655D186AFFDCFF1EF943FF5A4FA0" pageId="4" pageNumber="483" type="reference_group">
<paragraph id="8B8B36D6186AFFDCFF1EF943FF7A4EDA" blockId="4.[136,775,1670,1894]" pageId="4" pageNumber="483">
<treatmentCitationGroup id="AB2411F8186AFFDCFF1EF943FF7A4EDA" pageId="4" pageNumber="483">
<treatmentCitation id="0A9510C7186AFFDCFF1EF943FD874EFF" author="Tanaka H" box="[136,523,1670,1692]" page="245" pageId="4" pageNumber="483" year="2009">
<taxonomicName id="4C344D55186AFFDCFF1EF943FD874EFF" ID-CoL="4DJZ9" authority=": Tanaka, 2009: 245" authorityName="Tanaka" authorityPageNumber="245" authorityYear="2009" box="[136,523,1670,1692]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="pseudoannulata">
<emphasis id="B940EAC4186AFFDCFF1EF943FECC4EF8" box="[136,320,1670,1692]" italics="true" pageId="4" pageNumber="483">P. pseudoannulata</emphasis>
:
<bibRefCitation id="EFA54B27186AFFDCFED9F942FD874EFF" author="Tanaka H" box="[335,523,1670,1692]" pageId="4" pageNumber="483" refId="ref5027" refString="Tanaka H (2009) Lycosidae. In: Ono H (ed.) The spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa. Pp. 222 - 248." type="book" year="2009">Tanaka, 2009: 245</bibRefCitation>
</taxonomicName>
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, figs. 127128, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36D6186AFFDCFF1EF904FEAD4E90" blockId="4.[136,775,1670,1894]" pageId="4" pageNumber="483">
<treatmentCitationGroup id="AB2411F8186AFFDCFF1EF904FEAD4E90" pageId="4" pageNumber="483">
<treatmentCitation id="0A9510C7186AFFDCFF1EF904FDC94EB6" author="Zhu MS &amp; Zhang BS" box="[136,581,1728,1750]" page="279" pageId="4" pageNumber="483" year="2011">
<taxonomicName id="4C344D55186AFFDCFF1EF904FDC94EB6" ID-CoL="4DJZ9" authority=": Zhu &amp; Zhang, 2011: 279" authorityName="Zhu &amp; Zhang" authorityPageNumber="279" authorityYear="2011" box="[136,581,1728,1750]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="pseudoannulata">
<emphasis id="B940EAC4186AFFDCFF1EF904FEB24EB6" box="[136,318,1728,1749]" italics="true" pageId="4" pageNumber="483">P. pseudoannulata</emphasis>
:
<bibRefCitation id="EFA54B27186AFFDCFEDAF904FDC94EB6" author="Zhu MS &amp; Zhang BS" box="[332,581,1728,1750]" pageId="4" pageNumber="483" refId="ref5426" refString="Zhu MS &amp; Zhang BS (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, xxii + 558 pp." type="book" year="2011">Zhu &amp; Zhang, 2011: 279</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 202AD, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36D6186AFFDCFF1EF93EFEDB4F4F" blockId="4.[136,775,1670,1894]" pageId="4" pageNumber="483">
<treatmentCitationGroup id="AB2411F8186AFFDCFF1EF93EFEDB4F4F" pageId="4" pageNumber="483">
<treatmentCitation id="0A9510C7186AFFDCFF1EF93EFD974F6C" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" box="[136,539,1785,1807]" page="852" pageId="4" pageNumber="483" year="2012">
<taxonomicName id="4C344D55186AFFDCFF1EF93EFD974F6C" ID-CoL="4DJZ9" authority=": Yin et al., 2012: 852" authorityName="Yin" authorityPageNumber="852" authorityYear="2012" box="[136,539,1785,1807]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="pseudoannulata">
<emphasis id="B940EAC4186AFFDCFF1EF93EFEB24F6D" box="[136,318,1785,1807]" italics="true" pageId="4" pageNumber="483">P. pseudoannulata</emphasis>
:
<bibRefCitation id="EFA54B27186AFFDCFEDAF93DFD974F6C" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" box="[332,539,1785,1807]" pageId="4" pageNumber="483" refId="ref5347" refString="Yin CM, Peng XJ, Yan HM, Bao YH, Xu X, Tang G, Zhou QS &amp; Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">Yin et al., 2012: 852</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 427ah, 316c, d, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36D6186AFFDCFF1EF8F0FEAD4F05" blockId="4.[136,775,1670,1894]" pageId="4" pageNumber="483">
<treatmentCitationGroup id="AB2411F8186AFFDCFF1EF8F0FEAD4F05" pageId="4" pageNumber="483">
<treatmentCitation id="0A9510C7186AFFDCFF1EF8F0FDF74F2B" author="Baba YG &amp; Tanikawa A" box="[136,635,1843,1865]" page="74" pageId="4" pageNumber="483" year="2015">
<taxonomicName id="4C344D55186AFFDCFF1EF8F0FDF74F2B" ID-CoL="4DJZ9" authority=": Baba &amp; Tanikawa, 2015: 74" authorityName=": Baba &amp; Tanikawa" authorityPageNumber="74" authorityYear="2015" box="[136,635,1843,1865]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="pseudoannulata">
<emphasis id="B940EAC4186AFFDCFF1EF8F0FEC94F2B" box="[136,325,1843,1865]" italics="true" pageId="4" pageNumber="483">P. pseudoannulata</emphasis>
:
<bibRefCitation id="EFA54B27186AFFDCFEC3F8F7FDF74F2B" author="Baba YG &amp; Tanikawa A" box="[341,635,1843,1865]" pageId="4" pageNumber="483" refId="ref4580" refString="Baba YG &amp; Tanikawa A (2015) The handbook of spiders. Bun-ichi Sogo Shuppan, Tokyo, 112 pp." type="book" year="2015">Baba &amp; Tanikawa, 2015: 74</bibRefCitation>
</taxonomicName>
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, 7 figs., male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36D6186AFFDCFF1EF84FFF5A4FA0" blockId="4.[136,774,1931,1987]" pageId="4" pageNumber="483">For the complete list of taxonomic references see WSC (2020).</paragraph>
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<emphasis id="B940EAC4186AFFDCFF1EF82EFE0A4061" bold="true" box="[136,390,2026,2050]" pageId="4" pageNumber="483">Material examined.</emphasis>
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(
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),
<collectingCountry id="F3237646186AFFDCFF1EF7CFFF5F4040" box="[136,211,2059,2083]" name="Laos" pageId="4" pageNumber="483">LAOS</collectingCountry>
,
<collectingRegion id="49F0F834186AFFDCFF75F7CFFE164041" box="[227,410,2059,2083]" country="Laos" name="Viangchan" pageId="4" pageNumber="483">Vientiane Prov.</collectingRegion>
, env. of
<location id="8EEB600D186AFFDCFD91F7CFFC8E4040" LSID="urn:lsid:plazi:treatment:039D87C0186AFFDCFF01F9E3FA044023:8EEB600D186AFFDCFD91F7CFFC8E4040" box="[519,770,2059,2083]" country="Laos" latitude="18.614773" longLatPrecision="1" longitude="102.40885" name="Nam-Lik Eco-Village" pageId="4" pageNumber="483" stateProvince="Viangchan">Nam-Lik Eco-Village</location>
,
<geoCoordinate id="EE005011186AFFDCFF1EF7EDFEA04021" box="[136,300,2089,2114]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="483" precision="1" seconds="53.18" value="18.614773">18°3653.18″N</geoCoordinate>
<geoCoordinate id="EE005011186AFFDCFEA1F7EDFE644021" box="[311,488,2089,2114]" degrees="102" direction="east" minutes="24" orientation="longitude" pageId="4" pageNumber="483" precision="1" seconds="31.87" value="102.40885">102°2431.87″E</geoCoordinate>
, hand picking at bank of lake, coll.
<collectorName id="26C15300186AFFDCFC21F9ECFBDA4E23" box="[951,1110,1576,1600]" pageId="4" pageNumber="483">M.M. Omelko</collectorName>
,
<date id="FF8A1016186AFFDCFBF5F9ECFAED4E23" box="[1123,1377,1576,1600]" pageId="4" pageNumber="483" value="2013-11-19" valueMax="2013-11-20" valueMin="2013-11-19">
<collectingDate id="EFCEE9FE186AFFDCFBF5F9ECFAED4E23" box="[1123,1377,1576,1600]" pageId="4" pageNumber="483" value="2013-11-19" valueMax="2013-11-20" valueMin="2013-11-19">1920 November 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B5C3C8B186AFFDCFAF8F9ECFA814EE3" collectingDate="2013-11-21" collectingDateMax="2013-11-25" collectingDateMin="2013-11-21" collectionCode="FEFU" collectorName="M. M. Omelko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="4" pageNumber="483" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Viangchan">
<specimenCount id="9D32FD5F186AFFDCFAF8F9ECFA4E4E23" box="[1390,1474,1576,1600]" pageId="4" pageNumber="483" type="male">6 males</specimenCount>
<specimenCount id="9D32FD5F186AFFDCFCD5F98CFC274E03" box="[835,939,1608,1632]" pageId="4" pageNumber="483" type="female">2 females</specimenCount>
(
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), same locality,
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at river bank, coll.
<collectorName id="26C15300186AFFDCFCE1F9ACFB9C4EE3" box="[887,1040,1640,1664]" pageId="4" pageNumber="483">M.M. Omelko</collectorName>
,
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<collectingDate id="EFCEE9FE186AFFDCFB8FF9ACFA814EE3" box="[1049,1293,1640,1664]" pageId="4" pageNumber="483" value="2013-11-21" valueMax="2013-11-25" valueMin="2013-11-21">2125 November 2013</collectingDate>
</date>
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;
<materialsCitation id="3B5C3C8B186AFFDCFA80F9ACFBB64EA3" collectingDate="2013-11-19" collectingDateMax="2013-11-24" collectingDateMin="2013-11-19" collectionCode="FEFU" collectorName="M. M. Omelko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="4" pageNumber="483" specimenCount="2" specimenCount-male="2" stateProvince="Viangchan">
<specimenCount id="9D32FD5F186AFFDCFA80F9ACFAEA4EE3" box="[1302,1382,1640,1664]" pageId="4" pageNumber="483" type="male">2 males</specimenCount>
(
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), same locality,
<collectingMethod id="52754EC1186AFFDCFC76F94CFBD74EFC" box="[992,1115,1672,1696]" pageId="4" pageNumber="483">pitfall traps</collectingMethod>
at meadow, coll.
<collectorName id="26C15300186AFFDCFA89F94CFA314EC3" box="[1311,1469,1672,1696]" pageId="4" pageNumber="483">M.M. Omelko</collectorName>
,
<date id="FF8A1016186AFFDCFCD3F96CFBB64EA3" box="[837,1082,1704,1728]" pageId="4" pageNumber="483" value="2013-11-19" valueMax="2013-11-24" valueMin="2013-11-19">
<collectingDate id="EFCEE9FE186AFFDCFCD3F96CFBB64EA3" box="[837,1082,1704,1728]" pageId="4" pageNumber="483" value="2013-11-19" valueMax="2013-11-24" valueMin="2013-11-19">1924 November 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B5C3C8B186AFFDCFBD5F96CFAB44E83" collectingDate="2016-05-23" collectingDateMax="2013-11-24" collectingDateMin="2013-11-19" collectionCode="FEFU" collectorName="A. A. Komissarenko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="4" pageNumber="483" specimenCount="1" specimenCount-male="1" stateProvince="Viangchan">
<specimenCount id="9D32FD5F186AFFDCFBD5F96CFB064EA3" box="[1091,1162,1704,1728]" pageId="4" pageNumber="483" type="male">1 male</specimenCount>
(
<collectionCode id="ED25AE13186AFFDCFB00F96CFB504EA3" box="[1174,1244,1704,1728]" pageId="4" pageNumber="483">FEFU</collectionCode>
), same locality,
<collectingMethod id="52754EC1186AFFDCFA12F96CFCF44EBC" pageId="4" pageNumber="483">pitfall traps</collectingMethod>
, coll.
<collectorName id="26C15300186AFFDCFC2AF90CFB1A4E83" box="[956,1174,1736,1760]" pageId="4" pageNumber="483">A.A. Komissarenko</collectorName>
,
<date id="FF8A1016186AFFDCFB34F90CFAB44E83" box="[1186,1336,1736,1760]" pageId="4" pageNumber="483" value="2016-05-23">
<collectingDate id="EFCEE9FE186AFFDCFB34F90CFAB44E83" box="[1186,1336,1736,1760]" pageId="4" pageNumber="483" value="2016-05-23">23 May 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8B36D6186AFFDCFCD5F8CCFAEE4FC3" blockId="4.[835,1475,1800,1952]" pageId="4" pageNumber="483">
<emphasis id="B940EAC4186AFFDCFCD5F8CCFC064F43" bold="true" box="[835,906,1800,1824]" pageId="4" pageNumber="483">Notes.</emphasis>
This species belongs to the speciose species group
<taxonomicName id="4C344D55186AFFDCFCD2F8ECFC284F23" baseAuthorityName="Thorell" baseAuthorityYear="1872" box="[836,932,1832,1856]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B940EAC4186AFFDCFCD2F8ECFC284F23" box="[836,932,1832,1856]" italics="true" pageId="4" pageNumber="483">nebulosa</emphasis>
</taxonomicName>
. Although this species was treated in a number of taxonomic publications (
<collectionCode id="ED25AE13186AFFDCFBD9F88CFB014F03" box="[1103,1165,1864,1888]" country="0" httpUri="http://grbio.org/cool/cnv0-s4bi" name="Westfield State College, Museum and Herbarium" pageId="4" pageNumber="483">WSC</collectionCode>
, 2020) its copulatory organs have never been illustrated in detail. Therefore, we provide photos of the male palp and female epigyne here.
</paragraph>
</subSubSection>
<subSubSection id="C32E655D186AFFDCFCD5F80CFA044023" pageId="4" pageNumber="483" type="distribution">
<paragraph id="8B8B36D6186AFFDCFCD5F80CFA044023" blockId="4.[835,1476,1992,2112]" pageId="4" pageNumber="483">
<emphasis id="B940EAC4186AFFDCFCD5F80CFC5A4F83" bold="true" box="[835,982,1992,2016]" pageId="4" pageNumber="483">Distribution.</emphasis>
It is distributed from
<collectingCountry id="F3237646186AFFDCFB5FF80CFA8C4F83" box="[1225,1280,1992,2016]" name="India" pageId="4" pageNumber="483">India</collectingCountry>
to
<collectingCountry id="F3237646186AFFDCFAB3F80CFAEE4FBC" box="[1317,1378,1992,2015]" name="Japan" pageId="4" pageNumber="483">Japan</collectingCountry>
south to Java and is known from
<collectingCountry id="F3237646186AFFDCFBCCF82CFB1F4063" box="[1114,1171,2024,2048]" name="India" pageId="4" pageNumber="483">India</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFB37F82CFB724063" box="[1185,1278,2024,2048]" name="Pakistan" pageId="4" pageNumber="483">Pakistan</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFA9AF82CFAFA4063" box="[1292,1398,2024,2048]" name="Myanmar" pageId="4" pageNumber="483">Myanmar</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFA12F82CFA484063" box="[1412,1476,2024,2048]" name="China" pageId="4" pageNumber="483">China</collectingCountry>
(south, south-east and east provinces),
<collectingCountry id="F3237646186AFFDCFB7EF7CCFAFF407C" box="[1256,1395,2056,2080]" name="South Korea" pageId="4" pageNumber="483">South Korea</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFAE9F7CCFA32407C" box="[1407,1470,2056,2079]" name="Japan" pageId="4" pageNumber="483">Japan</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFCD3F7ECFC4C4023" box="[837,960,2088,2112]" name="Philippines" pageId="4" pageNumber="483">Philippines</collectingCountry>
,
<collectingCountry id="F3237646186AFFDCFC5AF7ECFBB84023" box="[972,1076,2088,2112]" name="Indonesia" pageId="4" pageNumber="483">Indonesia</collectingCountry>
(
<collectingRegion id="49F0F834186AFFDCFBD2F7ECFBF44023" box="[1092,1144,2088,2112]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="483">Java</collectingRegion>
), and
<collectingCountry id="F3237646186AFFDCFB2AF7ECFB7C405C" box="[1212,1264,2088,2111]" name="Laos" pageId="4" pageNumber="483">Laos</collectingCountry>
(first record).
</paragraph>
</subSubSection>
</treatment>
</document>