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<mods:title id="D97C59FE8E4242AC162DA016680AA1F9">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<paragraph id="8B8BCC4DFE5FFFB4FBC8FB30FD7FFAF2" blockId="22.[151,709,1259,1283]" box="[151,709,1259,1283]" pageId="22" pageNumber="23">
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<taxonomicName id="4C34B7CEFE5FFFB4FBC8FB30FD7FFAF2" authority="Bannikov &amp; Kotlyar, 2015" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[151,709,1259,1283]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5FFFB4FBC8FB30FD31FAF2" bold="true" box="[151,651,1259,1283]" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4FBC8FB30FE20FAF2" bold="true" box="[151,410,1259,1283]" italics="true" pageId="22" pageNumber="23">Pshekharus yesinorum</emphasis>
<bibRefCitation id="EFA5B1BCFE5FFFB4FAFFFB30FD31FAF2" author="Bannikov, A. F. &amp; Kotlyar, A. N." box="[416,651,1259,1283]" pageId="22" pageNumber="23" pagination="627 - 635" refId="ref19314" refString="Bannikov, A. F., &amp; Kotlyar, A. N. (2015). A new genus and species of Early Sarmatian porgies (Perciformes, Sparidae) from the Krasnodar region. Paleontological Journal, 49 (6), 627 - 635. https: // doi. org / 10.1134 / S 0031 030115060052" type="journal article" year="2015">Bannikov &amp; Kotlyar,</bibRefCitation>
</emphasis>
2015
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE5FFFB5F872FED0FDC2FAB2" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8B8BCC4DFE5FFFB4F872FED0FBB9FED2" blockId="22.[813,1027,267,291]" box="[813,1027,267,291]" pageId="22" pageNumber="23">
<figureCitation id="130FD0C8FE5FFFB4F872FED0FC0CFED2" box="[813,950,267,291]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="22" pageNumber="23">Figures 9d, e</figureCitation>
,
<figureCitation id="130FD0C8FE5FFFB4F89EFED0FBB9FED2" box="[961,1027,267,291]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="22" pageNumber="23">10ae</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F872FE90FC34FE72" blockId="22.[813,1437,331,387]" pageId="22" pageNumber="23">
1966
<taxonomicName id="4C34B7CEFE5FFFB4F834FE90FC5AFE92" authorityName="Roding" authorityYear="1798" box="[875,992,331,355]" family="Sparidae" genus="Cantharus" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE5FFFB4F834FE90FC5AFE92" box="[875,992,331,355]" italics="true" pageId="22" pageNumber="23">Cantharus</emphasis>
</taxonomicName>
?
<emphasis id="B940105FFE5FFFB4F8ACFE97FB8FFE92" box="[1011,1077,332,355]" italics="true" pageId="22" pageNumber="23">tietzei</emphasis>
<bibRefCitation id="EFA5B1BCFE5FFFB4FF60FE90FB5CFE92" author="Schubert, R. J." box="[1087,1254,331,355]" pageId="22" pageNumber="23" pagination="623 - 706" refId="ref23059" refString="Schubert, R. J. (1906). Die Fischotolithen des osterr. - ungar. Tertiars. III. Macruriden und Beryciden. Jahrbuch Der Kaiserlich-Koniglichen Geologischen Reichsanstalt, Wien, 56, 623 - 706." type="journal article" year="1906">Schubert, 1906</bibRefCitation>
—Smigielska: pl. 17, fig. 3.
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F872FE70FCC5FDF2" blockId="22.[813,1436,427,515]" pageId="22" pageNumber="23">
2015
<taxonomicName id="4C34B7CEFE5FFFB4F837FE70FBE6FE32" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[872,1116,427,451]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5FFFB4F837FE70FBE6FE32" box="[872,1116,427,451]" italics="true" pageId="22" pageNumber="23">Pshekharus yesinorum</emphasis>
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—Bannikov &amp; Kotlyar: pl. 10, fig. 1; pl. 11, figs. 13 (fig. 2 represents an otolith found in situ).
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F872FDF0FBEEFD92" blockId="22.[813,1436,555,611]" pageId="22" pageNumber="23">
2022?
<taxonomicName id="4C34B7CEFE5FFFB4F8D7FDF0FBA3FDB2" box="[904,1049,555,579]" family="Sparidae" genus="Pagellus" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE5FFFB4F8D7FDF0FC5BFDB2" box="[904,993,555,579]" italics="true" pageId="22" pageNumber="23">Pagellus</emphasis>
sp.
</taxonomicName>
—Brzobohatý, Zahradníkova &amp; Hudáčková: pl. 3, fig. M, N.
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F872FD50FA27FCF2" blockId="22.[813,1437,651,771]" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4F872FD50FC34FD52" box="[813,910,651,675]" italics="true" pageId="22" pageNumber="23">Material</emphasis>
<materialsCitation id="3B5CC610FE5FFFB4F8CBFD50FC26FD32" collectionCode="PI" pageId="22" pageNumber="23" specimenCode="PI 2577" specimenCount="1">
3 otoliths:
<specimenCount id="9D3207C4FE5FFFB4FF58FD50FB3FFD52" box="[1031,1157,651,675]" count="1" pageId="22" pageNumber="23" type="generic">1 specimen</specimenCount>
, Kozatskyi Yar NMNHU- P
<specimenCode id="DB926436FE5FFFB4F81AFD77FC26FD32" box="[837,924,683,707]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="22" pageNumber="23">PI 2577</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B5CC610FE5FFFB4F8F5FD70FCD9FD12" collectionCode="NMNHU-P" pageId="22" pageNumber="23" specimenCode="PI 2578" specimenCount="2">
<specimenCount id="9D3207C4FE5FFFB4F8F5FD70FB8FFD32" box="[938,1077,683,707]" count="2" pageId="22" pageNumber="23" type="generic">2 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE5FFFB4FFA4FD77FAC1FD32" box="[1275,1403,684,707]" pageId="22" pageNumber="23">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE5FFFB4FEDAFD77FCD9FD12" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="22" pageNumber="23">PI 2578</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE5FFFB4F830FD10FA22FD12" box="[879,1432,715,739]" collectionCode="NMNHU-P" pageId="22" pageNumber="23" specimenCode="PI 2588" specimenCount="1">
1 isolated jaw tooth, Mlyntsi,
<collectionCode id="ED255488FE5FFFB4FFE6FD17FA83FD12" box="[1209,1337,716,739]" pageId="22" pageNumber="23">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE5FFFB4FE1DFD17FA22FD12" box="[1346,1432,715,739]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="22" pageNumber="23">PI 2588</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE5FFFB4F872FD30FA23FCF2" box="[813,1433,747,771]" collectionCode="NMNHU-P" pageId="22" pageNumber="23" specimenCode="PI 2589" specimenCount="1">
4 isolated jaw teeth, Shydlivshchyna,
<collectionCode id="ED255488FE5FFFB4FFE1FD37FA84FCF2" box="[1214,1342,748,771]" pageId="22" pageNumber="23">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE5FFFB4FE1BFD37FA23FCF2" box="[1348,1433,747,771]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="22" pageNumber="23">PI 2589</specimenCode>
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.
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F872FCF7FB41FC32" blockId="22.[813,1437,811,1283]" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4F872FCF7FBF6FCB2" box="[813,1100,811,835]" italics="true" pageId="22" pageNumber="23">Description of otoliths</emphasis>
(specimen of
<figureCitation id="130FD0C8FE5FFFB4FE6EFCF0FA2FFCB2" box="[1329,1429,811,835]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="22" pageNumber="23">Fig. 9d</figureCitation>
) OL:OH = 1.65. Ventral rim regularly curved, rather smooth; dorsal rim intensely crenulated with pronounced postdorsal angle. Rostrum long, pointed, 21% of OL. Posterior tip pointed at same level as rostrum.
</paragraph>
<paragraph id="8B8BCC4DFE5FFFB4F81DFC17FAC8FB32" blockId="22.[813,1437,811,1283]" pageId="22" pageNumber="23">Inner face convex. Sulcus moderately long, deepened, slightly supramedian. OL:SuL = 1.3. Ostium ventrally widened, slightly shorter than cauda; cauda slightly flexed at tip. OsL:CaL = 0.9; OsH:CaH = 1.85. Ventral furrow indistinct, close to ventral rim of otolith; dorsal depression long, narrow, with crista superior to rear part of ostium and anterior part of cauda. Outer face flat to slightly concave, smooth except for some irregular radial furrows.</paragraph>
<paragraph id="8B8BCC4DFE5FFFB5F81DFB17FD50FD32" blockId="22.[813,1437,811,1283]" lastBlockId="23.[151,775,235,707]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4F81DFB17FBA6FB12" box="[834,1052,1227,1251]" italics="true" pageId="22" pageNumber="23">Description of teeth</emphasis>
In addition to otoliths, there is a series of small isolated jaw teeth (
<figureCitation id="130FD0C8FE5FFFB4FFE0FB30FA8DFAF2" box="[1215,1335,1259,1283]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="22" pageNumber="23">Fig. 10ae</figureCitation>
) sharing sparid morphology (e.g.
<bibRefCitation id="EFA5B1BCFE5EFFB5FAE5FF30FDD3FEF2" author="Bannikov, A. F." box="[442,617,235,259]" pageId="23" pageNumber="24" pagination="1 - 244" refId="ref19194" refString="Bannikov, A. F. (2010). Fossil acanthopterygian fishes (Teleostei, Acanthopterygii). In L. P. Tatarinov, E. I. Vorobyeva, &amp; E. N. Kurochkin (Eds.), Fossil vertebrates of Russia and adjacent countries (pp. 1 - 244). GEOS. (in Russian, with English summary)." type="book chapter" year="2010">Bannikov, 2010</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5EFFB5F921FF30FE90FED2" author="Bannikov, A. F. &amp; Kotlyar, A. N." pageId="23" pageNumber="24" pagination="627 - 635" refId="ref19314" refString="Bannikov, A. F., &amp; Kotlyar, A. N. (2015). A new genus and species of Early Sarmatian porgies (Perciformes, Sparidae) from the Krasnodar region. Paleontological Journal, 49 (6), 627 - 635. https: // doi. org / 10.1134 / S 0031 030115060052" type="journal article" year="2015">Bannikov &amp; Kotlyar, 2015</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5EFFB5FA64FED0FDAAFED2" author="Gol'din, P. &amp; Haiduc, B. - S. &amp; Kovalchuk, O. &amp; Gorka, M. &amp; Otryazhyi, P. &amp; Branzila, M. &amp; Paun, E. - I. &amp; Barkaszi, Z. &amp; Tibuleac, P. &amp; Ratoi, B. G." box="[315,528,267,291]" pageId="23" pageNumber="24" refId="ref20278" refString="Gol'din, P., Haiduc, B. - S., Kovalchuk, O., Gorka, M., Otryazhyi, P., Branzila, M., Paun, E. - I., Barkaszi, Z., Tibuleac, P., &amp; Ratoi, B. G. (2020). The Volhynian (late Middle Miocene) marine fishes and mammals as proxies for the onset of the Eastern Paratethys re-colonisation by vertebrate fauna. Palaeontologia Electronica, 23 (3), a 43. https: // doi. org / 10.26879 / 1091" type="journal volume" year="2020">Goldin et al., 2020</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE5EFFB5F97EFEDDFF71FEB2" author="Soster, A. &amp; Kovalchuk, O. M." pageId="23" pageNumber="24" pagination="415 - 422" refId="ref23677" refString="Soster, A., &amp; Kovalchuk, O. M. (2016). Late Neogene and Pleistocene porgy fishes (Teleostei, Sparidae) of the Eastern Paratethys, with comments on their palaeoecology. Vestnik Zoologii, 50 (5), 415 - 422. https: // doi. org / 10. 1515 / vzoo- 2016 - 0048" type="journal article" year="2016">Šoster &amp; Kovalchuk, 2016</bibRefCitation>
) and tentatively assigned here to
<taxonomicName id="4C34B7CEFE5EFFB5F909FEF0FF5DFE92" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5EFFB5F909FEF0FF5DFE92" italics="true" pageId="23" pageNumber="24">Pshekharus yesinorum</emphasis>
</taxonomicName>
. Most of these specimens (
<figureCitation id="130FD0C8FE5EFFB5F945FE90FD35FE92" box="[538,655,331,355]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="23" pageNumber="24">Fig. 10ad</figureCitation>
) represent posterior molariform teeth with symmetrical or partially asymmetrical crowns, which are round at the base and have a smooth surface. Tere is a less-pronounced blunt hook at the tip of
<specimenCount id="9D3207C4FE5EFFB5FA03FE17FE49FE12" box="[348,499,459,483]" count="1" pageId="23" pageNumber="24" type="generic">one specimen</specimenCount>
(
<figureCitation id="130FD0C8FE5EFFB5F95EFE10FDE1FE12" box="[513,603,459,483]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="23" pageNumber="24">Fig. 10d</figureCitation>
). Te specimen
<collectionCode id="ED255488FE5EFFB5FBC8FE37FEADFDF2" box="[151,279,492,515]" pageId="23" pageNumber="24">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE5EFFB5FA7DFE37FE80FDF2" box="[290,314,492,515]" country="Russia" name="Paleontological Institute" pageId="23" pageNumber="24">PI</collectionCode>
2589/4 (
<figureCitation id="130FD0C8FE5EFFB5FAFDFE30FE41FDF2" box="[418,507,491,515]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="23" pageNumber="24">Fig. 10e</figureCitation>
) represents an anterior tooth, which is narrow conical proximally and flattened distally. Crown width in the specimens considered (
<quantity id="4CCC61A8FE5EFFB5F996FDF0FF45FD92" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="11.0" metricValueMin="6.5" pageId="23" pageNumber="24" unit="mm" value="0.875" valueMax="1.1" valueMin="0.65">0.65 1.10 mm</quantity>
) is quite similar to each other despite their position in the jaw, while the crown height differs more substantially varying from
<quantity id="4CCC61A8FE5EFFB5FAE0FD50FDCEFD52" box="[447,628,651,675]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="23" pageNumber="24" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60 to 0.80 mm</quantity>
for posterior teeth and being equal to 2.0 mm for the anterior tooth.
</paragraph>
<caption id="DF4B9CC5FE5FFFB4FBF6F8C4FEDDF898" ID-DOI="http://doi.org/10.5281/zenodo.11999950" ID-Zenodo-Dep="11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="22" pageNumber="23" startId="22.[169,200,1823,1843]" targetBox="[153,1438,1393,1909]" targetPageId="22" targetType="figure">
<paragraph id="8B8BCC4DFE5FFFB4FBF6F8C4FEDDF898" blockId="22.[169,1415,1823,1897]" pageId="22" pageNumber="23">
<emphasis id="B940105FFE5FFFB4FBF6F8C4FF5DF8C2" bold="true" box="[169,231,1823,1843]" pageId="22" pageNumber="23">Fig. 10</emphasis>
Fish teeth from the studied localities:
<emphasis id="B940105FFE5FFFB4F971F8C4FDF4F8C2" bold="true" box="[558,590,1823,1843]" pageId="22" pageNumber="23">ae</emphasis>
<taxonomicName id="4C34B7CEFE5FFFB4F90CF8C4FC66F8C2" authority="Bannikov &amp; Kotlyar, 2015" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[595,988,1823,1843]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5FFFB4F90CF8C4FCB2F8C2" box="[595,776,1823,1843]" italics="true" pageId="22" pageNumber="23">Pshekharus yesinorum</emphasis>
<bibRefCitation id="EFA5B1BCFE5FFFB4F853F8C4FC66F8C2" author="Bannikov, A. F. &amp; Kotlyar, A. N." box="[780,988,1823,1843]" pageId="22" pageNumber="23" pagination="627 - 635" refId="ref19314" refString="Bannikov, A. F., &amp; Kotlyar, A. N. (2015). A new genus and species of Early Sarmatian porgies (Perciformes, Sparidae) from the Krasnodar region. Paleontological Journal, 49 (6), 627 - 635. https: // doi. org / 10.1134 / S 0031 030115060052" type="journal article" year="2015">Bannikov &amp; Kotlyar, 2015</bibRefCitation>
</taxonomicName>
, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions.
<emphasis id="B940105FFE5FFFB4F807F8E1FCE5F8BF" bold="true" box="[856,863,1850,1870]" pageId="22" pageNumber="23">f</emphasis>
<taxonomicName id="4C34B7CEFE5FFFB4F83CF8E1FC7DF8BF" box="[867,967,1850,1870]" class="Elasmobranchii" family="Dasyatidae" genus="Dasyatis" kingdom="Animalia" order="Myliobatiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE5FFFB4F83CF8E1FC12F8BF" box="[867,936,1850,1870]" italics="true" pageId="22" pageNumber="23">Dasyatis</emphasis>
sp.
</taxonomicName>
, anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)
</paragraph>
</caption>
<paragraph id="8B8BCC4DFE5EFFB5FBC8FD37FDC2FAB2" blockId="23.[151,776,747,1347]" pageId="23" pageNumber="24">
<emphasis id="B940105FFE5EFFB5FBC8FD37FEBFFCF2" box="[151,261,748,771]" italics="true" pageId="23" pageNumber="24">Discussion</emphasis>
<bibRefCitation id="EFA5B1BCFE5EFFB5FA4EFD30FDEAFCF2" author="Bannikov, A. F. &amp; Kotlyar, A. N." box="[273,592,747,771]" pageId="23" pageNumber="24" pagination="627 - 635" refId="ref19314" refString="Bannikov, A. F., &amp; Kotlyar, A. N. (2015). A new genus and species of Early Sarmatian porgies (Perciformes, Sparidae) from the Krasnodar region. Paleontological Journal, 49 (6), 627 - 635. https: // doi. org / 10.1134 / S 0031 030115060052" type="journal article" year="2015">Bannikov and Kotlyar (2015)</bibRefCitation>
used jaw dentition as a distinguishing character for the definition of their new genus
<taxonomicName id="4C34B7CEFE5EFFB5FA52FCF0FE39FCB2" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[269,387,811,835]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE5EFFB5FA52FCF0FE39FCB2" box="[269,387,811,835]" italics="true" pageId="23" pageNumber="24">Pshekharus</emphasis>
</taxonomicName>
, in particular the long anterior teeth, which are canine-like and cylindrical, such as those figured in
<figureCitation id="130FD0C8FE5EFFB5FBEEFCB0FEB9FC72" box="[177,259,875,899]" captionStart="Fig" captionStartId="22.[169,200,1823,1843]" captionTargetBox="[153,1438,1393,1909]" captionTargetId="figure-434@22.[168,1423,1409,1811]" captionTargetPageId="22" captionText="Fig. 10 Fish teeth from the studied localities: ae Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, isolated jaw teeth in lateral (a1, b1, c, d1, e) and dorsal view (a2, b2, d2), arranged in a sequence from distal to proximal positions. f Dasyatis sp., anterior tooth in dorsal (f1), occlusal view (f2), profile (f3) and labial view (f4)" figureDoi="http://doi.org/10.5281/zenodo.11999950" httpUri="https://zenodo.org/record/11999950/files/figure.png" pageId="23" pageNumber="24">Fig. 10e</figureCitation>
. Te sequence of teeth represented here and their association with otoliths that are similar to the in situ find in
<taxonomicName id="4C34B7CEFE5EFFB5FBC8FC77FEA7FC32" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[151,285,940,963]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5EFFB5FBC8FC77FEA7FC32" box="[151,285,940,963]" italics="true" pageId="23" pageNumber="24">P. yesinorum</emphasis>
</taxonomicName>
support the allocation of both teeth and otoliths to that species, which was originally described from the early Sarmatian s.l. of
<collectingCountry id="F3238CDDFE5EFFB5FADCFC30FE5EFBF2" box="[387,484,1003,1027]" name="Moldova" pageId="23" pageNumber="24">Moldova</collectingCountry>
. However, the otoliths tend to be slightly more compressed with a deeper ventral rim (OL:OH =1.65 vs. 1.852.05) than those from the Sarmatian s.l. (
<bibRefCitation id="EFA5B1BCFE5EFFB5FBB2FB90FE4DFB92" author="Bannikov, A. F. &amp; Kotlyar, A. N." box="[237,503,1099,1123]" pageId="23" pageNumber="24" pagination="627 - 635" refId="ref19314" refString="Bannikov, A. F., &amp; Kotlyar, A. N. (2015). A new genus and species of Early Sarmatian porgies (Perciformes, Sparidae) from the Krasnodar region. Paleontological Journal, 49 (6), 627 - 635. https: // doi. org / 10.1134 / S 0031 030115060052" type="journal article" year="2015">Bannikov &amp; Kotlyar, 2015</bibRefCitation>
), but these differences are considered to represent an expression of variability. Otoliths indicate that
<taxonomicName id="4C34B7CEFE5EFFB5FA7EFB57FE1FFB52" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[289,421,1164,1187]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5EFFB5FA7EFB57FE1FFB52" box="[289,421,1164,1187]" italics="true" pageId="23" pageNumber="24">P. yesinorum</emphasis>
</taxonomicName>
may have been rather widely distributed in the Paratethys in the late Badenian/Konkian and Sarmatian s.l. Specimens identified as
<taxonomicName id="4C34B7CEFE5EFFB5F91CFB10FD09FB12" authorityName="Roding" authorityYear="1798" box="[579,691,1227,1251]" family="Sparidae" genus="Cantharus" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE5EFFB5F91CFB10FD09FB12" box="[579,691,1227,1251]" italics="true" pageId="23" pageNumber="24">Cantharus</emphasis>
</taxonomicName>
?
<emphasis id="B940105FFE5EFFB5F997FB17FCBDFB12" box="[712,775,1228,1251]" italics="true" pageId="23" pageNumber="24">tietzei</emphasis>
by
<bibRefCitation id="EFA5B1BCFE5EFFB5FBE4FB30FEC2FAF2" author="Smigielska, T." box="[187,376,1259,1283]" pageId="23" pageNumber="24" pagination="205 - 275" refId="ref23645" refString="Smigielska, T. (1966). Otoliths of fishes from the Tortonian of Southern Poland. Annales de la Societe Geologique de Pologne, 36, 205 - 275." type="journal article" year="1966">Smigielska (1966)</bibRefCitation>
from the Pre-Carpathian Trough of
<collectingCountry id="F3238CDDFE5EFFB5FBC8FAD0FF5AFAD2" box="[151,224,1291,1315]" name="Poland" pageId="23" pageNumber="24">Poland</collectingCountry>
and?
<taxonomicName id="4C34B7CEFE5EFFB5FA7EFAD0FE26FAD2" box="[289,412,1291,1315]" family="Sparidae" genus="Pagellus" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE5EFFB5FA7EFAD0FECCFAD2" box="[289,374,1291,1315]" italics="true" pageId="23" pageNumber="24">Pagellus</emphasis>
sp.
</taxonomicName>
by Brzobohatý et. al. (2022) from the Vienna Basin likely represent
<taxonomicName id="4C34B7CEFE5EFFB5FAAFFAF7FDC8FAB2" authorityName="Bannikov &amp; Kotlyar" authorityYear="2015" box="[496,626,1324,1347]" family="Sparidae" genus="Pshekharus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="yesinorum">
<emphasis id="B940105FFE5EFFB5FAAFFAF7FDC8FAB2" box="[496,626,1324,1347]" italics="true" pageId="23" pageNumber="24">P. yesinorum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>