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<mods:title id="CFD0D31871109077F6220DA377FBE8AD">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="8DA98A8F03DD5BBDEC06EFA849A3A42B">Schwarzhans, Werner</mods:namePart>
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<mods:date id="E7132A5E6E2691E804B2AD0D663578A0">2022</mods:date>
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<treatment id="039D7D5BFE59FFB2FBC8FCB0FC64FED2" LSID="urn:lsid:plazi:treatment:039D7D5BFE59FFB2FBC8FCB0FC64FED2" httpUri="http://treatment.plazi.org/id/039D7D5BFE59FFB2FBC8FCB0FC64FED2" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C32E9FC6FE59FFB2FBC8FCB0FF60FC52" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8BCC4DFE59FFB2FBC8FCB0FF60FC52" blockId="16.[151,775,875,931]" pageId="16" pageNumber="17">
<taxonomicName id="4C34B7CEFE59FFB2FBC8FCB0FCBAFC72" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[151,768,875,899]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE59FFB2FBC8FCB0FD0CFC72" bold="true" box="[151,694,875,899]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8FCB0FE72FC72" bold="true" box="[151,456,875,899]" italics="true" pageId="16" pageNumber="17">Vanderhorstia prochazkai</emphasis>
<bibRefCitation id="EFA5B1BCFE59FFB2FA8DFCB0FD0CFC72" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[466,694,875,899]" pageId="16" pageNumber="17" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al.,</bibRefCitation>
</emphasis>
2020a
</taxonomicName>
,
<heading id="D0C37B21FE59FFB2FBC8FC50FF60FC52" box="[151,218,907,931]" fontSize="10" level="2" pageId="16" pageNumber="17" reason="7">
<bibRefCitation id="EFA5B1BCFE59FFB2FBC8FC50FF60FC52" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[151,218,907,931]" pageId="16" pageNumber="17" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB2FBC8FC10FD79F9F2" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8BCC4DFE59FFB2FBC8FC10FEAFFC12" blockId="16.[151,277,971,995]" box="[151,277,971,995]" pageId="16" pageNumber="17">
<figureCitation id="130FD0C8FE59FFB2FBC8FC10FEAFFC12" box="[151,277,971,995]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="16" pageNumber="17">Figure 6hj</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2FBC8FBD0FD9EFBB2" blockId="16.[151,775,1035,1091]" pageId="16" pageNumber="17">
2020a
<taxonomicName id="4C34B7CEFE59FFB2FB80FBD0FE40FBD2" box="[223,506,1035,1059]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE59FFB2FB80FBD0FE40FBD2" box="[223,506,1035,1059]" italics="true" pageId="16" pageNumber="17">Vanderhorstia prochazkai</emphasis>
</taxonomicName>
—Schwarzhans, Brzobohatý &amp; Radwańska: pl. 7, figs. 1218.
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2FBC8FBB0FDC9FB32" blockId="16.[151,774,1131,1219]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8FBB0FF42FB72" box="[151,248,1131,1155]" italics="true" pageId="16" pageNumber="17">Material</emphasis>
6 otoliths:
<specimenCount id="9D3207C4FE59FFB2FA2FFBB0FE57FB72" box="[368,493,1131,1155]" count="1" pageId="16" pageNumber="17" type="generic">1 specimen</specimenCount>
, Kozatskyi Yar, NMNHU- P
<collectionCode id="ED255488FE59FFB2FBEFFB57FF72FB52" box="[176,200,1164,1187]" country="Russia" name="Paleontological Institute" pageId="16" pageNumber="17">PI</collectionCode>
2581;
<specimenCount id="9D3207C4FE59FFB2FA49FB50FE19FB52" box="[278,419,1163,1187]" count="2" pageId="16" pageNumber="17" type="generic">2 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE59FFB2F94AFB57FDEBFB52" box="[533,593,1164,1187]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="16" pageNumber="17">NMB</collectionCode>
P1223;
<specimenCount id="9D3207C4FE59FFB2F9F1FB50FF6EFB32" count="3" pageId="16" pageNumber="17" type="generic">3 specimens</specimenCount>
, Shydlivshchyna,
<collectionCode id="ED255488FE59FFB2FACBFB77FDAEFB32" box="[404,532,1196,1219]" pageId="16" pageNumber="17">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE59FFB2F945FB77FD88FB32" box="[538,562,1196,1219]" country="Russia" name="Paleontological Institute" pageId="16" pageNumber="17">PI</collectionCode>
2582.
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2FBC8FB37FD79F9F2" blockId="16.[151,775,1259,1539]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8FB37FEB0FAF2" box="[151,266,1260,1283]" italics="true" pageId="16" pageNumber="17">Discussion</emphasis>
Tese are typical high-bodied and rather thick otoliths of
<taxonomicName id="4C34B7CEFE59FFB2FA4AFAD7FE17FAD2" box="[277,429,1291,1315]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE59FFB2FA4AFAD7FE17FAD2" box="[277,429,1291,1315]" italics="true" pageId="16" pageNumber="17">V. prochazkai</emphasis>
</taxonomicName>
that are further characterized by a very large subcaudal iugum.
<taxonomicName id="4C34B7CEFE59FFB2F95AFAF0FF03FA92" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE59FFB2F95AFAF0FF03FA92" italics="true" pageId="16" pageNumber="17">Vanderhorstia prochazkai</emphasis>
</taxonomicName>
has been described from the late Badenian and early Sarmatian s.s. of the Central Paratethys by Schwarzhans et. al. (2020a). Today, species of the
<taxonomicName id="4C34B7CEFE59FFB2F96FFA57FF01FA35" authority="Lineage" authorityName="Lineage" family="Gobiidae" genus="Asterropteryx" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B940105FFE59FFB2F96FFA57FD7EFA52" box="[560,708,1420,1443]" italics="true" pageId="16" pageNumber="17">Asterropteryx</emphasis>
Lineage
</taxonomicName>
live commensal with alpheid shrimps, and it has been suggested that this was also the case in the middle Miocene (
<bibRefCitation id="EFA5B1BCFE59FFB2FB89FA30FE29F9F2" author="Radwanska, U." box="[214,403,1515,1539]" pageId="16" pageNumber="17" pagination="597 - 605" refId="ref22708" refString="Radwanska, U. (2018). A commensal relationship between alpheid crustaceans and gobiid fish in the middle Miocene of southern Poland (Central Paratethys). Acta Geologica Polonica, 68, 597 - 605." type="journal article" year="2018">Radwańska, 2018</bibRefCitation>
;
<bibRefCitation id="EFA5B1BCFE59FFB2FAFFFA30FD0FF9F2" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[416,693,1515,1539]" pageId="16" pageNumber="17" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB2FBC8F9F0FED9F9B2" box="[151,355,1579,1603]" pageId="16" pageNumber="17" type="multiple">
<paragraph id="8B8BCC4DFE59FFB2FBC8F9F0FED9F9B2" blockId="16.[151,355,1579,1603]" box="[151,355,1579,1603]" pageId="16" pageNumber="17">
<heading id="D0C37B21FE59FFB2FBC8F9F0FED9F9B2" bold="true" box="[151,355,1579,1603]" fontSize="10" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C34B7CEFE59FFB2FBC8F9F0FED9F9B2" box="[151,355,1579,1603]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B940105FFE59FFB2FBC8F9F0FED9F9B2" bold="true" box="[151,355,1579,1603]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8F9F0FE87F9B2" bold="true" box="[151,317,1579,1603]" italics="true" pageId="16" pageNumber="17">Vanderhorstia</emphasis>
sp.
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E9FC6FE59FFB2FBC8F9B0FC64FED2" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8BCC4DFE59FFB2FBC8F9B0FF46F972" blockId="16.[151,252,1643,1667]" box="[151,252,1643,1667]" pageId="16" pageNumber="17">
<figureCitation id="130FD0C8FE59FFB2FBC8F9B0FF46F972" box="[151,252,1643,1667]" captionStart="Fig" captionStartId="14.[169,200,1796,1816]" captionTargetBox="[152,1438,540,1933]" captionTargetId="figure-92@14.[168,1423,556,1783]" captionTargetPageId="14" captionText="Fig. 6 Otoliths of Medoborichthys and Vanderhorstia: ad Medoborichthys podolicus n. gen. et n. sp., b holotype, NMNHU-P PI 2563, Mlyntsi, a, c, d paratypes,a Mlyntsi, NMNHU-P PI 2564,c Shydlivshchyna,NMNHU-P PI 2565, d (reversed) Kozatskyi Yar, NMB P1215. eg Medoborichthys renesulcis n. gen. et n. sp., e holotype, NMNHU-P PI 2566, Mlyntsi,f, g paratypes,f Kozatskyi Yar, NMB P1216, g (reversed) Mlyntsi, NMNHU-P PI 2567. hj Vanderhorstia prochazkai Schwarzhans et al., 2020a, 2020b, h (reversed), j (reversed) Mlyntsi, NMB P1223, h (reversed) Shydlivshchyna, NMNHU-P PI 2582. k Vanderhorstia sp., Shydlivshchyna, NMB P1224" figureDoi="http://doi.org/10.5281/zenodo.11999907" httpUri="https://zenodo.org/record/11999907/files/figure.png" pageId="16" pageNumber="17">Figure 6k</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2FBC8F970FEE8F912" blockId="16.[151,774,1707,1763]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8F970FF42F932" box="[151,248,1707,1731]" italics="true" pageId="16" pageNumber="17">Material</emphasis>
2 otoliths, Shydlivshchyna,
<collectionCode id="ED255488FE59FFB2F974F977FD11F932" box="[555,683,1708,1731]" pageId="16" pageNumber="17">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE59FFB2F9ECF977FD71F932" box="[691,715,1708,1731]" country="Russia" name="Paleontological Institute" pageId="16" pageNumber="17">PI</collectionCode>
2583 and
<collectionCode id="ED255488FE59FFB2FB99F917FEB8F912" box="[198,258,1740,1763]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="16" pageNumber="17">NMB</collectionCode>
P1224.
</paragraph>
<paragraph id="8B8BCC4DFE59FFB2FBC8F8D7FC64FED2" blockId="16.[151,776,1803,1924]" lastBlockId="16.[813,1436,235,291]" pageId="16" pageNumber="17">
<emphasis id="B940105FFE59FFB2FBC8F8D7FEB0F8D2" box="[151,266,1804,1827]" italics="true" pageId="16" pageNumber="17">Discussion</emphasis>
<specimenCount id="9D3207C4FE59FFB2FA4AF8D7FE7BF8D2" box="[277,449,1803,1827]" count="2" pageId="16" pageNumber="17" type="generic">Two specimens</specimenCount>
differ from
<taxonomicName id="4C34B7CEFE59FFB2F90FF8D7FD5DF8D2" box="[592,743,1803,1827]" family="Gobiidae" genus="Vanderhorstia" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="prochazkai">
<emphasis id="B940105FFE59FFB2F90FF8D7FD5DF8D2" box="[592,743,1803,1827]" italics="true" pageId="16" pageNumber="17">V. prochazkai</emphasis>
</taxonomicName>
in being distinctly thinner (OH:OT = 3.45 vs. mostly 3.0 3.3, rarely 3.5) and in showing a narrow subcaudal iugum. It may represent a second species, but the available specimens are not sufficiently well-preserved for a definite assessment.
</paragraph>
</subSubSection>
</treatment>
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