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<mods:title id="E596231B1DFFAE6E00DCD5E45508AB6D">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="F9B1BC0A24FCB20F3ACE657131A8EC62">Schwarzhans, Werner</mods:namePart>
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<treatment id="039D7D5BFE40FFABFBC8FA70FAEBFE12" LSID="urn:lsid:plazi:treatment:039D7D5BFE40FFABFBC8FA70FAEBFE12" httpUri="http://treatment.plazi.org/id/039D7D5BFE40FFABFBC8FA70FAEBFE12" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C32E9FC6FE40FFABFBC8FA70FD3BFA32" box="[151,641,1451,1475]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B8BCC4DFE40FFABFBC8FA70FD3BFA32" blockId="9.[151,641,1451,1475]" box="[151,641,1451,1475]" pageId="9" pageNumber="10">
<heading id="D0C37B21FE40FFABFBC8FA70FD3BFA32" box="[151,641,1451,1475]" fontSize="9" level="2" pageId="9" pageNumber="10" reason="8">
<taxonomicName id="4C34B7CEFE40FFABFBC8FA70FD3BFA32" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[151,641,1451,1475]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="reichenbacherae">
<emphasis id="B940105FFE40FFABFBC8FA70FE11FA32" bold="true" box="[151,427,1451,1475]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
<bibRefCitation id="EFA5B1BCFE40FFABFAEEFA70FD3BFA32" author="Schwarzhans, W." box="[433,641,1451,1475]" pageId="9" pageNumber="10" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C32E9FC6FE40FFABFBC8FA30FAEBFE12" pageId="9" pageNumber="10" type="description">
<paragraph id="8B8BCC4DFE40FFABFBC8FA30FF4DF9F2" blockId="9.[151,247,1515,1539]" box="[151,247,1515,1539]" pageId="9" pageNumber="10">
<figureCitation id="130FD0C8FE40FFABFBC8FA30FF4DF9F2" box="[151,247,1515,1539]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: ae Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, be paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,ce (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).gk Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, gi, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">
Figure 4
<bibRefCitation id="EFA5B1BCFE40FFABFBB0FA30FF4DF9F2" author="Bannikov, A. F." box="[239,247,1515,1539]" pageId="9" pageNumber="10" pagination="23 - 34" refId="ref19263" refString="Bannikov, A. F. (2014). The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV. Miscellanea Paleontologica, 12, 23 - 34." type="journal article" year="2014">f</bibRefCitation>
</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE40FFABFBC8F9F0FF40F992" blockId="9.[151,775,1579,1635]" pageId="9" pageNumber="10">
2014
<taxonomicName id="4C34B7CEFE40FFABFBBAF9F0FE43F9B2" authorityName="Schwarzhans" authorityYear="2014" box="[229,505,1579,1603]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="reichenbacherae">
<emphasis id="B940105FFE40FFABFBBAF9F0FE43F9B2" box="[229,505,1579,1603]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
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—Schwarzhans: pl. 9, figs. 29.
</paragraph>
<paragraph id="8B8BCC4DFE40FFABFBC8F950FE1BF912" blockId="9.[151,775,1675,1763]" pageId="9" pageNumber="10">
2020a
<taxonomicName id="4C34B7CEFE40FFABFBB3F950FD54F952" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[236,750,1675,1699]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="reichenbacherae">
<emphasis id="B940105FFE40FFABFBB3F950FE46F952" box="[236,508,1675,1699]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
<bibRefCitation id="EFA5B1BCFE40FFABF94FF950FD54F952" author="Schwarzhans, W." box="[528,750,1675,1699]" pageId="9" pageNumber="10" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
</taxonomicName>
— Schwarzhans, Brzobohatý &amp; Radwańska: pl. 4, figs. 69 (and references therein).
</paragraph>
<paragraph id="8B8BCC4DFE40FFABFBC8F8D0FD41F8D2" blockId="9.[151,763,1803,1827]" box="[151,763,1803,1827]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABFBC8F8D0FF42F8D2" box="[151,248,1803,1827]" italics="true" pageId="9" pageNumber="10">Material</emphasis>
1 otolith, Shydlivshchyna,
<collectionCode id="ED255488FE40FFABF944F8D7FD21F8D2" box="[539,667,1804,1827]" pageId="9" pageNumber="10">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE40FFABF9FDF8D7FD00F8D2" box="[674,698,1804,1827]" country="Russia" name="Paleontological Institute" pageId="9" pageNumber="10">PI</collectionCode>
2557.
</paragraph>
<paragraph id="8B8BCC4DFE40FFABFBC8F897FAEBFE12" blockId="9.[151,775,1867,1923]" lastBlockId="9.[813,1437,235,483]" pageId="9" pageNumber="10">
<emphasis id="B940105FFE40FFABFBC8F897FEB0F892" box="[151,266,1868,1891]" italics="true" pageId="9" pageNumber="10">Discussion</emphasis>
Te single, large otolith of
<quantity id="4CCC61A8FE40FFABF96BF890FD3DF892" box="[564,647,1867,1891]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="10" unit="mm" value="3.3">3.3 mm</quantity>
in length is readily recognized by the slightly convex inner face and the flat outer face, the parallelogram-like shape, and the postdorsal projection being markedly bent outward. Te sulcus shows a shoe sole shape with moderate ostial lobe, and the subcaudal iugum is clearly discernable despite erosion.
<taxonomicName id="4C34B7CEFE40FFABF8CCFEB0FB24FE72" authorityName="Schwarzhans" authorityYear="2014" box="[915,1182,363,387]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="reichenbacherae">
<emphasis id="B940105FFE40FFABF8CCFEB0FB24FE72" box="[915,1182,363,387]" italics="true" pageId="9" pageNumber="10">Gobius reichenbacherae</emphasis>
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was a common and widespread species in the middle Miocene of the Mediterranean and Paratethys and the late Miocene of the Mediterranean (
<bibRefCitation id="EFA5B1BCFE40FFABF882FE10FB4EFE12" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[989,1268,459,483]" pageId="9" pageNumber="10" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
,
<bibRefCitation id="EFA5B1BCFE40FFABFE5FFE10FAF9FE12" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[1280,1347,459,483]" pageId="9" pageNumber="10" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
).
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