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<document id="3DCB293DF8BC5B131EB96B5CB5D6B060" ID-CLB-Dataset="5910" ID-DOI="10.11646/zootaxa.3765.2.1" ID-GBIF-Dataset="300ad198-2377-4bfd-9381-6868ee9c5f7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5045725" ID-ZooBank="E7DDABDA-5DA6-4309-A26F-121FCB030EEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623111725456" checkinUser="felipe" docAuthor="Dyck, Steve Van" docDate="2014" docId="039D2A4709533E2AC6F812BAFE765164" docLanguage="en" docName="zt03765p133.pdf" docOrigin="Zootaxa 3765 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Antechinus subtropicus Van Dyck and Crowther" docType="treatment" docVersion="8" lastPageNumber="119" masterDocId="FFA4523F09423E38C66F1765FF84556A" masterDocTitle="The Black-tailed Antechinus, Antechinus arktos sp. nov.: a new species of carnivorous marsupial from montane regions of the Tweed Volcano caldera, eastern Australia" masterLastPageNumber="133" masterPageNumber="101" pageNumber="118" updateTime="1699002937234" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DAF5F6A2C8C32A97D4671F5DCB613A8C">The Black-tailed Antechinus, Antechinus arktos sp. nov.: a new species of carnivorous marsupial from montane regions of the Tweed Volcano caldera, eastern Australia</mods:title>
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<mods:namePart id="23E1CFB25234BACC6C141C41F15BD199">Dyck, Steve Van</mods:namePart>
<mods:affiliation id="1FA10C543DADC0DE9F7B4701B28A6803">Queensland Museum, Vertebrate Zoology (Mammals &amp; Birds), PO Box 3300, South Brisbane, Qld, 4101, Australia</mods:affiliation>
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<treatment id="039D2A4709533E2AC6F812BAFE765164" ID-DOI="http://doi.org/10.5281/zenodo.5046129" ID-GBIF-Taxon="183227950" ID-Zenodo-Dep="5046129" LSID="urn:lsid:plazi:treatment:039D2A4709533E2AC6F812BAFE765164" httpUri="http://treatment.plazi.org/id/039D2A4709533E2AC6F812BAFE765164" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118">
<subSubSection id="C32EC8DA09533E29C6F812BAFCC85092" box="[151,844,1503,1528]" pageId="17" pageNumber="118" type="nomenclature">
<paragraph id="8B8B9B5109533E29C6F812BAFCC85092" blockId="17.[151,1437,1503,2038]" box="[151,844,1503,1528]" pageId="17" pageNumber="118">
<emphasis id="B940474309533E29C6F812BAFE035092" bold="true" box="[151,391,1503,1528]" pageId="17" pageNumber="118">
(16)
<emphasis id="B940474309533E29C6A312BAFEB15092" bold="true" box="[204,309,1503,1528]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
versus
</emphasis>
<taxonomicName id="4C34E0D209533E29C7E012BAFCC85092" ID-CoL="66XSK" authority="Van Dyck and Crowther" box="[399,844,1503,1528]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C7E012BAFDB25092" bold="true" box="[399,566,1503,1528]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
Van Dyck and Crowther
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C32EC8DA09533E2AC6A81166FE765164" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118" type="description">
<paragraph id="8B8B9B5109533E29C6A81166FDF5530E" blockId="17.[151,1437,1503,2038]" pageId="17" pageNumber="118">
Pelage:
<taxonomicName id="4C34E0D209533E29C74C1161FE0E5376" authorityName="Dyck" authorityYear="2014" box="[291,394,1539,1564]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C74C1161FE0E5376" box="[291,394,1539,1564]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
has a brownish-grey head that changes markedly to an orange-brown rump, fuscous black hindfeet, a thick-based, finely-furred, black tail and an orange-yellow eye and cheek patch;
<taxonomicName id="4C34E0D209533E29C2B7114DFAFA532A" box="[1240,1406,1575,1600]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C2B7114DFAFA532A" box="[1240,1406,1575,1600]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
is uniformly brownish on the head and back.
</paragraph>
<paragraph id="8B8B9B5109533E29C6A8110AFBCB53C6" blockId="17.[151,1437,1503,2038]" pageId="17" pageNumber="118">
External Measurements:
<taxonomicName id="4C34E0D209533E29C78C1115FDCE53E2" authorityName="Dyck" authorityYear="2014" box="[483,586,1647,1672]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C78C1115FDCE53E2" box="[483,586,1647,1672]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
is larger than
<taxonomicName id="4C34E0D209533E29C49E1115FC1153E2" box="[753,917,1647,1672]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C49E1115FC1153E2" box="[753,917,1647,1672]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
in absolute measurement for hf in males and females.
<taxonomicName id="4C34E0D209533E29C69311F1FEE553C6" authorityName="Dyck" authorityYear="2014" box="[252,353,1683,1708]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C69311F1FEE553C6" box="[252,353,1683,1708]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
is significantly larger than
<taxonomicName id="4C34E0D209533E29C4FB11F1FCB153C6" box="[660,821,1683,1708]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C4FB11F1FCB153C6" box="[660,821,1683,1708]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
for males at tv (
<tableCitation id="C6B6AEEA09533E29C58411F6FBC553C6" box="[1003,1089,1683,1708]" captionStart="TABLE 5" captionStartId="13.[147,241,631,655]" captionTargetId="figure-113@13.[147,1770,777,1673]" captionTargetPageId="13" captionText="TABLE 5 ANOVA F-statistics (top two lines) for variation at each of the measured variables among all antechinus species and subspecies. Subsequent rows show significance values for ANOVA Post-Hoc tests of Antechinus arktos paired with each of its 12 congeners, for each measured variable. Shaded cells are significant at p=0.05, unshaded cells are not significant." pageId="17" pageNumber="118">Table 5</tableCitation>
).
</paragraph>
<paragraph id="8B8B9B5109533E29C6A811D2FC155228" blockId="17.[151,1437,1503,2038]" pageId="17" pageNumber="118">
Craniodental Characters:
<taxonomicName id="4C34E0D209533E29C79F11DDFDDF53BA" authorityName="Dyck" authorityYear="2014" box="[496,603,1719,1744]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C79F11DDFDDF53BA" box="[496,603,1719,1744]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
is larger than
<taxonomicName id="4C34E0D209533E29C57D11DDFC3C53BA" box="[786,952,1719,1744]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C57D11DDFC3C53BA" box="[786,952,1719,1744]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
in absolute measurement for a range of characters in males and females, as follows: APV, BL, Dent, IBW, IOW, IPV, HT, PL, NWR, PML, UML, HT-B, I
<superScript id="7C41361909533E29C6CF119BFF2C5266" attach="left" box="[160,168,1790,1804]" fontSize="6" pageId="17" pageNumber="118">1</superScript>
-P
<superScript id="7C41361909533E29C6AE119BFF4D5266" attach="left" box="[193,201,1790,1804]" fontSize="6" pageId="17" pageNumber="118">3</superScript>
, LML, I
<subScript id="17B0991409533E29C7441074FEB75275" attach="left" box="[299,307,1809,1823]" fontSize="6" pageId="17" pageNumber="118">1</subScript>
-P
<subScript id="17B0991409533E29C7231074FED05275" attach="left" box="[332,340,1809,1823]" fontSize="6" pageId="17" pageNumber="118">3</subScript>
and UPL.
<taxonomicName id="4C34E0D209533E29C7BF1066FDB25271" authorityName="Dyck" authorityYear="2014" box="[464,566,1794,1819]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C7BF1066FDB25271" box="[464,566,1794,1819]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
is significantly larger than
<taxonomicName id="4C34E0D209533E29C5011066FB945271" box="[878,1040,1794,1819]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C5011066FB945271" box="[878,1040,1794,1819]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
for a range of characters: in males NW, OBW and ZW, and in females at OBW, TC and BuL (
<tableCitation id="C6B6AEEA09533E29C543104CFC075228" box="[812,899,1833,1858]" captionStart="TABLE 5" captionStartId="13.[147,241,631,655]" captionTargetId="figure-113@13.[147,1770,777,1673]" captionTargetPageId="13" captionText="TABLE 5 ANOVA F-statistics (top two lines) for variation at each of the measured variables among all antechinus species and subspecies. Subsequent rows show significance values for ANOVA Post-Hoc tests of Antechinus arktos paired with each of its 12 congeners, for each measured variable. Shaded cells are significant at p=0.05, unshaded cells are not significant." pageId="17" pageNumber="118">Table 5</tableCitation>
).
</paragraph>
<paragraph id="8B8B9B5109533E29C6A81028FF62529C" blockId="17.[151,1437,1503,2038]" pageId="17" pageNumber="118">
Other Comments:
<taxonomicName id="4C34E0D209533E29C7FB102BFE7D520C" authorityName="Dyck" authorityYear="2014" box="[404,505,1869,1894]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C7FB102BFE7D520C" box="[404,505,1869,1894]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
occurs in south-east Qld and north-east NSW in areas of high elevation and rainfall on the Tweed Volcano caldera, whereas
<taxonomicName id="4C34E0D209533E29C40F1014FC8752E0" box="[608,771,1905,1930]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C40F1014FC8752E0" box="[608,771,1905,1930]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
appears to have a more subtropical distribution, occurring from Gympie south-east Qld south to Border Ranges NP in north-east NSW. The two species conceivably co-occur. Genetics: uncorrected pairwise difference at the mitochondrial gene CytB between
<taxonomicName id="4C34E0D209533E29C22D10DCFB2C52B8" authorityName="Dyck" authorityYear="2014" box="[1090,1192,1977,2002]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309533E29C22D10DCFB2C52B8" box="[1090,1192,1977,2002]" italics="true" pageId="17" pageNumber="118">A. arktos</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34E0D209533E29C28F10DCFA0652B8" box="[1248,1410,1977,2002]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="subtropicus">
<emphasis id="B940474309533E29C28F10DCFA0652B8" box="[1248,1410,1977,2002]" italics="true" pageId="17" pageNumber="118">A. subtropicus</emphasis>
</taxonomicName>
is 16.3%.
</paragraph>
<paragraph id="8B8B9B5109503E2AC6F817FDFE0855DA" blockId="18.[151,1436,151,395]" box="[151,396,151,176]" pageId="18" pageNumber="119">
<emphasis id="B940474309503E2AC6F817FDFE0855DA" box="[151,396,151,176]" italics="true" pageId="18" pageNumber="119">Bivariate Scatterplots</emphasis>
</paragraph>
<paragraph id="8B8B9B5109503E2AC6A817D9FEFC54E0" blockId="18.[151,1436,151,395]" pageId="18" pageNumber="119">
A range of scatterplots are shown for dental variables differentiating
<taxonomicName id="4C34E0D209503E2AC5BE17D9FBB355BF" authorityName="Dyck" authorityYear="2014" box="[977,1079,188,213]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC5BE17D9FBB355BF" box="[977,1079,188,213]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
from the three subspecies of
<taxonomicName id="4C34E0D209503E2AC3EB17D9FE8B559D" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">
<emphasis id="B940474309503E2AC3EB17D9FE8B559D" italics="true" pageId="18" pageNumber="119">A. swainsonii</emphasis>
</taxonomicName>
(
<figureCitation id="130F87D409503E2AC77117BAFE0E5592" box="[286,394,223,248]" captionStart="FIGURE 6" captionStartId="18.[151,249,1961,1983]" captionTargetBox="[224,1350,1097,1925]" captionTargetId="figure-457@18.[220,1352,1093,1927]" captionTargetPageId="18" captionText="FIGURE 6. Scatterplot of anterior palatal vacuity length (APV) versus inter palatal vacuity distance (IPV) measures for male A. arktos (closed circles), A. s. swainsonii (open triangles) and A. s. mimetes (open circles)." figureDoi="http://doi.org/10.5281/zenodo.5045746" httpUri="https://zenodo.org/record/5045746/files/figure.png" pageId="18" pageNumber="119">Figures 6</figureCitation>
and
<figureCitation id="130F87D409503E2AC7AC1785FE55559D" box="[451,465,224,247]" captionStart="FIGURE 7" captionStartId="19.[151,249,1031,1053]" captionTargetBox="[211,1337,185,1013]" captionTargetId="figure-80@19.[208,1340,181,1015]" captionTargetPageId="19" captionText="FIGURE 7. Scatterplot of anterior palatal vacuity length (APV) versus inter palatal vacuity distance (IPV) measures for female A. arktos (closed circles), A. s. swainsonii (open triangles) and A. s. mimetes (open circles)." figureDoi="http://doi.org/10.5281/zenodo.5045750" httpUri="https://zenodo.org/record/5045750/files/figure.png" pageId="18" pageNumber="119">7</figureCitation>
are
<taxonomicName id="4C34E0D209503E2AC46C1785FDED5592" authorityName="Dyck" authorityYear="2014" box="[515,617,223,248]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC46C1785FDED5592" box="[515,617,223,248]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
versus
<emphasis id="B940474309503E2AC4AE1785FCF0559D" box="[705,884,224,248]" italics="true" pageId="18" pageNumber="119">
A. s.
<taxonomicName id="4C34E0D209503E2AC4931785FCF0559D" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[764,884,224,247]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">swainsonii</taxonomicName>
</emphasis>
and
<emphasis id="B940474309503E2AC5C31785FBC7559D" box="[940,1091,224,248]" italics="true" pageId="18" pageNumber="119">A. s. mimetes</emphasis>
;
<figureCitation id="130F87D409503E2AC23D17BAFB385592" box="[1106,1212,223,248]" captionStart="FIGURE 8" captionStartId="19.[151,249,1975,1997]" captionTargetBox="[219,1348,1123,1951]" captionTargetId="figure-84@19.[215,1351,1119,1953]" captionTargetPageId="19" captionText="FIGURE 8. Scatterplot of anterior palatal vacuity length (APV) versus upper molar 2 width (M2W) measures for male A. arktos (closed circles) and A. s. insulanus (closed triangles)." figureDoi="http://doi.org/10.5281/zenodo.5045752" httpUri="https://zenodo.org/record/5045752/files/figure.png" pageId="18" pageNumber="119">Figures 8</figureCitation>
and
<figureCitation id="130F87D409503E2AC29A17BAFA875592" box="[1269,1283,223,248]" captionStart="FIGURE 9" captionStartId="20.[151,249,1034,1056]" captionTargetBox="[224,1344,189,1010]" captionTargetId="figure-91@20.[219,1348,185,1012]" captionTargetPageId="20" captionText="FIGURE 9. Scatterplot of anterior palatal vacuity length (APV) versus upper molar 2 width (M2W) measures for female A. arktos (closed circles) and A. s. insulanus (closed triangles)." figureDoi="http://doi.org/10.5281/zenodo.5045754" httpUri="https://zenodo.org/record/5045754/files/figure.png" pageId="18" pageNumber="119">9</figureCitation>
are
<taxonomicName id="4C34E0D209503E2AC35A1785FA1F5592" authorityName="Dyck" authorityYear="2014" box="[1333,1435,223,248]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC35A1785FA1F5592" box="[1333,1435,223,248]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
versus
<emphasis id="B940474309503E2AC6891661FE0F5477" box="[230,395,260,285]" italics="true" pageId="18" pageNumber="119">A. s. insulanus</emphasis>
). Compared with
<taxonomicName id="4C34E0D209503E2AC4361661FD745476" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[601,752,260,284]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">
<emphasis id="B940474309503E2AC4361661FD745476" box="[601,752,260,284]" italics="true" pageId="18" pageNumber="119">A. swainsonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34E0D209503E2AC4901661FCE15477" authorityName="Dyck" authorityYear="2014" box="[767,869,260,285]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC4901661FCE15477" box="[767,869,260,285]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
is most easily differentiated from
<emphasis id="B940474309503E2AC2841661FA185476" box="[1259,1436,260,284]" italics="true" pageId="18" pageNumber="119">
A. s.
<taxonomicName id="4C34E0D209503E2AC34B1660FA185476" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[1316,1436,261,284]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">swainsonii</taxonomicName>
</emphasis>
and
<emphasis id="B940474309503E2AC6A9164DFEDE5455" box="[198,346,296,320]" italics="true" pageId="18" pageNumber="119">A. s. mimetes</emphasis>
at numerous dental characters, particularly those associated with palatal vacuity length, skull and tooth breadth.
<taxonomicName id="4C34E0D209503E2AC753162AFE26540D" authorityName="Dyck" authorityYear="2014" box="[316,418,334,359]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC753162AFE26540D" box="[316,418,334,359]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
has smaller holes in the front of the skull and tends to have smaller 2
<superScript id="7C41361909503E2AC2D3162EFB4F5433" attach="left" box="[1212,1227,331,345]" fontSize="6" pageId="18" pageNumber="119">nd</superScript>
upper molar teeth than
<emphasis id="B940474309503E2AC6A11616FEF554E1" box="[206,369,370,395]" italics="true" pageId="18" pageNumber="119">A. s. insulanus</emphasis>
.
</paragraph>
<paragraph id="8B8B9B5109503E2AC6F816D0FEBE54A4" blockId="18.[151,1436,437,1038]" box="[151,314,437,462]" pageId="18" pageNumber="119">DFA and CVA</paragraph>
<paragraph id="8B8B9B5109503E2AC6A816BCFE765164" blockId="18.[151,1436,437,1038]" pageId="18" pageNumber="119">
Discriminant Function Analysis (DFA) indicated that 100% of females and males were clustered into the 4 taxon groups (
<taxonomicName id="4C34E0D209503E2AC755169BFE24577C" authorityName="Dyck" authorityYear="2014" box="[314,416,509,534]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC755169BFE24577C" box="[314,416,509,534]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
,
<emphasis id="B940474309503E2AC7C1169BFDE4577F" box="[430,608,510,533]" italics="true" pageId="18" pageNumber="119">
A. s.
<taxonomicName id="4C34E0D209503E2AC787169BFDE4577F" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[488,608,510,533]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">swainsonii</taxonomicName>
</emphasis>
,
<emphasis id="B940474309503E2AC401169BFC80577F" box="[622,772,510,533]" italics="true" pageId="18" pageNumber="119">A. s. mimetes</emphasis>
,
<emphasis id="B940474309503E2AC57D169BFC33577C" box="[786,951,509,534]" italics="true" pageId="18" pageNumber="119">A. s. insulanus</emphasis>
) correctly (posterior probabilities all equal to 1.00, not shown), based on the Mahalanobis distance of each individual from the centroid of the
<emphasis id="B940474309503E2AC2821547FAC75753" box="[1261,1347,546,569]" italics="true" pageId="18" pageNumber="119">a priori</emphasis>
species group. For CVA, 100% of the variation in dental characters was explained in the first three canonical roots for males and females. Variation was very well resolved for both sexes (although more clearly for females than males), as eigenvalues for the first three canonical roots were well above 1 (males: root 1 = 55.9; root 2 = 16.0; root 3 = 11.3; females: root 1 = 366.9; root 2 = 282.9; root 3 = 6.5) and about two-thirds of the variation was explained in the first root (67%) for males, whereas just over half (56%) was explained in the first root for females. However, cumulatively the first two roots explained 86% of variation in males and a massive 99% in females.
<figureCitation id="130F87D409503E2AC293159CFA185678" box="[1276,1436,761,786]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="20.[151,249,1978,2000]" captionStartId-1="21.[151,249,1158,1180]" captionTargetBox-0="[217,1335,1133,1954]" captionTargetBox-1="[156,1432,197,1135]" captionTargetId-0="figure-95@20.[212,1338,1129,1956]" captionTargetId-1="figure-436@21.[151,1435,193,1137]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURE 10. Scatterplot of canonical variates scores (roots 1 and 2) for males of A. arktos (closed circles) and the three subspecies of A. swainsonii: A. s. swainsonii (open triangles), A. s. mimetes (open circles) and A. s. insulanus (closed triangles)." captionText-1="FIGURE 11. Scatterplot of canonical variates scores (roots 1 and 2) for females of A. arktos (closed circles) and the three subspecies of A. swainsonii: A. s. swainsonii (open triangles), A. s. mimetes (open circles) and A. s. insulanus (closed triangles)." figureDoi-0="http://doi.org/10.5281/zenodo.5045758" figureDoi-1="http://doi.org/10.5281/zenodo.5045764" httpUri-0="https://zenodo.org/record/5045758/files/figure.png" httpUri-1="https://zenodo.org/record/5045764/files/figure.png" pageId="18" pageNumber="119">Figures 1011</figureCitation>
show scatterplots of canonical roots 12, for males and females, respectively; all four taxa are very tightly clustered within their taxon and very well separated between taxa, for both sexes, but particularly females. Canonical analysis suggests that
<taxonomicName id="4C34E0D209503E2AC7FF1403FE725614" authorityName="Dyck" authorityYear="2014" box="[400,502,869,894]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC7FF1403FE725614" box="[400,502,869,894]" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
most closely resembles
<emphasis id="B940474309503E2AC5651403FC1B5617" box="[778,927,870,893]" italics="true" pageId="18" pageNumber="119">A. s. mimetes</emphasis>
and
<emphasis id="B940474309503E2AC5B91403FBFE5614" box="[982,1146,869,894]" italics="true" pageId="18" pageNumber="119">A. s. insulanus</emphasis>
, and to a lesser degree
<emphasis id="B940474309503E2AC3EB1403FEAD56CB" italics="true" pageId="18" pageNumber="119">
A. s.
<taxonomicName id="4C34E0D209503E2AC6DE14EFFEAD56CB" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[177,297,906,929]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">swainsonii</taxonomicName>
</emphasis>
, which reflects their relative geographical proximities. This is a reflection of the large and robust
<taxonomicName id="4C34E0D209503E2AC3EB14EFFF5956AC" authorityName="Dyck" authorityYear="2014" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="arktos">
<emphasis id="B940474309503E2AC3EB14EFFF5956AC" italics="true" pageId="18" pageNumber="119">A. arktos</emphasis>
</taxonomicName>
skulls, compared with the generally narrow and slender
<emphasis id="B940474309503E2AC5CC14CBFBE056AF" box="[931,1124,942,965]" italics="true" pageId="18" pageNumber="119">
A. s.
<taxonomicName id="4C34E0D209503E2AC58314CBFBE056AF" baseAuthorityName="Waterhouse" baseAuthorityYear="1840" box="[1004,1124,942,965]" class="Mammalia" family="Dasyuridae" genus="Antechinus" kingdom="Animalia" order="Dasyuromorphia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="swainsonii">swainsonii</taxonomicName>
</emphasis>
skulls from
<collectingRegion id="49F055B309503E2AC36514C8FAF156AC" box="[1290,1397,941,966]" country="Australia" name="Tasmania" pageId="18" pageNumber="119">Tasmania</collectingRegion>
as differentiated from the generally more robust
<emphasis id="B940474309503E2AC4D214B7FCD95683" box="[701,861,978,1001]" italics="true" pageId="18" pageNumber="119">A. s. mimetes</emphasis>
from south-east
<collectingCountry id="F323DBC109503E2AC24214B4FB105680" box="[1069,1172,977,1002]" name="Australia" pageId="18" pageNumber="119">Australia</collectingCountry>
and particularly
<emphasis id="B940474309503E2AC30B14B7FE875164" italics="true" pageId="18" pageNumber="119">A. s. insulanus</emphasis>
from the Grampians.
</paragraph>
</subSubSection>
</treatment>
</document>