treatments-xml/data/03/9D/14/039D1412FFF9FFE64F9CCAF8FE3DFE99.xml

<document id="1E6767FE23B0153A583A1BADCE7D020A" ID-CLB-Dataset="171430" ID-DOI="10.11646/zootaxa.5196.4.5" ID-GBIF-Dataset="2c92fb8c-f0da-4e16-a919-c5f6c02c8472" ID-ISSN="1175-5326" ID-Zenodo-Dep="7235759" ID-ZooBank="9023DBC4-D8CC-4C45-ADD7-9C302B197BF6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666363031432" checkinUser="plazi" docAuthor="Nguyen, Sang Ngoc, Le, Manh Van, Vo, Thi-Dieu-Hien &amp; Murphy, Robert W." docDate="2022" docId="039D1412FFF9FFE64F9CCAF8FE3DFE99" docLanguage="en" docName="zootaxa.5169.4.5.pdf" docOrigin="Zootaxa 5196 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligodon tuani Nguyen &amp; Le &amp; Vo &amp; Murphy 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="564" masterDocId="FFA46C6AFFFBFFEF4F0BCE0AFFA0FFFC" masterDocTitle="A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Langbian Plateau, Vietnam" masterLastPageNumber="566" masterPageNumber="555" pageNumber="557" updateTime="1699408151037" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="34C943F5A250854B47329B580E85DD9A">A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Langbian Plateau, Vietnam</mods:title>
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<mods:namePart id="741242F6E80F2A6ECD3DC338AB075D67">Nguyen, Sang Ngoc</mods:namePart>
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<mods:affiliation id="7BAE77B89EC91474D60163513A98575B">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Dist. 3, Ho Chi Minh City, Vietnam. ngocsangitb @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1955 - 3329 &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="4D597EE67B6A6EFB906B791B07355023">Le, Manh Van</mods:namePart>
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<mods:affiliation id="F31D60C5B302A441578E32F5F8BA49A0">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Dist. 3, Ho Chi Minh City, Vietnam. ngocsangitb @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1955 - 3329 &amp; leemanh 94 @ gmail. com; https: // orcid. org / 0000 - 0002 - 7401 - 4830</mods:affiliation>
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<mods:namePart id="E00B13A0BADC1B06D8AF43AEB2F2418D">Vo, Thi-Dieu-Hien</mods:namePart>
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<mods:namePart id="7837821C5C38F8E3C8B9A6F7D8927204">Murphy, Robert W.</mods:namePart>
<mods:affiliation id="1E7A7FA8E7AB69708EBFC4CE3C06595D">Reptilia Zoo and Education Centre, 2501 Rutherford Rd Vaughan, ON, Canada L 4 K 2 N 6.</mods:affiliation>
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<taxonomicName id="4C34DE87FFF9FFED4F9CCAF8FEEFFAF1" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[151,335,1266,1293]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="557" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
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(
<figureCitation id="130FB981FFF9FFED4F94CB1DFEAAFACD" box="[159,266,1303,1329]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[152,255,1595,1619]" captionStartId-1="4.[152,255,1476,1500]" captionStartId-2="5.[152,255,1680,1704]" captionTargetBox-0="[151,1435,740,1571]" captionTargetBox-1="[153,1434,489,1453]" captionTargetBox-2="[151,1435,201,1655]" captionTargetId-0="figure-277@3.[151,1435,740,1571]" captionTargetId-1="figure-171@4.[153,1434,489,1453]" captionTargetId-2="figure-19@5.[151,1435,201,1656]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Holotype of Oligodon tuani sp. nov. in life, ITBCZ 7899." captionText-1="FIGURE 3. Head and hemipenis of the holotype of Oligodon tuani sp. nov. A–C, lateral, dorsal, and ventral views of head, respectively; D–F, line drawings of the head, scale bar 5 mm; G&amp;H, everted hemipenis of fresh specimen and in preservative, respectively." captionText-2="FIGURE 4. Paratypes of Oligodon tuani sp. nov. and habitat of the new species. A&amp;B, dorsal and ventral views of ITBCZ 7944; C, ITBCZ 7868; D, ITBCZ 8215; E, pine forest in Da Lat City, Lam Dong Province, Vietnam (habitat photo by T.M. Nguyen)." figureDoi-0="http://doi.org/10.5281/zenodo.7235765" figureDoi-1="http://doi.org/10.5281/zenodo.7235773" figureDoi-2="http://doi.org/10.5281/zenodo.7235777" httpUri-0="https://zenodo.org/record/7235765/files/figure.png" httpUri-1="https://zenodo.org/record/7235773/files/figure.png" httpUri-2="https://zenodo.org/record/7235777/files/figure.png" pageId="2" pageNumber="557">Figs. 2–4</figureCitation>
)
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<materialsCitation id="3B5CAF59FFF9FFED4F9CCB55FE81FA3D" ID-GBIF-Occurrence="3944669306" collectingDate="2020-09-15" collectionCode="ITBCZ" collectorName="Da Lat City" country="Vietnam" elevation="1490" latitude="11.948334" location="Lam Dong Province" longLatPrecision="21" longitude="108.41667" pageId="2" pageNumber="557" specimenCode="ITBCZ 7899" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
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. 
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, subadult male, collected near Cam Ly Stream, 
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, 
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, 
<collectingCountry id="F323E594FFF9FFED4A35CB55FA3DFA85" box="[1342,1437,1375,1401]" name="Vietnam" pageId="2" pageNumber="557">Vietnam</collectingCountry>
(coordinates 
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, 
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; 
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elevation 
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a.s.l.
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) by Tuan Minh Nguyen on 
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(
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&amp; 
<figureCitation id="130FB981FFF9FFED4E0CCBADFEB4FA3D" box="[263,276,1447,1473]" captionStart="FIGURE 3" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[153,1434,489,1453]" captionTargetId="figure-171@4.[153,1434,489,1453]" captionTargetPageId="4" captionText="FIGURE 3. Head and hemipenis of the holotype of Oligodon tuani sp. nov. A–C, lateral, dorsal, and ventral views of head, respectively; D–F, line drawings of the head, scale bar 5 mm; G&amp;H, everted hemipenis of fresh specimen and in preservative, respectively." figureDoi="http://doi.org/10.5281/zenodo.7235773" httpUri="https://zenodo.org/record/7235773/files/figure.png" pageId="2" pageNumber="557">3</figureCitation>
).
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<materialsCitation id="3B5CAF59FFF9FFED4FCCCBC1FE95F9D1" ID-GBIF-Occurrence="3944669303" collectingDate="2020-10-15" collectionCode="ITBCZ" collectorName="Cam Ly Road &amp; Da Lat City" country="Vietnam" elevation="1431" latitude="11.96" location="Tuan Minh Nguyen" longLatPrecision="21" longitude="108.39166" pageId="2" pageNumber="557" specimenCode="ITBCZ 7944" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
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<typeStatus id="548F1BA6FFF9FFED4FCCCBC1FE9FFA19" box="[199,319,1483,1509]" pageId="2" pageNumber="557" type="paratype">Paratypes</typeStatus>
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. 
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, adult male, collected on 
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<collectingDate id="EFCE7A2CFFF9FFED4C2FCBC0FC4CFA19" box="[804,1004,1482,1509]" pageId="2" pageNumber="557" value="2020-10-15">15 October 2020</collectingDate>
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by 
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, coordinates 
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, 
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, 
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elevation 
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a.s.l.
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, from 
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, vicinity of 
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, 
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, 
<collectingCountry id="F323E594FFF9FFED4FD8C819FE95F9D1" box="[211,309,1555,1581]" name="Vietnam" pageId="2" pageNumber="557">Vietnam</collectingCountry>
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; 
<materialsCitation id="3B5CAF59FFF9FFED4E4AC819FE0FF989" ID-GBIF-Occurrence="3944669302" collectingDate="2020-09-12" collectionCode="ITBCZ" collectorName="Da Sar &amp; Da Lat City" country="Vietnam" elevation="1520" latitude="12.017222" location="Tuan Minh Nguyen" longLatPrecision="21" longitude="108.491104" municipality="Lac Duong District" pageId="2" pageNumber="557" specimenCode="ITBCZ 7868" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCode id="DB920D7FFFF9FFED4E4AC819FE74F9D1" box="[321,468,1555,1581]" collectionCode="ITBCZ" pageId="2" pageNumber="557">ITBCZ 7868</specimenCode>
, subadult female, collected on 
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<collectingDate id="EFCE7A2CFFF9FFED4C25C819FBA8F9D1" box="[814,1032,1555,1581]" pageId="2" pageNumber="557" value="2020-09-12">12 September 2020</collectingDate>
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by 
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, coordinates 
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, 
<geoCoordinate id="EE00C3C3FFF9FFED4E24C83DFE66F9AD" box="[303,454,1591,1617]" degrees="108" direction="east" minutes="29" orientation="longitude" pageId="2" pageNumber="557" precision="15" seconds="28" value="108.491104">108°29’28”E</geoCoordinate>
; 
<elevation id="00194237FFF9FFED4ED9C83DFD71F9AD" box="[466,721,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="557" unit="m" value="1520.0">
elevation 
<quantity id="4CCC08E1FFF9FFED4D34C83CFD35F9AD" box="[575,661,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="557" unit="m" value="1520.0">1520 m</quantity>
a.s.l.
</elevation>
, from 
<collectorName id="26C1C0D2FFF9FFED4C17C83DFCCAF9AD" box="[796,874,1591,1617]" pageId="2" pageNumber="557">Da Sar</collectorName>
, 
<collectingMunicipality id="6BEF3F7EFFF9FFED4C7DC83DFBF1F9AD" box="[886,1105,1591,1617]" pageId="2" pageNumber="557">Lac Duong District</collectingMunicipality>
, vicinity of 
<collectorName id="26C1C0D2FFF9FFED4BD3C83DFAFDF9AD" box="[1240,1373,1591,1617]" pageId="2" pageNumber="557">Da Lat City</collectorName>
, 
<collectingRegion id="49F06BE6FFF9FFED4A62C83DFEE2F989" country="Vietnam" name="Lam Dong" pageId="2" pageNumber="557">Lam Dong Province</collectingRegion>
, 
<collectingCountry id="F323E594FFF9FFED4E46C851FE0FF989" box="[333,431,1627,1653]" name="Vietnam" pageId="2" pageNumber="557">Vietnam</collectingCountry>
</materialsCitation>
; and 
<materialsCitation id="3B5CAF59FFF9FFED4EE1C851FE97F965" ID-GBIF-Occurrence="3944669304" collectionCode="ITBCZ" country="Vietnam" location="Lam Dong Province" pageId="2" pageNumber="557" specimenCode="ITBCZ 8215" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCode id="DB920D7FFFF9FFED4EE1C851FDDCF989" box="[490,636,1626,1653]" collectionCode="ITBCZ" pageId="2" pageNumber="557">ITBCZ 8215</specimenCode>
, adult female, collected from Ward no. 10, Dalat City, 
<collectingRegion id="49F06BE6FFF9FFED4BD4C851FF68F965" country="Vietnam" name="Lam Dong" pageId="2" pageNumber="557">Lam Dong Province</collectingRegion>
, 
<collectingCountry id="F323E594FFF9FFED4FDEC875FE97F965" box="[213,311,1663,1689]" name="Vietnam" pageId="2" pageNumber="557">Vietnam</collectingCountry>
</materialsCitation>
, coordinates 
<geoCoordinate id="EE00C3C3FFF9FFED4EC5C874FDF9F965" box="[462,601,1662,1689]" degrees="11" direction="north" minutes="55" orientation="latitude" pageId="2" pageNumber="557" precision="15" seconds="56" value="11.932222">11°55’56”N</geoCoordinate>
, 
<geoCoordinate id="EE00C3C3FFF9FFED4D6DC874FD5CF965" box="[614,764,1662,1689]" degrees="108" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="557" precision="15" seconds="58" value="108.44945">108°26’58”E</geoCoordinate>
, elevation 
<quantity id="4CCC08E1FFF9FFED4C72C875FC70F965" box="[889,976,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="557" unit="m" value="1440.0">1440 m</quantity>
a.s.l., on 
<date id="FF8A83C4FFF9FFED4B49C875FB51F965" box="[1090,1265,1663,1689]" pageId="2" pageNumber="557" value="2022-03-24">24 March 2022</date>
by Tuan Minh Nguyen (
<figureCitation id="130FB981FFF9FFED4FF4C8A9FEE5F941" box="[255,325,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[152,255,1680,1704]" captionTargetBox="[151,1435,201,1655]" captionTargetId="figure-19@5.[151,1435,201,1656]" captionTargetPageId="5" captionText="FIGURE 4. Paratypes of Oligodon tuani sp. nov. and habitat of the new species. A&amp;B, dorsal and ventral views of ITBCZ 7944; C, ITBCZ 7868; D, ITBCZ 8215; E, pine forest in Da Lat City, Lam Dong Province, Vietnam (habitat photo by T.M. Nguyen)." figureDoi="http://doi.org/10.5281/zenodo.7235777" httpUri="https://zenodo.org/record/7235777/files/figure.png" pageId="2" pageNumber="557">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EF68FFFF9FFED4FCCC8CCFD78F821" pageId="2" pageNumber="557" type="diagnosis">
<paragraph id="8B8BA504FFF9FFED4FCCC8CCFD78F821" blockId="2.[151,1437,1375,2013]" pageId="2" pageNumber="557">
<emphasis id="B9407916FFF9FFED4FCCC8CCFE9AF91C" bold="true" box="[199,314,1734,1760]" pageId="2" pageNumber="557">Diagnosis</emphasis>
. 
<taxonomicName id="4C34DE87FFF9FFED4E43C8CDFE53F91C" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[328,499,1735,1761]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="557" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFF9FFED4E43C8CDFE53F91C" box="[328,499,1735,1761]" italics="true" pageId="2" pageNumber="557">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFF9FFED4EF1C8CDFDF4F91D" bold="true" box="[506,596,1735,1761]" pageId="2" pageNumber="557">
<taxonomicNameLabel id="A273C46DFFF9FFED4EF1C8CDFDF4F91D" box="[506,596,1735,1761]" pageId="2" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from its congeners by a combination of the following morphological characters: (1) large size in adults (TL ≤ 
<quantity id="4CCC08E1FFF9FFED4C12C8E1FCDCF8F9" box="[793,892,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="8.879999999999999" pageId="2" pageNumber="557" unit="mm" value="888.0">888 mm</quantity>
); (2) 19 dorsal scale rows at neck and midbody and 15 rows before vent; (3) TaL/TL ratio 
<quantity id="4CCC08E1FFF9FFED4D77C905FD7AF8D5" box="[636,730,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="4.6736" pageId="2" pageNumber="557" unit="in" value="0.184">0.184 in</quantity>
adult male and 
<quantity id="4CCC08E1FFF9FFED4C9AC905FC4EF8D5" box="[913,1006,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="3.1241999999999996" pageId="2" pageNumber="557" unit="in" value="0.123">0.123 in</quantity>
adult female; (4) ventrals 
<quantity id="4CCC08E1FFF9FFED4A17C905FA3CF8D5" box="[1308,1436,1807,1833]" metricMagnitude="0" metricUnit="m" metricValue="4.4704" metricValueMax="4.5466" metricValueMin="4.3942" pageId="2" pageNumber="557" unit="in" value="176.0" valueMax="179.0" valueMin="173.0">173–179 in</quantity>
males and 
<quantity id="4CCC08E1FFF9FFED4E04C939FE2EF8B1" box="[271,398,1843,1869]" metricMagnitude="0" metricUnit="m" metricValue="4.8260000000000005" metricValueMax="4.902200000000001" metricValueMin="4.7498000000000005" pageId="2" pageNumber="557" unit="in" value="190.0" valueMax="193.0" valueMin="187.0">187–193 in</quantity>
females; (5) subcaudals 58 or 
<quantity id="4CCC08E1FFF9FFED4DEDC938FCBFF8B1" box="[742,799,1842,1869]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" pageId="2" pageNumber="557" unit="in" value="59.0">59 in</quantity>
males and 44 or 
<quantity id="4CCC08E1FFF9FFED4CD4C938FBB8F8B1" box="[991,1048,1842,1869]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="2" pageNumber="557" unit="in" value="45.0">45 in</quantity>
females; (6) presubocular present; (7) 8 supralabials, fourth and fifth entering orbit; (8) cloacal plate undivided; (9) 10 maxillary teeth, posterior three enlarged; (10) vertebral stripe present; (11) temporal streak present but faint and interrupted; (12) long and deeply forked hemipenes, extending to 25 
<superScript id="7C41084CFFF9FFED4D12C994FD86F850" attach="left" box="[537,550,1950,1964]" fontSize="6" pageId="2" pageNumber="557">th</superScript>
subcaudal, without spines and papillae but with small and transversal folds and distal calyces; and (13) 13–15+3–4 dorsal blotches.
</paragraph>
</subSubSection>
<subSubSection id="C32EF68FFFF8FFEB4FCCCE9CFB6EF8DC" lastPageId="4" lastPageNumber="559" pageId="3" pageNumber="558" type="description">
<paragraph id="8B8BA504FFF8FFEC4FCCCE9CFCC2FF04" blockId="3.[151,1437,150,717]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCCE9CFE45FF4C" bold="true" box="[199,485,150,176]" pageId="3" pageNumber="558">
Description of 
<typeStatus id="548F1BA6FFF8FFEC4E7CCE9CFE40FF4C" box="[375,480,150,176]" pageId="3" pageNumber="558" type="holotype">holotype</typeStatus>
.
</emphasis>
Subadult male, total length 
<quantity id="4CCC08E1FFF8FFEC4C29CE9CFC22FF4D" box="[802,898,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="5.85" pageId="3" pageNumber="558" unit="mm" value="585.0">585 mm</quantity>
; head ovoid, faintly distinct from the poorly defined neck; body robust and rather elongate; SVL 
<quantity id="4CCC08E1FFF8FFEC4DCECEB0FC84FF29" box="[709,804,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.75" pageId="3" pageNumber="558" unit="mm" value="475.0">475 mm</quantity>
; tail quite long (TaL/TL = 0.188), 
<quantity id="4CCC08E1FFF8FFEC4BA8CEB0FB5EFF29" box="[1187,1278,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="558" unit="mm" value="110.0">110 mm</quantity>
in length; eye small with round pupil, eye diameter about half of snout length.
</paragraph>
<paragraph id="8B8BA504FFF8FFEC4FCCCF08FEEFFDC0" blockId="3.[151,1437,150,717]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCCF08FED5FEE0" box="[199,373,258,284]" italics="true" pageId="3" pageNumber="558">Head scalation</emphasis>
. Rostral thick, triangular and curved on to upper snout surface, visible from above, pointed posteriorly, touching internasals, nasals and first supralabial on both sides; the portion of rostral visible from above shorter than its distance from frontal; supralabials 8/8, the 4 
<superScript id="7C41084CFFF8FFEC4C26CF40FC9AFEA4" attach="left" box="[813,826,330,344]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
and 5 
<superScript id="7C41084CFFF8FFEC4C76CF40FC2AFEA4" attach="left" box="[893,906,330,344]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
bordering eye; 1/1 loreal; 1/1 presubocular; 1/1 preocular; 2/2 postoculars; nasal divided vertically; 2+2/2+2 temporals, anterior ones elongated; prefrontal wider than long and larger than internasal; frontal large and hexagonal, slightly longer than wide (
<quantity id="4CCC08E1FFF8FFEC4BA0CF98FAA4FE51" box="[1195,1284,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="558" unit="mm" value="5.7">5.7 mm</quantity>
vs. 
<quantity id="4CCC08E1FFF8FFEC4A3FCF98FA30FE50" box="[1332,1424,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="558" unit="mm" value="4.5">4.5 mm</quantity>
); parietals larger than the frontal, bordered laterally by the upper temporals and anteriorly by frontal, supraoculars and postoculars; 9/9 infralabials, first pair medially in contact with each other, first four in contact with anterior chin shield, the 5 
<superScript id="7C41084CFFF8FFEC4E15CFF4FE8BFDF0" attach="left" box="[286,299,510,524]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
largest and touching posterior chin shield; two pairs of gular scales between posterior chin shields and preventral scale.
</paragraph>
<paragraph id="8B8BA504FFF8FFEC4FCCCC4CFE94FD31" blockId="3.[151,1437,150,717]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCCC4CFECEFD9C" box="[199,366,582,608]" italics="true" pageId="3" pageNumber="558">Body scalation</emphasis>
. Dorsal scales smooth, in 19–19–15 rows; scale row reductions from 19 to 17 at ventrals 97 and 98 and from 17 to 15 at ventrals 124 and 132; vertebral scales similar to other dorsal scales in size and shape; ventrals 173 (plus 1 preventral), angulate; cloacal plate entire; subcaudals 59, all paired; terminal caudal scale forming a pointed cap.
</paragraph>
<caption id="DF4BF58CFFF8FFEC4F93C831FCC8F9AF" ID-DOI="http://doi.org/10.5281/zenodo.7235765" ID-Zenodo-Dep="7235765" box="[152,872,1595,1619]" httpUri="https://zenodo.org/record/7235765/files/figure.png" pageId="3" pageNumber="558" startId="3.[152,255,1595,1619]" targetBox="[151,1435,740,1571]" targetPageId="3">
<paragraph id="8B8BA504FFF8FFEC4F93C831FCC8F9AF" blockId="3.[152,872,1595,1619]" box="[152,872,1595,1619]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4F93C831FEB8F9AF" bold="true" box="[152,280,1595,1619]" pageId="3" pageNumber="558">FIGURE 2.</emphasis>
Holotype of 
<taxonomicName id="4C34DE87FFF8FFEC4E94C831FD99F9AF" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[415,569,1595,1619]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="558" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFF8FFEC4E94C831FD99F9AF" box="[415,569,1595,1619]" italics="true" pageId="3" pageNumber="558">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFF8FFEC4D34C831FD30F9AF" bold="true" box="[575,656,1595,1619]" pageId="3" pageNumber="558">
<taxonomicNameLabel id="A273C46DFFF8FFEC4D34C831FD30F9AF" box="[575,656,1595,1619]" pageId="3" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life, ITBCZ 7899.
</paragraph>
</caption>
<paragraph id="8B8BA504FFF8FFEC4FCCC888FEC2F943" blockId="3.[151,1437,1665,2016]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCC888FE91F967" box="[199,305,1666,1691]" italics="true" pageId="3" pageNumber="558">Dentition</emphasis>
. Maxillary teeth 10, curved posteriorly, smaller and shorter anteriorly; posterior three being enlarged and blade-shaped.
</paragraph>
<paragraph id="8B8BA504FFF8FFEC4FCCC8C0FD96F8AC" blockId="3.[151,1437,1665,2016]" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCC8C0FE9FF91F" box="[199,319,1738,1763]" italics="true" pageId="3" pageNumber="558">Hemipenis</emphasis>
. Hemipenis long and thin with two lobes, deeply forked at the level of 6 
<superScript id="7C41084CFFF8FFEC4B7AC8C3FBDEF92B" attach="left" box="[1137,1150,1737,1751]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
SC and tip of each organ reaching the SC 25, without spines and papillae; each lobe with numerous smooth, small and transversal folds, with a deep diagonal groove; smooth hexagonal calyces on the distal part of the organ. The sulcus is prominent and divided at the level of the fork (
<figureCitation id="130FB981FFF8FFEC4EEFC93FFD88F8AC" box="[484,552,1845,1872]" captionStart="FIGURE 3" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[153,1434,489,1453]" captionTargetId="figure-171@4.[153,1434,489,1453]" captionTargetPageId="4" captionText="FIGURE 3. Head and hemipenis of the holotype of Oligodon tuani sp. nov. A–C, lateral, dorsal, and ventral views of head, respectively; D–F, line drawings of the head, scale bar 5 mm; G&amp;H, everted hemipenis of fresh specimen and in preservative, respectively." figureDoi="http://doi.org/10.5281/zenodo.7235773" httpUri="https://zenodo.org/record/7235773/files/figure.png" pageId="3" pageNumber="558">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B8BA504FFF8FFEB4FCCC953FC56FE99" blockId="3.[151,1437,1665,2016]" lastBlockId="4.[151,1437,150,465]" lastPageId="4" lastPageNumber="559" pageId="3" pageNumber="558">
<emphasis id="B9407916FFF8FFEC4FCCC953FEE2F88F" box="[199,322,1881,1907]" italics="true" pageId="3" pageNumber="558">Coloration</emphasis>
. In life, overall dorsal coloration brown to olive with 13 darker blotches on body and 4 on tail. A dark brown band from eye to eye through preoculars, supraoculars, frontal, prefrontals and internasals, continues over the 4 
<superScript id="7C41084CFFF8FFEC4E00C9ABFEB8F853" attach="left" box="[267,280,1953,1967]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
and 5 
<superScript id="7C41084CFFF8FFEC4E54C9ABFECCF853" attach="left" box="[351,364,1953,1967]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
supralabials ending at lower part of the 6 
<superScript id="7C41084CFFF8FFEC4C40C9ABFCF8F853" attach="left" box="[843,856,1953,1967]" fontSize="6" pageId="3" pageNumber="558">th</superScript>
supralabial. Head with another dark brown arrowshaped marking, its apex pointing forward and reaching the posterior part of frontal, backwards obliquely crossing neck and reaching the 6 
<superScript id="7C41084CFFFFFFEB4E94CE9CFE0CFF58" attach="left" box="[415,428,150,164]" fontSize="6" pageId="4" pageNumber="559">th</superScript>
ventral. Temporal streak faint and discontinuous, distinct posteriorly, reaching the level of ventrals. Numerous scattered dorsal black spots confined to edges of the body scales. Edge of each body scale and intervening skin yellowish to white. Vertebral stripe distinct, broader and more distinct on the anterior part of body and on the tail. Ventral color cream anteriorly to pinkish centrally and posteriorly with black rectangular blotches. Ventral surface of tail white to pinkish with black spots. In alcohol, the color became paler, but the pattern remains; venter cream to white with black blotches and dots, pinkish color disappears.
</paragraph>
<paragraph id="8B8BA504FFFFFFEB4FCCCF64FD34FE2D" blockId="4.[151,1437,150,465]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4FCCCF64FE9FFE74" bold="true" box="[199,319,366,392]" pageId="4" pageNumber="559">Variation.</emphasis>
<materialsCitation id="3B5CAF59FFFFFFEB4E4ECF65FA39FE75" ID-GBIF-Occurrence="3944669305" box="[325,1433,367,393]" collectionCode="ITBCZ" pageId="4" pageNumber="559" specimenCode="ITBCZ 7944" specimenCount="1" typeStatus="paratype">
<typeStatus id="548F1BA6FFFFFFEB4E4ECF65FE07FE75" box="[325,423,367,393]" pageId="4" pageNumber="559" type="paratype">Paratype</typeStatus>
<specimenCode id="DB920D7FFFFFFFEB4EA6CF65FD9CFE75" box="[429,572,367,393]" collectionCode="ITBCZ" pageId="4" pageNumber="559">ITBCZ 7944</specimenCode>
has two preoculars on each side and its presubocular is absent on the left side
</materialsCitation>
. 
<materialsCitation id="3B5CAF59FFFFFFEB4F9CCF99FD30FE2D" ID-GBIF-Occurrence="3944669301" collectionCode="ITBCZ" pageId="4" pageNumber="559" specimenCode="ITBCZ 8215" specimenCount="1" typeStatus="paratype">
<typeStatus id="548F1BA6FFFFFFEB4F9CCF99FF59FE51" box="[151,249,403,429]" pageId="4" pageNumber="559" type="paratype">Paratype</typeStatus>
<specimenCode id="DB920D7FFFFFFFEB4FF4CF99FE2FFE51" box="[255,399,402,429]" collectionCode="ITBCZ" pageId="4" pageNumber="559">ITBCZ 8215</specimenCode>
has 15 blotches on body and faint a vertebral stripe. 
<tableCitation id="C6B690BFFFFFFFEB4CDCCF99FB89FE51" box="[983,1065,403,429]" captionStart="TABLE 1" captionStartId="6.[152,244,151,175]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) and scalation of the types series of Oligodon tuani sp. nov. See Material and Methods for abbreviations." httpUri="http://table.plazi.org/id/DF4BF58CFFFDFFE94F93CE9DFEF4FF2F" pageId="4" pageNumber="559" tableUuid="DF4BF58CFFFDFFE94F93CE9DFEF4FF2F">Table 1</tableCitation>
summarizes variation in size and scalation of the 
<typeStatus id="548F1BA6FFFFFFEB4E42CFBDFE0BFE2D" box="[329,427,439,465]" pageId="4" pageNumber="559" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D326E8DFFFFFFEB4EE9CFBDFD30FE2D" box="[482,656,439,465]" pageId="4" pageNumber="559" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
</materialsCitation>
.
</paragraph>
<caption id="DF4BF58CFFFFFFEB4F93CBCEFEB7F9D8" ID-DOI="http://doi.org/10.5281/zenodo.7235773" ID-Zenodo-Dep="7235773" httpUri="https://zenodo.org/record/7235773/files/figure.png" pageId="4" pageNumber="559" startId="4.[152,255,1476,1500]" targetBox="[153,1434,489,1453]" targetPageId="4">
<paragraph id="8B8BA504FFFFFFEB4F93CBCEFEB7F9D8" blockId="4.[151,1437,1476,1572]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4F93CBCEFEB9FA20" bold="true" box="[152,281,1476,1500]" pageId="4" pageNumber="559">FIGURE 3.</emphasis>
Head and hemipenis of the holotype of 
<taxonomicName id="4C34DE87FFFFFFEB4DB1CBCEFCF5FA20" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[698,853,1476,1500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="559" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFFFFEB4DB1CBCEFCF5FA20" box="[698,853,1476,1500]" italics="true" pageId="4" pageNumber="559">Oligodon tuani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273C46DFFFFFFEB4C56CBCEFC10FA20" box="[861,944,1476,1500]" pageId="4" pageNumber="559" rank="species">
<emphasis id="B9407916FFFFFFEB4C56CBCEFC09FA20" bold="true" box="[861,937,1476,1500]" pageId="4" pageNumber="559">sp. nov</emphasis>
.
</taxonomicNameLabel>
A–C, lateral, dorsal, and ventral views of head, respectively; D–F, line drawings of the head, scale bar 5 mm; G&amp;H, everted hemipenis of fresh specimen and in preservative, respectively.
</paragraph>
</caption>
<paragraph id="8B8BA504FFFFFFEB4FCCC859FCD4F925" blockId="4.[151,1437,1618,2041]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4FCCC859FE26F990" bold="true" box="[199,390,1618,1645]" pageId="4" pageNumber="559">Natural history.</emphasis>
All specimens were collected during the day. ITBCZ 7868 and 7944 were found dead on the road and the surrounding habitat is pine forest as well as flower and vegetable plantations. ITBCZ 7899 was found near 
<collectingRegion id="49F06BE6FFFFFFEB4FC5C891FEA3F949" box="[206,259,1691,1717]" country="Myanmar" name="Chin" pageId="4" pageNumber="559">Cam</collectingRegion>
Ly Stream on the pine hill bordered by a vegetable plantation and residential area. ITBCZ 8215 was collected on a pine-covered hill and near a residential area. (
<figureCitation id="130FB981FFFFFFEB4C13C8B4FCCCF925" box="[792,876,1726,1753]" captionStart="FIGURE 4" captionStartId="5.[152,255,1680,1704]" captionTargetBox="[151,1435,201,1655]" captionTargetId="figure-19@5.[151,1435,201,1656]" captionTargetPageId="5" captionText="FIGURE 4. Paratypes of Oligodon tuani sp. nov. and habitat of the new species. A&amp;B, dorsal and ventral views of ITBCZ 7944; C, ITBCZ 7868; D, ITBCZ 8215; E, pine forest in Da Lat City, Lam Dong Province, Vietnam (habitat photo by T.M. Nguyen)." figureDoi="http://doi.org/10.5281/zenodo.7235777" httpUri="https://zenodo.org/record/7235777/files/figure.png" pageId="4" pageNumber="559">Fig. 4E</figureCitation>
)
</paragraph>
<paragraph id="8B8BA504FFFFFFEB4FCCC8E8FB6EF8DC" blockId="4.[151,1437,1618,2041]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4FCCC8E8FE16F900" bold="true" box="[199,438,1762,1788]" pageId="4" pageNumber="559">Sexual dimorphism.</emphasis>
Tail of adult male longer than that of adult female (TaL/TL = 0.184 vs. 0.123). Males have more subcaudals (58 or 59 vs. 44 or 45) but fewer ventrals (173–179 vs. 187–193) than females.
</paragraph>
</subSubSection>
<subSubSection id="C32EF68FFFFFFFEB4FCCC920FEF8F895" pageId="4" pageNumber="559" type="distribution">
<paragraph id="8B8BA504FFFFFFEB4FCCC920FEF8F895" blockId="4.[151,1437,1618,2041]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4FCCC920FEC1F8B8" bold="true" box="[199,353,1834,1860]" pageId="4" pageNumber="559">Distribution.</emphasis>
The new species is currently known only from Da Lat City and its vicinity, 
<collectingRegion id="49F06BE6FFFFFFEB4BB9C921FA38F8B8" box="[1202,1432,1834,1861]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="559">Lam Dong Province</collectingRegion>
, 
<collectingCountry id="F323E594FFFFFFEB4F9CC944FF56F894" box="[151,246,1870,1896]" name="Vietnam" pageId="4" pageNumber="559">Vietnam</collectingCountry>
(
<figureCitation id="130FB981FFFFFFEB4E0EC944FEE7F895" box="[261,327,1870,1897]" captionStart="FIGURE 1" captionStartId="1.[152,255,1667,1691]" captionTargetBox="[151,1435,450,1640]" captionTargetId="figure-168@1.[151,1435,450,1640]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the currently known distribution of Oligodon tuani sp. nov. (triangle) and O. saintgironsi (square). 1 = Da Lat City and its vicinity, Lam Dong Province, Vietnam; 2 = Trang Bom, Dong Nai Province, Vietnam; 3 = Ho Chi Minh City, Vietnam; 4 = an imprecise locality in Cambodia." figureDoi="http://doi.org/10.5281/zenodo.7235763" httpUri="https://zenodo.org/record/7235763/files/figure.png" pageId="4" pageNumber="559">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EF68FFFFFFFE64FCCC979FE3DFE99" lastPageId="9" lastPageNumber="564" pageId="4" pageNumber="559" type="etymology">
<paragraph id="8B8BA504FFFFFFEB4FCCC979FC56F804" blockId="4.[151,1437,1618,2041]" pageId="4" pageNumber="559">
<emphasis id="B9407916FFFFFFEB4FCCC979FEE8F871" bold="true" box="[199,328,1907,1933]" pageId="4" pageNumber="559">Etymology</emphasis>
. We name the new species in honor of Mr. Tuan Minh Nguyen (
<collectingRegion id="49F06BE6FFFFFFEB4B29C979FAABF870" box="[1058,1291,1906,1933]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="559">Lam Dong Province</collectingRegion>
, 
<collectingCountry id="F323E594FFFFFFEB4A1CC978FAD8F870" box="[1303,1400,1906,1932]" name="Vietnam" pageId="4" pageNumber="559">Vietnam</collectingCountry>
) in recognition of his significant contribution to us including the 
<typeStatus id="548F1BA6FFFFFFEB4C43C99DFCD8F84D" box="[840,888,1943,1969]" pageId="4" pageNumber="559">type</typeStatus>
series of the new species and many other important samples of reptiles in recent years. We recommend “Langbian kukri snake” and “Rắn khiếm Langbian” for as the common English and Vietnamese names of the new species, respectively.
</paragraph>
<caption id="DF4BF58CFFFEFFEA4F93C89AFF57F90C" ID-DOI="http://doi.org/10.5281/zenodo.7235777" ID-Zenodo-Dep="7235777" httpUri="https://zenodo.org/record/7235777/files/figure.png" pageId="5" pageNumber="560" startId="5.[152,255,1680,1704]" targetBox="[151,1435,201,1655]" targetPageId="5">
<paragraph id="8B8BA504FFFEFFEA4F93C89AFF57F90C" blockId="5.[151,1437,1680,1776]" pageId="5" pageNumber="560">
<emphasis id="B9407916FFFEFFEA4F93C89AFEBAF954" bold="true" box="[152,282,1680,1704]" pageId="5" pageNumber="560">FIGURE 4.</emphasis>
Paratypes of 
<taxonomicName id="4C34DE87FFFEFFEA4EA2C89AFDE5F95B" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[425,581,1680,1704]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFEFFEA4EA2C89AFDE5F95B" box="[425,581,1680,1704]" italics="true" pageId="5" pageNumber="560">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFEFFEA4D46C89AFD3FF954" bold="true" box="[589,671,1680,1704]" pageId="5" pageNumber="560">
<taxonomicNameLabel id="A273C46DFFFEFFEA4D46C89AFD3FF954" box="[589,671,1680,1704]" pageId="5" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and habitat of the new species. A&amp;B, dorsal and ventral views of ITBCZ 7944; C, ITBCZ 7868; D, ITBCZ 8215; E, pine forest in Da Lat City, Lam Dong Province, Vietnam (habitat photo by T.M. Nguyen).
</paragraph>
</caption>
<paragraph id="8B8BA504FFFEFFEA4FCCC914FAB3F810" blockId="5.[151,1437,1822,2028]" pageId="5" pageNumber="560">
<emphasis id="B9407916FFFEFFEA4FCCC914FEC6F8C4" bold="true" box="[199,358,1822,1848]" pageId="5" pageNumber="560">Comparisons</emphasis>
. 
<taxonomicName id="4C34DE87FFFEFFEA4E7FC914FDBEF8C4" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[372,542,1822,1848]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFEFFEA4E7FC914FDBEF8C4" box="[372,542,1822,1848]" italics="true" pageId="5" pageNumber="560">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFEFFEA4D2EC914FDDEF8C4" bold="true" box="[549,638,1822,1848]" pageId="5" pageNumber="560">
<taxonomicNameLabel id="A273C46DFFFEFFEA4D2EC914FDDEF8C4" box="[549,638,1822,1848]" pageId="5" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has deeply forked hemipenis without both papillae and spines and thus differs from the following species that have conspicuous papillae on the hemipenes: 
<taxonomicName id="4C34DE87FFFEFFEA4B45C948FA85F8A0" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1916" box="[1102,1317,1858,1884]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="barroni">
<emphasis id="B9407916FFFEFFEA4B45C948FB66F8A0" box="[1102,1222,1858,1884]" italics="true" pageId="5" pageNumber="560">O. barroni</emphasis>
(Smith)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4A3AC948FE0FF87C" authority="David, Vogel &amp; Rooijen" authorityName="David, Vogel &amp; Rooijen" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="deuvei">
<emphasis id="B9407916FFFEFFEA4A3AC948FA3CF8A0" box="[1329,1436,1858,1884]" italics="true" pageId="5" pageNumber="560">O. deuvei</emphasis>
David, Vogel &amp; Rooijen
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4EB0C96CFD6EF87C" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[443,718,1894,1920]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="B9407916FFFEFFEA4EB0C96CFD9BF87C" box="[443,571,1894,1920]" italics="true" pageId="5" pageNumber="560">O. mouhoti</emphasis>
(Boulenger)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4DD1C96CFB74F87C" authority="David, Vogel &amp; Rooijen" authorityName="David, Vogel &amp; Rooijen" box="[730,1236,1894,1920]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="pseudotaeniatus">
<emphasis id="B9407916FFFEFFEA4DD1C96CFC14F87C" box="[730,948,1894,1920]" italics="true" pageId="5" pageNumber="560">O. pseudotaeniatus</emphasis>
David, Vogel &amp; Rooijen
</taxonomicName>
, and 
<taxonomicName id="4C34DE87FFFEFFEA4A19C96CFE5DF858" authority="(Gunther) (David et al. 2008 b)" authorityName="David et al." authorityYear="2008" baseAuthorityName="Gunther" baseAuthorityYear="1861" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="B9407916FFFEFFEA4A19C96CFA3DF87C" box="[1298,1437,1894,1920]" italics="true" pageId="5" pageNumber="560">O. taeniatus</emphasis>
(Günther) (
<bibRefCitation id="EFA5D8F5FFFEFFEA4E1CC980FE55F858" author="David, P. &amp; Vogel, G. &amp; van Rooijen, J." box="[279,501,1930,1956]" pageId="5" pageNumber="560" pagination="1 - 49" refId="ref7219" refString="David, P., Vogel, G. &amp; van Rooijen, J. (2008 b) A revision of the Oligodon taeniatus (Gunther, 1861) group (Squamata: Colubridae), with the description of three new species from the Indochinese region. Zootaxa, 1965 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 1965.1.1" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFEFFEA4E6DC981FE01F858" box="[358,417,1930,1956]" italics="true" pageId="5" pageNumber="560">et al.</emphasis>
2008b
</bibRefCitation>
)
</taxonomicName>
. The new species differs from 
<taxonomicName id="4C34DE87FFFEFFEA4C68C980FBF0F858" authority="(Blyth)" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[867,1104,1930,1956]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="catenatus">
<emphasis id="B9407916FFFEFFEA4C68C980FC53F858" box="[867,1011,1930,1956]" italics="true" pageId="5" pageNumber="560">O. catenatus</emphasis>
(Blyth)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4B55C980FAD4F858" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[1118,1396,1930,1956]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="cruentatus">
<emphasis id="B9407916FFFEFFEA4B55C980FB58F858" box="[1118,1272,1930,1956]" italics="true" pageId="5" pageNumber="560">O. cruentatus</emphasis>
(Günther)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4A8AC980FE49F834" authority="(Gray &amp; Hardwicke)" baseAuthorityName="Gray &amp; Hardwicke" baseAuthorityYear="1835" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="B9407916FFFEFFEA4A8AC980FF51F834" italics="true" pageId="5" pageNumber="560">O. dorsalis</emphasis>
(Gray &amp; Hardwicke)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4EFEC9A4FD5EF834" authority="Pellegrin" authorityName="Pellegrin" authorityYear="1910" box="[501,766,1966,1992]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="eberhardti">
<emphasis id="B9407916FFFEFFEA4EFEC9A4FD2EF834" box="[501,654,1966,1992]" italics="true" pageId="5" pageNumber="560">O. eberhardti</emphasis>
Pellegrin
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4C01C9A4FBB6F834" authority="Boulenger" authorityName="Boulenger" authorityYear="1918" box="[778,1046,1966,1992]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="hamptoni">
<emphasis id="B9407916FFFEFFEA4C01C9A4FC37F834" box="[778,919,1966,1992]" italics="true" pageId="5" pageNumber="560">O. hamptoni</emphasis>
Boulenger
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFEFFEA4B29C9A4FAE4F834" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[1058,1348,1966,1992]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="B9407916FFFEFFEA4B29C9A4FB11F834" box="[1058,1201,1966,1992]" italics="true" pageId="5" pageNumber="560">O. planiceps</emphasis>
(Boulenger)
</taxonomicName>
, and 
<taxonomicName id="4C34DE87FFFEFFEA4A8AC9A4FED7F810" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1868" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="560" phylum="Chordata" rank="species" species="theobaldi">
<emphasis id="B9407916FFFEFFEA4A8AC9A4FEA1F810" italics="true" pageId="5" pageNumber="560">O. theobaldi</emphasis>
(Günther)
</taxonomicName>
by having smooth and forked (vs. spinose and unforked) hemipenis (
<bibRefCitation id="EFA5D8F5FFFEFFEA4B75C9D8FAA6F810" author="Smith, M. A." box="[1150,1286,2002,2028]" pageId="5" pageNumber="560" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
).
</paragraph>
<caption id="DF4BF58CFFFDFFE94F93CE9DFEF4FF2F" ID-Table-UUID="DF4BF58CFFFDFFE94F93CE9DFEF4FF2F" httpUri="http://table.plazi.org/id/DF4BF58CFFFDFFE94F93CE9DFEF4FF2F" pageId="6" pageNumber="561" startId="6.[152,244,151,175]" targetBox="[166,1324,228,1379]" targetIsTable="true" targetPageId="6">
<paragraph id="8B8BA504FFFDFFE94F93CE9DFEF4FF2F" blockId="6.[151,1436,151,211]" pageId="6" pageNumber="561">
<emphasis id="B9407916FFFDFFE94F93CE9DFEAEFF53" bold="true" box="[152,270,151,175]" pageId="6" pageNumber="561">TABLE 1.</emphasis>
Measurements (in mm) and scalation of the types series of 
<taxonomicName id="4C34DE87FFFDFFE94C88CE9DFB85FF53" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[899,1061,151,175]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFDFFE94C88CE9DFB85FF53" box="[899,1061,151,175]" italics="true" pageId="6" pageNumber="561">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFDFFE94B20CE92FB20FF4C" bold="true" box="[1067,1152,152,176]" pageId="6" pageNumber="561">
<taxonomicNameLabel id="A273C46DFFFDFFE94B20CE92FB20FF4C" box="[1067,1152,152,176]" pageId="6" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
See Material and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="8B8BA504FFFDFFE94E87CEEEFB54FA9E" pageId="6" pageNumber="561">
<table id="F93457A4FFFD00104FADCEEEFA8CFA9F" box="[166,1324,228,1379]" gridcols="5" gridrows="29" pageId="6" pageNumber="561">
<tr id="3504A746FFFD00104FADCEEEFA8CFF00" box="[166,1324,228,252]" gridrow="0" pageId="6" pageNumber="561" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<th id="76D5CE3AFFFD00104E87CEEEFDBDFF00" box="[396,541,228,252]" gridcol="1" gridrow="0" pageId="6" pageNumber="561">Holotype</th>
<th id="76D5CE3AFFFD00104DAFCEEEFC86FF00" box="[676,806,228,252]" gridcol="2" gridrow="0" pageId="6" pageNumber="561">Paratypes</th>
</tr>
<tr id="3504A746FFFD00104FADCF06FA8CFED8" box="[166,1324,268,292]" gridrow="1" pageId="6" pageNumber="561" rowspan-0="1">
<td id="76D5CE3AFFFD00104E87CF06FDBDFED8" box="[396,541,268,292]" gridcol="1" gridrow="1" pageId="6" pageNumber="561">ITBCZ 7899</td>
<td id="76D5CE3AFFFD00104DAFCF06FC86FED8" box="[676,806,268,292]" gridcol="2" gridrow="1" pageId="6" pageNumber="561">ITBCZ 7944</td>
<td id="76D5CE3AFFFD00104CAECF06FBEBFED8" box="[933,1099,268,292]" gridcol="3" gridrow="1" pageId="6" pageNumber="561">ITBCZ 7868</td>
<td id="76D5CE3AFFFD00104BA2CF06FA8CFED8" box="[1193,1324,268,292]" gridcol="4" gridrow="1" pageId="6" pageNumber="561">ITBCZ 8215</td>
</tr>
<tr id="3504A746FFFD00104FADCF3EFA8CFEB0" box="[166,1324,308,332]" gridrow="2" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCF3EFE93FEB0" box="[166,307,308,332]" gridcol="0" gridrow="2" pageId="6" pageNumber="561">Sex</th>
<td id="76D5CE3AFFFD00104E87CF3EFDBDFEB0" box="[396,541,308,332]" gridcol="1" gridrow="2" pageId="6" pageNumber="561">Subadult male</td>
<td id="76D5CE3AFFFD00104DAFCF3EFC86FEB0" box="[676,806,308,332]" gridcol="2" gridrow="2" pageId="6" pageNumber="561">Male</td>
<td id="76D5CE3AFFFD00104CAECF3EFBEBFEB0" box="[933,1099,308,332]" gridcol="3" gridrow="2" pageId="6" pageNumber="561">Subadult female</td>
<td id="76D5CE3AFFFD00104BA2CF3EFA8CFEB0" box="[1193,1324,308,332]" gridcol="4" gridrow="2" pageId="6" pageNumber="561">Female</td>
</tr>
<tr id="3504A746FFFD00104FADCF56FA8CFE89" box="[166,1324,348,373]" gridrow="3" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCF56FE93FE89" box="[166,307,348,373]" gridcol="0" gridrow="3" pageId="6" pageNumber="561">SVL</th>
<td id="76D5CE3AFFFD00104E87CF56FDBDFE89" box="[396,541,348,373]" gridcol="1" gridrow="3" pageId="6" pageNumber="561">475</td>
<td id="76D5CE3AFFFD00104DAFCF56FC86FE89" box="[676,806,348,373]" gridcol="2" gridrow="3" pageId="6" pageNumber="561">720</td>
<td id="76D5CE3AFFFD00104CAECF56FBEBFE89" box="[933,1099,348,373]" gridcol="3" gridrow="3" pageId="6" pageNumber="561">458</td>
<td id="76D5CE3AFFFD00104BA2CF56FA8CFE89" box="[1193,1324,348,373]" gridcol="4" gridrow="3" pageId="6" pageNumber="561">779</td>
</tr>
<tr id="3504A746FFFD00104FADCF8FFA8CFE61" box="[166,1324,389,413]" gridrow="4" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCF8FFE93FE61" box="[166,307,389,413]" gridcol="0" gridrow="4" pageId="6" pageNumber="561">TaL</th>
<td id="76D5CE3AFFFD00104E87CF8FFDBDFE61" box="[396,541,389,413]" gridcol="1" gridrow="4" pageId="6" pageNumber="561">110</td>
<td id="76D5CE3AFFFD00104DAFCF8FFC86FE61" box="[676,806,389,413]" gridcol="2" gridrow="4" pageId="6" pageNumber="561">162</td>
<td id="76D5CE3AFFFD00104CAECF8FFBEBFE61" box="[933,1099,389,413]" gridcol="3" gridrow="4" pageId="6" pageNumber="561">76</td>
<td id="76D5CE3AFFFD00104BA2CF8FFA8CFE61" box="[1193,1324,389,413]" gridcol="4" gridrow="4" pageId="6" pageNumber="561">109</td>
</tr>
<tr id="3504A746FFFD00104FADCFA7FA8CFE39" box="[166,1324,429,453]" gridrow="5" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCFA7FE93FE39" box="[166,307,429,453]" gridcol="0" gridrow="5" pageId="6" pageNumber="561">TL</th>
<td id="76D5CE3AFFFD00104E87CFA7FDBDFE39" box="[396,541,429,453]" gridcol="1" gridrow="5" pageId="6" pageNumber="561">585</td>
<td id="76D5CE3AFFFD00104DAFCFA7FC86FE39" box="[676,806,429,453]" gridcol="2" gridrow="5" pageId="6" pageNumber="561">882</td>
<td id="76D5CE3AFFFD00104CAECFA7FBEBFE39" box="[933,1099,429,453]" gridcol="3" gridrow="5" pageId="6" pageNumber="561">534</td>
<td id="76D5CE3AFFFD00104BA2CFA7FA8CFE39" box="[1193,1324,429,453]" gridcol="4" gridrow="5" pageId="6" pageNumber="561">888</td>
</tr>
<tr id="3504A746FFFD00104FADCFDFFA8CFE11" box="[166,1324,469,493]" gridrow="6" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCFDFFE93FE11" box="[166,307,469,493]" gridcol="0" gridrow="6" pageId="6" pageNumber="561">TaL/TL</th>
<td id="76D5CE3AFFFD00104E87CFDFFDBDFE11" box="[396,541,469,493]" gridcol="1" gridrow="6" pageId="6" pageNumber="561">0.188</td>
<td id="76D5CE3AFFFD00104DAFCFDFFC86FE11" box="[676,806,469,493]" gridcol="2" gridrow="6" pageId="6" pageNumber="561">0.184</td>
<td id="76D5CE3AFFFD00104CAECFDFFBEBFE11" box="[933,1099,469,493]" gridcol="3" gridrow="6" pageId="6" pageNumber="561">0.142</td>
<td id="76D5CE3AFFFD00104BA2CFDFFA8CFE11" box="[1193,1324,469,493]" gridcol="4" gridrow="6" pageId="6" pageNumber="561">0.123</td>
</tr>
<tr id="3504A746FFFD00104FADCFF7FA8CFDEA" box="[166,1324,509,534]" gridrow="7" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCFF7FE93FDEA" box="[166,307,509,534]" gridcol="0" gridrow="7" pageId="6" pageNumber="561">HL</th>
<td id="76D5CE3AFFFD00104E87CFF7FDBDFDEA" box="[396,541,509,534]" gridcol="1" gridrow="7" pageId="6" pageNumber="561">20.3</td>
<td id="76D5CE3AFFFD00104DAFCFF7FC86FDEA" box="[676,806,509,534]" gridcol="2" gridrow="7" pageId="6" pageNumber="561">26.8</td>
<td id="76D5CE3AFFFD00104CAECFF7FBEBFDEA" box="[933,1099,509,534]" gridcol="3" gridrow="7" pageId="6" pageNumber="561">19.2</td>
<td id="76D5CE3AFFFD00104BA2CFF7FA8CFDEA" box="[1193,1324,509,534]" gridcol="4" gridrow="7" pageId="6" pageNumber="561">25.2</td>
</tr>
<tr id="3504A746FFFD00104FADCC2CFA8CFDC2" box="[166,1324,550,574]" gridrow="8" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCC2CFE93FDC2" box="[166,307,550,574]" gridcol="0" gridrow="8" pageId="6" pageNumber="561">HW</th>
<td id="76D5CE3AFFFD00104E87CC2CFDBDFDC2" box="[396,541,550,574]" gridcol="1" gridrow="8" pageId="6" pageNumber="561">13.3</td>
<td id="76D5CE3AFFFD00104DAFCC2CFC86FDC2" box="[676,806,550,574]" gridcol="2" gridrow="8" pageId="6" pageNumber="561">-</td>
<td id="76D5CE3AFFFD00104CAECC2CFBEBFDC2" box="[933,1099,550,574]" gridcol="3" gridrow="8" pageId="6" pageNumber="561">-</td>
<td id="76D5CE3AFFFD00104BA2CC2CFA8CFDC2" box="[1193,1324,550,574]" gridcol="4" gridrow="8" pageId="6" pageNumber="561">17.7</td>
</tr>
<tr id="3504A746FFFD00104FADCC44FA8CFD9A" box="[166,1324,590,614]" gridrow="9" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCC44FE93FD9A" box="[166,307,590,614]" gridcol="0" gridrow="9" pageId="6" pageNumber="561">HH</th>
<td id="76D5CE3AFFFD00104E87CC44FDBDFD9A" box="[396,541,590,614]" gridcol="1" gridrow="9" pageId="6" pageNumber="561">11.0</td>
<td id="76D5CE3AFFFD00104DAFCC44FC86FD9A" box="[676,806,590,614]" gridcol="2" gridrow="9" pageId="6" pageNumber="561">-</td>
<td id="76D5CE3AFFFD00104CAECC44FBEBFD9A" box="[933,1099,590,614]" gridcol="3" gridrow="9" pageId="6" pageNumber="561">-</td>
<td id="76D5CE3AFFFD00104BA2CC44FA8CFD9A" box="[1193,1324,590,614]" gridcol="4" gridrow="9" pageId="6" pageNumber="561">13.2</td>
</tr>
<tr id="3504A746FFFD00104FADCC7CFA8CFD72" box="[166,1324,630,654]" gridrow="10" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCC7CFE93FD72" box="[166,307,630,654]" gridcol="0" gridrow="10" pageId="6" pageNumber="561">ED</th>
<td id="76D5CE3AFFFD00104E87CC7CFDBDFD72" box="[396,541,630,654]" gridcol="1" gridrow="10" pageId="6" pageNumber="561">2.8</td>
<td id="76D5CE3AFFFD00104DAFCC7CFC86FD72" box="[676,806,630,654]" gridcol="2" gridrow="10" pageId="6" pageNumber="561">3.6</td>
<td id="76D5CE3AFFFD00104CAECC7CFBEBFD72" box="[933,1099,630,654]" gridcol="3" gridrow="10" pageId="6" pageNumber="561">3.1</td>
<td id="76D5CE3AFFFD00104BA2CC7CFA8CFD72" box="[1193,1324,630,654]" gridcol="4" gridrow="10" pageId="6" pageNumber="561">3.6</td>
</tr>
<tr id="3504A746FFFD00104FADCC94FA8CFD4B" box="[166,1324,670,695]" gridrow="11" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCC94FE93FD4B" box="[166,307,670,695]" gridcol="0" gridrow="11" pageId="6" pageNumber="561">SnL</th>
<td id="76D5CE3AFFFD00104E87CC94FDBDFD4B" box="[396,541,670,695]" gridcol="1" gridrow="11" pageId="6" pageNumber="561">6.0</td>
<td id="76D5CE3AFFFD00104DAFCC94FC86FD4B" box="[676,806,670,695]" gridcol="2" gridrow="11" pageId="6" pageNumber="561">6.9</td>
<td id="76D5CE3AFFFD00104CAECC94FBEBFD4B" box="[933,1099,670,695]" gridcol="3" gridrow="11" pageId="6" pageNumber="561">5.9</td>
<td id="76D5CE3AFFFD00104BA2CC94FA8CFD4B" box="[1193,1324,670,695]" gridcol="4" gridrow="11" pageId="6" pageNumber="561">8.2</td>
</tr>
<tr id="3504A746FFFD00104FADCCCDFA8CFD23" box="[166,1324,711,735]" gridrow="12" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCCCDFE93FD23" box="[166,307,711,735]" gridcol="0" gridrow="12" pageId="6" pageNumber="561">EN</th>
<td id="76D5CE3AFFFD00104E87CCCDFDBDFD23" box="[396,541,711,735]" gridcol="1" gridrow="12" pageId="6" pageNumber="561">3.7</td>
<td id="76D5CE3AFFFD00104DAFCCCDFC86FD23" box="[676,806,711,735]" gridcol="2" gridrow="12" pageId="6" pageNumber="561">3.8</td>
<td id="76D5CE3AFFFD00104CAECCCDFBEBFD23" box="[933,1099,711,735]" gridcol="3" gridrow="12" pageId="6" pageNumber="561">3.4</td>
<td id="76D5CE3AFFFD00104BA2CCCDFA8CFD23" box="[1193,1324,711,735]" gridcol="4" gridrow="12" pageId="6" pageNumber="561">3.9</td>
</tr>
<tr id="3504A746FFFD00104FADCCE5FA8CFCFB" box="[166,1324,751,775]" gridrow="13" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCCE5FE93FCFB" box="[166,307,751,775]" gridcol="0" gridrow="13" pageId="6" pageNumber="561">SL</th>
<td id="76D5CE3AFFFD00104E87CCE5FDBDFCFB" box="[396,541,751,775]" gridcol="1" gridrow="13" pageId="6" pageNumber="561">8/8</td>
<td id="76D5CE3AFFFD00104DAFCCE5FC86FCFB" box="[676,806,751,775]" gridcol="2" gridrow="13" pageId="6" pageNumber="561">8/8</td>
<td id="76D5CE3AFFFD00104CAECCE5FBEBFCFB" box="[933,1099,751,775]" gridcol="3" gridrow="13" pageId="6" pageNumber="561">8/8</td>
<td id="76D5CE3AFFFD00104BA2CCE5FA8CFCFB" box="[1193,1324,751,775]" gridcol="4" gridrow="13" pageId="6" pageNumber="561">8/8</td>
</tr>
<tr id="3504A746FFFD00104FADCD1DFA8CFCD3" box="[166,1324,791,815]" gridrow="14" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCD1DFE93FCD3" box="[166,307,791,815]" gridcol="0" gridrow="14" pageId="6" pageNumber="561">SL-Eye</th>
<td id="76D5CE3AFFFD00104E87CD1DFDBDFCD3" box="[396,541,791,815]" gridcol="1" gridrow="14" pageId="6" pageNumber="561">4–5/4–5</td>
<td id="76D5CE3AFFFD00104DAFCD1DFC86FCD3" box="[676,806,791,815]" gridcol="2" gridrow="14" pageId="6" pageNumber="561">4–5/4–5</td>
<td id="76D5CE3AFFFD00104CAECD1DFBEBFCD3" box="[933,1099,791,815]" gridcol="3" gridrow="14" pageId="6" pageNumber="561">4–5/4–5</td>
<td id="76D5CE3AFFFD00104BA2CD1DFA8CFCD3" box="[1193,1324,791,815]" gridcol="4" gridrow="14" pageId="6" pageNumber="561">4–5/4–5</td>
</tr>
<tr id="3504A746FFFD00104FADCD4AFA8CFCA4" box="[166,1324,832,856]" gridrow="15" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCD4AFE93FCA4" box="[166,307,832,856]" gridcol="0" gridrow="15" pageId="6" pageNumber="561">PreOc</th>
<td id="76D5CE3AFFFD00104E87CD4AFDBDFCA4" box="[396,541,832,856]" gridcol="1" gridrow="15" pageId="6" pageNumber="561">1/1</td>
<td id="76D5CE3AFFFD00104DAFCD4AFC86FCA4" box="[676,806,832,856]" gridcol="2" gridrow="15" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104CAECD4AFBEBFCA4" box="[933,1099,832,856]" gridcol="3" gridrow="15" pageId="6" pageNumber="561">1/1</td>
<td id="76D5CE3AFFFD00104BA2CD4AFA8CFCA4" box="[1193,1324,832,856]" gridcol="4" gridrow="15" pageId="6" pageNumber="561">1/1</td>
</tr>
<tr id="3504A746FFFD00104FADCD62FA8CFC7C" box="[166,1324,872,896]" gridrow="16" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCD62FE93FC7C" box="[166,307,872,896]" gridcol="0" gridrow="16" pageId="6" pageNumber="561">PreSubOc</th>
<td id="76D5CE3AFFFD00104E87CD62FDBDFC7C" box="[396,541,872,896]" gridcol="1" gridrow="16" pageId="6" pageNumber="561">1/1</td>
<td id="76D5CE3AFFFD00104DAFCD62FC86FC7C" box="[676,806,872,896]" gridcol="2" gridrow="16" pageId="6" pageNumber="561">0/1</td>
<td id="76D5CE3AFFFD00104CAECD62FBEBFC7C" box="[933,1099,872,896]" gridcol="3" gridrow="16" pageId="6" pageNumber="561">1/1</td>
<td id="76D5CE3AFFFD00104BA2CD62FA8CFC7C" box="[1193,1324,872,896]" gridcol="4" gridrow="16" pageId="6" pageNumber="561">1/1</td>
</tr>
<tr id="3504A746FFFD00104FADCD9AFA8CFC54" box="[166,1324,912,936]" gridrow="17" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCD9AFE93FC54" box="[166,307,912,936]" gridcol="0" gridrow="17" pageId="6" pageNumber="561">PosOc</th>
<td id="76D5CE3AFFFD00104E87CD9AFDBDFC54" box="[396,541,912,936]" gridcol="1" gridrow="17" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104DAFCD9AFC86FC54" box="[676,806,912,936]" gridcol="2" gridrow="17" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104CAECD9AFBEBFC54" box="[933,1099,912,936]" gridcol="3" gridrow="17" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104BA2CD9AFA8CFC54" box="[1193,1324,912,936]" gridcol="4" gridrow="17" pageId="6" pageNumber="561">2/2</td>
</tr>
<tr id="3504A746FFFD00104FADCDB2FA8CFC2C" box="[166,1324,952,976]" gridrow="18" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCDB2FE93FC2C" box="[166,307,952,976]" gridcol="0" gridrow="18" pageId="6" pageNumber="561">ATP</th>
<td id="76D5CE3AFFFD00104E87CDB2FDBDFC2C" box="[396,541,952,976]" gridcol="1" gridrow="18" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104DAFCDB2FC86FC2C" box="[676,806,952,976]" gridcol="2" gridrow="18" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104CAECDB2FBEBFC2C" box="[933,1099,952,976]" gridcol="3" gridrow="18" pageId="6" pageNumber="561">2/2</td>
<td id="76D5CE3AFFFD00104BA2CDB2FA8CFC2C" box="[1193,1324,952,976]" gridcol="4" gridrow="18" pageId="6" pageNumber="561">2/2</td>
</tr>
<tr id="3504A746FFFD00104FADCDEAFA8CFC04" box="[166,1324,992,1016]" gridrow="19" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCDEAFE93FC04" box="[166,307,992,1016]" gridcol="0" gridrow="19" pageId="6" pageNumber="561">VS (+ preVS)</th>
<td id="76D5CE3AFFFD00104E87CDEAFDBDFC04" box="[396,541,992,1016]" gridcol="1" gridrow="19" pageId="6" pageNumber="561">173 (+1)</td>
<td id="76D5CE3AFFFD00104DAFCDEAFC86FC04" box="[676,806,992,1016]" gridcol="2" gridrow="19" pageId="6" pageNumber="561">179 (+1)</td>
<td id="76D5CE3AFFFD00104CAECDEAFBEBFC04" box="[933,1099,992,1016]" gridcol="3" gridrow="19" pageId="6" pageNumber="561">187 (+1)</td>
<td id="76D5CE3AFFFD00104BA2CDEAFA8CFC04" box="[1193,1324,992,1016]" gridcol="4" gridrow="19" pageId="6" pageNumber="561">193 (+1)</td>
</tr>
<tr id="3504A746FFFD00104FADCA03FA8CFBDD" box="[166,1324,1033,1057]" gridrow="20" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCA03FE93FBDD" box="[166,307,1033,1057]" gridcol="0" gridrow="20" pageId="6" pageNumber="561">ASR</th>
<td id="76D5CE3AFFFD00104E87CA03FDBDFBDD" box="[396,541,1033,1057]" gridcol="1" gridrow="20" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104DAFCA03FC86FBDD" box="[676,806,1033,1057]" gridcol="2" gridrow="20" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104CAECA03FBEBFBDD" box="[933,1099,1033,1057]" gridcol="3" gridrow="20" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104BA2CA03FA8CFBDD" box="[1193,1324,1033,1057]" gridcol="4" gridrow="20" pageId="6" pageNumber="561">19</td>
</tr>
<tr id="3504A746FFFD00104FADCA3BFA8CFBB5" box="[166,1324,1073,1097]" gridrow="21" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCA3BFE93FBB5" box="[166,307,1073,1097]" gridcol="0" gridrow="21" pageId="6" pageNumber="561">MSR</th>
<td id="76D5CE3AFFFD00104E87CA3BFDBDFBB5" box="[396,541,1073,1097]" gridcol="1" gridrow="21" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104DAFCA3BFC86FBB5" box="[676,806,1073,1097]" gridcol="2" gridrow="21" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104CAECA3BFBEBFBB5" box="[933,1099,1073,1097]" gridcol="3" gridrow="21" pageId="6" pageNumber="561">19</td>
<td id="76D5CE3AFFFD00104BA2CA3BFA8CFBB5" box="[1193,1324,1073,1097]" gridcol="4" gridrow="21" pageId="6" pageNumber="561">19</td>
</tr>
<tr id="3504A746FFFD00104FADCA53FA8CFB8D" box="[166,1324,1113,1137]" gridrow="22" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCA53FE93FB8D" box="[166,307,1113,1137]" gridcol="0" gridrow="22" pageId="6" pageNumber="561">PSR</th>
<td id="76D5CE3AFFFD00104E87CA53FDBDFB8D" box="[396,541,1113,1137]" gridcol="1" gridrow="22" pageId="6" pageNumber="561">15</td>
<td id="76D5CE3AFFFD00104DAFCA53FC86FB8D" box="[676,806,1113,1137]" gridcol="2" gridrow="22" pageId="6" pageNumber="561">15</td>
<td id="76D5CE3AFFFD00104CAECA53FBEBFB8D" box="[933,1099,1113,1137]" gridcol="3" gridrow="22" pageId="6" pageNumber="561">15</td>
<td id="76D5CE3AFFFD00104BA2CA53FA8CFB8D" box="[1193,1324,1113,1137]" gridcol="4" gridrow="22" pageId="6" pageNumber="561">15</td>
</tr>
<tr id="3504A746FFFD00104FADCA8AFA8CFB65" box="[166,1324,1152,1177]" gridrow="23" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCA8AFE93FB65" box="[166,307,1152,1177]" gridcol="0" gridrow="23" pageId="6" pageNumber="561">1st reduction</th>
<td id="76D5CE3AFFFD00104E87CA8AFDBDFB65" box="[396,541,1152,1177]" gridcol="1" gridrow="23" pageId="6" pageNumber="561">97</td>
<td id="76D5CE3AFFFD00104DAFCA8AFC86FB65" box="[676,806,1152,1177]" gridcol="2" gridrow="23" pageId="6" pageNumber="561">111</td>
<td id="76D5CE3AFFFD00104CAECA8AFBEBFB65" box="[933,1099,1152,1177]" gridcol="3" gridrow="23" pageId="6" pageNumber="561">110</td>
<td id="76D5CE3AFFFD00104BA2CA8AFA8CFB65" box="[1193,1324,1152,1177]" gridcol="4" gridrow="23" pageId="6" pageNumber="561">111</td>
</tr>
<tr id="3504A746FFFD00104FADCAA3FA8CFB3E" box="[166,1324,1193,1218]" gridrow="24" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCAA3FE93FB3E" box="[166,307,1193,1218]" gridcol="0" gridrow="24" pageId="6" pageNumber="561">SC</th>
<td id="76D5CE3AFFFD00104E87CAA3FDBDFB3E" box="[396,541,1193,1218]" gridcol="1" gridrow="24" pageId="6" pageNumber="561">59</td>
<td id="76D5CE3AFFFD00104DAFCAA3FC86FB3E" box="[676,806,1193,1218]" gridcol="2" gridrow="24" pageId="6" pageNumber="561">58</td>
<td id="76D5CE3AFFFD00104CAECAA3FBEBFB3E" box="[933,1099,1193,1218]" gridcol="3" gridrow="24" pageId="6" pageNumber="561">45</td>
<td id="76D5CE3AFFFD00104BA2CAA3FA8CFB3E" box="[1193,1324,1193,1218]" gridcol="4" gridrow="24" pageId="6" pageNumber="561">44</td>
</tr>
<tr id="3504A746FFFD00104FADCAD8FA8CFB16" box="[166,1324,1234,1258]" gridrow="25" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCAD8FE93FB16" box="[166,307,1234,1258]" gridcol="0" gridrow="25" pageId="6" pageNumber="561">MT</th>
<td id="76D5CE3AFFFD00104E87CAD8FDBDFB16" box="[396,541,1234,1258]" gridcol="1" gridrow="25" pageId="6" pageNumber="561">10</td>
<td id="76D5CE3AFFFD00104DAFCAD8FC86FB16" box="[676,806,1234,1258]" gridcol="2" gridrow="25" pageId="6" pageNumber="561">10</td>
<td id="76D5CE3AFFFD00104CAECAD8FBEBFB16" box="[933,1099,1234,1258]" gridcol="3" gridrow="25" pageId="6" pageNumber="561">10</td>
<td id="76D5CE3AFFFD00104BA2CAD8FA8CFB16" box="[1193,1324,1234,1258]" gridcol="4" gridrow="25" pageId="6" pageNumber="561">10</td>
</tr>
<tr id="3504A746FFFD00104FADCAF0FA8CFAEE" box="[166,1324,1274,1298]" gridrow="26" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCAF0FE93FAEE" box="[166,307,1274,1298]" gridcol="0" gridrow="26" pageId="6" pageNumber="561">LHSC</th>
<td id="76D5CE3AFFFD00104E87CAF0FDBDFAEE" box="[396,541,1274,1298]" gridcol="1" gridrow="26" pageId="6" pageNumber="561">25</td>
<td id="76D5CE3AFFFD00104DAFCAF0FC86FAEE" box="[676,806,1274,1298]" gridcol="2" gridrow="26" pageId="6" pageNumber="561">25</td>
<td id="76D5CE3AFFFD00104CAECAF0FBEBFAEE" box="[933,1099,1274,1298]" gridcol="3" gridrow="26" pageId="6" pageNumber="561">-</td>
<td id="76D5CE3AFFFD00104BA2CAF0FA8CFAEE" box="[1193,1324,1274,1298]" gridcol="4" gridrow="26" pageId="6" pageNumber="561">-</td>
</tr>
<tr id="3504A746FFFD00104FADCB28FA8CFAC6" box="[166,1324,1314,1338]" gridrow="27" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCB28FE93FAC6" box="[166,307,1314,1338]" gridcol="0" gridrow="27" pageId="6" pageNumber="561">DB</th>
<td id="76D5CE3AFFFD00104E87CB28FDBDFAC6" box="[396,541,1314,1338]" gridcol="1" gridrow="27" pageId="6" pageNumber="561">13+4</td>
<td id="76D5CE3AFFFD00104DAFCB28FC86FAC6" box="[676,806,1314,1338]" gridcol="2" gridrow="27" pageId="6" pageNumber="561">14+3</td>
<td id="76D5CE3AFFFD00104CAECB28FBEBFAC6" box="[933,1099,1314,1338]" gridcol="3" gridrow="27" pageId="6" pageNumber="561">13+3</td>
<td id="76D5CE3AFFFD00104BA2CB28FA8CFAC6" box="[1193,1324,1314,1338]" gridcol="4" gridrow="27" pageId="6" pageNumber="561">15+3</td>
</tr>
<tr id="3504A746FFFD00104FADCB41FA8CFA9F" box="[166,1324,1355,1379]" gridrow="28" pageId="6" pageNumber="561">
<th id="76D5CE3AFFFD00104FADCB41FE93FA9F" box="[166,307,1355,1379]" gridcol="0" gridrow="28" pageId="6" pageNumber="561">VerS</th>
<td id="76D5CE3AFFFD00104E87CB41FDBDFA9F" box="[396,541,1355,1379]" gridcol="1" gridrow="28" pageId="6" pageNumber="561">Present</td>
<td id="76D5CE3AFFFD00104DAFCB41FC86FA9F" box="[676,806,1355,1379]" gridcol="2" gridrow="28" pageId="6" pageNumber="561">Present</td>
<td id="76D5CE3AFFFD00104CAECB41FBEBFA9F" box="[933,1099,1355,1379]" gridcol="3" gridrow="28" pageId="6" pageNumber="561">Present</td>
<td id="76D5CE3AFFFD00104BA2CB41FA8CFA9F" box="[1193,1324,1355,1379]" gridcol="4" gridrow="28" pageId="6" pageNumber="561">Present</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8BA504FFFDFFE94FCCCB98FC4FF99C" blockId="6.[151,1437,1426,2029]" pageId="6" pageNumber="561">
<taxonomicName id="4C34DE87FFFDFFE94FCCCB98FED3FA50" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[199,371,1426,1452]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFDFFE94FCCCB98FED3FA50" box="[199,371,1426,1452]" italics="true" pageId="6" pageNumber="561">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFDFFE94E70CB98FE76FA50" bold="true" box="[379,470,1426,1452]" pageId="6" pageNumber="561">
<taxonomicNameLabel id="A273C46DFFFDFFE94E70CB98FE76FA50" box="[379,470,1426,1452]" pageId="6" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has 19 dorsal scale rows at midbody and thus differs from the following species that have 13 or 15 dorsal scale rows: 
<taxonomicName id="4C34DE87FFFDFFE94D00CBBDFCB4FA2C" authority="Leviton" authorityName="Leviton" authorityYear="1953" box="[523,788,1462,1489]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="B9407916FFFDFFE94D00CBBDFD12FA2C" box="[523,690,1463,1489]" italics="true" pageId="6" pageNumber="561">O. annamensis</emphasis>
Leviton
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94C2BCBBDFAD5FA2C" authority="Pauwels, Wallach, David &amp; Chanhome" authorityName="Pauwels, Wallach, David &amp; Chanhome" box="[800,1397,1462,1489]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="jintakunei">
<emphasis id="B9407916FFFDFFE94C2BCBBDFC13FA2C" box="[800,947,1462,1489]" italics="true" pageId="6" pageNumber="561">O. jintakunei</emphasis>
Pauwels, Wallach, David &amp; Chanhome
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94A8ACBBDFE33FA08" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1914" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="B9407916FFFDFFE94A8ACBBDFEA2FA08" italics="true" pageId="6" pageNumber="561">O. inornatus</emphasis>
(Boulenger)
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94E95CBD1FC3CFA09" authority="Neang, Grismer &amp; Dattry" authorityName="Neang, Grismer &amp; Dattry" box="[414,924,1498,1525]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="kampucheaensis">
<emphasis id="B9407916FFFDFFE94E95CBD1FDD8FA08" box="[414,632,1498,1525]" italics="true" pageId="6" pageNumber="561">O. kampucheaensis</emphasis>
Neang, Grismer &amp; Dattry
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94CACCBD1FB43FA09" authority="Angel &amp; Bourret" authorityName="Angel &amp; Bourret" authorityYear="1933" box="[935,1251,1498,1525]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="lacroixi">
<emphasis id="B9407916FFFDFFE94CACCBD1FBBFFA08" box="[935,1055,1498,1525]" italics="true" pageId="6" pageNumber="561">O. lacroixi</emphasis>
Angel &amp; Bourret
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94BE5CBD1FF6EF9E4" authority="Wall" authorityName="Wall" authorityYear="1905" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="macdougalli">
<emphasis id="B9407916FFFDFFE94BE5CBD1FA3CFA08" box="[1262,1436,1498,1525]" italics="true" pageId="6" pageNumber="561">O. macdougalli</emphasis>
Wall
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94FD3CBF5FC12F9E4" authority="David, Nguyen, Nguyen, Jiang, Chen, Teynie &amp; Ziegler" authorityName="David, Nguyen, Nguyen, Jiang, Chen, Teynie &amp; Ziegler" authorityYear="2012" box="[216,946,1534,1561]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="nagao">
<emphasis id="B9407916FFFDFFE94FD3CBF5FE9FF9E4" box="[216,319,1535,1561]" italics="true" pageId="6" pageNumber="561">O. nagao</emphasis>
David, Nguyen, Nguyen, Jiang, Chen, Teynie &amp; Ziegler
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94CB7CBF5FE0BF9C0" authority="Nguyen, Tran, Nguyen, Neang, Yushchenki &amp; Poyarkov" authorityName="Nguyen, Tran, Nguyen, Neang, Yushchenki &amp; Poyarkov" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="rostralis">
<emphasis id="B9407916FFFDFFE94CB7CBF5FB9BF9E4" box="[956,1083,1534,1561]" italics="true" pageId="6" pageNumber="561">O. rostralis</emphasis>
Nguyen, Tran, Nguyen, Neang, Yushchenki &amp; Poyarkov
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94EBCC829FC97F9C0" authority="David, Hauser &amp; Vogel" authorityName="David, Hauser &amp; Vogel" box="[439,823,1570,1597]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="teyniei">
<emphasis id="B9407916FFFDFFE94EBCC829FD85F9C0" box="[439,549,1571,1597]" italics="true" pageId="6" pageNumber="561">O. teyniei</emphasis>
David, Hauser &amp; Vogel
</taxonomicName>
, 
<taxonomicName id="4C34DE87FFFDFFE94C49C829FBC1F9C0" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1888" box="[834,1121,1570,1597]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="B9407916FFFDFFE94C49C829FC6FF9C0" box="[834,975,1571,1597]" italics="true" pageId="6" pageNumber="561">O. torquatus</emphasis>
(Boulenger)
</taxonomicName>
, and 
<taxonomicName id="4C34DE87FFFDFFE94B96C829FA3CF9C0" authority="Gunther" authorityName="Gunther" authorityYear="1865" box="[1181,1436,1570,1597]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="B9407916FFFDFFE94B96C829FA99F9C0" box="[1181,1337,1570,1597]" italics="true" pageId="6" pageNumber="561">O. vertebralis</emphasis>
Günther
</taxonomicName>
(
<bibRefCitation id="EFA5D8F5FFFDFFE94F94C84CFE85F99C" author="Smith, M. A." box="[159,293,1606,1632]" pageId="6" pageNumber="561" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFDFFE94E3AC84CFE44F99D" author="Vassilieva, A. B." box="[305,484,1606,1633]" pageId="6" pageNumber="561" pagination="211 - 226" refId="ref8711" refString="Vassilieva, A. B. (2015) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from coastal southern Vietnam. Zootaxa, 4058 (2), 211 - 226. https: // doi. org / 10.11646 / zootaxa. 4058.2.4" type="journal article" year="2015">Vassilieva 2015</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFDFFE94EFBC84DFD6BF99C" author="Nguyen, H. N. &amp; Tran, B. V. &amp; Nguyen, L. H. &amp; Neang, T. &amp; Yushchenko, P. V. &amp; Poyarkov, N. A." box="[496,715,1606,1633]" pageId="6" pageNumber="561" pagination="8332" refId="ref8070" refString="Nguyen, H. N., Tran, B. V., Nguyen, L. H., Neang, T., Yushchenko, P. V. &amp; Poyarkov, N. A. (2020) A new species of Oligodon Fitzinger, 1826 from the Langbian Plateau, southern Vietnam, with additional information on Oligodon annamensis Leviton, 1953 (Squamata: Colubridae). PeerJ, 8, e 8332. https: // doi. org / 10.7717 / peerj. 8332" type="journal article" year="2020">
Nguyen 
<emphasis id="B9407916FFFDFFE94D5BC84DFD21F99C" box="[592,641,1606,1632]" italics="true" pageId="6" pageNumber="561">et al</emphasis>
. 2020
</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFDFFE94DDEC84CFC3BF99C" author="David, P. &amp; Nguyen, T. Q. &amp; Nguyen, T. T. &amp; Jiang, K. &amp; Chen, T. &amp; Teynie, A. &amp; Ziegler, T." box="[725,923,1606,1633]" pageId="6" pageNumber="561" pagination="45 - 62" refId="ref7291" refString="David, P., Nguyen, T. Q., Nguyen, T. T., Jiang, K., Chen, T., Teynie, A. &amp; Ziegler, T. (2012) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa, 3498 (1), 45 - 62. https: // doi. org / 10.11646 / zootaxa. 3498.1.3" type="journal article" year="2012">
David 
<emphasis id="B9407916FFFDFFE94C2AC84DFCF2F99C" box="[801,850,1606,1632]" italics="true" pageId="6" pageNumber="561">et al</emphasis>
. 2012
</bibRefCitation>
, 
<bibRefCitation id="EFA5D8F5FFFDFFE94CADC84CFC43F99C" author="David, P. &amp; Hauser, S. &amp; Vogel, G." box="[934,995,1606,1632]" pageId="6" pageNumber="561" pagination="12 - 24" refId="ref7097" refString="David, P., Hauser, S. &amp; Vogel, G. (2022) A new species of the genus Oligodon Fitzinger, 1826 (Reptilia: Colubridae) from southern Laos. Taprobanica, 11, 12 - 24. https: // doi. org / 10.47605 / tapro. v 11 i 1.273" type="journal article" year="2022">2022</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BA504FFFDFFE84FCCC860FBECFB39" blockId="6.[151,1437,1426,2029]" lastBlockId="7.[151,1437,150,2013]" lastPageId="7" lastPageNumber="562" pageId="6" pageNumber="561">
<taxonomicName id="4C34DE87FFFDFFE94FCCC860FED1F978" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[199,369,1642,1668]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFDFFE94FCCC860FED1F978" box="[199,369,1642,1668]" italics="true" pageId="6" pageNumber="561">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFDFFE94E7CC860FE70F978" bold="true" box="[375,464,1642,1668]" pageId="6" pageNumber="561">
<taxonomicNameLabel id="A273C46DFFFDFFE94E7CC860FE70F978" box="[375,464,1642,1668]" pageId="6" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C34DE87FFFDFFE94D6AC861FCC9F978" authority="(Cantor)" baseAuthorityName="Cantor" baseAuthorityYear="1839" box="[609,873,1642,1669]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="albocinctus">
<emphasis id="B9407916FFFDFFE94D6AC861FCA3F978" box="[609,771,1642,1669]" italics="true" pageId="6" pageNumber="561">O. albocinctus</emphasis>
(Cantor)
</taxonomicName>
by having more supralabials (8 vs. 7), supralabials 4 and 5 (vs. 3 and 4) touching the eye, dorsum with blotches (vs. cross-bars), and hemipenis without papillae (with short pointed papillae) (
<bibRefCitation id="EFA5D8F5FFFDFFE94EA3C8B8FD92F930" author="Smith, M. A." box="[424,562,1714,1740]" pageId="6" pageNumber="561" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
); 
<taxonomicName id="4C34DE87FFFDFFE94D43C8B9FCF2F930" authority="Vassilieva" authorityName="Vassilieva" box="[584,850,1714,1741]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="arenarius">
<emphasis id="B9407916FFFDFFE94D43C8B9FD78F930" box="[584,728,1715,1741]" italics="true" pageId="6" pageNumber="561">O. arenarius</emphasis>
Vassilieva
</taxonomicName>
by having a larger size (TL 
<quantity id="4CCC08E1FFFDFFE94B96C8B8FB5CF931" box="[1181,1276,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="8.879999999999999" pageId="6" pageNumber="561" unit="mm" value="888.0">888 mm</quantity>
vs. 
<quantity id="4CCC08E1FFFDFFE94A26C8B8FA30F930" box="[1325,1424,1714,1740]" metricMagnitude="-1" metricUnit="m" metricValue="3.8899999999999997" pageId="6" pageNumber="561" unit="mm" value="389.0">389 mm</quantity>
), more midbody scale rows (19 vs. 17), more maxillary teeth (10 vs 6–8), and more ventrals (173–193 vs. 131–144) (
<bibRefCitation id="EFA5D8F5FFFDFFE94F94C8F0FEF9F8E9" author="Vassilieva, A. B." box="[159,345,1786,1813]" pageId="6" pageNumber="561" pagination="211 - 226" refId="ref8711" refString="Vassilieva, A. B. (2015) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from coastal southern Vietnam. Zootaxa, 4058 (2), 211 - 226. https: // doi. org / 10.11646 / zootaxa. 4058.2.4" type="journal article" year="2015">Vassilieva 2015</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFDFFE94EA5C8F1FCA0F8E8" authority="Leong &amp; Grismer" authorityName="Leong &amp; Grismer" authorityYear="2004" box="[430,768,1786,1813]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="booliati">
<emphasis id="B9407916FFFDFFE94EA5C8F1FD8AF8E8" box="[430,554,1786,1813]" italics="true" pageId="6" pageNumber="561">O. booliati</emphasis>
Leong &amp; Grismer
</taxonomicName>
by having more supralabials (8 vs. 6 or 7), more ventrals (173–193 vs. 143–153), and dorsal color pattern (blotches vs. cross-bars) (
<bibRefCitation id="EFA5D8F5FFFDFFE94CE3C914FB59F8C4" author="Leong, T. M. &amp; Grismer, L. L." box="[1000,1273,1822,1848]" pageId="6" pageNumber="561" pagination="12 - 16" refId="ref7703" refString="Leong, T. M. &amp; Grismer, L. L. (2004) A new species of kukri snake, Oligodon (Colubridae) from Pulau Tioman, West Malaysia. Asiatic Herpetological Research, 10, 12 - 16." type="journal article" year="2004">Leong &amp; Grismer 2004</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFDFFE94A46C915FC0DF8A0" authority="Vassilieva, Geissler, Galoyan, Poyarkov, Devenger &amp; Bohme" authorityName="Vassilieva, Geissler, Galoyan, Poyarkov, Devenger &amp; Bohme" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="cattienensis">
<emphasis id="B9407916FFFDFFE94A46C915FF59F8A0" italics="true" pageId="6" pageNumber="561">O. cattienensis</emphasis>
Vassilieva, Geissler, Galoyan, Poyarkov, Devenger &amp; Bohme
</taxonomicName>
by having more midbody scale rows (19 vs. 17), more subcaudals in males (58 or 59 vs. 32–36) and in females (44 or 45 vs. 32); fewer maxillary teeth (10 vs. 11 or 12), fewer number of dorsal blotches (13–15+3–4 vs. 24–33+5), and remarkably longer hemipenis (extending to SC 25 vs. 11 or 12) (
<bibRefCitation id="EFA5D8F5FFFDFFE94ECFC9A4FD1AF834" author="Vassilieva, A. B. &amp; Geissler, P. &amp; Galoyan, E. A. &amp; Poyarkov, N. A. &amp; van Devender, R. W. &amp; Bohme, W." box="[452,698,1966,1992]" pageId="6" pageNumber="561" pagination="233 - 246" refId="ref8763" refString="Vassilieva, A. B., Geissler, P., Galoyan, E. A., Poyarkov, N. A., van Devender, R. W. &amp; Bohme, W. (2013) A new species of kukri snake (Oligodon Fitzinger, 1826; Squamata: Colubridae) from the Cat Tien National Park, southern Vietnam. Zootaxa, 3702 (3), 233 - 246. https: // doi. org / 10.11646 / zootaxa. 3702.3.2" type="journal article" year="2013">
Vassilieva 
<emphasis id="B9407916FFFDFFE94D34C9A5FDD7F834" box="[575,631,1966,1992]" italics="true" pageId="6" pageNumber="561">et al.</emphasis>
2013
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFDFFE94C00C9A5FBA0F834" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[779,1024,1966,1993]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="561" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="B9407916FFFDFFE94C00C9A5FC2BF834" box="[779,907,1967,1993]" italics="true" pageId="6" pageNumber="561">O. cinereus</emphasis>
(Günther)
</taxonomicName>
by having more midbody scale rows (19 vs. 17), more ventrals in females (187–193 vs. 165–185), more subcaudals (44–59 vs. 29–43), fewer maxillary teeth (10 vs. 11–14), remarkably longer hemipenis (extending to SC 25 vs. 11–14), and different dorsal color pattern (blotches vs. uniform or cross-bars or reticulations) (
<bibRefCitation id="EFA5D8F5FFFCFFE84C34CEB1FC67FF29" author="Smith, M. A." box="[831,967,187,213]" pageId="7" pageNumber="562" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84CD8CEB1FB3DFF29" author="David, P. &amp; Nguyen, T. Q. &amp; Nguyen, T. T. &amp; Jiang, K. &amp; Chen, T. &amp; Teynie, A. &amp; Ziegler, T." box="[979,1181,187,213]" pageId="7" pageNumber="562" pagination="45 - 62" refId="ref7291" refString="David, P., Nguyen, T. Q., Nguyen, T. T., Jiang, K., Chen, T., Teynie, A. &amp; Ziegler, T. (2012) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa, 3498 (1), 45 - 62. https: // doi. org / 10.11646 / zootaxa. 3498.1.3" type="journal article" year="2012">
David 
<emphasis id="B9407916FFFCFFE84B14CEB1FBF1FF29" box="[1055,1105,187,213]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2012
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84BE4CEB1FDA1FF05" authority="Nguyen, Nguyen, Le &amp; Murphy" authorityName="Nguyen, Nguyen, Le &amp; Murphy" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="condaoensis">
<emphasis id="B9407916FFFCFFE84BE4CEB1FA3CFF29" box="[1263,1436,187,213]" italics="true" pageId="7" pageNumber="562">O. condaoensis</emphasis>
Nguyen, Nguyen, Le &amp; Murphy
</taxonomicName>
by having more midbody scale rows (19 vs. 17), more subcaudals in males (58 or 59 vs. 37) and in females (44 or 45 vs. 33–34), fewer maxillary teeth (10 vs. 11–13), remarkably longer hemipenis (extending to SC 25 vs. 13 or 14), and different dorsal color pattern (blotches vs. uniform or faint stripe) (
<bibRefCitation id="EFA5D8F5FFFCFFE84A49CF2DFEB5FE99" author="Nguyen, S. N. &amp; Nguyen, V, D. H. &amp; Le, S. H. &amp; Murphy, R. W." pageId="7" pageNumber="562" pagination="261 - 273" refId="ref7900" refString="Nguyen, S. N., Nguyen, V, D. H., Le, S. H. &amp; Murphy, R. W. (2016) A new species of kukri snake (Squamata: Colubridae: Oligodon Fitzinger, 1826) from Con Dao Islands, southern Vietnam. Zootaxa, 4139 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4139.2.9" type="journal article" year="2016">
Nguyen 
<emphasis id="B9407916FFFCFFE84F9CCF41FF69FE99" box="[151,201,331,357]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2016
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84E6CCF41FB99FE99" authority="Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha" authorityName="Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha" box="[359,1081,331,357]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="huahin">
<emphasis id="B9407916FFFCFFE84E6CCF41FE78FE99" box="[359,472,331,357]" italics="true" pageId="7" pageNumber="562">O. huahin</emphasis>
Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha
</taxonomicName>
by having more midbody scale rows (19 vs. 17 or 15), more subcaudals in males (58 or 59 vs. 35–41), more maxillary teeth (10 vs. 6), remarkably longer hemipenis (extending to SC 25 vs. 14), and different dorsal color pattern (blotches vs. uniform or faint stripe) (
<bibRefCitation id="EFA5D8F5FFFCFFE84F94CFBDFE23FE2D" author="Pauwels, O. S. G. &amp; Larsen, H. &amp; Suthanthangjai, W. &amp; David, P. &amp; Sumontha, M." box="[159,387,439,465]" pageId="7" pageNumber="562" pagination="531 - 548" refId="ref8162" refString="Pauwels, O. S. G., Larsen, H., Suthanthangjai, W., David, P. &amp; Sumontha, M. (2017) a new kukri snake (Colubridae: Oligodon) from Hua Hin District, and the first record of O. deuvei from Thailand. Zootaxa, 4291 (3), 531 - 548. https: // doi. org / 10.11646 / zootaxa. 4291.3.6" type="journal article" year="2017">
Pauwels 
<emphasis id="B9407916FFFCFFE84E0DCFBDFE98FE2D" box="[262,312,439,465]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2017
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84EDDCFBDFD16FE2D" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1917" box="[470,694,439,465]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="joynsoni">
<emphasis id="B9407916FFFCFFE84EDDCFBDFDF8FE2C" box="[470,600,439,465]" italics="true" pageId="7" pageNumber="562">O. joynsoni</emphasis>
(Smith)
</taxonomicName>
by having more midbody scale rows (19 vs. 17), fewer maxillary teeth (10 vs. 11 or 12), remarkably longer hemipenis (extending to SC 25 vs. 14), and different dorsal color pattern (blotches vs. cross-bars or reticulations) (
<bibRefCitation id="EFA5D8F5FFFCFFE84D6ECFF5FD4BFDE5" author="Smith, M. A." box="[613,747,511,537]" pageId="7" pageNumber="562" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84DFCCFF5FC1CFDE5" author="David, P. &amp; Nguyen, T. Q. &amp; Nguyen, T. T. &amp; Jiang, K. &amp; Chen, T. &amp; Teynie, A. &amp; Ziegler, T." box="[759,956,511,537]" pageId="7" pageNumber="562" pagination="45 - 62" refId="ref7291" refString="David, P., Nguyen, T. Q., Nguyen, T. T., Jiang, K., Chen, T., Teynie, A. &amp; Ziegler, T. (2012) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from northern Vietnam, southern China and central Laos. Zootaxa, 3498 (1), 45 - 62. https: // doi. org / 10.11646 / zootaxa. 3498.1.3" type="journal article" year="2012">
David 
<emphasis id="B9407916FFFCFFE84C48CFF5FCD4FDE5" box="[835,884,511,537]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2012
</bibRefCitation>
, 
<bibRefCitation id="EFA5D8F5FFFCFFE84CCCCFF5FBA4FDE5" author="David, P. &amp; Hauser, S. &amp; Vogel, G." box="[967,1028,511,537]" pageId="7" pageNumber="562" pagination="12 - 24" refId="ref7097" refString="David, P., Hauser, S. &amp; Vogel, G. (2022) A new species of the genus Oligodon Fitzinger, 1826 (Reptilia: Colubridae) from southern Laos. Taprobanica, 11, 12 - 24. https: // doi. org / 10.47605 / tapro. v 11 i 1.273" type="journal article" year="2022">2022</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84B5ECFF5FAE1FDE5" authority="(Angel)" baseAuthorityName="Angel" baseAuthorityYear="1927" box="[1109,1345,511,537]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="B9407916FFFCFFE84B5ECFF5FB42FDE4" box="[1109,1250,511,537]" italics="true" pageId="7" pageNumber="562">O. macrurus</emphasis>
(Angel)
</taxonomicName>
by having a shorter tail (TaL/TL = 
<geoCoordinate id="EE00C3C3FFFCFFE84ED9CC29FDB1FDC1" box="[466,529,547,573]" degrees="0.184" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.184">0.184</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84D1ACC29FDFDFDC1" box="[529,605,547,573]" degrees="0.188" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.188">–0.188</geoCoordinate>
vs. 0.240), more midbody scale rows (19 vs. 17), more ventrals (173–193 vs. 143–152), fewer subcaudals (44–59 vs. 76–83), a shorter hemipenis (LHSC = 25 vs. 29) (
<bibRefCitation id="EFA5D8F5FFFCFFE84B9DCC4DFABCFD9D" author="Smith, M. A." box="[1174,1308,583,609]" pageId="7" pageNumber="562" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84A2DCC4DFF56FD79" author="Geissler, P. &amp; Krohn, A. R. &amp; Rennert, D." pageId="7" pageNumber="562" pagination="317 - 324" refId="ref7533" refString="Geissler, P., Krohn, A. R. &amp; Rennert, D. (2011) Herpetofauna records in coastal dune areas, Binh Thuan Province, southern Vietnam, with the rediscovery of Oligodon macrurus Angel, 1927. Russian Journal of Herpetology, 18, 317 - 324. https: // doi. org / 10.30906 / 1026 - 2296 - 2011 - 18 - 4 - 317 - 324" type="journal article" year="2011">
Geissler 
<emphasis id="B9407916FFFCFFE84A83CC4DFF0DFD79" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2011
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84E43CC61FD77FD79" authority="David, Vogel &amp; Rooijen" authorityName="David, Vogel &amp; Rooijen" box="[328,727,619,645]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="moricei">
<emphasis id="B9407916FFFCFFE84E43CC61FE60FD78" box="[328,448,619,645]" italics="true" pageId="7" pageNumber="562">O. moricei</emphasis>
David, Vogel &amp; Rooijen
</taxonomicName>
by having more midbody scale rows (19 vs. 17), more ventrals (187–193 vs. 
<quantity id="4CCC08E1FFFCFFE84E3ACC84FED8FD55" box="[305,376,654,681]" metricMagnitude="0" metricUnit="m" metricValue="4.445" pageId="7" pageNumber="562" unit="in" value="175.0">175 in</quantity>
females), more subcaudals (44 or 45 vs 
<quantity id="4CCC08E1FFFCFFE84C4BCC85FCD9FD55" box="[832,889,655,681]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="7" pageNumber="562" unit="in" value="41.0">41 in</quantity>
females), and dorsal blotches (vs. stripe) (
<bibRefCitation id="EFA5D8F5FFFCFFE84A5DCC85FE80FD31" author="David, P. &amp; Vogel, G. &amp; van Rooijen, J." pageId="7" pageNumber="562" pagination="1 - 49" refId="ref7219" refString="David, P., Vogel, G. &amp; van Rooijen, J. (2008 b) A revision of the Oligodon taeniatus (Gunther, 1861) group (Squamata: Colubridae), with the description of three new species from the Indochinese region. Zootaxa, 1965 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 1965.1.1" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84F9CCCB9FF6FFD31" box="[151,207,691,717]" italics="true" pageId="7" pageNumber="562">et al.</emphasis>
2008b
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84E7BCCB9FC40FD31" authority="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" box="[368,992,691,717]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="phangan">
<emphasis id="B9407916FFFCFFE84E7BCCB9FE54FD31" box="[368,500,691,717]" italics="true" pageId="7" pageNumber="562">O. phangan</emphasis>
Pauwels, Thongyai, Chantong &amp; Sumontha
</taxonomicName>
by having fewer maxillary teeth (10 vs. 12), more midbody scale rows (19 vs. 17), more ventrals (173–193 vs. 163–166), and more subcaudals (44–59 vs. 33–42) (
<bibRefCitation id="EFA5D8F5FFFCFFE84FFCCCF1FE7CFCE9" author="Pauwels, O. S. G. &amp; Thongyai, K. &amp; Chantong, P. &amp; Sumontha, M." box="[247,476,763,789]" pageId="7" pageNumber="562" pagination="537 - 557" refId="ref8241" refString="Pauwels, O. S. G., Thongyai, K., Chantong, P. &amp; Sumontha, M. (2021) Two new kukri snake species (Colubridae: Oligodon) from the Nakhon Si Thammarat Moutain Rang, and addition of O. ocellatus to the fauna of Thailand. Zootaxa, 4908 (4), 537 - 557. https: // doi. org / 10.11646 / zootaxa. 4908.4.7" type="journal article" year="2021">
Pauwels 
<emphasis id="B9407916FFFCFFE84E55CCF1FE31FCE9" box="[350,401,763,789]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2021
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84D3BCCF1FB7BFCE9" authority="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" box="[560,1243,763,789]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="promsombuti">
<emphasis id="B9407916FFFCFFE84D3BCCF1FD47FCE9" box="[560,743,763,789]" italics="true" pageId="7" pageNumber="562">O. promsombuti</emphasis>
Pauwels, Thongyai, Chantong &amp; Sumontha
</taxonomicName>
by having fewer maxillary teeth (10 vs. 12), more midbody scale rows (19 vs. 17), more subcaudals (44–59 vs. 40), and remarkably longer hemipenis (extending to SC 25 vs. 13) (
<bibRefCitation id="EFA5D8F5FFFCFFE84DBBCD49FC36FCA1" author="Pauwels, O. S. G. &amp; Thongyai, K. &amp; Chantong, P. &amp; Sumontha, M." box="[688,918,835,861]" pageId="7" pageNumber="562" pagination="537 - 557" refId="ref8241" refString="Pauwels, O. S. G., Thongyai, K., Chantong, P. &amp; Sumontha, M. (2021) Two new kukri snake species (Colubridae: Oligodon) from the Nakhon Si Thammarat Moutain Rang, and addition of O. ocellatus to the fauna of Thailand. Zootaxa, 4908 (4), 537 - 557. https: // doi. org / 10.11646 / zootaxa. 4908.4.7" type="journal article" year="2021">
Pauwels 
<emphasis id="B9407916FFFCFFE84C1CCD49FCEAFCA1" box="[791,842,835,861]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2021
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84CE0CD49FA83FCA1" authority="(Schlegel)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1003,1315,835,861]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="purpurascens">
<emphasis id="B9407916FFFCFFE84CE0CD49FB07FCA0" box="[1003,1191,835,861]" italics="true" pageId="7" pageNumber="562">O. purpurascens</emphasis>
(Schlegel)
</taxonomicName>
by having faint and interrupted temporal streak (vs. distinct and continuous temporal streak), different dorsal color pattern (blotches vs. conspicuous red and banded coloration or brownish dull coloration), and forked hemipenis without papillae (vs. unforked with papillae) (
<bibRefCitation id="EFA5D8F5FFFCFFE84D30CDA5FD53FC35" author="Boulenger, G. A." box="[571,755,943,969]" pageId="7" pageNumber="562" refId="ref6839" refString="Boulenger, G. A. (1894) Catalogue of the snakes in the British Museum. Vol. 2: Colubridae Aglyphae. British Museum of Natural History, London, 382 pp., 20 pls." type="book" year="1894">Boulenger 1894</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84DF5CDA5FC76FC35" author="Rooijen, J. V. &amp; Wood, P. L. &amp; Grismer, J. L. &amp; Grismer, L. L. &amp; Grossmann, W." box="[766,982,943,969]" pageId="7" pageNumber="562" pagination="215 - 220" refId="ref8453" refString="Rooijen, J. V., Wood, P. L., Grismer, J. L., Grismer, L. L. &amp; Grossmann, W. (2011) Color pattern dimorphism in the colubrid snake Oligodon purpurascens (Schlegel, 1837) (Reptilia: Squamata). Russian Journal of Herpetology, 18, 215 - 220." type="journal article" year="2011">
Rooijen 
<emphasis id="B9407916FFFCFFE84C56CDA5FC2EFC35" box="[861,910,943,969]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2011
</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84CEACDA5FB1EFC35" author="Pauwels, O. S. G. &amp; Larsen, H. &amp; Suthanthangjai, W. &amp; David, P. &amp; Sumontha, M." box="[993,1214,943,969]" pageId="7" pageNumber="562" pagination="531 - 548" refId="ref8162" refString="Pauwels, O. S. G., Larsen, H., Suthanthangjai, W., David, P. &amp; Sumontha, M. (2017) a new kukri snake (Colubridae: Oligodon) from Hua Hin District, and the first record of O. deuvei from Thailand. Zootaxa, 4291 (3), 531 - 548. https: // doi. org / 10.11646 / zootaxa. 4291.3.6" type="journal article" year="2017">
Pauwels 
<emphasis id="B9407916FFFCFFE84B4ECDA5FBD6FC35" box="[1093,1142,943,969]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84BC1CDA5FA30FC35" author="David, P. &amp; Hauser, S. &amp; Vogel, G." box="[1226,1424,943,969]" pageId="7" pageNumber="562" pagination="12 - 24" refId="ref7097" refString="David, P., Hauser, S. &amp; Vogel, G. (2022) A new species of the genus Oligodon Fitzinger, 1826 (Reptilia: Colubridae) from southern Laos. Taprobanica, 11, 12 - 24. https: // doi. org / 10.47605 / tapro. v 11 i 1.273" type="journal article" year="2022">
David 
<emphasis id="B9407916FFFCFFE84A1ECDA5FAE6FC35" box="[1301,1350,943,969]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2022
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84FDECDD9FD5AFC11" authority="Sumontha, Kunya, Dangsri &amp; Pauwels" authorityName="Sumontha, Kunya, Dangsri &amp; Pauwels" box="[213,762,979,1005]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="saiyok">
<emphasis id="B9407916FFFCFFE84FDECDD9FEE0FC11" box="[213,320,979,1005]" italics="true" pageId="7" pageNumber="562">O. saiyok</emphasis>
Sumontha, Kunya, Dangsri &amp; Pauwels
</taxonomicName>
by having fewer maxillary teeth (10 vs. 13), more midbody scale rows (19 vs. 17), more subcaudals (58 or 59 vs. 
<quantity id="4CCC08E1FFFCFFE84DEFCDFDFCBCFBED" box="[740,796,1015,1041]" metricMagnitude="0" metricUnit="m" metricValue="1.0922" pageId="7" pageNumber="562" unit="in" value="43.0">43 in</quantity>
males, 44 or 45 vs. 
<quantity id="4CCC08E1FFFCFFE84CFDCDFDFB8EFBED" box="[1014,1070,1015,1041]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="7" pageNumber="562" unit="in" value="38.0">38 in</quantity>
females), and remarkably longer hemipenis (extending to SC 25 vs. 18) (
<bibRefCitation id="EFA5D8F5FFFCFFE84D47CA11FC9FFBC9" author="Sumontha, M. &amp; Kunya, K. &amp; Dangsri, S. &amp; Pauwels, O. S. G." box="[588,831,1051,1077]" pageId="7" pageNumber="562" pagination="316 - 328" refId="ref8602" refString="Sumontha, M., Kunya, K., Dangsri, S. &amp; Pauwels, O. S. G. (2017) Oligodon saiyok, a new limestone-dwelling kukri snake (Serpentes: Colubridae) from Kanchanaburi Province, western Thailand. Zootaxa, 4294 (3), 316 - 328. https: // doi. org / 10.11646 / zootaxa. 4294.3.2" type="journal article" year="2017">
Sumontha 
<emphasis id="B9407916FFFCFFE84DCECA11FD55FBC9" box="[709,757,1051,1077]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2017
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84C86CA11FB20FBC9" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[909,1152,1051,1077]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="signatus">
<emphasis id="B9407916FFFCFFE84C86CA11FBABFBC8" box="[909,1035,1051,1077]" italics="true" pageId="7" pageNumber="562">O. signatus</emphasis>
(Gunther)
</taxonomicName>
by having more midbody scale rows (19 vs. 15–17), more ventrals (173–179 vs. 
<quantity id="4CCC08E1FFFCFFE84C06CA35FC2EFBA5" box="[781,910,1087,1113]" metricMagnitude="0" metricUnit="m" metricValue="3.683" metricValueMax="3.7845999999999997" metricValueMin="3.5814" pageId="7" pageNumber="562" unit="in" value="145.0" valueMax="149.0" valueMin="141.0">141–149 in</quantity>
males, 187–193 vs. 
<quantity id="4CCC08E1FFFCFFE84B71CA34FB5BFBA5" box="[1146,1275,1086,1113]" metricMagnitude="0" metricUnit="m" metricValue="3.9116" metricValueMax="3.9878" metricValueMin="3.8354" pageId="7" pageNumber="562" unit="in" value="154.0" valueMax="157.0" valueMin="151.0">151–157 in</quantity>
females), and dorsal blotches (vs. cross bands or transverse rhomboidal spots) (
<bibRefCitation id="EFA5D8F5FFFCFFE84C78CA69FB8CFB81" author="Boulenger, G. A." box="[883,1068,1123,1149]" pageId="7" pageNumber="562" refId="ref6839" refString="Boulenger, G. A. (1894) Catalogue of the snakes in the British Museum. Vol. 2: Colubridae Aglyphae. British Museum of Natural History, London, 382 pp., 20 pls." type="book" year="1894">Boulenger 1894</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84B33CA69FAA0FB81" author="David, P. &amp; Hauser, S. &amp; Vogel, G." box="[1080,1280,1123,1149]" pageId="7" pageNumber="562" pagination="12 - 24" refId="ref7097" refString="David, P., Hauser, S. &amp; Vogel, G. (2022) A new species of the genus Oligodon Fitzinger, 1826 (Reptilia: Colubridae) from southern Laos. Taprobanica, 11, 12 - 24. https: // doi. org / 10.47605 / tapro. v 11 i 1.273" type="journal article" year="2022">
David 
<emphasis id="B9407916FFFCFFE84B8FCA69FB15FB81" box="[1156,1205,1123,1149]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2022
</bibRefCitation>
); and from 
<taxonomicName id="4C34DE87FFFCFFE84A8ACA69FE24FB5D" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1875" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="splendidus">
<emphasis id="B9407916FFFCFFE84A8ACA69FEAFFB5D" italics="true" pageId="7" pageNumber="562">O. splendidus</emphasis>
(Günther)
</taxonomicName>
by having fewer midbody scale rows (19 vs. 21), a distinct vertebral stripe (vs. vertebral stripe absent), and remarkably longer hemipenis (extending to SC 25 vs. 19) (
<bibRefCitation id="EFA5D8F5FFFCFFE84CBCCAA1FB9FFB39" author="Smith, M. A." box="[951,1087,1195,1221]" pageId="7" pageNumber="562" refId="ref8555" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptiles and Amphibians. Vol. III - Serpentes. Taylor and Francis Ltd, London, 525 pp." type="book" year="1943">Smith 1943</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BA504FFFCFFE84FCCCAC5FE0DF821" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="562">
<taxonomicName id="4C34DE87FFFCFFE84FCCCAC5FED2FB14" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[199,370,1231,1257]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFFCFFE84FCCCAC5FED2FB14" box="[199,370,1231,1257]" italics="true" pageId="7" pageNumber="562">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFFCFFE84E72CAC5FE74FB15" bold="true" box="[377,468,1231,1257]" pageId="7" pageNumber="562">
<taxonomicNameLabel id="A273C46DFFFCFFE84E72CAC5FE74FB15" box="[377,468,1231,1257]" pageId="7" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to its congeners in the 
<taxonomicName id="4C34DE87FFFCFFE84C42CAC5FC6BFB15" baseAuthorityName="Cantor" baseAuthorityYear="1839" box="[841,971,1231,1257]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="B9407916FFFCFFE84C42CAC5FC6BFB15" box="[841,971,1231,1257]" italics="true" pageId="7" pageNumber="562">O. cyclurus</emphasis>
</taxonomicName>
species complex. However, the new species can be distinguished from 
<taxonomicName id="4C34DE87FFFCFFE84EF2CAF9FD24FAF1" baseAuthorityName="Gunther" baseAuthorityYear="1888" box="[505,644,1267,1293]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="B9407916FFFCFFE84EF2CAF9FD24FAF1" box="[505,644,1267,1293]" italics="true" pageId="7" pageNumber="562">O. chinensis</emphasis>
</taxonomicName>
by having more midbody scale rows (19 vs. 17), fewer subcaudals in females (44 or 45 vs. 47–53), remarkably longer hemipenis (extending to SC 25 vs. 12–14), and dorsal blotches (vs. cross bars) (
<bibRefCitation id="EFA5D8F5FFFCFFE84E17CB31FE4DFAA9" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[284,493,1339,1365]" pageId="7" pageNumber="562" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84E63CB31FE38FAA9" box="[360,408,1339,1365]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2008a
</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFFCFFE84EFDCB31FD6EFAA9" author="Nguyen, S. N. &amp; Nguyen, L. T. &amp; Nguyen, V, D. H. &amp; Phan, H. T. &amp; Jiang, K. &amp; Murphy, R. W." box="[502,718,1339,1365]" pageId="7" pageNumber="562" pagination="333 - 346" refId="ref7979" refString="Nguyen, S. N., Nguyen, L. T., Nguyen, V, D. H., Phan, H. T., Jiang, K. &amp; Murphy, R. W. (2017) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Cu Lao Cham Islands, central Vietnam. Zootaxa, 4286 (3), 333 - 346. https: // doi. org / 10.11646 / zootaxa. 4286.3.2" type="journal article" year="2017">
Nguyen 
<emphasis id="B9407916FFFCFFE84D5ECB31FD24FAA9" box="[597,644,1339,1365]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2017
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84C17CB31FC50FAA9" box="[796,1008,1339,1365]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="culaochamensis">
<emphasis id="B9407916FFFCFFE84C17CB31FC50FAA9" box="[796,1008,1339,1365]" italics="true" pageId="7" pageNumber="562">O. culaochamensis</emphasis>
</taxonomicName>
by having a shorter tail (TaL/TL 0.184 vs. 
<quantity id="4CCC08E1FFFCFFE84FB4CB55FEC9FA85" box="[191,361,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="5.5499" metricValueMax="5.5626" metricValueMin="5.5372" pageId="7" pageNumber="562" unit="in" value="0.2185" valueMax="0.219" valueMin="0.218">
<geoCoordinate id="EE00C3C3FFFCFFE84FB4CB55FF5EFA85" box="[191,254,1375,1401]" degrees="0.218" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.218">0.218</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84FF5CB55FEEBFA85" box="[254,331,1375,1401]" degrees="0.219" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.219">–0.219</geoCoordinate>
in
</quantity>
males, 0.123 vs. 
<quantity id="4CCC08E1FFFCFFE84D3ACB55FD7BFA85" box="[561,731,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="4.2545" metricValueMax="4.2926" metricValueMin="4.2164" pageId="7" pageNumber="562" unit="in" value="0.1675" valueMax="0.169" valueMin="0.166">
<geoCoordinate id="EE00C3C3FFFCFFE84D3ACB55FDD0FA85" box="[561,624,1375,1401]" degrees="0.166" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.166">0.166</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84D7BCB55FD1DFA85" box="[624,701,1375,1401]" degrees="0.169" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.169">–0.169</geoCoordinate>
in
</quantity>
females), more ventrals in both males (173–179 vs. 167–169) and females (187–193 vs. 179–182), fewer subcaudals in both males (58 or 59 vs. 63–66) and females (44 or 45 vs. 51 or 
<quantity id="4CCC08E1FFFCFFE84FD1CBACFEB3FA3D" box="[218,275,1446,1473]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" pageId="7" pageNumber="562" unit="in" value="52.0">52 in</quantity>
females), and remarkably longer hemipenis (extending to SC 25 vs. 20) (
<bibRefCitation id="EFA5D8F5FFFCFFE84B5BCBADFA8FFA3D" author="Nguyen, S. N. &amp; Nguyen, L. T. &amp; Nguyen, V, D. H. &amp; Phan, H. T. &amp; Jiang, K. &amp; Murphy, R. W." box="[1104,1327,1447,1473]" pageId="7" pageNumber="562" pagination="333 - 346" refId="ref7979" refString="Nguyen, S. N., Nguyen, L. T., Nguyen, V, D. H., Phan, H. T., Jiang, K. &amp; Murphy, R. W. (2017) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Cu Lao Cham Islands, central Vietnam. Zootaxa, 4286 (3), 333 - 346. https: // doi. org / 10.11646 / zootaxa. 4286.3.2" type="journal article" year="2017">
Nguyen 
<emphasis id="B9407916FFFCFFE84BB9CBADFB44FA3D" box="[1202,1252,1447,1473]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2017
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84A8ACBADFF56FA19" baseAuthorityName="Cantor" baseAuthorityYear="1839" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="B9407916FFFCFFE84A8ACBADFF56FA19" italics="true" pageId="7" pageNumber="562">O. cyclurus</emphasis>
</taxonomicName>
in having a longer tail (TaL/SVL 0.225 vs. 
<quantity id="4CCC08E1FFFCFFE84DD7CBC0FC24FA19" box="[732,900,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5941000000000005" metricValueMax="3.9623999999999997" metricValueMin="3.2258000000000004" pageId="7" pageNumber="562" unit="in" value="0.1415" valueMax="0.156" valueMin="0.127">
<geoCoordinate id="EE00C3C3FFFCFFE84DD7CBC0FCBBFA19" box="[732,795,1482,1509]" degrees="0.127" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.127">0.127</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84C11CBC0FCC8FA19" box="[794,872,1482,1509]" degrees="0.156" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.156">–0.156</geoCoordinate>
in
</quantity>
males, 0.140 vs. 
<quantity id="4CCC08E1FFFCFFE84B4DCBC1FB4EFA19" box="[1094,1262,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.1115" metricValueMax="3.5052" metricValueMin="2.7178" pageId="7" pageNumber="562" unit="in" value="0.1225" valueMax="0.138" valueMin="0.107">
<geoCoordinate id="EE00C3C3FFFCFFE84B4DCBC1FB25FA19" box="[1094,1157,1483,1509]" degrees="0.107" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.107">0.107</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84B8ECBC1FB72FA19" box="[1157,1234,1483,1509]" degrees="0.138" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.138">–0.138</geoCoordinate>
in
</quantity>
females), more ventrals in both males (173–179 vs. 160–173) and females (187–193 vs. 168–172), more subcaudals in both males (58 or 59 vs. 37–48) and females (44 or 45 vs. 
<quantity id="4CCC08E1FFFCFFE84DA0C819FCAFF9D1" box="[683,783,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="11.176" metricValueMin="7.62" pageId="7" pageNumber="562" unit="in" value="37.0" valueMax="44.0" valueMin="30.0">30–44 in</quantity>
females), and remarkably longer hemipenis (extending to SC 25 vs. 15–18) (
<bibRefCitation id="EFA5D8F5FFFCFFE84E65C83DFDE7F9AD" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[366,583,1591,1617]" pageId="7" pageNumber="562" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84EB7C83DFE4FF9AD" box="[444,495,1591,1617]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2008a
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84D90C83DFCE1F9AD" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[667,833,1591,1617]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="B9407916FFFCFFE84D90C83DFCE1F9AD" box="[667,833,1591,1617]" italics="true" pageId="7" pageNumber="562">O. formosanus</emphasis>
</taxonomicName>
by having a longer tail in males (TaL/SVL 0.225 vs. 
<geoCoordinate id="EE00C3C3FFFCFFE84F9CC850FF77F989" box="[151,215,1626,1653]" degrees="0.165" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.165">0.165</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84FDCC850FE85F989" box="[215,293,1626,1653]" degrees="0.195" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.195">–0.195</geoCoordinate>
) and shorter tail in females (TaL/SVL 0.140 vs. 
<geoCoordinate id="EE00C3C3FFFCFFE84C48C851FC24F989" box="[835,900,1627,1653]" degrees="0.149" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.149">0.149</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84C8FC851FC74F989" box="[900,980,1627,1653]" degrees="0.164" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.164">–0.164</geoCoordinate>
), more subcaudals in males (58 or 59 vs. 49–55), remarkably longer hemipenis (extending to SC 25 vs. 15–18), and dorsal blotches (vs. reticulate pattern) (
<bibRefCitation id="EFA5D8F5FFFCFFE84F94C8A9FED4F941" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[159,372,1699,1725]" pageId="7" pageNumber="562" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84FE7C8A9FEBDF941" box="[236,285,1699,1725]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2008a
</bibRefCitation>
); from 
<taxonomicName id="4C34DE87FFFCFFE84ECFC8A9FDEBF941" baseAuthorityName="Morice" baseAuthorityYear="1875" box="[452,587,1699,1725]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9407916FFFCFFE84ECFC8A9FDEBF941" box="[452,587,1699,1725]" italics="true" pageId="7" pageNumber="562">O. ocellatus</emphasis>
</taxonomicName>
by having a longer tail (TaL/SVL 0.225 vs. 
<quantity id="4CCC08E1FFFCFFE84B32C8A9FB41F941" box="[1081,1249,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="3.2131" metricValueMax="3.5813999999999995" metricValueMin="2.8448" pageId="7" pageNumber="562" unit="in" value="0.1265" valueMax="0.141" valueMin="0.112">
<geoCoordinate id="EE00C3C3FFFCFFE84B32C8A9FBD8F941" box="[1081,1144,1699,1725]" degrees="0.112" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.112">0.112</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84B73C8A9FB64F941" box="[1144,1220,1699,1725]" degrees="0.141" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.141">–0.141</geoCoordinate>
in
</quantity>
males, 0.140 vs. 
<quantity id="4CCC08E1FFFCFFE84F9CC8CDFEE0F91D" box="[151,320,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.6416" metricValueMax="2.8956" metricValueMin="2.3876" pageId="7" pageNumber="562" unit="in" value="0.104" valueMax="0.114" valueMin="0.094">
<geoCoordinate id="EE00C3C3FFFCFFE84F9CC8CDFF75F91D" box="[151,213,1735,1761]" degrees="0.094" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.094">0.094</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84FDDC8CDFE82F91D" box="[214,290,1735,1761]" degrees="0.114" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.114">–0.114</geoCoordinate>
in
</quantity>
females), more ventrals in both males (173–179 vs. 156–165) and females (187–193 vs. 152–180), more subcaudals in both males (58 or 59 vs. 32–44) and females (44 or 45 vs. 
<quantity id="4CCC08E1FFFCFFE84B19C8E1FBD6F8F9" box="[1042,1142,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="8.382" metricValueMin="6.604" pageId="7" pageNumber="562" unit="in" value="29.5" valueMax="33.0" valueMin="26.0">26–33 in</quantity>
females), and remarkably longer hemipenis (extending to SC 25 vs. 15–17) (
<bibRefCitation id="EFA5D8F5FFFCFFE84DC6C905FC03F8D5" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[717,931,1807,1833]" pageId="7" pageNumber="562" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84C11C905FCECF8D5" box="[794,844,1807,1833]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2008a
</bibRefCitation>
); and from 
<taxonomicName id="4C34DE87FFFCFFE84B28C905FB6BF8D4" box="[1059,1227,1807,1833]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="562" phylum="Chordata" rank="species" species="saintgironsi">
<emphasis id="B9407916FFFCFFE84B28C905FB6BF8D4" box="[1059,1227,1807,1833]" italics="true" pageId="7" pageNumber="562">O. saintgironsi</emphasis>
</taxonomicName>
by having a larger size (TL max 
<quantity id="4CCC08E1FFFCFFE84E3FC939FE32F8B1" box="[308,402,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="8.879999999999999" pageId="7" pageNumber="562" unit="mm" value="888.0">888 mm</quantity>
vs. 
<quantity id="4CCC08E1FFFCFFE84ECAC939FD82F8B1" box="[449,546,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="6.76" pageId="7" pageNumber="562" unit="mm" value="676.0">676 mm</quantity>
), a longer tail in males (TaL/SVL 0.225 vs. 
<geoCoordinate id="EE00C3C3FFFCFFE84B19C939FBF1F8B1" box="[1042,1105,1843,1869]" degrees="0.191" direction="north" orientation="latitude" pageId="7" pageNumber="562" precision="55" value="0.191">0.191</geoCoordinate>
<geoCoordinate id="EE00C3C3FFFCFFE84B5AC939FB00F8B1" box="[1105,1184,1843,1869]" degrees="0.203" direction="west" orientation="longitude" pageId="7" pageNumber="562" precision="55" value="-0.203">–0.203</geoCoordinate>
) and shorter tail in females (TaL/SVL 0.140 vs. 0.161), more midbody scale rows (19 vs. 17 or 18), first scale reduction occurring at the level of ventrals 97–111 (vs. 62–86), more ventrals in both males (173–179 vs. 166–170) and females (187–193 vs. 184), fewer subcaudals in females (44 or 45 vs. 53), and slightly shorter hemipenis (extending to SC 25 vs. 27 or 28) (
<bibRefCitation id="EFA5D8F5FFFCFFE84FC0C9C9FE01F821" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[203,417,1987,2013]" pageId="7" pageNumber="562" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFFCFFE84E13C9C9FEE9F821" box="[280,329,1987,2013]" italics="true" pageId="7" pageNumber="562">et al</emphasis>
. 2008a
</bibRefCitation>
).
</paragraph>
<caption id="DF4BF58CFFF3FFE74F93CC68FBF7FD5C" ID-Table-UUID="DF4BF58CFFF3FFE74F93CC68FBF7FD5C" httpUri="http://table.plazi.org/id/DF4BF58CFFF3FFE74F93CC68FBF7FD5C" pageId="8" pageNumber="563" startId="8.[152,244,610,636]" subCaptionStartIDs="8.[960,1107,646,672]" subCaptionStarts="abbr" targetBox="[166,2003,687,1580]" targetIsTable="true" targetPageId="8">
<paragraph id="8B8BA504FFF3FFE74F93CC68FBF7FD5C" blockId="8.[151,2039,610,672]" pageId="8" pageNumber="563">
<emphasis id="B9407916FFF3FFE74F93CC68FEAEFD80" bold="true" box="[152,270,610,636]" pageId="8" pageNumber="563">TABLE 2.</emphasis>
Main morphological characters of 
<taxonomicName id="4C34DE87FFF3FFE74D88CC68FC85FD80" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[643,805,610,636]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFF3FFE74D88CC68FC85FD80" box="[643,805,610,636]" italics="true" pageId="8" pageNumber="563">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFF3FFE74C21CC68FCDFFD80" bold="true" box="[810,895,610,636]" pageId="8" pageNumber="563">
<taxonomicNameLabel id="A273C46DFFF3FFE74C21CC68FCDFFD80" box="[810,895,610,636]" pageId="8" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in comparison with other 
<taxonomicName id="4C34DE87FFF3FFE74B9ECC68FB5AFD80" authorityName="Fitzinger" authorityYear="1826" box="[1173,1274,610,636]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="genus">
<emphasis id="B9407916FFF3FFE74B9ECC68FB5AFD80" box="[1173,1274,610,636]" italics="true" pageId="8" pageNumber="563">Oligodon</emphasis>
</taxonomicName>
in the 
<taxonomicName id="4C34DE87FFF3FFE74A4ACC68F9A6FD80" baseAuthorityName="Cantor" baseAuthorityYear="1839" box="[1345,1542,610,636]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="B9407916FFF3FFE74A4ACC68F9A6FD80" box="[1345,1542,610,636]" italics="true" pageId="8" pageNumber="563">Oligodon cyclurus</emphasis>
</taxonomicName>
species complex. Data taken from David 
<emphasis id="B9407916FFF3FFE748CACC68F850FD80" box="[1985,2032,610,636]" italics="true" pageId="8" pageNumber="563">et al</emphasis>
. (2008a), 
<bibRefCitation id="EFA5D8F5FFF3FFE74FFCCC8CFE75FD5C" author="Nguyen, S. N. &amp; Nguyen, L. T. &amp; Nguyen, V, D. H. &amp; Phan, H. T. &amp; Jiang, K. &amp; Murphy, R. W." box="[247,469,646,672]" pageId="8" pageNumber="563" pagination="333 - 346" refId="ref7979" refString="Nguyen, S. N., Nguyen, L. T., Nguyen, V, D. H., Phan, H. T., Jiang, K. &amp; Murphy, R. W. (2017) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Cu Lao Cham Islands, central Vietnam. Zootaxa, 4286 (3), 333 - 346. https: // doi. org / 10.11646 / zootaxa. 4286.3.2" type="journal article" year="2017">
Nguyen 
<emphasis id="B9407916FFF3FFE74E59CC8CFE21FD5C" box="[338,385,646,672]" italics="true" pageId="8" pageNumber="563">et al</emphasis>
. (2017)
</bibRefCitation>
and this work. See Material and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="8B8BA504FFF3FFE74FADCCA5F8DBF9F4" pageId="8" pageNumber="563">
<table id="F93457A4FFF300104FADCCA5F873F9D0" box="[166,2003,687,1580]" gridcols="9" gridrows="19" pageId="8" pageNumber="563">
<tr id="3504A746FFF300104FADCCA5F873FD10" box="[166,2003,687,748]" gridrow="0" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCCA5FEDBFD10" box="[166,379,687,748]" gridcol="0" gridrow="0" pageId="8" pageNumber="563">Characters</th>
<th id="76D5CE3AFFF300104EBECCA5FDEFFD10" box="[437,591,687,748]" gridcol="1" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74EBECCA5FDEFFD34" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[437,591,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFF3FFE74EBECCA5FDEFFD34" box="[437,591,687,712]" italics="true" pageId="8" pageNumber="563">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFF3FFE74EBECCD9FDA6FD10" bold="true" box="[437,518,723,748]" pageId="8" pageNumber="563">
<taxonomicNameLabel id="A273C46DFFF3FFE74EBECCD9FDA6FD10" box="[437,518,723,748]" pageId="8" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76D5CE3AFFF300104D85CCA5FC85FD10" box="[654,805,687,748]" gridcol="2" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74D85CCA5FCABFD34" baseAuthorityName="Gunther" baseAuthorityYear="1888" box="[654,779,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="chinensis">
<emphasis id="B9407916FFF3FFE74D85CCA5FCABFD34" box="[654,779,687,712]" italics="true" pageId="8" pageNumber="563">O. chinensis</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104C46CCA5FBAEFD10" box="[845,1038,687,748]" gridcol="3" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74C46CCA5FBAEFD34" box="[845,1038,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="culaochamensis">
<emphasis id="B9407916FFF3FFE74C46CCA5FBAEFD34" box="[845,1038,687,712]" italics="true" pageId="8" pageNumber="563">O. culaochamensis</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104B4ACCA5FB43FD10" box="[1089,1251,687,748]" gridcol="4" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74B4ACCA5FB16FD34" baseAuthorityName="Cantor" baseAuthorityYear="1839" box="[1089,1206,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="B9407916FFF3FFE74B4ACCA5FB16FD34" box="[1089,1206,687,712]" italics="true" pageId="8" pageNumber="563">O. cyclurus</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104A06CCA5FA3AFD10" box="[1293,1434,687,748]" gridcol="5" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74A06CCA5FA3AFD34" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[1293,1434,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="B9407916FFF3FFE74A06CCA5FA3AFD34" box="[1293,1434,687,712]" italics="true" pageId="8" pageNumber="563">O. fasciolatus</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104AB5CCA5F9F3FD10" box="[1470,1619,687,748]" gridcol="6" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74AB5CCA5F9F3FD34" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[1470,1619,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="formosanus">
<emphasis id="B9407916FFF3FFE74AB5CCA5F9F3FD34" box="[1470,1619,687,712]" italics="true" pageId="8" pageNumber="563">O. formosanus</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104976CCA5F8A9FD10" box="[1661,1801,687,748]" gridcol="7" gridrow="0" pageId="8" pageNumber="563">
<taxonomicName id="4C34DE87FFF3FFE74976CCA5F957FD34" baseAuthorityName="Morice" baseAuthorityYear="1875" box="[1661,1783,687,712]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="563" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B9407916FFF3FFE74976CCA5F957FD34" box="[1661,1783,687,712]" italics="true" pageId="8" pageNumber="563">O. ocellatus</emphasis>
</taxonomicName>
</th>
<th id="76D5CE3AFFF300104837CCA5F873FD10" box="[1852,2003,687,748]" gridcol="8" gridrow="0" pageId="8" pageNumber="563">
<emphasis id="B9407916FFF3FFE74837CCA5F873FD34" box="[1852,2003,687,712]" italics="true" pageId="8" pageNumber="563">O. saintgironsi</emphasis>
</th>
</tr>
<tr id="3504A746FFF300104FADCCF1F873FCE8" box="[166,2003,763,788]" gridrow="1" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCCF1FEDBFCE8" box="[166,379,763,788]" gridcol="0" gridrow="1" pageId="8" pageNumber="563">TL max (mm)</th>
<td id="76D5CE3AFFF300104EBECCF1FDEFFCE8" box="[437,591,763,788]" gridcol="1" gridrow="1" pageId="8" pageNumber="563">888</td>
<td id="76D5CE3AFFF300104D85CCF1FC85FCE8" box="[654,805,763,788]" gridcol="2" gridrow="1" pageId="8" pageNumber="563">729</td>
<td id="76D5CE3AFFF300104C46CCF1FBAEFCE8" box="[845,1038,763,788]" gridcol="3" gridrow="1" pageId="8" pageNumber="563">582</td>
<td id="76D5CE3AFFF300104B4ACCF1FB43FCE8" box="[1089,1251,763,788]" gridcol="4" gridrow="1" pageId="8" pageNumber="563">710</td>
<td id="76D5CE3AFFF300104A06CCF1FA3AFCE8" box="[1293,1434,763,788]" gridcol="5" gridrow="1" pageId="8" pageNumber="563">807</td>
<td id="76D5CE3AFFF300104AB5CCF1F9F3FCE8" box="[1470,1619,763,788]" gridcol="6" gridrow="1" pageId="8" pageNumber="563">942</td>
<td id="76D5CE3AFFF300104976CCF1F8A9FCE8" box="[1661,1801,763,788]" gridcol="7" gridrow="1" pageId="8" pageNumber="563">852</td>
<td id="76D5CE3AFFF300104837CCF1F873FCE8" box="[1852,2003,763,788]" gridcol="8" gridrow="1" pageId="8" pageNumber="563">676</td>
</tr>
<tr id="3504A746FFF300104FADCD29F873FCC0" box="[166,2003,803,828]" gridrow="2" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCD29FEDBFCC0" box="[166,379,803,828]" gridcol="0" gridrow="2" pageId="8" pageNumber="563">TaL / TL ♂</th>
<td id="76D5CE3AFFF300104EBECD29FDEFFCC0" box="[437,591,803,828]" gridcol="1" gridrow="2" pageId="8" pageNumber="563">0.184–0.188</td>
<td id="76D5CE3AFFF300104D85CD29FC85FCC0" box="[654,805,803,828]" gridcol="2" gridrow="2" pageId="8" pageNumber="563">0.187–0.195</td>
<td id="76D5CE3AFFF300104C46CD29FBAEFCC0" box="[845,1038,803,828]" gridcol="3" gridrow="2" pageId="8" pageNumber="563">0.218–0.219</td>
<td id="76D5CE3AFFF300104B4ACD29FB43FCC0" box="[1089,1251,803,828]" gridcol="4" gridrow="2" pageId="8" pageNumber="563">0.127–0.156</td>
<td id="76D5CE3AFFF300104A06CD29FA3AFCC0" box="[1293,1434,803,828]" gridcol="5" gridrow="2" pageId="8" pageNumber="563">0.155–0.216</td>
<td id="76D5CE3AFFF300104AB5CD29F9F3FCC0" box="[1470,1619,803,828]" gridcol="6" gridrow="2" pageId="8" pageNumber="563">0.165–0.195</td>
<td id="76D5CE3AFFF300104976CD29F8A9FCC0" box="[1661,1801,803,828]" gridcol="7" gridrow="2" pageId="8" pageNumber="563">0.112–0.141</td>
<td id="76D5CE3AFFF300104837CD29F873FCC0" box="[1852,2003,803,828]" gridcol="8" gridrow="2" pageId="8" pageNumber="563">0.191–0.203</td>
</tr>
<tr id="3504A746FFF300104FADCD46F873FC99" box="[166,2003,844,869]" gridrow="3" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCD46FEDBFC99" box="[166,379,844,869]" gridcol="0" gridrow="3" pageId="8" pageNumber="563">TaL / TL ♀</th>
<td id="76D5CE3AFFF300104EBECD46FDEFFC99" box="[437,591,844,869]" gridcol="1" gridrow="3" pageId="8" pageNumber="563">0.123–0.142</td>
<td id="76D5CE3AFFF300104D85CD46FC85FC99" box="[654,805,844,869]" gridcol="2" gridrow="3" pageId="8" pageNumber="563">0.152–0.158</td>
<td id="76D5CE3AFFF300104C46CD46FBAEFC99" box="[845,1038,844,869]" gridcol="3" gridrow="3" pageId="8" pageNumber="563">0.166–0.169</td>
<td id="76D5CE3AFFF300104B4ACD46FB43FC99" box="[1089,1251,844,869]" gridcol="4" gridrow="3" pageId="8" pageNumber="563">0.107–0.138</td>
<td id="76D5CE3AFFF300104A06CD46FA3AFC99" box="[1293,1434,844,869]" gridcol="5" gridrow="3" pageId="8" pageNumber="563">0.116–0.158</td>
<td id="76D5CE3AFFF300104AB5CD46F9F3FC99" box="[1470,1619,844,869]" gridcol="6" gridrow="3" pageId="8" pageNumber="563">0.149–0.164</td>
<td id="76D5CE3AFFF300104976CD46F8A9FC99" box="[1661,1801,844,869]" gridcol="7" gridrow="3" pageId="8" pageNumber="563">0.094–0.114</td>
<td id="76D5CE3AFFF300104837CD46F873FC99" box="[1852,2003,844,869]" gridcol="8" gridrow="3" pageId="8" pageNumber="563">0.161</td>
</tr>
<tr id="3504A746FFF300104FADCD7EF873FC71" box="[166,2003,884,909]" gridrow="4" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCD7EFEDBFC71" box="[166,379,884,909]" gridcol="0" gridrow="4" pageId="8" pageNumber="563">ASR</th>
<td id="76D5CE3AFFF300104EBECD7EFDEFFC71" box="[437,591,884,909]" gridcol="1" gridrow="4" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104D85CD7EFC85FC71" box="[654,805,884,909]" gridcol="2" gridrow="4" pageId="8" pageNumber="563">17</td>
<td id="76D5CE3AFFF300104C46CD7EFBAEFC71" box="[845,1038,884,909]" gridcol="3" gridrow="4" pageId="8" pageNumber="563">17–19</td>
<td id="76D5CE3AFFF300104B4ACD7EFB43FC71" box="[1089,1251,884,909]" gridcol="4" gridrow="4" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104A06CD7EFA3AFC71" box="[1293,1434,884,909]" gridcol="5" gridrow="4" pageId="8" pageNumber="563">21–23</td>
<td id="76D5CE3AFFF300104AB5CD7EF9F3FC71" box="[1470,1619,884,909]" gridcol="6" gridrow="4" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104976CD7EF8A9FC71" box="[1661,1801,884,909]" gridcol="7" gridrow="4" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104837CD7EF873FC71" box="[1852,2003,884,909]" gridcol="8" gridrow="4" pageId="8" pageNumber="563">19</td>
</tr>
<tr id="3504A746FFF300104FADCD96F873FC49" box="[166,2003,924,949]" gridrow="5" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCD96FEDBFC49" box="[166,379,924,949]" gridcol="0" gridrow="5" pageId="8" pageNumber="563">MSR</th>
<td id="76D5CE3AFFF300104EBECD96FDEFFC49" box="[437,591,924,949]" gridcol="1" gridrow="5" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104D85CD96FC85FC49" box="[654,805,924,949]" gridcol="2" gridrow="5" pageId="8" pageNumber="563">17</td>
<td id="76D5CE3AFFF300104C46CD96FBAEFC49" box="[845,1038,924,949]" gridcol="3" gridrow="5" pageId="8" pageNumber="563">17</td>
<td id="76D5CE3AFFF300104B4ACD96FB43FC49" box="[1089,1251,924,949]" gridcol="4" gridrow="5" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104A06CD96FA3AFC49" box="[1293,1434,924,949]" gridcol="5" gridrow="5" pageId="8" pageNumber="563">21</td>
<td id="76D5CE3AFFF300104AB5CD96F9F3FC49" box="[1470,1619,924,949]" gridcol="6" gridrow="5" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104976CD96F8A9FC49" box="[1661,1801,924,949]" gridcol="7" gridrow="5" pageId="8" pageNumber="563">19</td>
<td id="76D5CE3AFFF300104837CD96F873FC49" box="[1852,2003,924,949]" gridcol="8" gridrow="5" pageId="8" pageNumber="563">17 or 18</td>
</tr>
<tr id="3504A746FFF300104FADCDCEF873FC21" box="[166,2003,964,989]" gridrow="6" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCDCEFEDBFC21" box="[166,379,964,989]" gridcol="0" gridrow="6" pageId="8" pageNumber="563">PSR</th>
<td id="76D5CE3AFFF300104EBECDCEFDEFFC21" box="[437,591,964,989]" gridcol="1" gridrow="6" pageId="8" pageNumber="563">15</td>
<td id="76D5CE3AFFF300104D85CDCEFC85FC21" box="[654,805,964,989]" gridcol="2" gridrow="6" pageId="8" pageNumber="563">15</td>
<td id="76D5CE3AFFF300104C46CDCEFBAEFC21" box="[845,1038,964,989]" gridcol="3" gridrow="6" pageId="8" pageNumber="563">15</td>
<td id="76D5CE3AFFF300104B4ACDCEFB43FC21" box="[1089,1251,964,989]" gridcol="4" gridrow="6" pageId="8" pageNumber="563">15–17</td>
<td id="76D5CE3AFFF300104A06CDCEFA3AFC21" box="[1293,1434,964,989]" gridcol="5" gridrow="6" pageId="8" pageNumber="563">17</td>
<td id="76D5CE3AFFF300104AB5CDCEF9F3FC21" box="[1470,1619,964,989]" gridcol="6" gridrow="6" pageId="8" pageNumber="563">17 (15)</td>
<td id="76D5CE3AFFF300104976CDCEF8A9FC21" box="[1661,1801,964,989]" gridcol="7" gridrow="6" pageId="8" pageNumber="563">15 (13)</td>
<td id="76D5CE3AFFF300104837CDCEF873FC21" box="[1852,2003,964,989]" gridcol="8" gridrow="6" pageId="8" pageNumber="563">15</td>
</tr>
<tr id="3504A746FFF300104FADCDE1F873FBFA" box="[166,2003,1003,1030]" gridrow="7" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCDE1FEDBFBFA" box="[166,379,1003,1030]" gridcol="0" gridrow="7" pageId="8" pageNumber="563">1st reduction (VS)</th>
<td id="76D5CE3AFFF300104EBECDE1FDEFFBFA" box="[437,591,1003,1030]" gridcol="1" gridrow="7" pageId="8" pageNumber="563">97–111</td>
<td id="76D5CE3AFFF300104D85CDE1FC85FBFA" box="[654,805,1003,1030]" gridcol="2" gridrow="7" pageId="8" pageNumber="563">116–148</td>
<td id="76D5CE3AFFF300104C46CDE1FBAEFBFA" box="[845,1038,1003,1030]" gridcol="3" gridrow="7" pageId="8" pageNumber="563">78–135</td>
<td id="76D5CE3AFFF300104B4ACDE1FB43FBFA" box="[1089,1251,1003,1030]" gridcol="4" gridrow="7" pageId="8" pageNumber="563">82–115</td>
<td id="76D5CE3AFFF300104A06CDE1FA3AFBFA" box="[1293,1434,1003,1030]" gridcol="5" gridrow="7" pageId="8" pageNumber="563">74–112</td>
<td id="76D5CE3AFFF300104AB5CDE1F9F3FBFA" box="[1470,1619,1003,1030]" gridcol="6" gridrow="7" pageId="8" pageNumber="563">94–114</td>
<td id="76D5CE3AFFF300104976CDE1F8A9FBFA" box="[1661,1801,1003,1030]" gridcol="7" gridrow="7" pageId="8" pageNumber="563">79–107</td>
<td id="76D5CE3AFFF300104837CDE1F873FBFA" box="[1852,2003,1003,1030]" gridcol="8" gridrow="7" pageId="8" pageNumber="563">62–86</td>
</tr>
<tr id="3504A746FFF300104FADCA1FF873FBD2" box="[166,2003,1045,1070]" gridrow="8" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCA1FFEDBFBD2" box="[166,379,1045,1070]" gridcol="0" gridrow="8" pageId="8" pageNumber="563">VS ♂</th>
<td id="76D5CE3AFFF300104EBECA1FFDEFFBD2" box="[437,591,1045,1070]" gridcol="1" gridrow="8" pageId="8" pageNumber="563">173–179</td>
<td id="76D5CE3AFFF300104D85CA1FFC85FBD2" box="[654,805,1045,1070]" gridcol="2" gridrow="8" pageId="8" pageNumber="563">175–184</td>
<td id="76D5CE3AFFF300104C46CA1FFBAEFBD2" box="[845,1038,1045,1070]" gridcol="3" gridrow="8" pageId="8" pageNumber="563">167–169</td>
<td id="76D5CE3AFFF300104B4ACA1FFB43FBD2" box="[1089,1251,1045,1070]" gridcol="4" gridrow="8" pageId="8" pageNumber="563">160–173</td>
<td id="76D5CE3AFFF300104A06CA1FFA3AFBD2" box="[1293,1434,1045,1070]" gridcol="5" gridrow="8" pageId="8" pageNumber="563">160–186</td>
<td id="76D5CE3AFFF300104AB5CA1FF9F3FBD2" box="[1470,1619,1045,1070]" gridcol="6" gridrow="8" pageId="8" pageNumber="563">155–176</td>
<td id="76D5CE3AFFF300104976CA1FF8A9FBD2" box="[1661,1801,1045,1070]" gridcol="7" gridrow="8" pageId="8" pageNumber="563">156–165</td>
<td id="76D5CE3AFFF300104837CA1FF873FBD2" box="[1852,2003,1045,1070]" gridcol="8" gridrow="8" pageId="8" pageNumber="563">166–170</td>
</tr>
<tr id="3504A746FFF300104FADCA37F873FBAA" box="[166,2003,1085,1110]" gridrow="9" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCA37FEDBFBAA" box="[166,379,1085,1110]" gridcol="0" gridrow="9" pageId="8" pageNumber="563">VS ♀</th>
<td id="76D5CE3AFFF300104EBECA37FDEFFBAA" box="[437,591,1085,1110]" gridcol="1" gridrow="9" pageId="8" pageNumber="563">187–193</td>
<td id="76D5CE3AFFF300104D85CA37FC85FBAA" box="[654,805,1085,1110]" gridcol="2" gridrow="9" pageId="8" pageNumber="563">182–206</td>
<td id="76D5CE3AFFF300104C46CA37FBAEFBAA" box="[845,1038,1085,1110]" gridcol="3" gridrow="9" pageId="8" pageNumber="563">179–182</td>
<td id="76D5CE3AFFF300104B4ACA37FB43FBAA" box="[1089,1251,1085,1110]" gridcol="4" gridrow="9" pageId="8" pageNumber="563">168–172</td>
<td id="76D5CE3AFFF300104A06CA37FA3AFBAA" box="[1293,1434,1085,1110]" gridcol="5" gridrow="9" pageId="8" pageNumber="563">163–196</td>
<td id="76D5CE3AFFF300104AB5CA37F9F3FBAA" box="[1470,1619,1085,1110]" gridcol="6" gridrow="9" pageId="8" pageNumber="563">164–189</td>
<td id="76D5CE3AFFF300104976CA37F8A9FBAA" box="[1661,1801,1085,1110]" gridcol="7" gridrow="9" pageId="8" pageNumber="563">152–180</td>
<td id="76D5CE3AFFF300104837CA37F873FBAA" box="[1852,2003,1085,1110]" gridcol="8" gridrow="9" pageId="8" pageNumber="563">184</td>
</tr>
<tr id="3504A746FFF300104FADCA6FF873FB82" box="[166,2003,1125,1150]" gridrow="10" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCA6FFEDBFB82" box="[166,379,1125,1150]" gridcol="0" gridrow="10" pageId="8" pageNumber="563">SC ♂</th>
<td id="76D5CE3AFFF300104EBECA6FFDEFFB82" box="[437,591,1125,1150]" gridcol="1" gridrow="10" pageId="8" pageNumber="563">58–59</td>
<td id="76D5CE3AFFF300104D85CA6FFC85FB82" box="[654,805,1125,1150]" gridcol="2" gridrow="10" pageId="8" pageNumber="563">60–64</td>
<td id="76D5CE3AFFF300104C46CA6FFBAEFB82" box="[845,1038,1125,1150]" gridcol="3" gridrow="10" pageId="8" pageNumber="563">63–66</td>
<td id="76D5CE3AFFF300104B4ACA6FFB43FB82" box="[1089,1251,1125,1150]" gridcol="4" gridrow="10" pageId="8" pageNumber="563">47–48</td>
<td id="76D5CE3AFFF300104A06CA6FFA3AFB82" box="[1293,1434,1125,1150]" gridcol="5" gridrow="10" pageId="8" pageNumber="563">43–61</td>
<td id="76D5CE3AFFF300104AB5CA6FF9F3FB82" box="[1470,1619,1125,1150]" gridcol="6" gridrow="10" pageId="8" pageNumber="563">49–55</td>
<td id="76D5CE3AFFF300104976CA6FF8A9FB82" box="[1661,1801,1125,1150]" gridcol="7" gridrow="10" pageId="8" pageNumber="563">32–44</td>
<td id="76D5CE3AFFF300104837CA6FF873FB82" box="[1852,2003,1125,1150]" gridcol="8" gridrow="10" pageId="8" pageNumber="563">55–59</td>
</tr>
<tr id="3504A746FFF300104FADCA84F873FB5B" box="[166,2003,1166,1191]" gridrow="11" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCA84FEDBFB5B" box="[166,379,1166,1191]" gridcol="0" gridrow="11" pageId="8" pageNumber="563">SC ♀</th>
<td id="76D5CE3AFFF300104EBECA84FDEFFB5B" box="[437,591,1166,1191]" gridcol="1" gridrow="11" pageId="8" pageNumber="563">44–45</td>
<td id="76D5CE3AFFF300104D85CA84FC85FB5B" box="[654,805,1166,1191]" gridcol="2" gridrow="11" pageId="8" pageNumber="563">47–53</td>
<td id="76D5CE3AFFF300104C46CA84FBAEFB5B" box="[845,1038,1166,1191]" gridcol="3" gridrow="11" pageId="8" pageNumber="563">51–52</td>
<td id="76D5CE3AFFF300104B4ACA84FB43FB5B" box="[1089,1251,1166,1191]" gridcol="4" gridrow="11" pageId="8" pageNumber="563">30–44</td>
<td id="76D5CE3AFFF300104A06CA84FA3AFB5B" box="[1293,1434,1166,1191]" gridcol="5" gridrow="11" pageId="8" pageNumber="563">34–48</td>
<td id="76D5CE3AFFF300104AB5CA84F9F3FB5B" box="[1470,1619,1166,1191]" gridcol="6" gridrow="11" pageId="8" pageNumber="563">43–48</td>
<td id="76D5CE3AFFF300104976CA84F8A9FB5B" box="[1661,1801,1166,1191]" gridcol="7" gridrow="11" pageId="8" pageNumber="563">26–33</td>
<td id="76D5CE3AFFF300104837CA84F873FB5B" box="[1852,2003,1166,1191]" gridcol="8" gridrow="11" pageId="8" pageNumber="563">53</td>
</tr>
<tr id="3504A746FFF300104FADCABCF873FB33" box="[166,2003,1206,1231]" gridrow="12" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCABCFEDBFB33" box="[166,379,1206,1231]" gridcol="0" gridrow="12" pageId="8" pageNumber="563">SL</th>
<td id="76D5CE3AFFF300104EBECABCFDEFFB33" box="[437,591,1206,1231]" gridcol="1" gridrow="12" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104D85CABCFC85FB33" box="[654,805,1206,1231]" gridcol="2" gridrow="12" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104C46CABCFBAEFB33" box="[845,1038,1206,1231]" gridcol="3" gridrow="12" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104B4ACABCFB43FB33" box="[1089,1251,1206,1231]" gridcol="4" gridrow="12" pageId="8" pageNumber="563">8 (7)</td>
<td id="76D5CE3AFFF300104A06CABCFA3AFB33" box="[1293,1434,1206,1231]" gridcol="5" gridrow="12" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104AB5CABCF9F3FB33" box="[1470,1619,1206,1231]" gridcol="6" gridrow="12" pageId="8" pageNumber="563">7 or 8 (6)</td>
<td id="76D5CE3AFFF300104976CABCF8A9FB33" box="[1661,1801,1206,1231]" gridcol="7" gridrow="12" pageId="8" pageNumber="563">8 (7)</td>
<td id="76D5CE3AFFF300104837CABCF873FB33" box="[1852,2003,1206,1231]" gridcol="8" gridrow="12" pageId="8" pageNumber="563">8</td>
</tr>
<tr id="3504A746FFF300104FADCAD4F873FB0B" box="[166,2003,1246,1271]" gridrow="13" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCAD4FEDBFB0B" box="[166,379,1246,1271]" gridcol="0" gridrow="13" pageId="8" pageNumber="563">SL-Eye</th>
<td id="76D5CE3AFFF300104EBECAD4FDEFFB0B" box="[437,591,1246,1271]" gridcol="1" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104D85CAD4FC85FB0B" box="[654,805,1246,1271]" gridcol="2" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104C46CAD4FBAEFB0B" box="[845,1038,1246,1271]" gridcol="3" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104B4ACAD4FB43FB0B" box="[1089,1251,1246,1271]" gridcol="4" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104A06CAD4FA3AFB0B" box="[1293,1434,1246,1271]" gridcol="5" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104AB5CAD4F9F3FB0B" box="[1470,1619,1246,1271]" gridcol="6" gridrow="13" pageId="8" pageNumber="563">3+4 / 4+5</td>
<td id="76D5CE3AFFF300104976CAD4F8A9FB0B" box="[1661,1801,1246,1271]" gridcol="7" gridrow="13" pageId="8" pageNumber="563">4+5</td>
<td id="76D5CE3AFFF300104837CAD4F873FB0B" box="[1852,2003,1246,1271]" gridcol="8" gridrow="13" pageId="8" pageNumber="563">4+5</td>
</tr>
<tr id="3504A746FFF300104FADCB0CF873FAE3" box="[166,2003,1286,1311]" gridrow="14" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCB0CFEDBFAE3" box="[166,379,1286,1311]" gridcol="0" gridrow="14" pageId="8" pageNumber="563">PreSubOc</th>
<td id="76D5CE3AFFF300104EBECB0CFDEFFAE3" box="[437,591,1286,1311]" gridcol="1" gridrow="14" pageId="8" pageNumber="563">1 (0)</td>
<td id="76D5CE3AFFF300104D85CB0CFC85FAE3" box="[654,805,1286,1311]" gridcol="2" gridrow="14" pageId="8" pageNumber="563">1 (0)</td>
<td id="76D5CE3AFFF300104C46CB0CFBAEFAE3" box="[845,1038,1286,1311]" gridcol="3" gridrow="14" pageId="8" pageNumber="563">1</td>
<td id="76D5CE3AFFF300104B4ACB0CFB43FAE3" box="[1089,1251,1286,1311]" gridcol="4" gridrow="14" pageId="8" pageNumber="563">0 or 1</td>
<td id="76D5CE3AFFF300104A06CB0CFA3AFAE3" box="[1293,1434,1286,1311]" gridcol="5" gridrow="14" pageId="8" pageNumber="563">1 (0)</td>
<td id="76D5CE3AFFF300104AB5CB0CF9F3FAE3" box="[1470,1619,1286,1311]" gridcol="6" gridrow="14" pageId="8" pageNumber="563">1 (0)</td>
<td id="76D5CE3AFFF300104976CB0CF8A9FAE3" box="[1661,1801,1286,1311]" gridcol="7" gridrow="14" pageId="8" pageNumber="563">1 (0)</td>
<td id="76D5CE3AFFF300104837CB0CF873FAE3" box="[1852,2003,1286,1311]" gridcol="8" gridrow="14" pageId="8" pageNumber="563">1</td>
</tr>
<tr id="3504A746FFF300104FADCB24F873FABB" box="[166,2003,1326,1351]" gridrow="15" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCB24FEDBFABB" box="[166,379,1326,1351]" gridcol="0" gridrow="15" pageId="8" pageNumber="563">IL</th>
<td id="76D5CE3AFFF300104EBECB24FDEFFABB" box="[437,591,1326,1351]" gridcol="1" gridrow="15" pageId="8" pageNumber="563">9</td>
<td id="76D5CE3AFFF300104D85CB24FC85FABB" box="[654,805,1326,1351]" gridcol="2" gridrow="15" pageId="8" pageNumber="563">9 (8)</td>
<td id="76D5CE3AFFF300104C46CB24FBAEFABB" box="[845,1038,1326,1351]" gridcol="3" gridrow="15" pageId="8" pageNumber="563">9</td>
<td id="76D5CE3AFFF300104B4ACB24FB43FABB" box="[1089,1251,1326,1351]" gridcol="4" gridrow="15" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104A06CB24FA3AFABB" box="[1293,1434,1326,1351]" gridcol="5" gridrow="15" pageId="8" pageNumber="563">8</td>
<td id="76D5CE3AFFF300104AB5CB24F9F3FABB" box="[1470,1619,1326,1351]" gridcol="6" gridrow="15" pageId="8" pageNumber="563">9</td>
<td id="76D5CE3AFFF300104976CB24F8A9FABB" box="[1661,1801,1326,1351]" gridcol="7" gridrow="15" pageId="8" pageNumber="563">9 or 10</td>
<td id="76D5CE3AFFF300104837CB24F873FABB" box="[1852,2003,1326,1351]" gridcol="8" gridrow="15" pageId="8" pageNumber="563">9</td>
</tr>
<tr id="3504A746FFF300104FADCB5DF873FA68" box="[166,2003,1367,1428]" gridrow="16" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCB5DFEDBFA68" box="[166,379,1367,1428]" gridcol="0" gridrow="16" pageId="8" pageNumber="563">Dorsal pattern</th>
<td id="76D5CE3AFFF300104EBECB5DFDEFFA68" box="[437,591,1367,1428]" gridcol="1" gridrow="16" pageId="8" pageNumber="563">Blotched</td>
<td id="76D5CE3AFFF300104D85CB5DFC85FA68" box="[654,805,1367,1428]" gridcol="2" gridrow="16" pageId="8" pageNumber="563">Blotched or transverse bars</td>
<td id="76D5CE3AFFF300104C46CB5DFBAEFA68" box="[845,1038,1367,1428]" gridcol="3" gridrow="16" pageId="8" pageNumber="563">Blotched</td>
<td id="76D5CE3AFFF300104B4ACB5DFB43FA68" box="[1089,1251,1367,1428]" gridcol="4" gridrow="16" pageId="8" pageNumber="563">Reticulate or narrow blotches</td>
<td id="76D5CE3AFFF300104A06CB5DFA3AFA68" box="[1293,1434,1367,1428]" gridcol="5" gridrow="16" pageId="8" pageNumber="563">Reticulate or large blotches</td>
<td id="76D5CE3AFFF300104AB5CB5DF9F3FA68" box="[1470,1619,1367,1428]" gridcol="6" gridrow="16" pageId="8" pageNumber="563">Reticulate</td>
<td id="76D5CE3AFFF300104976CB5DF8A9FA68" box="[1661,1801,1367,1428]" gridcol="7" gridrow="16" pageId="8" pageNumber="563">Reticulate or large blotches</td>
<td id="76D5CE3AFFF300104837CB5DF873FA68" box="[1852,2003,1367,1428]" gridcol="8" gridrow="16" pageId="8" pageNumber="563">Blotched</td>
</tr>
<tr id="3504A746FFF300104FADCBA9F873FA1C" box="[166,2003,1443,1504]" gridrow="17" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCBA9FEDBFA1C" box="[166,379,1443,1504]" gridcol="0" gridrow="17" pageId="8" pageNumber="563">Vertebral stripe at midbody</th>
<td id="76D5CE3AFFF300104EBECBA9FDEFFA1C" box="[437,591,1443,1504]" gridcol="1" gridrow="17" pageId="8" pageNumber="563">Narrow</td>
<td id="76D5CE3AFFF300104D85CBA9FC85FA1C" box="[654,805,1443,1504]" gridcol="2" gridrow="17" pageId="8" pageNumber="563">Broad</td>
<td id="76D5CE3AFFF300104C46CBA9FBAEFA1C" box="[845,1038,1443,1504]" gridcol="3" gridrow="17" pageId="8" pageNumber="563">None or faint</td>
<td id="76D5CE3AFFF300104B4ACBA9FB43FA1C" box="[1089,1251,1443,1504]" gridcol="4" gridrow="17" pageId="8" pageNumber="563">None</td>
<td id="76D5CE3AFFF300104A06CBA9FA3AFA1C" box="[1293,1434,1443,1504]" gridcol="5" gridrow="17" pageId="8" pageNumber="563">None</td>
<td id="76D5CE3AFFF300104AB5CBA9F9F3FA1C" box="[1470,1619,1443,1504]" gridcol="6" gridrow="17" pageId="8" pageNumber="563">Broad</td>
<td id="76D5CE3AFFF300104976CBA9F8A9FA1C" box="[1661,1801,1443,1504]" gridcol="7" gridrow="17" pageId="8" pageNumber="563">None</td>
<td id="76D5CE3AFFF300104837CBA9F873FA1C" box="[1852,2003,1443,1504]" gridcol="8" gridrow="17" pageId="8" pageNumber="563">Narrow</td>
</tr>
<tr id="3504A746FFF300104FADCBE5F873F9D0" box="[166,2003,1519,1580]" gridrow="18" pageId="8" pageNumber="563">
<th id="76D5CE3AFFF300104FADCBE5FEDBF9D0" box="[166,379,1519,1580]" gridcol="0" gridrow="18" pageId="8" pageNumber="563">Length of hemipenes (SC)</th>
<td id="76D5CE3AFFF300104EBECBE5FDEFF9D0" box="[437,591,1519,1580]" gridcol="1" gridrow="18" pageId="8" pageNumber="563">25</td>
<td id="76D5CE3AFFF300104D85CBE5FC85F9D0" box="[654,805,1519,1580]" gridcol="2" gridrow="18" pageId="8" pageNumber="563">12–13</td>
<td id="76D5CE3AFFF300104C46CBE5FBAEF9D0" box="[845,1038,1519,1580]" gridcol="3" gridrow="18" pageId="8" pageNumber="563">20</td>
<td id="76D5CE3AFFF300104B4ACBE5FB43F9D0" box="[1089,1251,1519,1580]" gridcol="4" gridrow="18" pageId="8" pageNumber="563">15–18</td>
<td id="76D5CE3AFFF300104A06CBE5FA3AF9D0" box="[1293,1434,1519,1580]" gridcol="5" gridrow="18" pageId="8" pageNumber="563">14–21</td>
<td id="76D5CE3AFFF300104AB5CBE5F9F3F9D0" box="[1470,1619,1519,1580]" gridcol="6" gridrow="18" pageId="8" pageNumber="563">15–18</td>
<td id="76D5CE3AFFF300104976CBE5F8A9F9D0" box="[1661,1801,1519,1580]" gridcol="7" gridrow="18" pageId="8" pageNumber="563">15–17</td>
<td id="76D5CE3AFFF300104837CBE5F873F9D0" box="[1852,2003,1519,1580]" gridcol="8" gridrow="18" pageId="8" pageNumber="563">27–28</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8BA504FFF2FFE64FCCCE9DFE3DFE99" blockId="9.[151,1436,151,357]" pageId="9" pageNumber="564">
<taxonomicName id="4C34DE87FFF2FFE64FCCCE9DFED3FF4C" authority="Nguyen &amp; Le &amp; Vo &amp; Murphy, 2022" authorityName="Nguyen &amp; Le &amp; Vo &amp; Murphy" authorityYear="2022" box="[199,371,151,177]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="564" phylum="Chordata" rank="species" species="tuani" status="sp. nov.">
<emphasis id="B9407916FFF2FFE64FCCCE9DFED3FF4C" box="[199,371,151,177]" italics="true" pageId="9" pageNumber="564">Oligodon tuani</emphasis>
</taxonomicName>
<emphasis id="B9407916FFF2FFE64E70CE9DFE76FF4D" bold="true" box="[379,470,151,177]" pageId="9" pageNumber="564">
<taxonomicNameLabel id="A273C46DFFF2FFE64E70CE9DFE76FF4D" box="[379,470,151,177]" pageId="9" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also morphologically similar to 
<taxonomicName id="4C34DE87FFF2FFE64C6FCE9DFBA1FF4D" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[868,1025,151,177]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="564" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="B9407916FFF2FFE64C6FCE9DFBA1FF4D" box="[868,1025,151,177]" italics="true" pageId="9" pageNumber="564">O. fasciolatus</emphasis>
</taxonomicName>
. However, it differs from 
<taxonomicName id="4C34DE87FFF2FFE64A27CE9DFF6DFF29" baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="564" phylum="Chordata" rank="species" species="fasciolatus">
<emphasis id="B9407916FFF2FFE64A27CE9DFF6DFF29" italics="true" pageId="9" pageNumber="564">O. fasciolatus</emphasis>
</taxonomicName>
by having fewer dorsal scale rows at neck (19 vs. 21–23), at midbody (19 vs. 21) and before the vent (15 vs. 17), remarkably longer hemipenis (extending to SC 25 vs. 14–21), temporal streak faint and discontinuous (vs. distinct and continuous), ventral surface with black rectangular blotches (vs. pure white or immaculate) (
<bibRefCitation id="EFA5D8F5FFF2FFE64A32CF09FF72FEBD" author="Gunther, A." pageId="9" pageNumber="564" refId="ref7609" refString="Gunther, A. (1864) The Reptiles of British India. Taylor &amp; Francis, London, xxvii + 452 pp." type="book" year="1864">Gunther, 1864</bibRefCitation>
; 
<bibRefCitation id="EFA5D8F5FFF2FFE64FD4CF2DFE16FEBD" author="David, P. &amp; Vogel, G. &amp; Pauwels, O. S. G." box="[223,438,295,321]" pageId="9" pageNumber="564" pagination="19 - 37" refId="ref7156" refString="David, P., Vogel, G. &amp; Pauwels, O. S. G. (2008 a) A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from southern Vietnam and Cambodia. Zootaxa, 1939, 19 - 37. https: // doi. org / 10.11646 / zootaxa. 1939.1.3" type="journal article" year="2008">
David 
<emphasis id="B9407916FFF2FFE64E27CF2DFEFFFEBD" box="[300,351,295,321]" italics="true" pageId="9" pageNumber="564">et al</emphasis>
. 2008a
</bibRefCitation>
; this study). The main morphological characters of the 
<taxonomicName id="4C34DE87FFF2FFE64B26CF2DFB10FEBD" baseAuthorityName="Cantor" baseAuthorityYear="1839" box="[1069,1200,295,321]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="564" phylum="Chordata" rank="species" species="cyclurus">
<emphasis id="B9407916FFF2FFE64B26CF2DFB10FEBD" box="[1069,1200,295,321]" italics="true" pageId="9" pageNumber="564">O. cyclurus</emphasis>
</taxonomicName>
species complex are summarized in 
<tableCitation id="C6B690BFFFF2FFE64E48CF41FE39FE99" box="[323,409,331,357]" captionStart="TABLE 2" captionStartId="8.[152,244,610,636]" captionTargetBox="[166,2003,687,1580]" captionText="TABLE 2. Main morphological characters of Oligodon tuani sp. nov. in comparison with other Oligodon in the Oligodon cyclurus species complex. Data taken from David et al. (2008a), Nguyen et al. (2017) and this work. See Material and Methods for abbreviations." httpUri="http://table.plazi.org/id/DF4BF58CFFF3FFE74F93CC68FBF7FD5C" pageId="9" pageNumber="564" tableUuid="DF4BF58CFFF3FFE74F93CC68FBF7FD5C">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
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