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<mods:title id="B2CD0DA75481EA0EFAF0293C286539D0">A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)</mods:title>
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<mods:namePart id="7D9B5EA8CE14ED3E6D1EBB10392F1F3D">Sartori, Michel</mods:namePart>
<mods:affiliation id="CB68E876BDD3A901CF8A110255C7C283">Museum of zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland. E-mail: michel. sartori @ vd. ch Laboratory of Aquatic Entomology, Florida A &amp; M University, Tallahassee, FL 32307, USA</mods:affiliation>
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<mods:namePart id="EDCEED256DC7AEBF627D4D2A2EB79AF5">Peters, Janice G.</mods:namePart>
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<heading id="D0C4EE11442AFFD2CE8E75C8FD12FBD9" bold="true" box="[151,689,1018,1044]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<taxonomicName id="4C3322FE442AFFD2CE8E75C8FD80FBD9" ID-CoL="6DPDB" authority="(Navas, 1933)" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[151,547,1018,1044]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lugens" status="comb. nov.">
<emphasis id="B947856F442AFFD2CE8E75C8FED0FBD9" bold="true" box="[151,371,1018,1044]" italics="true" pageId="34" pageNumber="35">Dudgeodes lugens</emphasis>
(
<bibRefCitation id="EFA2248C442AFFD2CF9A75C8FDBFFBD9" author="Navas, L." box="[387,540,1018,1044]" pageId="34" pageNumber="35" pagination="1 - 22" refId="ref26222" refString="Navas, L. (1933) Nevropteres et insectes voisins - Chine et pays environnants - 4 e Serie. Notes d'entomologie chinoise, 1, 1 - 22." type="journal article" year="1933">Navás, 1933</bibRefCitation>
)
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<taxonomicNameLabel id="A2743814442AFFD2CC3275C8FD12FBD9" box="[555,689,1018,1044]" pageId="34" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
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<treatmentCitation id="0A927F6C442AFFD2CE8E7272FDBEFB9A" author="Navas, L." box="[151,541,1088,1111]" page="17" pageId="34" pageNumber="35" year="1933">
<taxonomicName id="4C3322FE442AFFD2CE8E7272FDBEFB9A" authority="Navas, 1933: 17" authorityName="Navas" authorityPageNumber="17" authorityYear="1933" box="[151,541,1088,1111]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B947856F442AFFD2CE8E7272FECBFB9A" box="[151,360,1088,1111]" italics="true" pageId="34" pageNumber="35">Teloganodes lugens</emphasis>
<bibRefCitation id="EFA2248C442AFFD2CF767272FDBEFB9A" author="Navas, L." box="[367,541,1088,1111]" pageId="34" pageNumber="35" pagination="1 - 22" refId="ref26222" refString="Navas, L. (1933) Nevropteres et insectes voisins - Chine et pays environnants - 4 e Serie. Notes d'entomologie chinoise, 1, 1 - 22." type="journal article" year="1933">Navás, 1933: 17</bibRefCitation>
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<emphasis id="B947856F442AFFD2CE8E72B0FE28FB51" bold="true" box="[151,395,1154,1180]" pageId="34" pageNumber="35">Material examined.</emphasis>
None
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<paragraph id="8B8C597D442AFFD2CEDF7298FC5CF969" blockId="34.[151,1436,1154,1700]" pageId="34" pageNumber="35">
<emphasis id="B947856F442AFFD2CEDF7298FEFDFB09" bold="true" box="[198,350,1194,1220]" pageId="34" pageNumber="35">Description.</emphasis>
This species was described from a single female subimago from Southeast
<collectingCountry id="F32419ED442AFFD2CAF07298FA91FB09" box="[1257,1330,1194,1220]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
, “Chekiang, Chusan”, captured
<date id="FF8D7FBD442AFFD2CFB772E0FD91FB21" box="[430,562,1234,1260]" pageId="34" pageNumber="35" value="1931-06-11">11.VI.1931</date>
by O. Piel. This location is nowadays known as Zhou Shan Island, in
<collectingRegion id="49F7979F442AFFD2CB7C72E0FEE6FAD9" country="China" name="Zhejiang" pageId="34" pageNumber="35">Zhejiang province</collectingRegion>
. The
<typeStatus id="5488E7DF442AFFD2CF9072C8FE18FAD9" box="[393,443,1274,1300]" pageId="34" pageNumber="35">type</typeStatus>
material could not be located, and this specimen is neither housed in the Museum national dHistoire Naturelle de Paris (
<bibRefCitation id="EFA2248C442AFFD2CC767310FCE1FAF1" author="Berthelemy, C." box="[623,834,1314,1340]" pageId="34" pageNumber="35" pagination="2 - 3" refId="ref25159" refString="Berthelemy, C. (1965) Types of pinned Ephemeroptera deposited in the Natural History Museum in Paris. Eatonia, 7, 2 - 3." type="journal article" year="1965">Berthelemy 1965</bibRefCitation>
, J. Legrand, comm. pers.) nor in the Museum of Zoology of Barcelona (
<bibRefCitation id="EFA2248C442AFFD2CFA87378FCB8FAA9" author="Alba-Tercedor, J. &amp; Peters, W. L." box="[433,795,1354,1380]" pageId="34" pageNumber="35" pagination="217 - 227" refId="ref25050" refString="Alba-Tercedor, J. &amp; Peters, W. L. (1985) Types and additional specimens of Ephemeroptera studied by Longinos Navas in the Museo de Zoologia del Ayuntamiento, Barcelona, Spain. Aquatic Insects, 7, 217 - 227." type="journal article" year="1985">Alba-Tercedor &amp; Peters 1985</bibRefCitation>
). Based on the drawing of the hindwing, we think it plausible that this species belongs to the genus
<taxonomicName id="4C3322FE442AFFD2CCCF7341FCF9FA41" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[726,858,1395,1420]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B947856F442AFFD2CCCF7341FCF9FA41" box="[726,858,1395,1420]" italics="true" pageId="34" pageNumber="35">Dudgeodes</emphasis>
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; the shape of the costal process and the length/ width ratio (ca. 1.8) is similar to that of
<taxonomicName id="4C3322FE442AFFD2CC7173A9FD41FA79" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[616,738,1435,1460]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hutanis">
<emphasis id="B947856F442AFFD2CC7173A9FD41FA79" box="[616,738,1435,1460]" italics="true" pageId="34" pageNumber="35">D. hutanis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3322FE442AFFD2CCE973A9FC64FA79" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[752,967,1435,1460]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B947856F442AFFD2CCE973A9FC64FA79" box="[752,967,1435,1460]" italics="true" pageId="34" pageNumber="35">Dudgeodes lugens</emphasis>
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would differ from
<taxonomicName id="4C3322FE442AFFD2CABE73A9FA83FA79" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[1191,1312,1435,1460]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hutanis">
<emphasis id="B947856F442AFFD2CABE73A9FA83FA79" box="[1191,1312,1435,1460]" italics="true" pageId="34" pageNumber="35">D. hutanis</emphasis>
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by having more numerous intercalary veins in the cubital field. The only other mention of
<taxonomicName id="4C3322FE442AFFD2CA2373F0FB45FA11" baseAuthorityName="Allen" baseAuthorityYear="1965" box="[1082,1254,1474,1500]" class="Insecta" family="Teloganodidae" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Teloganodidae</taxonomicName>
from continental
<collectingCountry id="F32419ED442AFFD2CEA473D8FEA0F9C9" box="[189,259,1514,1540]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
was made by
<bibRefCitation id="EFA2248C442AFFD2CFA973D8FD75F9C9" author="Tong, X. &amp; Dudgeon, D." box="[432,726,1514,1540]" pageId="34" pageNumber="35" pagination="197 - 207" refId="ref26602" refString="Tong, X. &amp; Dudgeon, D. (2000) Ephemerellidae (Insecta: Ephemeroptera) from Hong-Kong, China, with description of two new species. Aquatic Insects, 22, 197 - 207." type="journal article" year="2000">Tong &amp; Dudgeon (2000)</bibRefCitation>
who mentioned two nymphs of
<emphasis id="B947856F442AFFD2CA4273D9FAF9F9C9" box="[1115,1370,1515,1540]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C3322FE442AFFD2CA7C73D9FAEDF9C9" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[1125,1358,1515,1540]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tristis">Teloganodes tristis</taxonomicName>
</emphasis>
from Tai Pau Forest,
<collectingCountry id="F32419ED442AFFD2CF4A7020FE41F9E1" box="[339,482,1554,1580]" name="Hong Kong" pageId="34" pageNumber="35">Hong Kong</collectingCountry>
. The drawings clearly indicate that this species belongs to the genus
<taxonomicName id="4C3322FE442AFFD2CB017021FA3FF9E1" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[1304,1436,1555,1580]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B947856F442AFFD2CB017021FA3FF9E1" box="[1304,1436,1555,1580]" italics="true" pageId="34" pageNumber="35">Dudgeodes</emphasis>
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and is closely related to
<taxonomicName id="4C3322FE442AFFD2CFAF7009FD93F999" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[438,560,1595,1620]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hutanis">
<emphasis id="B947856F442AFFD2CFAF7009FD93F999" box="[438,560,1595,1620]" italics="true" pageId="34" pageNumber="35">D. hutanis</emphasis>
</taxonomicName>
. The status of Chinese
<taxonomicName id="4C3322FE442AFFD2CD5E7009FC68F999" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[839,971,1595,1620]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B947856F442AFFD2CD5E7009FC68F999" box="[839,971,1595,1620]" italics="true" pageId="34" pageNumber="35">Dudgeodes</emphasis>
</taxonomicName>
, whether the nymphs from
<collectingCountry id="F32419ED442AFFD2CB0B7008FA3FF999" box="[1298,1436,1594,1620]" name="Hong Kong" pageId="34" pageNumber="35">Hong Kong</collectingCountry>
are the immature stage of
<taxonomicName id="4C3322FE442AFFD2CFD57051FD9FF9B1" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[460,572,1635,1660]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="B947856F442AFFD2CFD57051FD9FF9B1" box="[460,572,1635,1660]" italics="true" pageId="34" pageNumber="35">D. lugens</emphasis>
</taxonomicName>
or are in fact conspecific with those of another species described here or even a new species may be solved when more material becomes available.
</paragraph>
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