treatments-xml/data/03/9A/87/039A87C6C062FFE4FF3AFDA4FB14F964.xml

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<mods:title id="096F69E4598680A65424226D02938732">New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature</mods:title>
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<paragraph id="8B8C36D0C062FFEFFF3AFDA4FE25FA6F" blockId="2.[151,449,1489,1584]" box="[151,449,1489,1515]" pageId="2" pageNumber="203">
<heading id="D0C481BCC062FFEFFF3AFDA4FE25FA6F" bold="true" box="[151,449,1489,1515]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
<emphasis id="B947EAC2C062FFEFFF3AFDA4FE25FA6F" bold="true" box="[151,449,1489,1515]" pageId="2" pageNumber="203">
<emphasis id="B947EAC2C062FFEFFF3AFDA4FEB9FA6F" bold="true" box="[151,349,1489,1515]" italics="true" pageId="2" pageNumber="203">
<taxonomicName id="4C334D53C062FFEFFF3AFDA4FEB3FA6F" ID-CoL="58V4M" box="[151,343,1489,1515]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="aturirii" status="sp. nov.">Trimma aturirii</taxonomicName>
,
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<taxonomicNameLabel id="A27457B9C062FFEFFECEFDA4FE25FA6F" box="[355,449,1489,1515]" pageId="2" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8C36D0C062FFEFFF3AFD81FEA1F9B4" blockId="2.[151,449,1489,1584]" pageId="2" pageNumber="203">
Aturiri’s pygmy goby 
<figureCitation id="13082A55C062FFEFFF3AFE62FF07F9B4" box="[151,227,1559,1584]" captionStart="FIGURE 1" captionStartId="2.[151,250,1261,1283]" captionTargetBox="[151,1436,375,1239]" captionTargetId="figure@2.[151,1436,375,1239]" captionTargetPageId="2" captionText="FIGURE 1. Trimma aturirii, 16.2 F, ROM 98809, lateral (A) and dorsal (B) views of head papillae. Note paths of sensory nerves (narrow purple lines linking papillae); specimen somewhat distorted by a fully splayed hyoid arch. Specimen stained with cyanine blue. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/240777/files/figure.png" pageId="2" pageNumber="203">Figs. 1</figureCitation>
, 
<figureCitation id="13082A55C062FFEFFF5CFE62FEFAF9B4" box="[241,286,1559,1584]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="4.[151,250,1059,1081]" captionStartId-1="6.[151,250,1951,1973]" captionStartId-2="7.[151,250,1942,1964]" captionStartId-3="8.[151,250,1050,1072]" captionTargetBox-0="[151,1435,193,1037]" captionTargetBox-1="[157,1429,193,1932]" captionTargetBox-2="[151,1435,193,1919]" captionTargetBox-3="[152,1434,179,1035]" captionTargetId-0="figure@4.[151,1435,193,1038]" captionTargetId-1="figure@6.[157,1429,193,1934]" captionTargetId-2="figure@7.[151,1435,193,1921]" captionTargetId-3="figure@8.[152,1435,179,1035]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." captionText-1="FIGURE 4. Trimma aturirii, 16.2 mm SL male, ROM 95269, lateral (A) and dorsal (B) views of head papillae. Red arrows show supernumerary papillae, thin white lines indicate lines of papillae. Specimen stained with cyanine blue. Photos by R. Winterbottom." captionText-2="FIGURE 5. Trimma aturirii, left lateral views of live specimens from Mapia Atoll, ROM 94889, holotype (A—image reversed); Ayau Besar Atoll, ROM 98853 (B—image reversed); and a freshly collected specimen from Mof Island, ROM 87934 (C). Photos by M. V. Erdmann." captionText-3="FIGURE 6. Trimma aturirii, 14.0 mm SL female, ROM 87934, Mof Island, Raja Ampat, lateral view of head (unstained). Photo by R. Winterbottom." httpUri-0="https://zenodo.org/record/240779/files/figure.png" httpUri-1="https://zenodo.org/record/240780/files/figure.png" httpUri-2="https://zenodo.org/record/240781/files/figure.png" httpUri-3="https://zenodo.org/record/240782/files/figure.png" pageId="2" pageNumber="203">3–6</figureCitation>
, 
<figureCitation id="13082A55C062FFEFFE84FE6DFEA1F9B4" box="[297,325,1560,1584]" captionStart="FIGURE 18" captionStartId="23.[151,250,1897,1919]" captionTargetBox="[174,1412,963,1865]" captionTargetId="figure@23.[166,1420,949,1876]" captionTargetPageId="23" captionText="FIGURE 18. Distribution of the three new species based on material examined for this study. Trimma aturirii, black inverted triangles. Trimma kardium, black diamonds. Trimma trioculatum, coloured pentagons. Blue fill, type locality, Group 1 haplogroup; green fill, Palau, Group 2 haplogroup; yellow fill, Rabaul, New Britain, Group 3 haplogroup; red fill, unknown haplogroup." httpUri="https://zenodo.org/record/240794/files/figure.png" pageId="2" pageNumber="203">18</figureCitation>
</paragraph>
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<emphasis id="B947EAC2C062FFEFFF3AFE2AFE6FF9FC" bold="true" box="[151,395,1631,1656]" pageId="2" pageNumber="203">Material Examined.</emphasis>
All 
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specimens are from the northern part of West 
<collectingCountry id="F3247640C062FFEFFB86FE15FB94F9F3" box="[1067,1136,1632,1655]" name="Papua New Guinea" pageId="2" pageNumber="203">Papua</collectingCountry>
province, 
<collectingCountry id="F3247640C062FFEFFB5CFE2AFA86F9FC" box="[1265,1378,1631,1656]" name="Indonesia" pageId="2" pageNumber="203">Indonesia</collectingCountry>
. The description is based on up to 
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, 14.0– 
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culled from all five lots.
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.
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<collectionCode id="ED22AE15C062FFEFFEE4FEDDFE6DF944" LSID="urn:lsid:biocol.org:col:34778" box="[329,393,1704,1728]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="2" pageNumber="203">ROM</collectionCode>
94889, (
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female), Mapia Atoll, south end of 
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Pegun, (
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, 
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), steep wall covered with coarse white 
<taxonomicName id="4C334D53C062FFEFFD45FEB9FCB2F961" box="[744,854,1740,1765]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="2" pageNumber="203" phylum="Chlorophyta" rank="genus">
<emphasis id="B947EAC2C062FFEFFD45FEB9FCB2F961" box="[744,854,1740,1765]" italics="true" pageId="2" pageNumber="203">Halimeda</emphasis>
</taxonomicName>
sand, 
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, clove oil, 
<date id="FF8D1010C062FFEFFBFEFEB9FB44F960" box="[1107,1184,1740,1764]" pageId="2" pageNumber="203">2 Nov.</date>
, 2013, M.V. Erdmann.
</paragraph>
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<typeStatus id="54888872C062FFEFFF6AFE9AFEA5F88C" box="[199,321,1775,1800]" pageId="2" pageNumber="203" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED22AE15C062FFEFFEF8FE85FE76F883" box="[341,402,1776,1799]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="203">MZB</collectionCode>
22713, 7(13.5–14.7, ex-ROM 87934,), 
<collectingCountry id="F3247640C062FFEFFCD3FE9AFC5AF88C" box="[894,958,1775,1800]" name="Iceland" pageId="2" pageNumber="203">Pulau</collectingCountry>
Mof (north of Waigeo), (
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, 
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), 
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, clove oil, 
<date id="FF8D1010C062FFEFFDACFF60FDAAF8A8" box="[513,590,1813,1836]" pageId="2" pageNumber="203">7 Mar.</date>
, 2011, M.V. Erdmann. 
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95269, 5(14.4–16.6), NW side of Ayau Besar Atoll, (
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, 
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), exposed oceanic atoll with clear water, steep outer wall with small crevices and overhangs, coarse white coralline sand, 
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, clove oil, 
<date id="FF8D1010C062FFEFFCE6FF29FC75F8F1" box="[843,913,1884,1909]" pageId="2" pageNumber="203">1 July</date>
, 2014, M.V. Erdmann. 
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98809, 10(11.0– 16.8), Ayau Besar Atoll, (
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, 
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), steep outer reef slope with white calcareous sand, 
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, clove oil, 
<date id="FF8D1010C062FFEFFEABFFD1FEBFF838" box="[262,347,1956,1980]" pageId="2" pageNumber="203" value="1922-10">22 Oct.</date>
, 2014, M.V. Erdmann. 
<collectionCode id="ED22AE15C062FFEFFDCDFFD1FD44F838" LSID="urn:lsid:biocol.org:col:34778" box="[608,672,1956,1980]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="2" pageNumber="203">ROM</collectionCode>
98853, 3(14.1–18.8),NW side of Ayau Besar Atoll, (
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, 
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,), 
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, 
<date id="FF8D1010C062FFEFFE36FFBDFE16F865" box="[411,498,1992,2017]" pageId="2" pageNumber="203" value="1919-02">19 Feb.</date>
, 2015, MVE-15-010, M.V. Erdmann.
</paragraph>
<paragraph id="8B8C36D0C063FFEEFF6AF8E2FC27FF51" blockId="3.[151,1436,151,213]" pageId="3" pageNumber="204">
Tissue specimens (all from Ayau Besar Atoll, collected by M.V. Erdmann): MB0614201, 
<date id="FF8D1010C063FFEEFAA9F8EDFAAEFF34" box="[1284,1354,151,176]" pageId="3" pageNumber="204">1 Jul.</date>
, 2014; MB0615401, MB0615402, 
<date id="FF8D1010C063FFEEFE63F8C9FDC7FF50" box="[462,547,188,212]" pageId="3" pageNumber="204" value="1922-10">22 Oct.</date>
, 2014; 
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T19515, 
<date id="FF8D1010C063FFEEFC8CF8C9FC9CFF51" box="[801,888,188,213]" pageId="3" pageNumber="204" value="1919-02">19 Feb.</date>
, 2015.
</paragraph>
</subSubSection>
<caption id="DF4C6658C063FFEEFF3AFE80FDE7F8CC" httpUri="https://zenodo.org/record/240778/files/figure.png" pageId="3" pageNumber="204" targetBox="[210,1436,278,1758]" targetPageId="3">
<paragraph id="8B8C36D0C063FFEEFF3AFE80FDE7F8CC" blockId="3.[151,1436,1781,1864]" pageId="3" pageNumber="204">
<emphasis id="B947EAC2C063FFEEFF3AFE80FEE9F88E" bold="true" box="[151,269,1781,1803]" pageId="3" pageNumber="204">FIGURE 2</emphasis>
. Head of a hypothetical 
<taxonomicName id="4C334D53C063FFEEFDABFE83FDB1F88F" box="[518,597,1782,1803]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
<emphasis id="B947EAC2C063FFEEFDABFE83FDB1F88F" box="[518,597,1782,1803]" italics="true" pageId="3" pageNumber="204">Trimma</emphasis>
</taxonomicName>
in lateral (above) and dorsal (below) views, showing papillae and papillae rows as redefined in this paper. Thin red lines = nerve pathways; papillae lines accented by green dotted lines = those mentioned in the redefinitions proposed in this paper.
</paragraph>
</caption>
<caption id="DF4C6658C064FFE9FF3AFC56FA7EFBF3" httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="4" pageNumber="205" targetBox="[151,1435,193,1037]" targetPageId="4">
<paragraph id="8B8C36D0C064FFE9FF3AFC56FA7EFBF3" blockId="4.[151,1436,1059,1143]" pageId="4" pageNumber="205">
<emphasis id="B947EAC2C064FFE9FF3AFC56FEE8FBBC" bold="true" box="[151,268,1059,1081]" pageId="4" pageNumber="205">FIGURE 3</emphasis>
. 
<emphasis id="B947EAC2C064FFE9FEB5FC51FE58FBBD" box="[280,444,1060,1081]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="4C334D53C064FFE9FEB5FC51FE5CFBBD" box="[280,440,1060,1081]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="aturirii">Trimma aturirii</taxonomicName>
,
</emphasis>
15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom.
</paragraph>
</caption>
<subSubSection id="C329655BC064FFE9FF6AFCEBFF24FA0B" pageId="4" pageNumber="205" type="diagnosis">
<paragraph id="8B8C36D0C064FFE9FF6AFCEBFF24FA0B" blockId="4.[151,1436,1182,2035]" pageId="4" pageNumber="205">
<emphasis id="B947EAC2C064FFE9FF6AFCEBFEA5FB33" bold="true" box="[199,321,1182,1207]" pageId="4" pageNumber="205">Diagnosis.</emphasis>
A species of 
<taxonomicName id="4C334D53C064FFE9FE75FCEAFDCBFB32" box="[472,559,1183,1206]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="genus">
<emphasis id="B947EAC2C064FFE9FE75FCEAFDCBFB32" box="[472,559,1183,1206]" italics="true" pageId="4" pageNumber="205">Trimma</emphasis>
</taxonomicName>
unique in the possession of two dark oblique stripes on either side of the pupil (blue, edged with red in life, dark brown in preservative), the lower of which continues posteriorly across the dorsal margin of the opercle. The dorsal half of the body is red and the ventral half abruptly white in life. The species has a narrow bony interorbital (≤50 % pupil width), a moderate interorbital trench with a slight groove posterodorsal to the eye, no scales on the cheek, opercle or in the predorsal midline, no elongate spines in the first dorsal fin, 9–10 dorsal and 9 anal fin rays, pectoral fin rays 17–18 of which 7–11 are branched, and an unbranched fifth pelvic fin ray.
</paragraph>
</subSubSection>
<subSubSection id="C329655BC064FFE4FF6AFDEFFC74FF34" lastPageId="9" lastPageNumber="210" pageId="4" pageNumber="205" type="description">
<paragraph id="8B8C36D0C064FFE8FF6AFDEFFE45FCFB" blockId="4.[151,1436,1182,2035]" lastBlockId="5.[151,1437,151,896]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<emphasis id="B947EAC2C064FFE9FF6AFDEFFEBDFA37" bold="true" box="[199,345,1434,1459]" pageId="4" pageNumber="205">Description.</emphasis>
Dorsal fins VI + 
<date id="FF8D1010C064FFE9FD85FDEEFD96FA37" box="[552,626,1434,1459]" pageId="4" pageNumber="205" value="2010-01-09">
I 
<emphasis id="B947EAC2C064FFE9FD97FDEFFDACFA37" bold="true" box="[570,584,1434,1459]" pageId="4" pageNumber="205">9</emphasis>
–10
</date>
(mean = 9.6), second or third spine longest but not elongate, reaching posteriorly to between bases of sixth spine of D1 and spine of D2 when adpressed, first ray of second dorsal fin unbranched (branched in 
<specimenCount id="9D35FD59C064FFE9FE61FD96FD86FA7F" box="[460,610,1506,1531]" pageId="4" pageNumber="205" type="generic">3 specimens</specimenCount>
), posterior element of last dorsal ray unbranched, last ray reaching posteriorly 50– 
<emphasis id="B947EAC2C064FFE9FEECFE73FEB9F99B" bold="true" box="[321,349,1542,1567]" pageId="4" pageNumber="205">67</emphasis>
–72% (mean = 65.8%) of distance between its base and first dorsal procurrent caudal ray; anal fin 
<date id="FF8D1010C064FFE9FF3AFE5EFF51F9C7" box="[151,181,1579,1603]" pageId="4" pageNumber="205">I 9</date>
(once 10), first ray unbranched as is posterior element of last ray, which reaches posteriorly 45– 
<emphasis id="B947EAC2C064FFE9FB5CFE5FFAE9F9C7" bold="true" box="[1265,1293,1578,1603]" pageId="4" pageNumber="205">62</emphasis>
–65% (mean = 57.4%) distance between its base and first ventral procurrent caudal ray; pectoral fin 
<emphasis id="B947EAC2C064FFE9FBC7FE3BFB62F9E3" bold="true" box="[1130,1158,1614,1639]" pageId="4" pageNumber="205">17</emphasis>
–18 (mean = 17.2), 3– 
<emphasis id="B947EAC2C064FFE9FA2DFE3BFA6AF9E3" bold="true" box="[1408,1422,1614,1639]" pageId="4" pageNumber="205">4</emphasis>
– 6 (mean = 4.1) dorsal and 2– 
<emphasis id="B947EAC2C064FFE9FE4CFE07FE0BF90F" bold="true" box="[481,495,1650,1675]" pageId="4" pageNumber="205">5</emphasis>
(mean = 4.1) ventral unbranched rays separated by 
<date id="FF8D1010C064FFE9FB92FE06FB77F90F" box="[1087,1171,1650,1675]" pageId="4" pageNumber="205" value="2012-08-07">
7– 
<emphasis id="B947EAC2C064FFE9FBF6FE07FB8DF90F" bold="true" box="[1115,1129,1650,1675]" pageId="4" pageNumber="205">8</emphasis>
–12
</date>
(mean = 9.1) branched rays, fin reaching posteriorly to a point above area between anus to urogenital papilla; pelvic fin 
<date id="FF8D1010C064FFE9FB7BFEE2FB13F92B" box="[1238,1271,1686,1711]" pageId="4" pageNumber="205">I 5</date>
, first four rays with one sequential branch, fifth ray unbranched (
<figureCitation id="13082A55C064FFE9FD68FECFFCE3F957" box="[709,775,1722,1747]" captionStart="FIGURE 3" captionStartId="4.[151,250,1059,1081]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@4.[151,1435,193,1038]" captionTargetPageId="4" captionText="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="4" pageNumber="205">Fig. 3</figureCitation>
) and 52–67% the length of fourth (mean = 57.0, fourth ray damaged in 
<typeStatus id="54888872C064FFE9FEB2FEABFE61F973" box="[287,389,1758,1783]" pageId="4" pageNumber="205" type="holotype">holotype</typeStatus>
), fourth ray reaching posteriorly to a point between posterior margin of urogenital papilla and base of second anal ray; no fraenum; basal membrane varies from absent, where membrane from fifth ray joins to abdomen lateral to midline to 9% length of fourth ray (
<figureCitation id="13082A55C064FFE9FCB6FF53FC87F8BB" box="[795,867,1830,1855]" captionStart="FIGURE 3" captionStartId="4.[151,250,1059,1081]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@4.[151,1435,193,1038]" captionTargetPageId="4" captionText="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="4" pageNumber="205">Fig. 3</figureCitation>
, red arrow). Lateral scales 23– 
<emphasis id="B947EAC2C064FFE9FB66FF53FB03F8BB" bold="true" box="[1227,1255,1830,1855]" pageId="4" pageNumber="205">24</emphasis>
(mean = 23.4), where 24 count due to additional small anteriormost scale; anterior transverse scales 
<date id="FF8D1010C064FFE9FBF6FF3EFB4BF8E7" box="[1115,1199,1866,1891]" pageId="4" pageNumber="205" value="2010-09-08">
8– 
<emphasis id="B947EAC2C064FFE9FBDBFF3FFB60F8E7" bold="true" box="[1142,1156,1866,1891]" pageId="4" pageNumber="205">9</emphasis>
–10
</date>
(mean = 8.9; yellow arrow in 
<figureCitation id="13082A55C064FFE9FF50FF1BFEDBF803" box="[253,319,1902,1927]" captionStart="FIGURE 3" captionStartId="4.[151,250,1059,1081]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@4.[151,1435,193,1038]" captionTargetPageId="4" captionText="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="4" pageNumber="205">Fig. 3</figureCitation>
shows direction of count); posterior transverse scales 7– 
<emphasis id="B947EAC2C064FFE9FC6CFF1BFC2BF803" bold="true" box="[961,975,1902,1927]" pageId="4" pageNumber="205">8</emphasis>
(mean = 7.9; blue arrow in 
<figureCitation id="13082A55C064FFE9FAA1FF1BFAAAF803" box="[1292,1358,1902,1927]" captionStart="FIGURE 3" captionStartId="4.[151,250,1059,1081]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@4.[151,1435,193,1038]" captionTargetPageId="4" captionText="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="4" pageNumber="205">Fig. 3</figureCitation>
shows direction of count); no scales on median predorsal, cheek or opercle; scales extending anteriorly on sides of body from upper pectoral base curving up to middle of first dorsal fin, a few specimens with 1–2 small isolated cycloid scales on side of nape; pectoral-fin base usually with 3 vertical rows of scales, with 1– 
<emphasis id="B947EAC2C064FFE9FBD5FFAFFB62F877" bold="true" box="[1144,1158,2010,2035]" pageId="4" pageNumber="205">2</emphasis>
, 2– 
<emphasis id="B947EAC2C064FFE9FB1FFFAFFB24F877" bold="true" box="[1202,1216,2010,2035]" pageId="4" pageNumber="205">4</emphasis>
and 3– 
<emphasis id="B947EAC2C064FFE9FABBFFAFFAC0F877" bold="true" box="[1302,1316,2010,2035]" pageId="4" pageNumber="205">5</emphasis>
scales per row respectively; scales in midline anterior to pelvic fin 4– 
<emphasis id="B947EAC2C065FFE8FCEFF8E2FCB4FF34" bold="true" box="[834,848,151,176]" pageId="5" pageNumber="206">5</emphasis>
(mean = 4.5), where 4, usually an additional scale anteriorly just lateral to midline; scales on pectoral-fin base, breast, anterior midline of belly cycloid (
<figureCitation id="13082A55C065FFE8FAA0F8C9FAB5FF50" box="[1293,1361,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1059,1081]" captionTargetBox="[151,1435,193,1037]" captionTargetId="figure@4.[151,1435,193,1038]" captionTargetPageId="4" captionText="FIGURE 3. Trimma aturirii, 15.3 mm SL female, ROM 95269, ventral view of region between pelvic and anal fins. Red arrow indicates basal membrane; green arrows show anterior extent of ctenoid scales on belly; yellow and blue arrows show direction for anterior and posterior transverse scale counts respectively. Specimen stained with cyanine blue. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240779/files/figure.png" pageId="5" pageNumber="206">Fig. 3</figureCitation>
, green arrows show anteriormost ctenoid scales), scales on sides of nape and rest of body ctenoid; 12 circumpeduncular scales, 7– 
<emphasis id="B947EAC2C065FFE8FEA9F971FEF6FE99" bold="true" box="[260,274,260,285]" pageId="5" pageNumber="206">8</emphasis>
(mean = 7.2) scales between base of last anal fin ray and first ventral procurrent caudal ray. Upper jaw teeth with an outer row of 6–8 evenly spaced, curved, enlarged canines decreasing in size posteriorly and extending to bend of premaxilla, short inner row of straight conical teeth slightly larger than teeth anterior to them, ending before bend of premaxilla, 2–3 irregular rows of small conical teeth between these, grading to single row on posterior part of premaxilla. Lower jaw teeth with outer row of about 6 evenly spaced, curved, enlarged canines ending at bend of dentary, inner row of enlarged, slightly curved teeth about one-third length of outer row, decreasing in size posteriorly, ending at beginning of coronoid process, 2–3 irregular rows of small conical teeth between these, grading posteriorly to single row and ending near anterodorsal tip of coronoid process of dentary. Tongue broadly rounded, sometimes with small central tip. Gill opening extending anteroventrally to below mid- to posterior third of pupil; outer gill rakers on first gill arch 3– 
<emphasis id="B947EAC2C065FFE8FC9BFA32FCA0FDE4" bold="true" box="[822,836,583,608]" pageId="5" pageNumber="206">4</emphasis>
(mean = 3.3) + 
<date id="FF8D1010C065FFE8FC57FA3DFB8EFDE4" box="[1018,1130,583,608]" pageId="5" pageNumber="206" value="2013-12-14">
12– 
<emphasis id="B947EAC2C065FFE8FB89FA32FBA4FDE4" bold="true" box="[1060,1088,583,608]" pageId="5" pageNumber="206">13</emphasis>
–14
</date>
(mean = 13.1), total rakers 15– 
<emphasis id="B947EAC2C065FFE8FF6CFA19FF39FD01" bold="true" box="[193,221,620,645]" pageId="5" pageNumber="206">17</emphasis>
(mean = 16.4). Anterior naris a short tube extending half way over upper lip, posterior naris a pore with raised rim, separated from eye by 
<emphasis id="B947EAC2C065FFE8FDB4FAFAFDD8FD2C" bold="true" box="[537,572,655,680]" pageId="5" pageNumber="206">1.5</emphasis>
–2 times its diameter, nasal apparatus occupying anterior two-thirds of snout (
<figureCitation id="13082A55C065FFE8FF32FAC1FF00FD48" box="[159,228,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[157,1429,193,1932]" captionTargetId="figure@6.[157,1429,193,1934]" captionTargetPageId="6" captionText="FIGURE 4. Trimma aturirii, 16.2 mm SL male, ROM 95269, lateral (A) and dorsal (B) views of head papillae. Red arrows show supernumerary papillae, thin white lines indicate lines of papillae. Specimen stained with cyanine blue. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/240780/files/figure.png" pageId="5" pageNumber="206">Fig. 4</figureCitation>
). Bony interorbital width 30– 
<emphasis id="B947EAC2C065FFE8FD80FAC1FDADFD49" bold="true" box="[557,585,692,717]" pageId="5" pageNumber="206">33</emphasis>
–43% (mean = 36.6) of pupil diameter in width, interorbital with moderately developed ‘U’-shaped trough about as wide as deep, a shallow postorbital groove but not trench-like. Cephalic sensory papillae as in 
<figureCitation id="13082A55C065FFE8FE3CFA89FE16FC90" box="[401,498,764,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[157,1429,193,1932]" captionTargetId="figure@6.[157,1429,193,1934]" captionTargetPageId="6" captionText="FIGURE 4. Trimma aturirii, 16.2 mm SL male, ROM 95269, lateral (A) and dorsal (B) views of head papillae. Red arrows show supernumerary papillae, thin white lines indicate lines of papillae. Specimen stained with cyanine blue. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/240780/files/figure.png" pageId="5" pageNumber="206">Figure 4</figureCitation>
, number of papillae in each row given in 
<tableCitation id="C6B1036BC065FFE8FC61FA89FBC4FC90" box="[972,1056,764,789]" captionStart="TABLE 1" captionStartId="5.[151,239,940,963]" captionTargetBox="[159,1349,1086,2018]" captionTargetPageId="5" captionText="TABLE 1. Counts of cephalic sensory papillae in the different rows of the three new species of Trimma. Data given as minimum to maximum values, with the mean, Standard Deviation and number of specimens for which data were available respectively given in parentheses, except where the value was invariant, where only the number of specimens is given. Terminology follows that used in Winterbottom (2011) as emended here." httpUri="http://table.plazi.org/id/DF4C6658C065FFE8FF3AFBD9FC0CFBA7" pageId="5" pageNumber="206" tableUuid="DF4C6658C065FFE8FF3AFBD9FC0CFBA7">Table 1</tableCitation>
. Ridge of tissue (or dermal crest) may extend anterior to first dorsal spine for up to two-thirds distance to interorbital. Epaxialis extending anteriorly to above a vertical with posterior margin of pupil. Abdominal/caudal vertebral transition not examined, but presumed to be 
<typeStatus id="54888872C065FFE8FEE4FB1DFE64FC04" box="[329,384,872,896]" pageId="5" pageNumber="206">Type</typeStatus>
B.
</paragraph>
<caption id="DF4C6658C065FFE8FF3AFBD9FC0CFBA7" ID-Table-UUID="DF4C6658C065FFE8FF3AFBD9FC0CFBA7" httpUri="http://table.plazi.org/id/DF4C6658C065FFE8FF3AFBD9FC0CFBA7" pageId="5" pageNumber="206" targetBox="[159,1349,1086,2018]" targetIsTable="true" targetPageId="5">
<paragraph id="8B8C36D0C065FFE8FF3AFBD9FC0CFBA7" blockId="5.[151,1436,940,1059]" pageId="5" pageNumber="206">
<emphasis id="B947EAC2C065FFE8FF3AFBD9FEEEFC47" bold="true" box="[151,266,940,963]" pageId="5" pageNumber="206">TABLE 1.</emphasis>
Counts of cephalic sensory papillae in the different rows of the three new species of 
<taxonomicName id="4C334D53C065FFE8FB0AFBD9FB1EFC47" box="[1191,1274,940,963]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="genus">
<emphasis id="B947EAC2C065FFE8FB0AFBD9FB1EFC47" box="[1191,1274,940,963]" italics="true" pageId="5" pageNumber="206">Trimma</emphasis>
</taxonomicName>
. Data given as minimum to maximum values, with the mean, Standard Deviation and number of specimens for which data were available respectively given in parentheses, except where the value was invariant, where only the number of specimens is given. Terminology follows that used in Winterbottom (2011) as emended here.
</paragraph>
</caption>
<paragraph id="8B8C36D0C065FFE8FF32FC4BFAC4F866" pageId="5" pageNumber="206">
<table id="F933C470C0650012FF32FC4BFAA1F866" box="[159,1349,1086,2018]" gridcols="4" gridrows="23" pageId="5" pageNumber="206">
<tr id="35033492C0650012FF32FC4BFAA1FBD1" box="[159,1349,1086,1109]" gridrow="0" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FC4BFEC2FBD1" box="[159,294,1086,1109]" gridcol="0" gridrow="0" pageId="5" pageNumber="206">Papillae row</th>
<th id="76D25DEEC0650012FE09FC4BFD76FBD1" box="[420,658,1086,1109]" gridcol="1" gridrow="0" pageId="5" pageNumber="206">
<taxonomicName id="4C334D53C065FFE8FE09FC35FDE2FBD1" box="[420,518,1088,1109]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="aturirii">
<emphasis id="B947EAC2C065FFE8FE09FC35FDE2FBD1" box="[420,518,1088,1109]" italics="true" pageId="5" pageNumber="206">T. aturirii</emphasis>
</taxonomicName>
</th>
<th id="76D25DEEC0650012FCA5FC4BFC0EFBD1" box="[776,1002,1086,1109]" gridcol="2" gridrow="0" pageId="5" pageNumber="206">
<taxonomicName id="4C334D53C065FFE8FCA5FC35FC90FBD0" box="[776,884,1086,1109]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="kardium">
<emphasis id="B947EAC2C065FFE8FCA5FC35FC90FBD0" box="[776,884,1086,1109]" italics="true" pageId="5" pageNumber="206">T. kardium</emphasis>
</taxonomicName>
</th>
<th id="76D25DEEC0650012FBFAFC4BFAA1FBD1" box="[1111,1349,1086,1109]" gridcol="3" gridrow="0" pageId="5" pageNumber="206">
<taxonomicName id="4C334D53C065FFE8FBFAFC35FB02FBD0" box="[1111,1254,1086,1109]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="trioculatum">
<emphasis id="B947EAC2C065FFE8FBFAFC35FB02FBD0" box="[1111,1254,1086,1109]" italics="true" pageId="5" pageNumber="206">T. trioculatum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35033492C0650012FF32FC1DFAA1FBFA" box="[159,1349,1128,1150]" gridrow="1" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FC1DFEC2FBFA" box="[159,294,1128,1150]" gridcol="0" gridrow="1" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FC1CFF06FBFA" box="[213,226,1129,1150]" italics="true" pageId="5" pageNumber="206">a</emphasis>
</th>
<td id="76D25DEEC0650012FE09FC1DFD76FBFA" box="[420,658,1128,1150]" gridcol="1" gridrow="1" pageId="5" pageNumber="206">6 (17)</td>
<td id="76D25DEEC0650012FCA5FC1DFC0EFBFA" box="[776,1002,1128,1150]" gridcol="2" gridrow="1" pageId="5" pageNumber="206">6 (4)</td>
<td id="76D25DEEC0650012FBFAFC1DFAA1FBFA" box="[1111,1349,1128,1150]" gridcol="3" gridrow="1" pageId="5" pageNumber="206">6 (11)</td>
</tr>
<tr id="35033492C0650012FF32FCE4FAA1FB23" box="[159,1349,1169,1191]" gridrow="2" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FCE4FEC2FB23" box="[159,294,1169,1191]" gridcol="0" gridrow="2" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FCE4FF06FB23" box="[213,226,1169,1191]" italics="true" pageId="5" pageNumber="206">b</emphasis>
</th>
<td id="76D25DEEC0650012FE09FCE4FD76FB23" box="[420,658,1169,1191]" gridcol="1" gridrow="2" pageId="5" pageNumber="206">5–7 (5.9, 0.484, 16)</td>
<td id="76D25DEEC0650012FCA5FCE4FC0EFB23" box="[776,1002,1169,1191]" gridcol="2" gridrow="2" pageId="5" pageNumber="206">5–7 (5.8, 0.839, 4)</td>
<td id="76D25DEEC0650012FBFAFCE4FAA1FB23" box="[1111,1349,1169,1191]" gridcol="3" gridrow="2" pageId="5" pageNumber="206">6–8 (6.3, 0.617, 11)</td>
</tr>
<tr id="35033492C0650012FF32FCCFFAA1FB54" box="[159,1349,1210,1232]" gridrow="3" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FCCFFEC2FB54" box="[159,294,1210,1232]" gridcol="0" gridrow="3" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FCCEFF04FB54" box="[213,224,1211,1232]" italics="true" pageId="5" pageNumber="206">c</emphasis>
</th>
<td id="76D25DEEC0650012FE09FCCFFD76FB54" box="[420,658,1210,1232]" gridcol="1" gridrow="3" pageId="5" pageNumber="206">5 (16)</td>
<td id="76D25DEEC0650012FCA5FCCFFC0EFB54" box="[776,1002,1210,1232]" gridcol="2" gridrow="3" pageId="5" pageNumber="206">5 (4)</td>
<td id="76D25DEEC0650012FBFAFCCFFAA1FB54" box="[1111,1349,1210,1232]" gridcol="3" gridrow="3" pageId="5" pageNumber="206">5 (11)</td>
</tr>
<tr id="35033492C0650012FF32FC91FAA1FB7E" box="[159,1349,1252,1274]" gridrow="4" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FC91FEC2FB7E" box="[159,294,1252,1274]" gridcol="0" gridrow="4" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FC90FF0AFB7E" box="[213,238,1253,1274]" italics="true" pageId="5" pageNumber="206">cp</emphasis>
</th>
<td id="76D25DEEC0650012FE09FC91FD76FB7E" box="[420,658,1252,1274]" gridcol="1" gridrow="4" pageId="5" pageNumber="206">1 (16)</td>
<td id="76D25DEEC0650012FCA5FC91FC0EFB7E" box="[776,1002,1252,1274]" gridcol="2" gridrow="4" pageId="5" pageNumber="206">1 (4)</td>
<td id="76D25DEEC0650012FBFAFC91FAA1FB7E" box="[1111,1349,1252,1274]" gridcol="3" gridrow="4" pageId="5" pageNumber="206">1 (11)</td>
</tr>
<tr id="35033492C0650012FF32FD78FAA1FAA7" box="[159,1349,1293,1315]" gridrow="5" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FD78FEC2FAA7" box="[159,294,1293,1315]" gridcol="0" gridrow="5" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FD78FF06FAA7" box="[213,226,1293,1315]" italics="true" pageId="5" pageNumber="206">d</emphasis>
</th>
<td id="76D25DEEC0650012FE09FD78FD76FAA7" box="[420,658,1293,1315]" gridcol="1" gridrow="5" pageId="5" pageNumber="206">4–6 (5.5, 0.707, 16)</td>
<td id="76D25DEEC0650012FCA5FD78FC0EFAA7" box="[776,1002,1293,1315]" gridcol="2" gridrow="5" pageId="5" pageNumber="206">6–7 (6.5, 0.500, 4)</td>
<td id="76D25DEEC0650012FBFAFD78FAA1FAA7" box="[1111,1349,1293,1315]" gridcol="3" gridrow="5" pageId="5" pageNumber="206">5–8 (6.4, 0.979, 11)</td>
</tr>
<tr id="35033492C0650012FF32FD43FAA1FAC8" box="[159,1349,1334,1356]" gridrow="6" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FD43FEC2FAC8" box="[159,294,1334,1356]" gridcol="0" gridrow="6" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FD43FF03FAC8" box="[213,231,1334,1356]" italics="true" pageId="5" pageNumber="206">d'</emphasis>
</th>
<td id="76D25DEEC0650012FE09FD43FD76FAC8" box="[420,658,1334,1356]" gridcol="1" gridrow="6" pageId="5" pageNumber="206">5–7 (5.8, 0.634, 16)</td>
<td id="76D25DEEC0650012FCA5FD43FC0EFAC8" box="[776,1002,1334,1356]" gridcol="2" gridrow="6" pageId="5" pageNumber="206">6–7 (6.5, 0.500, 4)</td>
<td id="76D25DEEC0650012FBFAFD43FAA1FAC8" box="[1111,1349,1334,1356]" gridcol="3" gridrow="6" pageId="5" pageNumber="206">6–9 (7.0, 0.853, 11)</td>
</tr>
<tr id="35033492C0650012FF32FD15FAA1FAF2" box="[159,1349,1376,1398]" gridrow="7" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FD15FEC2FAF2" box="[159,294,1376,1398]" gridcol="0" gridrow="7" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FD14FEFFFAF2" box="[213,283,1377,1398]" italics="true" pageId="5" pageNumber="206">e –ant.</emphasis>
</th>
<td id="76D25DEEC0650012FE09FD15FD76FAF2" box="[420,658,1376,1398]" gridcol="1" gridrow="7" pageId="5" pageNumber="206">10–13 (11.1, 0.872, 17)</td>
<td id="76D25DEEC0650012FCA5FD15FC0EFAF2" box="[776,1002,1376,1398]" gridcol="2" gridrow="7" pageId="5" pageNumber="206">11–13 (11.5, 0.866, 4)</td>
<td id="76D25DEEC0650012FBFAFD15FAA1FAF2" box="[1111,1349,1376,1398]" gridcol="3" gridrow="7" pageId="5" pageNumber="206">11–14 (12.5, 0.782, 11)</td>
</tr>
<tr id="35033492C0650012FF32FDFCFAA1FA1B" box="[159,1349,1417,1439]" gridrow="8" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FDFCFEC2FA1B" box="[159,294,1417,1439]" gridcol="0" gridrow="8" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FDFFFEF9FA1B" box="[213,285,1418,1439]" italics="true" pageId="5" pageNumber="206">e –post</emphasis>
.
</th>
<td id="76D25DEEC0650012FE09FDFCFD76FA1B" box="[420,658,1417,1439]" gridcol="1" gridrow="8" pageId="5" pageNumber="206">10–14 (11.3, 1.250, 16)</td>
<td id="76D25DEEC0650012FCA5FDFCFC0EFA1B" box="[776,1002,1417,1439]" gridcol="2" gridrow="8" pageId="5" pageNumber="206">12–13 (12.3, 0.433, 4)</td>
<td id="76D25DEEC0650012FBFAFDFCFAA1FA1B" box="[1111,1349,1417,1439]" gridcol="3" gridrow="8" pageId="5" pageNumber="206">12–15 (13.1, 0.966, 11)</td>
</tr>
<tr id="35033492C0650012FF32FDC7FAA1FA4C" box="[159,1349,1458,1480]" gridrow="9" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FDC7FEC2FA4C" box="[159,294,1458,1480]" gridcol="0" gridrow="9" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FDC6FEF9FA4C" box="[213,285,1459,1480]" italics="true" pageId="5" pageNumber="206">i – ant.</emphasis>
</th>
<td id="76D25DEEC0650012FE09FDC7FD76FA4C" box="[420,658,1458,1480]" gridcol="1" gridrow="9" pageId="5" pageNumber="206">7–8 (7.5, 0.499, 17)</td>
<td id="76D25DEEC0650012FCA5FDC7FC0EFA4C" box="[776,1002,1458,1480]" gridcol="2" gridrow="9" pageId="5" pageNumber="206">7–8 (7.8, 0.433, 4)</td>
<td id="76D25DEEC0650012FBFAFDC7FAA1FA4C" box="[1111,1349,1458,1480]" gridcol="3" gridrow="9" pageId="5" pageNumber="206">6–8 (7.2, 0.833, 11)</td>
</tr>
<tr id="35033492C0650012FF32FDA9FAA1FA76" box="[159,1349,1500,1522]" gridrow="10" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FDA9FEC2FA76" box="[159,294,1500,1522]" gridcol="0" gridrow="10" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF79FDA8FEC2FA76" box="[212,294,1501,1522]" italics="true" pageId="5" pageNumber="206">i – post.</emphasis>
</th>
<td id="76D25DEEC0650012FE09FDA9FD76FA76" box="[420,658,1500,1522]" gridcol="1" gridrow="10" pageId="5" pageNumber="206">6–8 (7.4, 0.704, 16)</td>
<td id="76D25DEEC0650012FCA5FDA9FC0EFA76" box="[776,1002,1500,1522]" gridcol="2" gridrow="10" pageId="5" pageNumber="206">7–8 (7.5, 0.500, 4)</td>
<td id="76D25DEEC0650012FBFAFDA9FAA1FA76" box="[1111,1349,1500,1522]" gridcol="3" gridrow="10" pageId="5" pageNumber="206">6–8, 7.2, 0.575, 11)</td>
</tr>
<tr id="35033492C0650012FF32FE70FAA1F99F" box="[159,1349,1541,1563]" gridrow="11" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FE70FEC2F99F" box="[159,294,1541,1563]" gridcol="0" gridrow="11" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FE73FF06F99F" box="[213,226,1542,1563]" italics="true" pageId="5" pageNumber="206">p</emphasis>
</th>
<td id="76D25DEEC0650012FE09FE70FD76F99F" box="[420,658,1541,1563]" gridcol="1" gridrow="11" pageId="5" pageNumber="206">6 (17)</td>
<td id="76D25DEEC0650012FCA5FE70FC0EF99F" box="[776,1002,1541,1563]" gridcol="2" gridrow="11" pageId="5" pageNumber="206">6 (4)</td>
<td id="76D25DEEC0650012FBFAFE70FAA1F99F" box="[1111,1349,1541,1563]" gridcol="3" gridrow="11" pageId="5" pageNumber="206">6–7 (6.1, 0.287, 11)</td>
</tr>
<tr id="35033492C0650012FF32FE5BFAA1F9C1" box="[159,1349,1582,1605]" gridrow="12" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FE5BFEC2F9C1" box="[159,294,1582,1605]" gridcol="0" gridrow="12" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FE45FF3BF9C1" box="[213,223,1584,1605]" italics="true" pageId="5" pageNumber="206">r</emphasis>
</th>
<td id="76D25DEEC0650012FE09FE5BFD76F9C1" box="[420,658,1582,1605]" gridcol="1" gridrow="12" pageId="5" pageNumber="206">2 (17)</td>
<td id="76D25DEEC0650012FCA5FE5BFC0EF9C1" box="[776,1002,1582,1605]" gridcol="2" gridrow="12" pageId="5" pageNumber="206">2 (4)</td>
<td id="76D25DEEC0650012FBFAFE5BFAA1F9C1" box="[1111,1349,1582,1605]" gridcol="3" gridrow="12" pageId="5" pageNumber="206">1–2 (1.9, 0.287, 11)</td>
</tr>
<tr id="35033492C0650012FF32FE2DFAA1F9EA" box="[159,1349,1624,1646]" gridrow="13" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FE2DFEC2F9EA" box="[159,294,1624,1646]" gridcol="0" gridrow="13" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FE2DFF38F9EA" box="[213,220,1624,1646]" italics="true" pageId="5" pageNumber="206">f</emphasis>
</th>
<td id="76D25DEEC0650012FE09FE2DFD76F9EA" box="[420,658,1624,1646]" gridcol="1" gridrow="13" pageId="5" pageNumber="206">2–4 (2.9, 0.428, 16)</td>
<td id="76D25DEEC0650012FCA5FE2DFC0EF9EA" box="[776,1002,1624,1646]" gridcol="2" gridrow="13" pageId="5" pageNumber="206">3 (4)</td>
<td id="76D25DEEC0650012FBFAFE2DFAA1F9EA" box="[1111,1349,1624,1646]" gridcol="3" gridrow="13" pageId="5" pageNumber="206">2–3 (2.9, 0.287, 11)</td>
</tr>
<tr id="35033492C0650012FF32FEF4FAA1F913" box="[159,1349,1665,1687]" gridrow="14" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FEF4FEC2F913" box="[159,294,1665,1687]" gridcol="0" gridrow="14" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FEF7FF11F913" box="[213,245,1666,1687]" italics="true" pageId="5" pageNumber="206">cs&quot;</emphasis>
</th>
<td id="76D25DEEC0650012FE09FEF4FD76F913" box="[420,658,1665,1687]" gridcol="1" gridrow="14" pageId="5" pageNumber="206">3 (17)</td>
<td id="76D25DEEC0650012FCA5FEF4FC0EF913" box="[776,1002,1665,1687]" gridcol="2" gridrow="14" pageId="5" pageNumber="206">3 (4)</td>
<td id="76D25DEEC0650012FBFAFEF4FAA1F913" box="[1111,1349,1665,1687]" gridcol="3" gridrow="14" pageId="5" pageNumber="206">3 (11)</td>
</tr>
<tr id="35033492C0650012FF32FEDFFAA1F944" box="[159,1349,1706,1728]" gridrow="15" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FEDFFEC2F944" box="[159,294,1706,1728]" gridcol="0" gridrow="15" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FEDEFF06F944" box="[213,226,1707,1728]" italics="true" pageId="5" pageNumber="206">g</emphasis>
</th>
<td id="76D25DEEC0650012FE09FEDFFD76F944" box="[420,658,1706,1728]" gridcol="1" gridrow="15" pageId="5" pageNumber="206">6–8 (7.2, 0.716, 11)</td>
<td id="76D25DEEC0650012FCA5FEDFFC0EF944" box="[776,1002,1706,1728]" gridcol="2" gridrow="15" pageId="5" pageNumber="206">4–6 (5.0, 0,707, 4)</td>
<td id="76D25DEEC0650012FBFAFEDFFAA1F944" box="[1111,1349,1706,1728]" gridcol="3" gridrow="15" pageId="5" pageNumber="206">4–6 (5.0, 0.577, 6)</td>
</tr>
<tr id="35033492C0650012FF32FEA1FAA1F96E" box="[159,1349,1748,1770]" gridrow="16" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FEA1FEC2F96E" box="[159,294,1748,1770]" gridcol="0" gridrow="16" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FEA0FF06F96E" box="[213,226,1749,1770]" italics="true" pageId="5" pageNumber="206">n</emphasis>
</th>
<td id="76D25DEEC0650012FE09FEA1FD76F96E" box="[420,658,1748,1770]" gridcol="1" gridrow="16" pageId="5" pageNumber="206">1 (5)</td>
<td id="76D25DEEC0650012FCA5FEA1FC0EF96E" box="[776,1002,1748,1770]" gridcol="2" gridrow="16" pageId="5" pageNumber="206">1 (1)</td>
<td id="76D25DEEC0650012FBFAFEA1FAA1F96E" box="[1111,1349,1748,1770]" gridcol="3" gridrow="16" pageId="5" pageNumber="206">1-3 (2)</td>
</tr>
<tr id="35033492C0650012FF32FE88FAA1F897" box="[159,1349,1789,1811]" gridrow="17" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FE88FEC2F897" box="[159,294,1789,1811]" gridcol="0" gridrow="17" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FE8BFF04F897" box="[213,224,1790,1811]" italics="true" pageId="5" pageNumber="206">x</emphasis>
</th>
<td id="76D25DEEC0650012FE09FE88FD76F897" box="[420,658,1789,1811]" gridcol="1" gridrow="17" pageId="5" pageNumber="206">5–6 (5.5, 0.500, 16)</td>
<td id="76D25DEEC0650012FCA5FE88FC0EF897" box="[776,1002,1789,1811]" gridcol="2" gridrow="17" pageId="5" pageNumber="206">6–7 (6.5, 0.500, 4)</td>
<td id="76D25DEEC0650012FBFAFE88FAA1F897" box="[1111,1349,1789,1811]" gridcol="3" gridrow="17" pageId="5" pageNumber="206">7–8 (7.6, 0.484, 8)</td>
</tr>
<tr id="35033492C0650012FF32FF53FAA1F8B9" box="[159,1349,1830,1853]" gridrow="18" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FF53FEC2F8B9" box="[159,294,1830,1853]" gridcol="0" gridrow="18" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FF5DFF06F8B9" box="[213,226,1832,1853]" italics="true" pageId="5" pageNumber="206">u</emphasis>
</th>
<td id="76D25DEEC0650012FE09FF53FD76F8B9" box="[420,658,1830,1853]" gridcol="1" gridrow="18" pageId="5" pageNumber="206">4–5 (4.8, 0.433, 4)</td>
<td id="76D25DEEC0650012FCA5FF53FC0EF8B9" box="[776,1002,1830,1853]" gridcol="2" gridrow="18" pageId="5" pageNumber="206">5 (1)</td>
<td id="76D25DEEC0650012FBFAFF53FAA1F8B9" box="[1111,1349,1830,1853]" gridcol="3" gridrow="18" pageId="5" pageNumber="206">5 (1)</td>
</tr>
<tr id="35033492C0650012FF32FF25FAA1F8E2" box="[159,1349,1872,1894]" gridrow="19" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FF25FEC2F8E2" box="[159,294,1872,1894]" gridcol="0" gridrow="19" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FF24FF3BF8E2" box="[213,223,1873,1894]" italics="true" pageId="5" pageNumber="206">z</emphasis>
</th>
<td id="76D25DEEC0650012FE09FF25FD76F8E2" box="[420,658,1872,1894]" gridcol="1" gridrow="19" pageId="5" pageNumber="206">4–6 (4.5, 0.612, 16)</td>
<td id="76D25DEEC0650012FCA5FF25FC0EF8E2" box="[776,1002,1872,1894]" gridcol="2" gridrow="19" pageId="5" pageNumber="206">5–6 (5.3, 0.433, 4)</td>
<td id="76D25DEEC0650012FBFAFF25FAA1F8E2" box="[1111,1349,1872,1894]" gridcol="3" gridrow="19" pageId="5" pageNumber="206">5–6 (5.4, 0.490, 10)</td>
</tr>
<tr id="35033492C0650012FF32FF0CFAA1F80B" box="[159,1349,1913,1935]" gridrow="20" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FF0CFEC2F80B" box="[159,294,1913,1935]" gridcol="0" gridrow="20" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FF0FFF0DF80B" box="[213,233,1914,1935]" italics="true" pageId="5" pageNumber="206">ot</emphasis>
</th>
<td id="76D25DEEC0650012FE09FF0CFD76F80B" box="[420,658,1913,1935]" gridcol="1" gridrow="20" pageId="5" pageNumber="206">10–13 (11.1, 1.053, 16)</td>
<td id="76D25DEEC0650012FCA5FF0CFC0EF80B" box="[776,1002,1913,1935]" gridcol="2" gridrow="20" pageId="5" pageNumber="206">9-13 (11.8, 1.639,6)</td>
<td id="76D25DEEC0650012FBFAFF0CFAA1F80B" box="[1111,1349,1913,1935]" gridcol="3" gridrow="20" pageId="5" pageNumber="206">12–16 (13.5, 1.076, 11)</td>
</tr>
<tr id="35033492C0650012FF32FFD7FAA1F83C" box="[159,1349,1954,1976]" gridrow="21" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FFD7FEC2F83C" box="[159,294,1954,1976]" gridcol="0" gridrow="21" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FFD6FF08F83C" box="[213,236,1955,1976]" italics="true" pageId="5" pageNumber="206">os</emphasis>
</th>
<td id="76D25DEEC0650012FE09FFD7FD76F83C" box="[420,658,1954,1976]" gridcol="1" gridrow="21" pageId="5" pageNumber="206">4–6 (4.8, 0.6534, 15)</td>
<td id="76D25DEEC0650012FCA5FFD7FC0EF83C" box="[776,1002,1954,1976]" gridcol="2" gridrow="21" pageId="5" pageNumber="206">4 (2)</td>
<td id="76D25DEEC0650012FBFAFFD7FAA1F83C" box="[1111,1349,1954,1976]" gridcol="3" gridrow="21" pageId="5" pageNumber="206">4–7 (6.1, 0.875, 9)</td>
</tr>
<tr id="35033492C0650012FF32FFB9FAA1F866" box="[159,1349,1996,2018]" gridrow="22" pageId="5" pageNumber="206">
<th id="76D25DEEC0650012FF32FFB9FEC2F866" box="[159,294,1996,2018]" gridcol="0" gridrow="22" pageId="5" pageNumber="206">
Row 
<emphasis id="B947EAC2C065FFE8FF78FFB8FF0DF866" box="[213,233,1997,2018]" italics="true" pageId="5" pageNumber="206">oi</emphasis>
</th>
<td id="76D25DEEC0650012FE09FFB9FD76F866" box="[420,658,1996,2018]" gridcol="1" gridrow="22" pageId="5" pageNumber="206">3–4 (3.8, 0.390, 16)</td>
<td id="76D25DEEC0650012FCA5FFB9FC0EF866" box="[776,1002,1996,2018]" gridcol="2" gridrow="22" pageId="5" pageNumber="206">4 (4)</td>
<td id="76D25DEEC0650012FBFAFFB9FAA1F866" box="[1111,1349,1996,2018]" gridcol="3" gridrow="22" pageId="5" pageNumber="206">3–5 (3.9, 0.539, 10)</td>
</tr>
</table>
</paragraph>
<caption id="DF4C6658C066FFEBFF3AFFEAFECFF876" httpUri="https://zenodo.org/record/240780/files/figure.png" pageId="6" pageNumber="207" targetBox="[157,1429,193,1932]" targetPageId="6">
<paragraph id="8B8C36D0C066FFEBFF3AFFEAFECFF876" blockId="6.[151,1436,1951,2034]" pageId="6" pageNumber="207">
<emphasis id="B947EAC2C066FFEBFF3AFFEAFEEAF830" bold="true" box="[151,270,1951,1973]" pageId="6" pageNumber="207">FIGURE 4</emphasis>
. 
<emphasis id="B947EAC2C066FFEBFEB1FFD5FE26F831" box="[284,450,1952,1973]" italics="true" pageId="6" pageNumber="207">
<taxonomicName id="4C334D53C066FFEBFEB1FFD5FE5AF831" box="[284,446,1952,1973]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="aturirii">Trimma aturirii</taxonomicName>
,
</emphasis>
16.2 mm SL male, ROM 95269, lateral (A) and dorsal (B) views of head papillae. Red arrows show supernumerary papillae, thin white lines indicate lines of papillae. Specimen stained with cyanine blue. Photos by R. Winterbottom.
</paragraph>
</caption>
<caption id="DF4C6658C067FFEAFF3AFFE3FDF6F86E" httpUri="https://zenodo.org/record/240781/files/figure.png" pageId="7" pageNumber="208" targetBox="[151,1435,193,1919]" targetPageId="7">
<paragraph id="8B8C36D0C067FFEAFF3AFFE3FDF6F86E" blockId="7.[151,1436,1942,2026]" pageId="7" pageNumber="208">
<emphasis id="B947EAC2C067FFEAFF3AFFE3FEF6F828" bold="true" box="[151,274,1942,1964]" pageId="7" pageNumber="208">FIGURE 5</emphasis>
. 
<emphasis id="B947EAC2C067FFEAFE8EFFEDFE2AF829" box="[291,462,1944,1965]" italics="true" pageId="7" pageNumber="208">
<taxonomicName id="4C334D53C067FFEAFE8EFFEDFE2EF829" box="[291,458,1944,1965]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="aturirii">Trimma aturirii</taxonomicName>
,
</emphasis>
left lateral views of live specimens from Mapia Atoll, ROM 94889, holotype (A—image reversed); Ayau Besar Atoll, ROM 98853 (B—image reversed); and a freshly collected specimen from Mof Island, ROM 87934 (C). Photos by M.V. Erdmann.
</paragraph>
</caption>
<caption id="DF4C6658C068FFE5FF3AFC6FFE4EFBCB" httpUri="https://zenodo.org/record/240782/files/figure.png" pageId="8" pageNumber="209" targetBox="[152,1434,179,1035]" targetPageId="8">
<paragraph id="8B8C36D0C068FFE5FF3AFC6FFE4EFBCB" blockId="8.[151,1437,1050,1103]" pageId="8" pageNumber="209">
<emphasis id="B947EAC2C068FFE5FF3AFC6FFEF4FBAB" bold="true" box="[151,272,1050,1072]" pageId="8" pageNumber="209">FIGURE 6</emphasis>
. 
<emphasis id="B947EAC2C068FFE5FEB2FC6EFE22FBB4" box="[287,454,1051,1072]" italics="true" pageId="8" pageNumber="209">
<taxonomicName id="4C334D53C068FFE5FEB2FC6EFE27FBB4" box="[287,451,1051,1072]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="aturirii">Trimma aturirii</taxonomicName>
,
</emphasis>
14.0mm SL female, ROM 87934, Mof Island, Raja Ampat, lateral view of head (unstained). Photo by R. Winterbottom.
</paragraph>
</caption>
<paragraph id="8B8C36D0C068FFE5FF6AFC09FC94F97D" blockId="8.[151,1437,1148,2037]" pageId="8" pageNumber="209">
<emphasis id="B947EAC2C068FFE5FF6AFC09FE43FB11" box="[199,423,1148,1173]" italics="true" pageId="8" pageNumber="209">Colour pattern, live</emphasis>
, based on images of 
<typeStatus id="54888872C068FFE5FD36FC09FCE4FB11" box="[667,768,1148,1173]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
, 
<quantity id="4CCB9B35C068FFE5FCA0FC09FC96FB10" box="[781,882,1148,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="8" pageNumber="209" unit="mm" value="18.2">18.2 mm</quantity>
SL female, ROM 94889, Mapia Atoll, 
<figureCitation id="13082A55C068FFE5FA9BFC09FA9DFB11" box="[1334,1401,1148,1173]" captionStart="FIGURE 5" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[151,1435,193,1919]" captionTargetId="figure@7.[151,1435,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Trimma aturirii, left lateral views of live specimens from Mapia Atoll, ROM 94889, holotype (A—image reversed); Ayau Besar Atoll, ROM 98853 (B—image reversed); and a freshly collected specimen from Mof Island, ROM 87934 (C). Photos by M. V. Erdmann." httpUri="https://zenodo.org/record/240781/files/figure.png" pageId="8" pageNumber="209">Fig. 5</figureCitation>
A, and 
<typeStatus id="54888872C068FFE5FF6BFCD4FECDFB3D" box="[198,297,1185,1209]" pageId="8" pageNumber="209" type="paratype">paratype</typeStatus>
, Ayau Besar Atoll, ROM 98809, 
<figureCitation id="13082A55C068FFE5FD0DFCD5FD05FB3D" box="[672,737,1184,1209]" captionStart="FIGURE 5" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[151,1435,193,1919]" captionTargetId="figure@7.[151,1435,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Trimma aturirii, left lateral views of live specimens from Mapia Atoll, ROM 94889, holotype (A—image reversed); Ayau Besar Atoll, ROM 98853 (B—image reversed); and a freshly collected specimen from Mof Island, ROM 87934 (C). Photos by M. V. Erdmann." httpUri="https://zenodo.org/record/240781/files/figure.png" pageId="8" pageNumber="209">Fig. 5</figureCitation>
B. 
<typeStatus id="54888872C068FFE5FCA4FCD5FC95FB3D" box="[777,881,1184,1209]" pageId="8" pageNumber="209" type="holotype">Holotype</typeStatus>
bicoloured, dark red dorsally and white ventrally below midlateral line continuing anteriorly across cheek just below eye. Midline of dorsum with diffuse white stripe from above posterior margin of pupil to anteriormost dorsal procurrent caudal fin ray. Iris dark gold above pupil, red margined purple stripe across top of pupil which continues anteriorly on snout as diffuse, and much wider, red stripe. White stripe margined dorsally with red across bottom of pupil, continuing anteriorly to maxilla and, after short disjunction behind eye, posteriorly across top of opercle, where white grades to blue margined dorsally with red. Area of snout between stripes and above dorsal stripe yellow, anterior snout and lips red. Rays of dorsal, pectoral and upper half of caudal fins light red (pelvic and anal fin not visible), ventral half of caudal lighter. Upper pectoral fin base with white spot more intense than surroundings. Internal lighter stripe (spinal cord?) anteriorly, followed by three internal lighter spots beneath dorsal fins. Specimen from Ayau Besar Atoll essentially similar, but upper body more orange than red, lower half of body very pale pink, anterior half of lower eye stripe blue rather than white, and spot at upper pectoral fin base more obvious and bluish-white. In other live specimens, dorsum reddish-orange (possibly juveniles). Freshly collected specimen from 
<collectingCountry id="F3247640C068FFE5FBA1FE25FBA8F9ED" box="[1036,1100,1616,1641]" name="Iceland" pageId="8" pageNumber="209">Pulau</collectingCountry>
Mof (ROM 87934, 
<figureCitation id="13082A55C068FFE5FA98FE25FA93F9ED" box="[1333,1399,1616,1641]" captionStart="FIGURE 5" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[151,1435,193,1919]" captionTargetId="figure@7.[151,1435,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Trimma aturirii, left lateral views of live specimens from Mapia Atoll, ROM 94889, holotype (A—image reversed); Ayau Besar Atoll, ROM 98853 (B—image reversed); and a freshly collected specimen from Mof Island, ROM 87934 (C). Photos by M. V. Erdmann." httpUri="https://zenodo.org/record/240781/files/figure.png" pageId="8" pageNumber="209">Fig. 5</figureCitation>
C) with yellow head grading to orange on nape and then to orange-red behind this with light greenish-yellow cheek, pale pinkish white abdomen and posterior body, blue or white of iris stripes dark purple, dorsal fins with basal, half-pupil diameter wide basal white stripe bordered dorsally by yellow stripe half this width, then hyaline, caudal fin with outer edges whitish and central rays dirty orange-brown.
</paragraph>
<paragraph id="8B8C36D0C068FFE4FF6AFF71FC74FF34" blockId="8.[151,1437,1148,2037]" lastBlockId="9.[151,1437,151,1760]" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<emphasis id="B947EAC2C068FFE5FF6AFF71FE61F899" box="[199,389,1796,1821]" italics="true" pageId="8" pageNumber="209">Preserved colour</emphasis>
. Background pale straw yellow, with heavy concentrations of dark brown chromatophores on dorsal half of nape and body, decreasing in number ventrally, where absent from about two scale rows from ventral midline. Moderate number of similar chromatophores on cheek with some on dorsal part of opercle, and on upper and lower lips. Stripes across iris evident as lines of dark brown chromatophores (
<figureCitation id="13082A55C068FFE5FB25FF05FB37F80D" box="[1160,1235,1904,1929]" captionStart="FIGURE 6" captionStartId="8.[151,250,1050,1072]" captionTargetBox="[152,1434,179,1035]" captionTargetId="figure@8.[152,1435,179,1035]" captionTargetPageId="8" captionText="FIGURE 6. Trimma aturirii, 14.0 mm SL female, ROM 87934, Mof Island, Raja Ampat, lateral view of head (unstained). Photo by R. Winterbottom." httpUri="https://zenodo.org/record/240782/files/figure.png" pageId="8" pageNumber="209">Fig. 6</figureCitation>
) which continue anterodorsally onto snout. Lower line continues posterodorsally as dark ovoid patch over dorsal tip of preopercle and from anterodorsal process of opercle half way to its posterodorsal tip; in some specimens dark line continuous from snout to opercle. Fins hyaline except for scattered brown pigment cells on distal half of second dorsal fin, some in caudal fin, and along distal margin of last few anal fin rays.
</paragraph>
</subSubSection>
<subSubSection id="C329655BC069FFE4FF6AF8C9FE6DFEC4" pageId="9" pageNumber="210" type="etymology">
<paragraph id="8B8C36D0C069FFE4FF6AF8C9FE6DFEC4" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
<emphasis id="B947EAC2C069FFE4FF6AF8C9FEAAFF51" bold="true" box="[199,334,188,213]" pageId="9" pageNumber="210">Etymology.</emphasis>
Named in honor of Abraham Aturiri, Governor of West 
<collectingCountry id="F3247640C069FFE4FC54F8C8FBD9FF50" box="[1017,1085,189,212]" name="Papua New Guinea" pageId="9" pageNumber="210">Papua</collectingCountry>
province from 
<date id="FF8D1010C069FFE4FB57F8C9FA92FF51" box="[1274,1398,188,213]" pageId="9" pageNumber="210" value="2006" valueMax="2016">2006-2016</date>
, in recognition of his energetic efforts to ensure the conservation and wise, sustainable use of the hugely diverse coral reefs of the Bird's Head Seascape for the benefit of the Papuan people. The species has been informally referred to as 
<taxonomicName id="4C334D53C069FFE4FF18F95CFEE8FEC4" box="[181,268,297,320]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B947EAC2C069FFE4FF18F95CFEE8FEC4" box="[181,268,297,320]" italics="true" pageId="9" pageNumber="210">Trimma</emphasis>
</taxonomicName>
RW sp 99.
</paragraph>
</subSubSection>
<subSubSection id="C329655BC069FFE4FF6AF939FEB9F98C" pageId="9" pageNumber="210" type="distribution">
<paragraph id="8B8C36D0C069FFE4FF6AF939FE97FE0C" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
<emphasis id="B947EAC2C069FFE4FF6AF939FE84FEE1" bold="true" box="[199,352,332,357]" pageId="9" pageNumber="210">Distribution.</emphasis>
Currently known with certainty only from the northern oceanic atolls of West 
<collectingCountry id="F3247640C069FFE4FB4BF938FACFFEE0" box="[1254,1323,333,356]" name="Papua New Guinea" pageId="9" pageNumber="210">Papua</collectingCountry>
province, 
<collectingCountry id="F3247640C069FFE4FF3AF91AFEE0FE0C" box="[151,260,367,392]" name="Indonesia" pageId="9" pageNumber="210">Indonesia</collectingCountry>
(
<figureCitation id="13082A55C069FFE4FEBEF91AFE82FE0C" box="[275,358,367,392]" captionStart="FIGURE 18" captionStartId="23.[151,250,1897,1919]" captionTargetBox="[174,1412,963,1865]" captionTargetId="figure@23.[166,1420,949,1876]" captionTargetPageId="23" captionText="FIGURE 18. Distribution of the three new species based on material examined for this study. Trimma aturirii, black inverted triangles. Trimma kardium, black diamonds. Trimma trioculatum, coloured pentagons. Blue fill, type locality, Group 1 haplogroup; green fill, Palau, Group 2 haplogroup; yellow fill, Rabaul, New Britain, Group 3 haplogroup; red fill, unknown haplogroup." httpUri="https://zenodo.org/record/240794/files/figure.png" pageId="9" pageNumber="210">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8B8C36D0C069FFE4FF6AF9E1FEB9F98C" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
<emphasis id="B947EAC2C069FFE4FF6AF9E1FE81FE29" bold="true" box="[199,357,404,429]" pageId="9" pageNumber="210">Comparisons</emphasis>
. 
<taxonomicName id="4C334D53C069FFE4FEDAF9E0FDCDFE28" box="[375,553,405,428]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="aturirii">
<emphasis id="B947EAC2C069FFE4FEDAF9E0FDCDFE28" box="[375,553,405,428]" italics="true" pageId="9" pageNumber="210">Trimma aturirii</emphasis>
</taxonomicName>
is unique among species in the genus in its bipartite colour pattern (red or orange above, white or pale pink below). The two oblique dark stripes on either side of the pupil (blue-edged with red margins in life, dark brown in preservative), the lower of which continues after a short hiatus posteriorly along the dorsal margin of the opercle, are shared only with an undescribed species (
<emphasis id="B947EAC2C069FFE4FBA5FA74FBF1FD9C" box="[1032,1045,513,536]" italics="true" pageId="9" pageNumber="210">T</emphasis>
. RW sp 49) from the Line Islands, which lie some 7,500 kilometers due east of the easternmost sample from 
<collectingCountry id="F3247640C069FFE4FC5BFA51FB83FDB9" box="[1014,1127,548,573]" name="Indonesia" pageId="9" pageNumber="210">Indonesia</collectingCountry>
. Specimens from the latter locality invariably have 10 (vs. 9) anal rays, and the red colouration of the body extends ventrally almost to the ventral midline (vs. confined above midlateral septum). Ten other described species of 
<taxonomicName id="4C334D53C069FFE4FBC7FA18FB24FD00" box="[1130,1216,621,644]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B947EAC2C069FFE4FBC7FA18FB24FD00" box="[1130,1216,621,644]" italics="true" pageId="9" pageNumber="210">Trimma</emphasis>
</taxonomicName>
share with the new species the lack of scales in the predorsal midline and an unbranched fifth pelvic fin ray. Of these, 
<taxonomicName id="4C334D53C069FFE4FA84FAE4FE99FD48" authority="Winterbottom, 2000" authorityName="Winterbottom" authorityYear="2000" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="anaima">
<emphasis id="B947EAC2C069FFE4FA84FAE4FA78FD2C" box="[1321,1436,657,680]" italics="true" pageId="9" pageNumber="210">T. anaima</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FF3AFAC1FE99FD48" author="Winterbottom" box="[151,381,692,717]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom, 2000</bibRefCitation>
</taxonomicName>
, has 8 dorsal and anal fin rays, unbranched pectoral fin rays, and less than 15 total gill rakers (vs. 9–10, 9, middle rays branched and 15 or more gill rakers). 
<taxonomicName id="4C334D53C069FFE4FC0FFAACFA8EFD74" authority="Tomiyama, 1936" authorityName="Tomiyama" authorityYear="1936" box="[930,1386,727,752]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="grammistes">
<emphasis id="B947EAC2C069FFE4FC0FFAACFB63FD74" box="[930,1159,729,752]" italics="true" pageId="9" pageNumber="210">Trimma grammistes</emphasis>
(
<bibRefCitation id="EFA24B21C069FFE4FB36FAA2FA85FD74" author="Tomiyama" box="[1179,1377,727,752]" pageId="9" pageNumber="210" refString="Tomiyama, I. (1936) Gobiidae of Japan. Japanese Journal of Zoology, 7, 37 - 112." type="journal article" year="1936">Tomiyama, 1936</bibRefCitation>
)
</taxonomicName>
has unbranched pectoral fin rays and a second dorsal spine that reaches posteriorly beyond the first element of the second dorsal fin when adpressed (vs. branched, and not reaching beyond spine). That species also has a pupildiameter wide black stripe from the upper lip through the eye and along the body above the midlateral line until it curves gradually to the center of the caudal peduncle and onto the caudal fin (vs. stripe absent). Three other species, 
<taxonomicName id="4C334D53C069FFE4FF3AFBF8FE1EFC20" authority="Winterbottom, 2000" authorityName="Winterbottom" authorityYear="2000" box="[151,506,908,933]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="bisella">
<emphasis id="B947EAC2C069FFE4FF3AFBF8FEE7FC21" box="[151,259,908,933]" italics="true" pageId="9" pageNumber="210">T. bisella</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FEA2FBF9FE1EFC20" author="Winterbottom" box="[271,506,908,933]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom, 2000</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C334D53C069FFE4FDA6FBF8FCBDFC20" authority="Winterbottom, 2004" authorityName="Winterbottom" authorityYear="2004" box="[523,857,908,933]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="cana">
<emphasis id="B947EAC2C069FFE4FDA6FBF8FD87FC20" box="[523,611,909,932]" italics="true" pageId="9" pageNumber="210">T. cana</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FDC3FBF9FCBDFC20" author="Winterbottom" box="[622,857,908,933]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2004) Three new species of Trimma (Pisces; Gobiidae) from the central, western and south Pacific. aqua, Journal of Ichthyology and aquatic Biology, 9 (1), 7 - 16" type="journal article" year="2004">Winterbottom, 2004</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C334D53C069FFE4FC32FBF8FB18FC20" authority="Winterbottom, 2004" authorityName="Winterbottom" authorityYear="2004" box="[927,1276,908,933]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="sostra">
<emphasis id="B947EAC2C069FFE4FC32FBF8FBE0FC20" box="[927,1028,909,932]" italics="true" pageId="9" pageNumber="210">T. sostra</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FBBCFBF9FB18FC20" author="Winterbottom" box="[1041,1276,908,933]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2004) Three new species of Trimma (Pisces; Gobiidae) from the central, western and south Pacific. aqua, Journal of Ichthyology and aquatic Biology, 9 (1), 7 - 16" type="journal article" year="2004">Winterbottom, 2004</bibRefCitation>
</taxonomicName>
, have greatly elongate second dorsal spines (vs. not elongate) and lack a dermal crest anterior to the first dorsal fin (vs. present). The former species has four red bars on the head and white saddles on the peduncle (vs. both absent), while the latter two have a pale body with seven broad red bars or blotches (vs. bars absent). 
<taxonomicName id="4C334D53C069FFE4FB04FB8CFDEDFBB0" authority="Winterbottom &amp; Southcott, 2007" authorityName="Winterbottom &amp; Southcott" authorityYear="2007" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="sanguinellus">
<emphasis id="B947EAC2C069FFE4FB04FB8CFA7FFB94" box="[1193,1435,1015,1040]" italics="true" pageId="9" pageNumber="210">Trimma sanguinellus</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FF3AFC69FDEDFBB0" author="Winterbottom" box="[151,521,1052,1077]" pageId="9" pageNumber="210" refString="Winterbottom, R. &amp; Southcott, L. (2007) Two new species of the genus Trimma from western Thailand. aqua, Journal of Ichthyology and aquatic Biology, 13 (2), 69 - 76." type="journal article" year="2007">Winterbottom &amp; Southcott, 2007</bibRefCitation>
</taxonomicName>
has an entirely red body, unbranched pectoral fin rays, and a second dorsal spine that extends posteriorly beyond the first element of the second dorsal fin when adpressed (vs. bicoloured, branched, and second spine anterior first spine of second dorsal fin respectively). 
<taxonomicName id="4C334D53C069FFE4FC1CFC10FABDFBF8" authority="Hoese et al., 2011" authorityName="Hoese et al." authorityYear="2011" box="[945,1369,1124,1149]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="maiandros">
<emphasis id="B947EAC2C069FFE4FC1CFC10FB62FBF9" box="[945,1158,1124,1149]" italics="true" pageId="9" pageNumber="210">Trimma maiandros</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FB21FC10FABDFBF8" author="Hoese" box="[1164,1369,1124,1149]" pageId="9" pageNumber="210" refString="Hoese, D. F., Winterbottom, R. &amp; Reader, S. (2011) Trimma maiandros, a new species of pygmy goby (Gobiidae) from the Indo-west Pacific. aqua, International Journal of Ichthyology, 72 (2), 103 - 110." type="journal article" year="2011">
Hoese 
<emphasis id="B947EAC2C069FFE4FB74FC10FAEEFBF9" box="[1241,1290,1124,1149]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
, has a pale grey body with large yellow blotches, 5 or fewer branched pectoral rays, and no opercular stripe (vs. bicoloured, &lt;5 branched pectoral rays, and dark opercular stripe respectively). 
<taxonomicName id="4C334D53C069FFE4FBABFCD8FA78FB40" authority="Winterbottom, 2001" authorityName="Winterbottom" authorityYear="2001" box="[1030,1436,1196,1221]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="stobbsi">
<emphasis id="B947EAC2C069FFE4FBABFCD8FB57FB41" box="[1030,1203,1196,1221]" italics="true" pageId="9" pageNumber="210">Trimma stobbsi</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FB17FCD9FA78FB40" author="Winterbottom" box="[1210,1436,1196,1221]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2001) Two new gobiid fish species in Trimma and Trimmatom (Teleostei: Gobiidae) from the Indian and Western Pacific Oceans. aqua, Journal of Ichthyology and aquatic Biology, 5 (1), 19 - 24." type="journal article" year="2001">Winterbottom, 2001</bibRefCitation>
</taxonomicName>
has a yellow to brown body with scale pockets outlined and a dark, half pupil diameter sized, roughly ovoid spot on the upper margin of the opercle near its posterior margin (vs. bicoloured with stripe along dorsal margin of opercle). 
<taxonomicName id="4C334D53C069FFE4FEACFD6CFD0CFAB4" authority="Winterbottom, 2009" authorityName="Winterbottom" authorityYear="2009" box="[257,744,1303,1328]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="hotsarihiensis">
<emphasis id="B947EAC2C069FFE4FEACFD6CFE18FAB4" box="[257,508,1303,1328]" italics="true" pageId="9" pageNumber="210">Trimma hotsarihiensis</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FDAEFD62FD0CFAB4" author="Winterbottom" box="[515,744,1303,1328]" pageId="9" pageNumber="210" refString="Winterbottom, R. (2009) A new species of the genus Trimma (Percomorpha; Gobiidae) from Helen Reef, South-West Islands of Palau. aqua, Journal of Ichthyology and aquatic Biology, 15 (2), 109 - 116." type="journal article" year="2009">Winterbottom, 2009</bibRefCitation>
</taxonomicName>
, has a yellow body with scale margins more strongly marked, yellow spots in the caudal fin and dark basal stripes in the dorsal and anal fins. It has a dark diffuse stripe across the top of the opercle, but the stripe is horizontal and ends anteriorly at the level of the mid-pupil rather than being obliquely oriented downwards to pass below the eye. 
<taxonomicName id="4C334D53C069FFE4FD51FDF0FB43FA18" authority="Suzuki &amp; Senou, 2008" authorityName="Suzuki &amp; Senou" authorityYear="2008" box="[764,1191,1412,1437]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="kudoi">
<emphasis id="B947EAC2C069FFE4FD51FDF0FC7DFA19" box="[764,921,1412,1437]" italics="true" pageId="9" pageNumber="210">Trimma kudoi</emphasis>
<bibRefCitation id="EFA24B21C069FFE4FC0EFDF1FB43FA18" author="Suzuki" box="[931,1191,1412,1437]" pageId="9" pageNumber="210" refString="Suzuki, T. &amp; Senou, H. (2008) Two new species of the gobiid fish genus Trimma (Perciformes: Gobioidei) from Southern Japan. Bulletin of the National Museum of Natural Science, Series A, 2 (Supplement), 97 - 106." type="journal article" year="2008">Suzuki &amp; Senou, 2008</bibRefCitation>
</taxonomicName>
, has a similar overall colouration as 
<taxonomicName id="4C334D53C069FFE4FE92FDDCFE1DFA44" box="[319,505,1447,1472]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="hotsarihiensis">
<emphasis id="B947EAC2C069FFE4FE92FDDCFE1DFA44" box="[319,505,1447,1472]" italics="true" pageId="9" pageNumber="210">T. hotsarihiensis</emphasis>
</taxonomicName>
, but lacks the dark stripe over the opercle. It has two oblique purple-blue stripes, one at the dorsal margin of the iris and a second that passes across the middle of the pupil (vs. two stripes on either side of the pupil).
</paragraph>
</subSubSection>
<subSubSection id="C329655BC069FFE4FF6AFE61FB14F964" pageId="9" pageNumber="210" type="discussion">
<paragraph id="8B8C36D0C069FFE4FF6AFE61FB66F91C" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
<emphasis id="B947EAC2C069FFE4FF6AFE61FEA1F9A9" bold="true" box="[199,325,1556,1581]" pageId="9" pageNumber="210">Discussion</emphasis>
. Specimens of this species were usually found perched individually on vertical surfaces at the back of crevices and overhangs in depths ranging between 
<quantity id="4CCB9B35C069FFE4FD41FE4DFCB5F9D4" box="[748,849,1592,1616]" metricMagnitude="1" metricUnit="m" metricValue="5.4" metricValueMax="7.0" metricValueMin="3.8" pageId="9" pageNumber="210" unit="m" value="54.0" valueMax="70.0" valueMin="38.0">
38– 
<specimenCount id="9D35FD59C069FFE4FCB8FE4DFCB5F9D4" box="[789,849,1592,1616]" pageId="9" pageNumber="210" type="generic">70 m</specimenCount>
</quantity>
. Despite extensive sampling of these depths on reefs throughout eastern 
<collectingCountry id="F3247640C069FFE4FEDEFE29FE07F9F1" box="[371,483,1628,1653]" name="Indonesia" pageId="9" pageNumber="210">Indonesia</collectingCountry>
, this species has only been found on exposed outer reef slopes of the northernmost oceanic atolls and islands of West 
<collectingCountry id="F3247640C069FFE4FDB5FEF4FDB9F91C" box="[536,605,1665,1688]" name="Papua New Guinea" pageId="9" pageNumber="210">Papua</collectingCountry>
(Mapia Atoll, Ayau Besar Atoll and 
<collectingCountry id="F3247640C069FFE4FC57FE0AFBDEF91C" box="[1018,1082,1663,1688]" name="Iceland" pageId="9" pageNumber="210">Pulau</collectingCountry>
Mof).
</paragraph>
<paragraph id="8B8C36D0C069FFE4FF6AFED1FB14F964" blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="210">
An analysis of the CO1 gene of the three tissue specimens of this species suggests that they are phenetically closest to 
<taxonomicName id="4C334D53C069FFE4FEAAFEBCFE5BF964" box="[263,447,1735,1760]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="hotsarihiensis">
<emphasis id="B947EAC2C069FFE4FEAAFEBCFE5BF964" box="[263,447,1735,1760]" italics="true" pageId="9" pageNumber="210">T. hotsarihiensis</emphasis>
</taxonomicName>
, from which they are separated by a minimum of 9.9% of the base pairs.
</paragraph>
</subSubSection>
</treatment>
</document>