treatments-xml/data/03/9A/87/039A87C10D629B0AFF12F9C7FB2EF9A1.xml

<document ID-DOI="10.26107/RBZ-2023-0021" ID-ISSN="2345-7600" ID-Zenodo-Dep="7816093" ID-ZooBank="887FBA2D-60F5-4CEE-B5F0-047EE6B9709D" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1681182249975" checkinUser="felipe" docAuthor="Huang, Wen-Chien, Liao, Te-Yu &amp; Tan, Heok Hui" docDate="2023" docId="039A87C10D629B0AFF12F9C7FB2EF9A1" docLanguage="en" docName="RafflesBZool.71.268–278.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.4:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="4" docTitle="Uropterygius cyamommatus Huang &amp; Liao &amp; Tan 2023, new species" docType="treatment" docVersion="2" lastPageNumber="273" masterDocId="FFA3FFB90D609B0FFFD9FF84FFEAFFD3" masterDocTitle="Uropterygius cyamommatus, a new moray eel (Anguilliformes: Muraenidae) from anchialine caves in Christmas Island, Australia, and Panglao Island, the Philippines" masterLastPageNumber="278" masterPageNumber="268" pageNumber="270" updateTime="1681184308295" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Uropterygius cyamommatus, a new moray eel (Anguilliformes: Muraenidae) from anchialine caves in Christmas Island, Australia, and Panglao Island, the Philippines</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Wen-Chien</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liao, Te-Yu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tan, Heok Hui</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Raffles Bulletin of Zoology</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>71</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>268</mods:start>
<mods:end>278</mods:end>
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<mods:identifier type="DOI">10.26107/RBZ-2023-0021</mods:identifier>
<mods:identifier type="ISSN">2345-7600</mods:identifier>
<mods:identifier type="Zenodo-Dep">7816093</mods:identifier>
<mods:identifier type="ZooBank">887FBA2D-60F5-4CEE-B5F0-047EE6B9709D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039A87C10D629B0AFF12F9C7FB2EF9A1" httpUri="http://treatment.plazi.org/id/039A87C10D629B0AFF12F9C7FB2EF9A1" lastPageId="5" lastPageNumber="273" pageId="2" pageNumber="270">
<subSubSection pageId="2" pageNumber="270" type="nomenclature">
<paragraph blockId="2.[203,663,1603,1691]" box="[203,663,1603,1627]" pageId="2" pageNumber="270">
<heading bold="true" box="[203,663,1603,1627]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="270" reason="2">
<emphasis bold="true" box="[203,663,1603,1627]" pageId="2" pageNumber="270">
<taxonomicName authority="Huang &amp; Liao &amp; Tan, 2023" authorityName="Huang &amp; Liao &amp; Tan" authorityYear="2023" box="[203,514,1603,1627]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="2" pageNumber="270" phylum="Chordata" rank="species" species="cyamommatus" status="new species">
<emphasis bold="true" box="[203,514,1603,1627]" italics="true" pageId="2" pageNumber="270">Uropterygius cyamommatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[529,663,1603,1627]" pageId="2" pageNumber="270" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[203,663,1603,1691]" pageId="2" pageNumber="270">
Common name: Bean-eyed snake moray (
<figureCitation box="[332,437,1667,1691]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="3.[136,171,1949,1971]" captionStartId-1="5.[138,173,811,833]" captionStartId-2="6.[813,848,950,972]" captionStartId-3="7.[136,171,1140,1162]" captionStartId-4="8.[113,148,1532,1554]" captionTargetBox-0="[136,1474,302,1933]" captionTargetBox-1="[136,1475,189,795]" captionTargetBox-2="[813,1452,189,935]" captionTargetBox-3="[143,1469,189,1123]" captionTargetBox-4="[113,1451,189,1517]" captionTargetId-0="figure-8@3.[136,1474,302,1933]" captionTargetId-1="figure-568@5.[136,1475,189,795]" captionTargetId-2="figure-486@6.[813,1452,189,935]" captionTargetId-3="figure-8@7.[75,1511,169,1127]" captionTargetId-4="figure-12@8.[113,1451,189,1517]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="Fig. 3. Uropterygius cyamommatus, new species, holotype, WAM P.35403.001, 361 mm TL. A, fresh colouration (photographed by HHT); B, preserved colouration (photographed by WCH). Arrows indicate the position of anus." captionText-1="Fig. 4. Uropterygius cyamommatus, new species, lateral view of head marked with cephalic sensory pores: red for supraorbital pores; green for infraorbital pores; blue for preoperculo-mandibular pores; yellow for the branchial pore. Arrow indicates the gill opening. Photo of WAM P.35403.001, holotype, 361 mm TL (photographed by HHT)." captionText-2="Fig.5. Uropterygius cyamommatus, new species, lateral view of the head region of a specimen without the left eye. A, external view; B, radiograph showing the skeletal structure of orbital region. Photo of NMMB-P37200, paratype, 380 mm TL (photographed by HHT)." captionText-3="Fig. 6. Dentition of Uropterygius cyamommatus, new species, holotype, WAM P.35403.001. Upper jaw (left) and lower jaw (right). Dotted circles represent the sockets of missing teeth. Schematic drawings by WCH." captionText-4="Fig. 7. Radiograph showing the skeletal structure of Uropterygius cyamommatus, new species, ZRC 46849-1, 352 mm TL (photographed by HHT). Arrow indicates the position of anus." figureDoi-0="http://doi.org/10.5281/zenodo.7816101" figureDoi-1="http://doi.org/10.5281/zenodo.7816103" figureDoi-2="http://doi.org/10.5281/zenodo.7816105" figureDoi-3="http://doi.org/10.5281/zenodo.7816107" figureDoi-4="http://doi.org/10.5281/zenodo.7816111" httpUri-0="https://zenodo.org/record/7816101/files/figure.png" httpUri-1="https://zenodo.org/record/7816103/files/figure.png" httpUri-2="https://zenodo.org/record/7816105/files/figure.png" httpUri-3="https://zenodo.org/record/7816107/files/figure.png" httpUri-4="https://zenodo.org/record/7816111/files/figure.png" pageId="2" pageNumber="270">Figs. 3–7</figureCitation>
, 
<tableCitation box="[450,531,1667,1691]" captionStart="Table 1" captionStartId="4.[113,168,189,211]" captionText="Table 1. Morphometric measurements and meristic counts of Uropterygius cyamommatus, new species. The means of morphometric values and vertebral number are given in parentheses." pageId="2" pageNumber="270">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="273" pageId="2" pageNumber="270" type="materials_examined">
<paragraph blockId="2.[112,753,1731,1883]" pageId="2" pageNumber="270">
<materialsCitation collectingDate="2010-02-01" collectionCode="WAM" collectorName="H. H. Tan" country="Christmas Island" location="Whip Cave" pageId="2" pageNumber="270" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[113,222,1731,1755]" pageId="2" pageNumber="270">
<typeStatus box="[113,216,1731,1755]" pageId="2" pageNumber="270">Holotype</typeStatus>
.
</emphasis>
<collectionCode box="[226,293,1731,1755]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="270" type="Museum">WAM</collectionCode>
P.35403.001 (
<quantity box="[442,528,1731,1755]" metricMagnitude="-1" metricUnit="m" metricValue="3.61" pageId="2" pageNumber="270" unit="mm" value="361.0">361 mm</quantity>
TL, male); 
<collectingCountry name="Christmas Island" pageId="2" pageNumber="270">Christmas Island</collectingCountry>
, 
<collectingCountry box="[189,291,1763,1787]" name="Australia" pageId="2" pageNumber="270">Australia</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:039A87C10D629B0AFF12F9C7FB2EF9A1:8EEC600C0D629B0DFEF5F967FE4EF928" box="[300,420,1763,1787]" country="Christmas Island" name="Whip Cave" pageId="2" pageNumber="270">Whip Cave</location>
(10°25.377ʹS 105°42.081ʹE), a cave system with a narrow cave mouth and lower chambers filled with anchialine water affected by tidal changes; 
<collectingMethod pageId="2" pageNumber="270">baited fish trap</collectingMethod>
, 
<date box="[217,414,1859,1883]" pageId="2" pageNumber="270" value="2010-02-01">
<collectingDate box="[217,414,1859,1883]" pageId="2" pageNumber="270" value="2010-02-01">01 February 2010</collectingDate>
</date>
, coll. 
<collectorName box="[482,583,1860,1883]" pageId="2" pageNumber="270">H.H. Tan</collectorName>
et al.
</materialsCitation>
</paragraph>
<paragraph blockId="2.[113,753,1923,2107]" lastBlockId="2.[811,1453,861,1205]" pageId="2" pageNumber="270">
<materialsCitation collectionCode="WAM" country="Christmas Island" county="Pre-anus" location="Freshwater Cave" municipality="Vertebral" pageId="2" pageNumber="270" specimenCode="NMMB-P37200, ZRC 56488-1, ZRC 56488- 2" specimenCount="4" typeStatus="paratype">
<emphasis bold="true" box="[113,259,1923,1947]" pageId="2" pageNumber="270">
<typeStatus box="[113,249,1923,1947]" pageId="2" pageNumber="270">Paratypes</typeStatus>
.
</emphasis>
Four specimens (
<quantity box="[510,680,1923,1947]" metricMagnitude="-1" metricUnit="m" metricValue="3.67" metricValueMax="4.01" metricValueMin="3.3299999999999996" pageId="2" pageNumber="270" unit="mm" value="367.0" valueMax="401.0" valueMin="333.0">333 – 401 mm</quantity>
TL). 
<specimenCode box="[113,288,1955,1979]" collectionCode="NMMB-P" pageId="2" pageNumber="270">NMMB-P37200</specimenCode>
(
<quantity box="[304,397,1955,1979]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="270" unit="mm" value="380.0">380 mm</quantity>
, male), 
<collectionCode box="[484,552,1955,1979]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="270" type="Museum">WAM</collectionCode>
P.35404.001 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="270" unit="mm" value="390.0">390 mm</quantity>
, female), 
<specimenCode box="[263,411,1987,2011]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="270">ZRC 56488-1</specimenCode>
(
<quantity box="[424,515,1987,2011]" metricMagnitude="-1" metricUnit="m" metricValue="4.01" pageId="2" pageNumber="270" unit="mm" value="401.0">401 mm</quantity>
, female), 
<specimenCode collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="270">ZRC 56488- 2</specimenCode>
(333+ mm*, male; * for tail tip damaged but hypural plate intact); 
<collectingCountry box="[258,443,2051,2075]" name="Christmas Island" pageId="2" pageNumber="270">Christmas Island</collectingCountry>
, 
<collectingCountry box="[456,558,2051,2075]" name="Australia" pageId="2" pageNumber="270">Australia</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:039A87C10D629B0AFF12F9C7FB2EF9A1:8EEC600C0D629B0DFDE3F787FD1AF7C8" box="[570,752,2051,2075]" country="Christmas Island" county="Pre-anus" municipality="Vertebral" name="Freshwater Cave" pageId="2" pageNumber="270">Freshwater Cave</location>
(10°30.796ʹS 105°37.448ʹE), a cave with a large but steep 
<emphasis bold="true" box="[813,952,861,885]" pageId="2" pageNumber="270">Description.</emphasis>
Data is presented for 
<typeStatus box="[1192,1285,861,885]" pageId="2" pageNumber="270">holotype</typeStatus>
first, 
<typeStatus box="[1349,1451,862,886]" pageId="2" pageNumber="270">paratypes</typeStatus>
and 
<typeStatus box="[864,1092,893,918]" isOtherTypeStatus="true" pageId="2" pageNumber="270">non-type specimens</typeStatus>
in parentheses. 
<location LSID="urn:lsid:plazi:treatment:039A87C10D629B0AFF12F9C7FB2EF9A1:8EEC600C0D629B0DFADEFCF9FA61FC46" box="[1287,1419,893,917]" country="Christmas Island" county="Pre-anus" municipality="Vertebral" name="Proportions" pageId="2" pageNumber="270">Proportions</location>
in percentage of TL: tail length 59.0 (56.3–61.1), trunk length 31.0 (29.7–33.8), head length 10.0 (8.9–9.9), depth at gill opening 3.5 (3.2–5.2), depth at anus 3.2 (2.6–4.6). 
<location LSID="urn:lsid:plazi:treatment:039A87C10D629B0AFF12F9C7FB2EF9A1:8EEC600C0D629B0DFAEBFC59FA46FC26" box="[1330,1452,989,1013]" country="Christmas Island" county="Pre-anus" municipality="Vertebral" name="Proportions" pageId="2" pageNumber="270">Proportions</location>
in percentage of HL: length of upper jaw 28.7 (26.8–33.1), length of lower jaw 28.3 (25.5–32.6), interorbital width 11.1 (12.1–14.4), snout length 13.1 (14.0–17.0), eye diameter 3.9 (3.0–4.6). 
<collectingMunicipality box="[924,1025,1117,1141]" pageId="2" pageNumber="270">Vertebral</collectingMunicipality>
counts: 
<collectingCounty box="[1117,1210,1118,1141]" pageId="2" pageNumber="270">Pre-anus</collectingCounty>
vertebrae 58 (56–62), pre-dorsal fin vertebrae 137 (130–138), pre-anal fin vertebrae 138 (131–139), total vertebrae 146 (141–149)
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[811,1453,1245,1686]" pageId="2" pageNumber="270">
<materialsCitation collectionCode="WAM" pageId="2" pageNumber="270" specimenCode="NMMB-P37200" specimenCount="1" typeStatus="paratype">
A small-sized slender moray eel, anus well anterior to midpoint of body, tail long, body laterally compressed with depth nearly consistent throughout whole fish, tail tip blunt, caudal fin short (
<figureCitation box="[990,1055,1341,1365]" captionStart="Fig" captionStartId="3.[136,171,1949,1971]" captionTargetBox="[136,1474,302,1933]" captionTargetId="figure-8@3.[136,1474,302,1933]" captionTargetPageId="3" captionText="Fig. 3. Uropterygius cyamommatus, new species, holotype, WAM P.35403.001, 361 mm TL. A, fresh colouration (photographed by HHT); B, preserved colouration (photographed by WCH). Arrows indicate the position of anus." figureDoi="http://doi.org/10.5281/zenodo.7816101" httpUri="https://zenodo.org/record/7816101/files/figure.png" pageId="2" pageNumber="270">Fig. 3</figureCitation>
). Dorsal and anal fins inconspicuous, restricted to the posterior end of tail. Gill opening small, oval, below lateral midline of body, about same size with eye. Eyes very small, above mid-jaw (
<figureCitation box="[1218,1284,1437,1461]" captionStart="Fig" captionStartId="5.[138,173,811,833]" captionTargetBox="[136,1475,189,795]" captionTargetId="figure-568@5.[136,1475,189,795]" captionTargetPageId="5" captionText="Fig. 4. Uropterygius cyamommatus, new species, lateral view of head marked with cephalic sensory pores: red for supraorbital pores; green for infraorbital pores; blue for preoperculo-mandibular pores; yellow for the branchial pore. Arrow indicates the gill opening. Photo of WAM P.35403.001, holotype, 361 mm TL (photographed by HHT)." figureDoi="http://doi.org/10.5281/zenodo.7816103" httpUri="https://zenodo.org/record/7816103/files/figure.png" pageId="2" pageNumber="270">Fig. 4</figureCitation>
), 
<specimenCount box="[1303,1451,1438,1462]" pageId="2" pageNumber="270" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(
<specimenCode box="[819,993,1469,1493]" collectionCode="NMMB-P" pageId="2" pageNumber="270">NMMB-P37200</specimenCode>
and 
<collectionCode box="[1041,1109,1470,1494]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="270" type="Museum">WAM</collectionCode>
P.35404.001) with a reduced left eye embedded in skin (
<figureCitation box="[1068,1157,1501,1525]" captionStart="Fig" captionStartId="6.[813,848,950,972]" captionTargetBox="[813,1452,189,935]" captionTargetId="figure-486@6.[813,1452,189,935]" captionTargetPageId="6" captionText="Fig.5. Uropterygius cyamommatus, new species, lateral view of the head region of a specimen without the left eye. A, external view; B, radiograph showing the skeletal structure of orbital region. Photo of NMMB-P37200, paratype, 380 mm TL (photographed by HHT)." figureDoi="http://doi.org/10.5281/zenodo.7816105" httpUri="https://zenodo.org/record/7816105/files/figure.png" pageId="2" pageNumber="270">Fig. 5A</figureCitation>
). Snout blunt, nasal cavity somewhat enlarged. Jaws short, subequal in length, teeth not visible when mouth closed. Anterior nostril a stubby tube, shorter than eye diameter, its base in a depression close to tip of snout. Posterior nostril an oval hole located above anterior margin of eye, without a conspicuous raised rim
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[812,1453,1725,2101]" lastBlockId="4.[113,653,1853,1904]" lastPageId="4" lastPageNumber="272" pageId="2" pageNumber="270">
Three supraorbital pores, first on tip of snout immediately above lip; second just next to upper base of anterior nostril, at the level of horizontal middle line of eye; third on upper margin of snout at the same level of posterior nostril, above and posterior to the first infraorbital pore. Four infraorbital pores arranging equidistantly, first just below base of anterior nostril; second on the midpoint between anterior nostril base and anterior margin of eye; third below middle point of eye; fourth below and posterior to eye. Second supraorbital and first infraorbital pores in depression of anterior nostril base. Typically six preoperculo-mandibular pores but with a variation from four to seven pores, all along lower jaw, 
<superScript attach="right" box="[113,164,1853,1875]" fontSize="5" pageId="4" pageNumber="272">aData</superScript>
not including 
<collectionCode box="[309,356,1854,1875]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="272">ZRC</collectionCode>
56488-2 due to damaged tail.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816101" ID-Zenodo-Dep="7816101" httpUri="https://zenodo.org/record/7816101/files/figure.png" pageId="3" pageNumber="271" startId="3.[136,171,1949,1971]" targetBox="[136,1474,302,1933]" targetPageId="3">
<paragraph blockId="3.[136,1474,1949,2000]" pageId="3" pageNumber="271">
Fig. 3. 
<taxonomicName authority="Huang &amp; Liao &amp; Tan, 2023" authorityName="Huang &amp; Liao &amp; Tan" authorityYear="2023" box="[204,475,1949,1971]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="3" pageNumber="271" phylum="Chordata" rank="species" species="cyamommatus" status="new species">
<emphasis box="[204,475,1949,1971]" italics="true" pageId="3" pageNumber="271">Uropterygius cyamommatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[487,605,1949,1971]" pageId="3" pageNumber="271" rank="species">new species</taxonomicNameLabel>
, holotype, WAM P.35403.001, 361 mm TL. A, fresh colouration (photographed by HHT); B, preserved colouration (photographed by WCH). Arrows indicate the position of anus.
</paragraph>
</caption>
<caption pageId="4" pageNumber="272" startId="4.[113,168,189,211]">
<paragraph blockId="4.[113,1451,189,240]" pageId="4" pageNumber="272">
Table 1. Morphometric measurements and meristic counts of 
<taxonomicName authority="Huang &amp; Liao &amp; Tan, 2023" authorityName="Huang &amp; Liao &amp; Tan" authorityYear="2023" box="[702,973,190,211]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="272" phylum="Chordata" rank="species" species="cyamommatus" status="new species">
<emphasis box="[702,973,190,211]" italics="true" pageId="4" pageNumber="272">Uropterygius cyamommatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[985,1103,189,211]" pageId="4" pageNumber="272" rank="species">new species</taxonomicNameLabel>
. The means of morphometric values and vertebral number are given in parentheses.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="272">
<table box="[128,1422,289,1819]" gridcols="5" gridrows="33" pageId="4" pageNumber="272">
<tr box="[128,1422,289,311]" gridrow="0" pageId="4" pageNumber="272" rowspan-0="1">
<th box="[558,937,289,311]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="4" pageNumber="272">
<emphasis bold="true" box="[636,815,289,311]" pageId="4" pageNumber="272">Christmas Island</emphasis>
</th>
<th box="[1004,1173,289,311]" gridcol="3" gridrow="0" pageId="4" pageNumber="272">
<emphasis bold="true" box="[1011,1166,290,311]" pageId="4" pageNumber="272">Panglao Island</emphasis>
</th>
<th box="[1239,1422,289,311]" gridcol="4" gridrow="0" pageId="4" pageNumber="272">
<emphasis bold="true" box="[1303,1358,290,311]" pageId="4" pageNumber="272">Total</emphasis>
</th>
</tr>
<tr box="[128,1422,336,358]" gridrow="1" pageId="4" pageNumber="272" rowspan-0="1">
<td box="[558,651,336,358]" gridcol="1" gridrow="1" pageId="4" pageNumber="272">
<emphasis bold="true" box="[558,651,336,357]" pageId="4" pageNumber="272">Holotype</emphasis>
</td>
<td box="[756,937,336,358]" gridcol="2" gridrow="1" pageId="4" pageNumber="272">
<emphasis bold="true" box="[756,937,336,358]" pageId="4" pageNumber="272">Paratypes (n = 4)</emphasis>
</td>
<td box="[1004,1173,336,358]" gridcol="3" gridrow="1" pageId="4" pageNumber="272">
<emphasis bold="true" box="[1053,1123,336,358]" pageId="4" pageNumber="272">(n = 4)</emphasis>
</td>
<td box="[1239,1422,336,358]" gridcol="4" gridrow="1" pageId="4" pageNumber="272">
<emphasis bold="true" box="[1295,1365,336,358]" pageId="4" pageNumber="272">(n = 9)</emphasis>
</td>
</tr>
<tr box="[128,1422,390,412]" gridrow="2" pageId="4" pageNumber="272">
<th box="[128,414,390,412]" gridcol="0" gridrow="2" pageId="4" pageNumber="272">TL (mm)</th>
<td box="[558,651,390,412]" gridcol="1" gridrow="2" pageId="4" pageNumber="272">361</td>
<td box="[756,937,390,412]" gridcol="2" gridrow="2" pageId="4" pageNumber="272">333–401</td>
<td box="[1004,1173,390,412]" gridcol="3" gridrow="2" pageId="4" pageNumber="272">291–352</td>
<td box="[1239,1422,390,412]" gridcol="4" gridrow="2" pageId="4" pageNumber="272">291–401</td>
</tr>
<tr box="[128,1422,438,459]" gridrow="3" pageId="4" pageNumber="272">
<th box="[128,1422,438,459]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="4" pageNumber="272">% TL</th>
</tr>
<tr box="[128,1422,484,506]" gridrow="4" pageId="4" pageNumber="272">
<th box="[128,414,484,506]" gridcol="0" gridrow="4" pageId="4" pageNumber="272">Tail length</th>
<td box="[558,651,484,506]" gridcol="1" gridrow="4" pageId="4" pageNumber="272">59.0</td>
<td box="[756,937,484,506]" gridcol="2" gridrow="4" pageId="4" pageNumber="272">57.7–61.1 (59.5)a</td>
<td box="[1004,1173,484,506]" gridcol="3" gridrow="4" pageId="4" pageNumber="272">56.3–57.7 (56.9)b</td>
<td box="[1239,1422,484,506]" gridcol="4" gridrow="4" pageId="4" pageNumber="272">56.3–61.1 (58.3)a, b</td>
</tr>
<tr box="[128,1422,530,553]" gridrow="5" pageId="4" pageNumber="272">
<th box="[128,414,530,553]" gridcol="0" gridrow="5" pageId="4" pageNumber="272">Preanal length</th>
<td box="[558,651,530,553]" gridcol="1" gridrow="5" pageId="4" pageNumber="272">41.0</td>
<td box="[756,937,530,553]" gridcol="2" gridrow="5" pageId="4" pageNumber="272">38.9–42.3 (40.5)a</td>
<td box="[1004,1173,530,553]" gridcol="3" gridrow="5" pageId="4" pageNumber="272">42.3–43.7 (43.1)b</td>
<td box="[1239,1422,530,553]" gridcol="4" gridrow="5" pageId="4" pageNumber="272">38.9–43.7 (41.7)a, b</td>
</tr>
<tr box="[128,1422,577,600]" gridrow="6" pageId="4" pageNumber="272">
<th box="[128,414,577,600]" gridcol="0" gridrow="6" pageId="4" pageNumber="272">Trunk length</th>
<td box="[558,651,577,600]" gridcol="1" gridrow="6" pageId="4" pageNumber="272">31.0</td>
<td box="[756,937,577,600]" gridcol="2" gridrow="6" pageId="4" pageNumber="272">29.7–33.3 (31.2)a</td>
<td box="[1004,1173,577,600]" gridcol="3" gridrow="6" pageId="4" pageNumber="272">33.3–33.8 (33.6)b</td>
<td box="[1239,1422,577,600]" gridcol="4" gridrow="6" pageId="4" pageNumber="272">29.7–33.8 (32.2)a, b</td>
</tr>
<tr box="[128,1422,624,647]" gridrow="7" pageId="4" pageNumber="272">
<th box="[128,414,624,647]" gridcol="0" gridrow="7" pageId="4" pageNumber="272">Head length</th>
<td box="[558,651,624,647]" gridcol="1" gridrow="7" pageId="4" pageNumber="272">10.0</td>
<td box="[756,937,624,647]" gridcol="2" gridrow="7" pageId="4" pageNumber="272">9.0–9.7 (9.3)a</td>
<td box="[1004,1173,624,647]" gridcol="3" gridrow="7" pageId="4" pageNumber="272">8.9–9.9 (9.5)b</td>
<td box="[1239,1422,624,647]" gridcol="4" gridrow="7" pageId="4" pageNumber="272">8.9–10.0 (9.5)a, b</td>
</tr>
<tr box="[128,1422,671,694]" gridrow="8" pageId="4" pageNumber="272">
<th box="[128,414,671,694]" gridcol="0" gridrow="8" pageId="4" pageNumber="272">Body depth at gill opening</th>
<td box="[558,651,671,694]" gridcol="1" gridrow="8" pageId="4" pageNumber="272">3.5</td>
<td box="[756,937,671,694]" gridcol="2" gridrow="8" pageId="4" pageNumber="272">3.2–5.2 (4.1)a</td>
<td box="[1004,1173,671,694]" gridcol="3" gridrow="8" pageId="4" pageNumber="272">3.2–4.2 (3.6)b</td>
<td box="[1239,1422,671,694]" gridcol="4" gridrow="8" pageId="4" pageNumber="272">3.2–5.2 (3.8)a, b</td>
</tr>
<tr box="[128,1422,718,741]" gridrow="9" pageId="4" pageNumber="272">
<th box="[128,414,718,741]" gridcol="0" gridrow="9" pageId="4" pageNumber="272">Body depth at anus</th>
<td box="[558,651,718,741]" gridcol="1" gridrow="9" pageId="4" pageNumber="272">3.2</td>
<td box="[756,937,718,741]" gridcol="2" gridrow="9" pageId="4" pageNumber="272">2.6–4.6 (3.5)a</td>
<td box="[1004,1173,718,741]" gridcol="3" gridrow="9" pageId="4" pageNumber="272">3.5–3.8 (3.6)b</td>
<td box="[1239,1422,718,741]" gridcol="4" gridrow="9" pageId="4" pageNumber="272">2.6–4.6 (3.5)a, b</td>
</tr>
<tr box="[128,1422,766,787]" gridrow="10" pageId="4" pageNumber="272">
<th box="[128,1422,766,787]" colspan="5" colspanRight="4" gridcol="0" gridrow="10" pageId="4" pageNumber="272">% HL</th>
</tr>
<tr box="[128,1422,812,834]" gridrow="11" pageId="4" pageNumber="272">
<th box="[128,414,812,834]" gridcol="0" gridrow="11" pageId="4" pageNumber="272">Length of upper jaw</th>
<td box="[558,651,812,834]" gridcol="1" gridrow="11" pageId="4" pageNumber="272">28.7</td>
<td box="[756,937,812,834]" gridcol="2" gridrow="11" pageId="4" pageNumber="272">30.3–33.1 (32.2)</td>
<td box="[1004,1173,812,834]" gridcol="3" gridrow="11" pageId="4" pageNumber="272">26.8–30.1 (28.4)</td>
<td box="[1239,1422,812,834]" gridcol="4" gridrow="11" pageId="4" pageNumber="272">26.8–33.1 (30.1)</td>
</tr>
<tr box="[128,1422,859,881]" gridrow="12" pageId="4" pageNumber="272">
<th box="[128,414,859,881]" gridcol="0" gridrow="12" pageId="4" pageNumber="272">Length of lower jaw</th>
<td box="[558,651,859,881]" gridcol="1" gridrow="12" pageId="4" pageNumber="272">28.3</td>
<td box="[756,937,859,881]" gridcol="2" gridrow="12" pageId="4" pageNumber="272">29.9–32.6 (31.5)</td>
<td box="[1004,1173,859,881]" gridcol="3" gridrow="12" pageId="4" pageNumber="272">25.5–29.1 (27.3)</td>
<td box="[1239,1422,859,881]" gridcol="4" gridrow="12" pageId="4" pageNumber="272">25.5–32.6 (29.3)</td>
</tr>
<tr box="[128,1422,906,928]" gridrow="13" pageId="4" pageNumber="272">
<th box="[128,414,906,928]" gridcol="0" gridrow="13" pageId="4" pageNumber="272">Snout length</th>
<td box="[558,651,906,928]" gridcol="1" gridrow="13" pageId="4" pageNumber="272">13.1</td>
<td box="[756,937,906,928]" gridcol="2" gridrow="13" pageId="4" pageNumber="272">15.3–17.0 (16.0)</td>
<td box="[1004,1173,906,928]" gridcol="3" gridrow="13" pageId="4" pageNumber="272">14.0–15.2 (14.4)</td>
<td box="[1239,1422,906,928]" gridcol="4" gridrow="13" pageId="4" pageNumber="272">13.1–17.0 (15.0)</td>
</tr>
<tr box="[128,1422,953,975]" gridrow="14" pageId="4" pageNumber="272">
<th box="[128,414,953,975]" gridcol="0" gridrow="14" pageId="4" pageNumber="272">Interorbital width</th>
<td box="[558,651,953,975]" gridcol="1" gridrow="14" pageId="4" pageNumber="272">11.1</td>
<td box="[756,937,953,975]" gridcol="2" gridrow="14" pageId="4" pageNumber="272">12.1–14.4 (13.4)</td>
<td box="[1004,1173,953,975]" gridcol="3" gridrow="14" pageId="4" pageNumber="272">12.1–12.5 (12.2)</td>
<td box="[1239,1422,953,975]" gridcol="4" gridrow="14" pageId="4" pageNumber="272">11.1–14.4 (12.6)</td>
</tr>
<tr box="[128,1422,1000,1022]" gridrow="15" pageId="4" pageNumber="272">
<th box="[128,414,1000,1022]" gridcol="0" gridrow="15" pageId="4" pageNumber="272">Eye diameter</th>
<td box="[558,651,1000,1022]" gridcol="1" gridrow="15" pageId="4" pageNumber="272">3.9</td>
<td box="[756,937,1000,1022]" gridcol="2" gridrow="15" pageId="4" pageNumber="272">3.0–4.6 (3.9)</td>
<td box="[1004,1173,1000,1022]" gridcol="3" gridrow="15" pageId="4" pageNumber="272">4.0–4.3 (4.1)</td>
<td box="[1239,1422,1000,1022]" gridcol="4" gridrow="15" pageId="4" pageNumber="272">3.0–4.6 (4.0)</td>
</tr>
<tr box="[128,1422,1047,1069]" gridrow="16" pageId="4" pageNumber="272">
<th box="[128,1422,1047,1069]" colspan="5" colspanRight="4" gridcol="0" gridrow="16" pageId="4" pageNumber="272">Teeth</th>
</tr>
<tr box="[128,1422,1094,1116]" gridrow="17" pageId="4" pageNumber="272">
<th box="[128,414,1094,1116]" gridcol="0" gridrow="17" pageId="4" pageNumber="272">Intermaxillary + Vomerine</th>
<td box="[558,651,1094,1116]" gridcol="1" gridrow="17" pageId="4" pageNumber="272">38</td>
<td box="[756,937,1094,1116]" gridcol="2" gridrow="17" pageId="4" pageNumber="272">39–44</td>
<td box="[1004,1173,1094,1116]" gridcol="3" gridrow="17" pageId="4" pageNumber="272">30–45</td>
<td box="[1239,1422,1094,1116]" gridcol="4" gridrow="17" pageId="4" pageNumber="272">30–45</td>
</tr>
<tr box="[128,1422,1140,1163]" gridrow="18" pageId="4" pageNumber="272">
<th box="[128,414,1140,1163]" gridcol="0" gridrow="18" pageId="4" pageNumber="272">Maxillary outer</th>
<td box="[558,651,1140,1163]" gridcol="1" gridrow="18" pageId="4" pageNumber="272">20–22</td>
<td box="[756,937,1140,1163]" gridcol="2" gridrow="18" pageId="4" pageNumber="272">22–26</td>
<td box="[1004,1173,1140,1163]" gridcol="3" gridrow="18" pageId="4" pageNumber="272">20–24</td>
<td box="[1239,1422,1140,1163]" gridcol="4" gridrow="18" pageId="4" pageNumber="272">20–26</td>
</tr>
<tr box="[128,1422,1187,1209]" gridrow="19" pageId="4" pageNumber="272">
<th box="[128,414,1187,1209]" gridcol="0" gridrow="19" pageId="4" pageNumber="272">Maxillary inner</th>
<td box="[558,651,1187,1209]" gridcol="1" gridrow="19" pageId="4" pageNumber="272">11–12</td>
<td box="[756,937,1187,1209]" gridcol="2" gridrow="19" pageId="4" pageNumber="272">9–13</td>
<td box="[1004,1173,1187,1209]" gridcol="3" gridrow="19" pageId="4" pageNumber="272">8–13</td>
<td box="[1239,1422,1187,1209]" gridcol="4" gridrow="19" pageId="4" pageNumber="272">8–13</td>
</tr>
<tr box="[128,1422,1234,1256]" gridrow="20" pageId="4" pageNumber="272">
<th box="[128,414,1234,1256]" gridcol="0" gridrow="20" pageId="4" pageNumber="272">Dentary outer</th>
<td box="[558,651,1234,1256]" gridcol="1" gridrow="20" pageId="4" pageNumber="272">29–30</td>
<td box="[756,937,1234,1256]" gridcol="2" gridrow="20" pageId="4" pageNumber="272">29–32</td>
<td box="[1004,1173,1234,1256]" gridcol="3" gridrow="20" pageId="4" pageNumber="272">22–29</td>
<td box="[1239,1422,1234,1256]" gridcol="4" gridrow="20" pageId="4" pageNumber="272">22–32</td>
</tr>
<tr box="[128,1422,1281,1303]" gridrow="21" pageId="4" pageNumber="272">
<th box="[128,414,1281,1303]" gridcol="0" gridrow="21" pageId="4" pageNumber="272">Dentary middle</th>
<td box="[558,651,1281,1303]" gridcol="1" gridrow="21" pageId="4" pageNumber="272">2–5</td>
<td box="[756,937,1281,1303]" gridcol="2" gridrow="21" pageId="4" pageNumber="272">0–6</td>
<td box="[1004,1173,1281,1303]" gridcol="3" gridrow="21" pageId="4" pageNumber="272">0–7</td>
<td box="[1239,1422,1281,1303]" gridcol="4" gridrow="21" pageId="4" pageNumber="272">0–7</td>
</tr>
<tr box="[128,1422,1328,1350]" gridrow="22" pageId="4" pageNumber="272">
<th box="[128,414,1328,1350]" gridcol="0" gridrow="22" pageId="4" pageNumber="272">Dentary inner</th>
<td box="[558,651,1328,1350]" gridcol="1" gridrow="22" pageId="4" pageNumber="272">12</td>
<td box="[756,937,1328,1350]" gridcol="2" gridrow="22" pageId="4" pageNumber="272">9–19</td>
<td box="[1004,1173,1328,1350]" gridcol="3" gridrow="22" pageId="4" pageNumber="272">10–16</td>
<td box="[1239,1422,1328,1350]" gridcol="4" gridrow="22" pageId="4" pageNumber="272">9–19</td>
</tr>
<tr box="[128,1422,1375,1397]" gridrow="23" pageId="4" pageNumber="272">
<th box="[128,1422,1375,1397]" colspan="5" colspanRight="4" gridcol="0" gridrow="23" pageId="4" pageNumber="272">Cephalic sensory pores</th>
</tr>
<tr box="[128,1422,1422,1444]" gridrow="24" pageId="4" pageNumber="272">
<th box="[128,414,1422,1444]" gridcol="0" gridrow="24" pageId="4" pageNumber="272">Supraorbital</th>
<td box="[558,651,1422,1444]" gridcol="1" gridrow="24" pageId="4" pageNumber="272">3</td>
<td box="[756,937,1422,1444]" gridcol="2" gridrow="24" pageId="4" pageNumber="272">3</td>
<td box="[1004,1173,1422,1444]" gridcol="3" gridrow="24" pageId="4" pageNumber="272">3</td>
<td box="[1239,1422,1422,1444]" gridcol="4" gridrow="24" pageId="4" pageNumber="272">3</td>
</tr>
<tr box="[128,1422,1469,1491]" gridrow="25" pageId="4" pageNumber="272">
<th box="[128,414,1469,1491]" gridcol="0" gridrow="25" pageId="4" pageNumber="272">Infraorbital</th>
<td box="[558,651,1469,1491]" gridcol="1" gridrow="25" pageId="4" pageNumber="272">4</td>
<td box="[756,937,1469,1491]" gridcol="2" gridrow="25" pageId="4" pageNumber="272">4</td>
<td box="[1004,1173,1469,1491]" gridcol="3" gridrow="25" pageId="4" pageNumber="272">4</td>
<td box="[1239,1422,1469,1491]" gridcol="4" gridrow="25" pageId="4" pageNumber="272">4</td>
</tr>
<tr box="[128,1422,1515,1538]" gridrow="26" pageId="4" pageNumber="272">
<th box="[128,414,1515,1538]" gridcol="0" gridrow="26" pageId="4" pageNumber="272">Preoperculo-mandibular</th>
<td box="[558,651,1515,1538]" gridcol="1" gridrow="26" pageId="4" pageNumber="272">6</td>
<td box="[756,937,1515,1538]" gridcol="2" gridrow="26" pageId="4" pageNumber="272">5–7</td>
<td box="[1004,1173,1515,1538]" gridcol="3" gridrow="26" pageId="4" pageNumber="272">4–7</td>
<td box="[1239,1422,1515,1538]" gridcol="4" gridrow="26" pageId="4" pageNumber="272">4–7</td>
</tr>
<tr box="[128,1422,1562,1584]" gridrow="27" pageId="4" pageNumber="272">
<th box="[128,414,1562,1584]" gridcol="0" gridrow="27" pageId="4" pageNumber="272">Branchial</th>
<td box="[558,651,1562,1584]" gridcol="1" gridrow="27" pageId="4" pageNumber="272">1</td>
<td box="[756,937,1562,1584]" gridcol="2" gridrow="27" pageId="4" pageNumber="272">1</td>
<td box="[1004,1173,1562,1584]" gridcol="3" gridrow="27" pageId="4" pageNumber="272">1</td>
<td box="[1239,1422,1562,1584]" gridcol="4" gridrow="27" pageId="4" pageNumber="272">1</td>
</tr>
<tr box="[128,1422,1609,1631]" gridrow="28" pageId="4" pageNumber="272">
<th box="[128,1422,1609,1631]" colspan="5" colspanRight="4" gridcol="0" gridrow="28" pageId="4" pageNumber="272">Vertebrae</th>
</tr>
<tr box="[128,1422,1656,1678]" gridrow="29" pageId="4" pageNumber="272">
<th box="[128,414,1656,1678]" gridcol="0" gridrow="29" pageId="4" pageNumber="272">Pre-anus</th>
<td box="[558,651,1656,1678]" gridcol="1" gridrow="29" pageId="4" pageNumber="272">58</td>
<td box="[756,937,1656,1678]" gridcol="2" gridrow="29" pageId="4" pageNumber="272">56–59 (57)</td>
<td box="[1004,1173,1656,1678]" gridcol="3" gridrow="29" pageId="4" pageNumber="272">59–62 (60)</td>
<td box="[1239,1422,1656,1678]" gridcol="4" gridrow="29" pageId="4" pageNumber="272">56–62 (59)</td>
</tr>
<tr box="[128,1422,1703,1725]" gridrow="30" pageId="4" pageNumber="272">
<th box="[128,414,1703,1725]" gridcol="0" gridrow="30" pageId="4" pageNumber="272">Pre-dorsal fin</th>
<td box="[558,651,1703,1725]" gridcol="1" gridrow="30" pageId="4" pageNumber="272">137</td>
<td box="[756,937,1703,1725]" gridcol="2" gridrow="30" pageId="4" pageNumber="272">132–137 (133)</td>
<td box="[1004,1173,1703,1725]" gridcol="3" gridrow="30" pageId="4" pageNumber="272">130–138 (134)</td>
<td box="[1239,1422,1703,1725]" gridcol="4" gridrow="30" pageId="4" pageNumber="272">130–138 (134)</td>
</tr>
<tr box="[128,1422,1750,1772]" gridrow="31" pageId="4" pageNumber="272">
<th box="[128,414,1750,1772]" gridcol="0" gridrow="31" pageId="4" pageNumber="272">Pre-anal fin</th>
<td box="[558,651,1750,1772]" gridcol="1" gridrow="31" pageId="4" pageNumber="272">138</td>
<td box="[756,937,1750,1772]" gridcol="2" gridrow="31" pageId="4" pageNumber="272">133–139 (135)</td>
<td box="[1004,1173,1750,1772]" gridcol="3" gridrow="31" pageId="4" pageNumber="272">131–139 (135)</td>
<td box="[1239,1422,1750,1772]" gridcol="4" gridrow="31" pageId="4" pageNumber="272">131–139 (135)</td>
</tr>
<tr box="[128,1422,1797,1819]" gridrow="32" pageId="4" pageNumber="272">
<th box="[128,414,1797,1819]" gridcol="0" gridrow="32" pageId="4" pageNumber="272">Total</th>
<td box="[558,651,1797,1819]" gridcol="1" gridrow="32" pageId="4" pageNumber="272">146</td>
<td box="[756,937,1797,1819]" gridcol="2" gridrow="32" pageId="4" pageNumber="272">144–146 (145)</td>
<td box="[1004,1173,1797,1819]" gridcol="3" gridrow="32" pageId="4" pageNumber="272">141–149 (145)</td>
<td box="[1239,1422,1797,1819]" gridcol="4" gridrow="32" pageId="4" pageNumber="272">141–149 (145)</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[113,653,1853,1904]" box="[113,633,1881,1904]" pageId="4" pageNumber="272">
<superScript attach="right" box="[113,165,1881,1904]" fontSize="5" pageId="4" pageNumber="272">bData</superScript>
not including 
<collectionCode box="[310,357,1882,1903]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="272">ZRC</collectionCode>
63039 due to damaged tail.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816103" ID-Zenodo-Dep="7816103" httpUri="https://zenodo.org/record/7816103/files/figure.png" pageId="5" pageNumber="273" startId="5.[138,173,811,833]" targetBox="[136,1475,189,795]" targetPageId="5">
<paragraph blockId="5.[136,1475,811,890]" pageId="5" pageNumber="273">
Fig. 4. 
<taxonomicName authority="Huang &amp; Liao &amp; Tan, 2023" authorityName="Huang &amp; Liao &amp; Tan" authorityYear="2023" box="[210,486,812,833]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="cyamommatus" status="new species">
<emphasis box="[210,486,812,833]" italics="true" pageId="5" pageNumber="273">Uropterygius cyamommatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[500,621,811,833]" pageId="5" pageNumber="273" rank="species">new species</taxonomicNameLabel>
, lateral view of head marked with cephalic sensory pores: red for supraorbital pores; green for infraorbital pores; blue for preoperculo-mandibular pores; yellow for the branchial pore. Arrow indicates the gill opening. Photo of WAM P.35403.001, holotype, 361 mm TL (photographed by HHT).
</paragraph>
</caption>
<paragraph blockId="5.[136,775,954,1043]" pageId="5" pageNumber="273">
last (generally sixth) pore posterior to corner of mouth. One branchial pore on posterior-dorsal area of head, closer to corner of mouth rather than gill opening (
<figureCitation box="[593,661,1018,1043]" captionStart="Fig" captionStartId="5.[138,173,811,833]" captionTargetBox="[136,1475,189,795]" captionTargetId="figure-568@5.[136,1475,189,795]" captionTargetPageId="5" captionText="Fig. 4. Uropterygius cyamommatus, new species, lateral view of head marked with cephalic sensory pores: red for supraorbital pores; green for infraorbital pores; blue for preoperculo-mandibular pores; yellow for the branchial pore. Arrow indicates the gill opening. Photo of WAM P.35403.001, holotype, 361 mm TL (photographed by HHT)." figureDoi="http://doi.org/10.5281/zenodo.7816103" httpUri="https://zenodo.org/record/7816103/files/figure.png" pageId="5" pageNumber="273">Fig. 4</figureCitation>
).
</paragraph>
<paragraph blockId="5.[135,775,1082,1714]" pageId="5" pageNumber="273">
<materialsCitation collectionCode="ZRC" pageId="5" pageNumber="273" specimenCode="NMMB-P37200, ZRC 46849-2, ZRC 56488-1" specimenCount="3" typeStatus="paratype">
Teeth (
<figureCitation box="[214,280,1082,1106]" captionStart="Fig" captionStartId="7.[136,171,1140,1162]" captionTargetBox="[143,1469,189,1123]" captionTargetId="figure-8@7.[75,1511,169,1127]" captionTargetPageId="7" captionText="Fig. 6. Dentition of Uropterygius cyamommatus, new species, holotype, WAM P.35403.001. Upper jaw (left) and lower jaw (right). Dotted circles represent the sockets of missing teeth. Schematic drawings by WCH." figureDoi="http://doi.org/10.5281/zenodo.7816107" httpUri="https://zenodo.org/record/7816107/files/figure.png" pageId="5" pageNumber="273">Fig. 6</figureCitation>
) conical, slightly elongated, end pointed and slightly retrorse, edge smooth. Intermaxillary, maxillary, and vomerine teeth continuously; peripheral series small, numerous, and equal-sized; inner series longer, fewer, and spaced. Intermaxillary and vomerine plates with 30–45 teeth, approximately arranged in 7–8 rows on intermaxilla and becoming single row on vomerine, except for 
<specimenCount box="[637,775,1275,1299]" pageId="5" pageNumber="273" type="generic" typeStatus="paratype">one paratype</specimenCount>
(
<specimenCode box="[143,322,1307,1331]" collectionCode="NMMB-P" pageId="5" pageNumber="273">NMMB-P37200</specimenCode>
) without teeth on vomerine. Maxilla with 20–26 teeth on outer row; 8–13 teeth on inner row, extending to posterior end of outer row. Dentary teeth arranging in 2–3 rows anteriorly and 2 rows posteriorly; teeth on outer row small and equal-sized, 
<quantity box="[387,484,1434,1459]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="8.128" metricValueMin="5.588" pageId="5" pageNumber="273" unit="in" value="27.0" valueMax="32.0" valueMin="22.0">22–32 in</quantity>
number; inner teeth 9–19, larger and more spaced than those of outer, decreasing in size posteriorly, extending to or slightly before the posterior end of outer row; most examined specimens with a middle row of 2–7 teeth on anterior part of dentary, except for 
<specimenCount pageId="5" pageNumber="273" type="generic">two specimens</specimenCount>
(
<specimenCode box="[264,441,1595,1619]" collectionCode="NMMB-P" pageId="5" pageNumber="273">NMMB-P37200</specimenCode>
and 
<specimenCode box="[500,653,1594,1619]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="273">ZRC 46849-2</specimenCode>
) with only two rows of dentary teeth; 
<specimenCount box="[448,592,1627,1651]" pageId="5" pageNumber="273" type="generic" typeStatus="paratype">one paratype</specimenCount>
(
<specimenCode box="[610,767,1626,1651]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="273">ZRC 56488-1</specimenCode>
) with an additional one row of two very small teeth on the anteriormost middle part of lower jaw
</materialsCitation>
.
</paragraph>
<paragraph blockId="5.[135,775,1754,2003]" pageId="5" pageNumber="273">
Body uniform brown, slightly lighter ventrally, most pale on venter below lower jaw. Anterior nostril (including its base depression), posterior nostril opening, gill opening, oral cavity, and head pores whitish. Numerous small, inconspicuous whitish neuromasts arranged in irregular lines on anterior head region and in one row along lateral line (
<figureCitation box="[144,213,1946,1971]" captionStart="Fig" captionStartId="5.[138,173,811,833]" captionTargetBox="[136,1475,189,795]" captionTargetId="figure-568@5.[136,1475,189,795]" captionTargetPageId="5" captionText="Fig. 4. Uropterygius cyamommatus, new species, lateral view of head marked with cephalic sensory pores: red for supraorbital pores; green for infraorbital pores; blue for preoperculo-mandibular pores; yellow for the branchial pore. Arrow indicates the gill opening. Photo of WAM P.35403.001, holotype, 361 mm TL (photographed by HHT)." figureDoi="http://doi.org/10.5281/zenodo.7816103" httpUri="https://zenodo.org/record/7816103/files/figure.png" pageId="5" pageNumber="273">Fig. 4</figureCitation>
). Preserved colouration similar to fresh fish except faded (
<figureCitation box="[212,297,1978,2003]" captionStart="Fig" captionStartId="3.[136,171,1949,1971]" captionTargetBox="[136,1474,302,1933]" captionTargetId="figure-8@3.[136,1474,302,1933]" captionTargetPageId="3" captionText="Fig. 3. Uropterygius cyamommatus, new species, holotype, WAM P.35403.001, 361 mm TL. A, fresh colouration (photographed by HHT); B, preserved colouration (photographed by WCH). Arrows indicate the position of anus." figureDoi="http://doi.org/10.5281/zenodo.7816101" httpUri="https://zenodo.org/record/7816101/files/figure.png" pageId="5" pageNumber="273">Fig. 3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="273" type="distribution">
<paragraph blockId="5.[136,775,2042,2098]" pageId="5" pageNumber="273">
<emphasis bold="true" box="[136,284,2042,2066]" pageId="5" pageNumber="273">Distribution.</emphasis>
Currently, this species is only known from anchialine caves in 
<collectingCountry box="[352,533,2074,2098]" name="Christmas Island" pageId="5" pageNumber="273">Christmas Island</collectingCountry>
and Panglao Island.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="273" type="etymology">
<paragraph blockId="5.[835,1474,954,1011]" pageId="5" pageNumber="273">
<emphasis bold="true" box="[835,965,955,979]" pageId="5" pageNumber="273">Etymology.</emphasis>
From Greek words kúamos (a bean) and ómma (the eye), in reference to its tiny bean-shaped eyes.
</paragraph>
<paragraph blockId="5.[835,1475,1050,1331]" pageId="5" pageNumber="273">
<emphasis bold="true" box="[835,1005,1050,1074]" pageId="5" pageNumber="273">Comparisons.</emphasis>
The new species belongs to the genus 
<taxonomicName authorityName="Ruppell" authorityYear="1838" box="[835,977,1083,1106]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="genus">
<emphasis box="[835,977,1083,1106]" italics="true" pageId="5" pageNumber="273">Uropterygius</emphasis>
</taxonomicName>
according to a combination of the following characters: (1) three supraorbital pores (vs. four in 
<taxonomicName box="[1363,1472,1114,1138]" class="Actinopterygii" family="Muraenidae" genus="Anarchias" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="genus">
<emphasis box="[1363,1472,1114,1138]" italics="true" pageId="5" pageNumber="273">Anarchias</emphasis>
</taxonomicName>
<collectingCountry box="[835,906,1146,1170]" name="Jordan" pageId="5" pageNumber="273">Jordan</collectingCountry>
&amp; Starks, 1906); (2) the anus anterior to the midpoint of body (vs. anus far behind mid-body in 
<taxonomicName authorityName="Jordan &amp; Snyder" authorityYear="1901" box="[1364,1474,1179,1202]" class="Actinopterygii" family="Muraenidae" genus="Scuticaria" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="genus">
<emphasis box="[1364,1474,1179,1202]" italics="true" pageId="5" pageNumber="273">Scuticaria</emphasis>
</taxonomicName>
<bibRefCitation author="Jordan DS &amp; Snyder JO" box="[835,1103,1210,1235]" pageId="5" pageNumber="273" pagination="837 - 890" refId="ref5620" refString="Jordan DS &amp; Snyder JO (1901) A review of the apodal fishes or eels of Japan, with descriptions of nineteen new species. Proceedings of the United States National Museum, 23: 837 - 890." type="journal article" year="1901">
<collectingCountry box="[835,908,1210,1234]" name="Jordan" pageId="5" pageNumber="273">Jordan</collectingCountry>
&amp; Snyder, 1901
</bibRefCitation>
); (3) jaws short and subequal in length (vs. gape very long, lower jaw longer than upper jaw in 
<taxonomicName authority="Richardson, 1848" authorityName="Richardson" authorityYear="1848" box="[911,1286,1274,1299]" class="Actinopterygii" family="Muraenidae" genus="Channomuraena" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="genus">
<emphasis box="[911,1090,1274,1298]" italics="true" pageId="5" pageNumber="273">Channomuraena</emphasis>
<bibRefCitation author="Richardson J" box="[1097,1286,1274,1299]" pageId="5" pageNumber="273" pagination="1844 - 48" refId="ref6246" refString="Richardson J (1848) [1844 - 48] Ichthyology of the voyage of H. M. S. Erebus &amp; Terror. In: Richardson J &amp; Gray JE (eds.) The zoology of the voyage of H. M. S. Erebus &amp; Terror, under the command of Captain Sir James Clark Ross, R. N., F. R. S., during the years 1839 to 1843. Volume 2. Janson, London, 139 pp." type="book chapter" year="1848">Richardson, 1848</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Chen &amp; Shao, 1995" authorityName="Chen &amp; Shao" authorityYear="1995" class="Actinopterygii" family="Muraenidae" genus="Cirrimaxilla" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="genus">
<emphasis box="[1342,1475,1274,1298]" italics="true" pageId="5" pageNumber="273">Cirrimaxilla</emphasis>
<bibRefCitation author="Chen HM &amp; Shao KT" box="[835,1055,1306,1331]" pageId="5" pageNumber="273" pagination="328 - 332" refId="ref5063" refString="Chen HM &amp; Shao KT (1995) New eel genus, Cirrimaxilla, and description of the type species, Cirrimaxilla formosa (Pisces: Muraenidae) from southern Taiwan. Bulletin of Marine Science, 57 (2): 328 - 332." type="journal article" year="1995">Chen &amp; Shao, 1995</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph blockId="5.[834,1475,1370,1651]" pageId="5" pageNumber="273">
<taxonomicName authorityName="Huang &amp; Liao &amp; Tan" authorityYear="2023" box="[835,1132,1371,1394]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="cyamommatus">
<emphasis box="[835,1132,1371,1394]" italics="true" pageId="5" pageNumber="273">Uropterygius cyamommatus</emphasis>
</taxonomicName>
can be easily distinguished from the other five uniform brown-coloured species of the genus, viz. 
<taxonomicName authorityName="Ruppell" authorityYear="1838" box="[882,1010,1434,1458]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="concolor">
<emphasis box="[882,1010,1434,1458]" italics="true" pageId="5" pageNumber="273">U. concolor</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Randall &amp; Golani" authorityYear="1995" box="[1024,1116,1435,1458]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="genie">
<emphasis box="[1024,1116,1435,1458]" italics="true" pageId="5" pageNumber="273">U. genie</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="McCosker &amp; Smith" authorityYear="1997" box="[1129,1239,1434,1458]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="golanii">
<emphasis box="[1129,1239,1434,1458]" italics="true" pageId="5" pageNumber="273">U. golanii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Gosline" authorityYear="1958" box="[1252,1388,1435,1458]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="inornatus">
<emphasis box="[1252,1388,1435,1458]" italics="true" pageId="5" pageNumber="273">U. inornatus</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="McCosker &amp; Smith" authorityYear="1997" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="xenodontus">
<emphasis italics="true" pageId="5" pageNumber="273">U. xenodontus</emphasis>
</taxonomicName>
, by the combination of eye diameter, tail length, vertebrae count, and dentition (Table 2). In comparison, it has distinctively small eyes; its tail length is exclusively long except for 
<taxonomicName authorityName="Ruppell" authorityYear="1838" box="[1008,1137,1562,1586]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="concolor">
<emphasis box="[1008,1137,1562,1586]" italics="true" pageId="5" pageNumber="273">U. concolor</emphasis>
</taxonomicName>
(56.3–61.1, vs. 52.4–60.0% of TL); and its total vertebral number is only overlapped with 
<taxonomicName authorityName="McCosker &amp; Smith" authorityYear="1997" box="[837,947,1626,1650]" class="Actinopterygii" family="Muraenidae" genus="Uropterygius" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="273" phylum="Chordata" rank="species" species="golanii">
<emphasis box="[837,947,1626,1650]" italics="true" pageId="5" pageNumber="273">U. golanii</emphasis>
</taxonomicName>
(141–149, vs. 145–148).
</paragraph>
</subSubSection>
</treatment>
</document>