441 lines
57 KiB
XML
441 lines
57 KiB
XML
<document id="7A5156D3AC293313F5E5CEFA6A24DFAA" ID-DOI="10.11646/zootaxa.4103.5.4" ID-GBIF-Dataset="b9b4b63c-d1af-47a8-8e0a-3c63a6cbcc91" ID-ISSN="1175-5326" ID-Zenodo-Dep="263213" ID-ZooBank="D31571A3-8049-423F-AF40-37E4D5FA18D2" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460701761610" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="039A87B6C03EFFB2A7B100DC446FFE49" docLanguage="en" docName="zootaxa.4103.5.4.pdf.imf" docOrigin="Zootaxa 4103 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus" docType="treatment" docVersion="7" lastPageNumber="467" masterDocId="FFA3FFCEC03FFFB6A72603304003FF86" masterDocTitle="Taxonomy of ' Euconnus complex'. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepaloconnus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="472" masterPageNumber="463" pageNumber="464" updateTime="1698354660124" updateUser="plazi">
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<mods:mods id="96301843CE83674F553F12BC757F977E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6B3766AC1D2EC1B84DAC75F9E23A5AEF">
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<mods:title id="A6757B7AE7C5E7782B75B8A119BEFE21">Taxonomy of ' Euconnus complex'. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepaloconnus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6DC6748664C086C4A6DC79A0354FEF7E" type="personal">
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<mods:role id="DA9F258451A915FF87737806E058E23C">
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<mods:roleTerm id="960C996A51B6C39163C04A13C6B3436A">Author</mods:roleTerm>
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<mods:namePart id="085E9DCFC2D1EE9E5D45514F7DA6D3DA">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="A4443E6BF8628727E24BC2F24179EB8E">text</mods:typeOfResource>
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<mods:titleInfo id="E6F81F9FEE34A63248CABA5789F02D34">
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<mods:title id="DAFCC6773F5B8D1C73F5167FE2978896">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="323E409FF28BEA2824EE95A03B8DC52E">
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<mods:date id="9BA8EB1EF06D427699D6C5206FE7EF2C">2016</mods:date>
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<mods:detail id="59043B84B9C7888D1A26DCF9B84CD543" type="volume">
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<mods:number id="6B580EE18779C3A2BF8A486F0F47DD62">4103</mods:number>
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</mods:detail>
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<mods:detail id="D574936FCF7F33571C4B06AA0E80590D" type="issue">
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<mods:number id="83BE4964978C6EB903E874C36C08C222">5</mods:number>
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<mods:end id="DFEE83C620D93EA40745D2EE8675A41B">472</mods:end>
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<mods:classification id="966450F2DACE83207AEDB7027EE9F5CB">journal article</mods:classification>
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<mods:identifier id="68F6BB87C4DF78FAAAA86F1815617AC5" type="DOI">10.11646/zootaxa.4103.5.4</mods:identifier>
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<mods:identifier id="3E72700468FF1E424D1CA6565F13FB66" type="GBIF-Dataset">b9b4b63c-d1af-47a8-8e0a-3c63a6cbcc91</mods:identifier>
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<mods:identifier id="27F35CA1BF1254276BBB0DBFD0E91163" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="53763953D597A9220848187FD53B4605" type="Zenodo-Dep">263213</mods:identifier>
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</mods:mods>
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<subSection id="E2BC2F4BC03EFFB2A7B100DC446FFE49" lastPageId="4" lastPageNumber="467" pageId="1" pageNumber="464" type="multiple">
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<treatment id="039A87B6C03EFFB2A7B100DC446FFE49" ID-DOI="http://doi.org/10.5281/zenodo.6073094" ID-GBIF-Taxon="127711643" ID-Zenodo-Dep="6073094" LSID="urn:lsid:plazi:treatment:039A87B6C03EFFB2A7B100DC446FFE49" httpUri="http://treatment.plazi.org/id/039A87B6C03EFFB2A7B100DC446FFE49" lastPageId="4" lastPageNumber="467" pageId="1" pageNumber="464">
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<subSubSection id="C329652BC03EFFB7A7B100DC44A8FB80" box="[151,1195,1003,1030]" pageId="1" pageNumber="464" type="nomenclature">
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<paragraph id="8B8C36A0C03EFFB7A7B100DC44A8FB80" blockId="1.[151,1195,1003,1030]" box="[151,1195,1003,1030]" pageId="1" pageNumber="464">
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<heading id="D0C481CCC03EFFB7A7B100DC44A8FB80" bold="true" box="[151,1195,1003,1030]" fontSize="11" level="1" pageId="1" pageNumber="464" reason="1">
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<keyStep id="30C72E05C03EFFB7A7B100DC44A8FB80" box="[151,1195,1003,1030]" pageId="1" pageNumber="464">
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<key id="FFA26191C03EFFB7A7B100DC44A8FB80" box="[151,1195,1003,1030]" pageId="1" pageNumber="464">
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<keyLead id="30C29595C03EFFB7A7B100DC44A8FB80" box="[151,1195,1003,1030]" pageId="1" pageNumber="464">
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<emphasis id="B947EAB2C03EFFB7A7B100DC44A8FB80" bold="true" box="[151,1195,1003,1030]" pageId="1" pageNumber="464">
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1. Morphological structures and taxonomic status of
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<taxonomicName id="4C334D23C03EFFB7A40F00DB43A6FB83" box="[809,933,1003,1029]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A40F00DB43A6FB83" bold="true" box="[809,933,1003,1029]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C334D23C03EFFB7A49000DB44A7FB80" authority="Franz" authorityName="Franz" box="[950,1188,1003,1030]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A49000DB444CFB83" bold="true" box="[950,1103,1003,1029]" italics="true" pageId="1" pageNumber="464">Napoconnus</emphasis>
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Franz
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</taxonomicName>
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)
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</emphasis>
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</keyLead>
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</key>
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</keyStep>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C329652BC03EFFB7A7B107024299FBE8" pageId="1" pageNumber="464" type="diagnosis">
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<paragraph id="8B8C36A0C03EFFB7A7B107024299FBE8" blockId="1.[151,1436,1074,2035]" pageId="1" pageNumber="464">
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The general body shape of
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<taxonomicName id="4C334D23C03EFFB7A6F207044357FBCD" box="[468,852,1074,1099]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
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<emphasis id="B947EAB2C03EFFB7A6F207044241FBCD" box="[468,578,1076,1099]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
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(
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<emphasis id="B947EAB2C03EFFB7A573070442E2FBCD" box="[597,737,1076,1099]" italics="true" pageId="1" pageNumber="464">Napoconnus</emphasis>
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)
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<emphasis id="B947EAB2C03EFFB7A5D207024357FBCD" box="[756,852,1074,1099]" italics="true" pageId="1" pageNumber="464">kubienai</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13082A25C03EFFB7A441070243A9FBCD" box="[871,938,1074,1099]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="1" pageNumber="464">Fig. 1</figureCitation>
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) is very similar to that of many species of
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<taxonomicName id="4C334D23C03EFFB7A7B107674106FBE8" box="[151,261,1111,1134]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A7B107674106FBE8" box="[151,261,1111,1134]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
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</taxonomicName>
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, especially the subgenus
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<taxonomicName id="4C334D23C03EFFB7A50507664290FBE9" box="[547,659,1110,1135]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A50507664290FBE9" box="[547,659,1110,1135]" italics="true" pageId="1" pageNumber="464">Napochus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C329652BC03EFFB5A7E1074A41C7F92C" lastPageId="3" lastPageNumber="466" pageId="1" pageNumber="464" type="description">
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<paragraph id="8B8C36A0C03EFFB7A7E1074A4112F9E1" blockId="1.[151,1436,1074,2035]" pageId="1" pageNumber="464">
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The head capsule (
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<figureCitation id="13082A25C03EFFB7A6BF074A41E2FB15" box="[409,481,1146,1171]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="1" pageNumber="464">Figs 1</figureCitation>
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,
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<figureCitation id="13082A25C03EFFB7A6C9074B420FFB15" box="[495,524,1147,1171]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">10</figureCitation>
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) is divided into the posterior part ('neck' region) and anterior, exposed part by an occipital constriction broader than half width of head. The anterior part of the head is slightly elongate and distinctly flattened; vertex distinctly demarcated from the 'neck' region and not bulging posterodorsally; frons subtrapezoidal; antennal insertions moderately broadly separated; compound eyes located anterolaterally, so that the tempora are long. The tempora and genae are covered with thick bristles (
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<figureCitation id="13082A25C03EFFB7A320063A4459FAA5" box="[1030,1114,1290,1315]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">Fig. 10</figureCitation>
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). The posterior tentorial pits (
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<figureCitation id="13082A25C03EFFB7A7B9061E40F5FAC1" box="[159,246,1326,1351]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">Fig. 10</figureCitation>
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;
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<emphasis id="B947EAB2C03EFFB7A620061F4129FAC0" box="[262,298,1327,1350]" italics="true" pageId="1" pageNumber="464">ptp</emphasis>
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) are elongate and located slightly in front of the transverse impression demarcating ventrally the anterior and posterior parts of the head capsule; the gular plate (
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<figureCitation id="13082A25C03EFFB7A44A066243C3FAED" box="[876,960,1362,1387]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">Fig. 10</figureCitation>
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;
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<emphasis id="B947EAB2C03EFFB7A4EA066443EBFAED" box="[972,1000,1364,1387]" italics="true" pageId="1" pageNumber="464">gp</emphasis>
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) is broad and subtrapezoidal, but with its anterior portion adjacent to the tentorial pits narrowed and elongate. The submentum (
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<figureCitation id="13082A25C03EFFB7A3940646450AFA09" box="[1202,1289,1398,1423]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">Fig. 10</figureCitation>
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;
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<emphasis id="B947EAB2C03EFFB7A23F06474545FA08" box="[1305,1350,1399,1422]" italics="true" pageId="1" pageNumber="464">smn</emphasis>
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) is not demarcated laterally by sutures, and hypostomal ridges (
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<figureCitation id="13082A25C03EFFB7A43F06AA436DFA35" box="[793,878,1434,1459]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">Fig. 10</figureCitation>
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;
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<emphasis id="B947EAB2C03EFFB7A45D06AA4390FA35" box="[891,915,1434,1459]" italics="true" pageId="1" pageNumber="464">hr</emphasis>
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) are short, strongly bent mesally and running nearly parallel to the posterior margins of cardines. Most of these characters can be found in
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<taxonomicName id="4C334D23C03EFFB7A388068F451FFA50" box="[1198,1308,1471,1494]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B947EAB2C03EFFB7A388068F451FFA50" box="[1198,1308,1471,1494]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
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</taxonomicName>
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, except for the unusually short and strongly bent mesally hypostomal ridges (in
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<taxonomicName id="4C334D23C03EFFB7A49506D44422FA7D" box="[947,1057,1508,1531]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B947EAB2C03EFFB7A49506D44422FA7D" box="[947,1057,1508,1531]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27457C9C03EFFB7A30A06D3446FFA7C" box="[1068,1132,1507,1530]" pageId="1" pageNumber="464" sensu="stricto">s. str.</taxonomicNameLabel>
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,
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<taxonomicName id="4C334D23C03EFFB7A35C06D244E9FA7D" box="[1146,1258,1506,1531]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A35C06D244E9FA7D" box="[1146,1258,1506,1531]" italics="true" pageId="1" pageNumber="464">Napochus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C334D23C03EFFB7A3DD06D24596FA7D" box="[1275,1429,1506,1531]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03EFFB7A3DD06D24596FA7D" box="[1275,1429,1506,1531]" italics="true" pageId="1" pageNumber="464">Filonapochus</emphasis>
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</taxonomicName>
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,
|
||
<taxonomicName id="4C334D23C03EFFB7A7B105364110F999" box="[151,275,1542,1567]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A7B105364110F999" box="[151,275,1542,1567]" italics="true" pageId="1" pageNumber="464">Tetramelus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03EFFB7A606053641CDF999" box="[288,462,1542,1567]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A606053641CDF999" box="[288,462,1542,1567]" italics="true" pageId="1" pageNumber="464">Paratetramelus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03EFFB7A6FA0537427BF998" box="[476,632,1543,1566]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A6FA0537427BF998" box="[476,632,1543,1566]" italics="true" pageId="1" pageNumber="464">Heteroconnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B947EAB2C03EFFB7A5880536434BF999" box="[686,840,1542,1567]" italics="true" pageId="1" pageNumber="464">Glabriconnus</emphasis>
|
||
the ridges are long and extend posteromesally nearly to the posterior tentorial pits; in
|
||
<taxonomicName id="4C334D23C03EFFB7A6DA051A42AAF9C5" box="[508,681,1578,1603]" class="Insecta" family="Staphylinidae" genus="Rhomboconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A6DA051A42AAF9C5" box="[508,681,1578,1603]" italics="true" pageId="1" pageNumber="464">Rhomboconnus</emphasis>
|
||
</taxonomicName>
|
||
the posterior ends of ridges are far from tentorial pits but connected at middle).
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03EFFB7A7E1054244F3F8E5" blockId="1.[151,1436,1074,2035]" pageId="1" pageNumber="464">
|
||
The antennae (
|
||
<figureCitation id="13082A25C03EFFB7A654054241BFF90D" box="[370,444,1650,1675]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="1" pageNumber="464">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13082A25C03EFFB7A6EB054341EBF90D" box="[461,488,1651,1675]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="1" pageNumber="464">11</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C334D23C03EFFB7A5380544439CF90D" box="[542,927,1650,1675]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03EFFB7A5380544428FF90D" box="[542,652,1652,1675]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03EFFB7A5860544432FF90D" box="[672,812,1652,1675]" italics="true" pageId="1" pageNumber="464">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03EFFB7A4190542439CF90D" box="[831,927,1650,1675]" italics="true" pageId="1" pageNumber="464">kubienai</emphasis>
|
||
</taxonomicName>
|
||
are relatively short, with scape and pedicel elongate and proximal flagellomeres subcylindrical and compactly assembled; the club is composed of three antennomeres, of which the antennomere IX is only slightly broader than VIII and therefore the club appears as two-segmented. Antennomeres X and XI are much broader and longer than IX. Such composition of the antennal club is not known in any of the hitherto reviewed subgenera of
|
||
<taxonomicName id="4C334D23C03EFFB7A47E043443C5F89D" box="[856,966,1796,1819]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A47E043443C5F89D" box="[856,966,1796,1819]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, but in other species of
|
||
<taxonomicName id="4C334D23C03EFFB7A3F60434455FF89D" box="[1232,1372,1796,1819]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A3F60434455FF89D" box="[1232,1372,1796,1819]" italics="true" pageId="1" pageNumber="464">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EFA24B51C03EFFB7A7B10417411BF8B9" author="Franz" box="[151,280,1830,1855]" pageId="1" pageNumber="464" refString="Franz, H. (1957) Neue Scydmaenidae (Coleoptera) aus Belgisch-Congo. I. Acta Zoologica Cracoviensia, 2, 17 - 60." type="journal article" year="1957">Franz 1957</bibRefCitation>
|
||
) the club is clearly three-segmented, with the antennomere IX about as broad as X and XI. Antennal clubs in such species resemble those of
|
||
<taxonomicName id="4C334D23C03EFFB7A572047A42C7F8E5" box="[596,708,1866,1891]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A572047A42C7F8E5" box="[596,708,1866,1891]" italics="true" pageId="1" pageNumber="464">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, except for being three-, and not four-segmented.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03EFFB7A7E1045E4296F874" blockId="1.[151,1436,1074,2035]" pageId="1" pageNumber="464">
|
||
The prothorax of
|
||
<taxonomicName id="4C334D23C03EFFB7A6AD045F4306F801" box="[395,773,1902,1927]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03EFFB7A6AD045F41FAF800" box="[395,505,1903,1926]" italics="true" pageId="1" pageNumber="464">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03EFFB7A52F045F4296F800" box="[521,661,1903,1926]" italics="true" pageId="1" pageNumber="464">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03EFFB7A583045E4306F801" box="[677,773,1902,1927]" italics="true" pageId="1" pageNumber="464">kubienai</emphasis>
|
||
</taxonomicName>
|
||
in dorsal view (
|
||
<figureCitation id="13082A25C03EFFB7A49F045E43FAF801" box="[953,1017,1902,1927]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="1" pageNumber="464">Fig. 1</figureCitation>
|
||
) is elongate and subtrapezoidal, with sides nearly straight and strongly convergent anteriorly, the pronotum is broadest near base. The pronotal base bears a faint transverse impression and a pair of feebly marked lateral antebasal pits. The general shape of pronotum is very similar to that of
|
||
<taxonomicName id="4C334D23C03EFFB7A53804EA428EF875" box="[542,653,2010,2035]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="464" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03EFFB7A53804EA428EF875" box="[542,653,2010,2035]" italics="true" pageId="1" pageNumber="464">Napochus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4C6628C03DFFB4A7B1052241A3F9E0" httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="2" pageNumber="465" targetBox="[158,1426,180,1516]" targetPageId="2">
|
||
<paragraph id="8B8C36A0C03DFFB4A7B1052241A3F9E0" blockId="2.[151,1435,1554,1638]" pageId="2" pageNumber="465">
|
||
<emphasis id="B947EAB2C03DFFB4A7B105224142F9AE" bold="true" box="[151,321,1554,1577]" pageId="2" pageNumber="465">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="4C334D23C03DFFB4A668052442F9F9AF" authority="Franz" authorityName="Franz" box="[334,762,1555,1577]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03DFFB4A668052441B1F9AF" box="[334,434,1556,1577]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03DFFB4A6E105244246F9AF" box="[455,581,1556,1577]" italics="true" pageId="2" pageNumber="465">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03DFFB4A57A052342B0F9AF" box="[604,691,1555,1577]" italics="true" pageId="2" pageNumber="465">kubienai</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
(1–3);
|
||
<taxonomicName id="4C334D23C03DFFB4A4740524451CF9AF" authority="Franz" authorityName="Franz" box="[850,1311,1555,1577]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C03DFFB4A474052443B5F9AF" box="[850,950,1556,1577]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03DFFB4A4ED0523445CF9AF" box="[971,1119,1555,1577]" italics="true" pageId="2" pageNumber="465">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03DFFB4A352052344DBF9AF" box="[1140,1240,1555,1577]" italics="true" pageId="2" pageNumber="465">shutjensis</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
(4–6); and
|
||
<taxonomicName id="4C334D23C03DFFB4A7B10502426FF9C1" authority="Franz" authorityName="Franz" box="[151,620,1585,1607]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="khumbuensis" subGenus="Nepaloconnus">
|
||
<emphasis id="B947EAB2C03DFFB4A7B1050240F8F9C1" box="[151,251,1586,1607]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03DFFB4A62F05014199F9C1" box="[265,410,1585,1607]" italics="true" pageId="2" pageNumber="465">Nepaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03DFFB4A68F0501422FF9C1" box="[425,556,1585,1607]" italics="true" pageId="2" pageNumber="465">khumbuensis</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
(7–9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C36A0C03DFFB4A7E105BD4574F96B" blockId="2.[151,1436,1677,2026]" pageId="2" pageNumber="465">
|
||
The prosternum (
|
||
<figureCitation id="13082A25C03DFFB4A6AA05BD41DCF920" box="[396,479,1677,1702]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="2" pageNumber="465">Fig. 10</figureCitation>
|
||
) is short, lacking prosternal process or carina; the notosternal sutures (
|
||
<figureCitation id="13082A25C03DFFB4A22A05BD4562F920" box="[1292,1377,1677,1702]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="2" pageNumber="465">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03DFFB4A24905BE4590F923" box="[1391,1427,1678,1701]" italics="true" pageId="2" pageNumber="465">nss</emphasis>
|
||
) and hypomeral ridges (
|
||
<figureCitation id="13082A25C03DFFB4A6BA058141F3F94C" box="[412,496,1713,1738]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="2" pageNumber="465">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03DFFB4A6DB05814221F94C" box="[509,546,1713,1738]" italics="true" pageId="2" pageNumber="465">hyr</emphasis>
|
||
) seem complete (but they are only partly visible in the studied specimen), and the sides of prothorax are covered with thick bristles. These are characters similar to those known in
|
||
<taxonomicName id="4C334D23C03DFFB4A22605E54573F968" box="[1280,1392,1749,1774]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03DFFB4A22605E54573F968" box="[1280,1392,1749,1774]" italics="true" pageId="2" pageNumber="465">Napochus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03DFFB4A7E105C941A0F8DF" blockId="2.[151,1436,1677,2026]" pageId="2" pageNumber="465">
|
||
The mesoventral structures, especially the carinate, narrow and highly elevated mesoventral intercoxal process (
|
||
<figureCitation id="13082A25C03DFFB4A7B9042D40F1F8B0" box="[159,242,1821,1846]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="2" pageNumber="465">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03DFFB4A7D8042E4134F8B3" box="[254,311,1822,1845]" italics="true" pageId="2" pageNumber="465">msvp</emphasis>
|
||
) of
|
||
<taxonomicName id="4C334D23C03DFFB4A640042E42E2F8B0" box="[358,737,1821,1846]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03DFFB4A640042E41D7F8B3" box="[358,468,1822,1845]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03DFFB4A6C2042E4273F8B3" box="[484,624,1822,1845]" italics="true" pageId="2" pageNumber="465">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03DFFB4A5A7042D42E2F8B0" box="[641,737,1821,1846]" italics="true" pageId="2" pageNumber="465">kubienai</emphasis>
|
||
</taxonomicName>
|
||
do not deviate from those known in
|
||
<taxonomicName id="4C334D23C03DFFB4A35A042E44E9F8B3" box="[1148,1258,1822,1845]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B947EAB2C03DFFB4A35A042E44E9F8B3" box="[1148,1258,1822,1845]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27457C9C03DFFB4A3D4042E452EF8B3" box="[1266,1325,1822,1845]" pageId="2" pageNumber="465" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
and other subgenera of
|
||
<taxonomicName id="4C334D23C03DFFB4A60B04724198F8DF" box="[301,411,1858,1881]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03DFFB4A60B04724198F8DF" box="[301,411,1858,1881]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03DFFB4A7E104554397F86C" blockId="2.[151,1436,1677,2026]" pageId="2" pageNumber="465">
|
||
The metaventral intercoxal process of
|
||
<taxonomicName id="4C334D23C03DFFB4A5A804564417F8F8" box="[654,1044,1893,1918]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03DFFB4A5A8045642FFF8FB" box="[654,764,1894,1917]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03DFFB4A4340456439DF8FB" box="[786,926,1894,1917]" italics="true" pageId="2" pageNumber="465">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03DFFB4A49204554417F8F8" box="[948,1044,1893,1918]" italics="true" pageId="2" pageNumber="465">kubienai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13082A25C03DFFB4A30E04554482F8F8" box="[1064,1153,1893,1918]" captionStart="FIGURES 10 – 12" captionStartId="3.[151,264,1210,1233]" captionTargetBox="[151,1436,181,1189]" captionTargetId="figure@3.[151,1436,181,1189]" captionTargetPageId="3" captionText="FIGURES 10 – 12. Morphological structures of Euconnus (Napoconnus) kubienai Franz. Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v 3, metaventrite." httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="2" pageNumber="465">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03DFFB4A3B4045644CBF8FB" box="[1170,1224,1894,1917]" italics="true" pageId="2" pageNumber="465">mtvp</emphasis>
|
||
) is not similar to structures found in previously studied subgenera of
|
||
<taxonomicName id="4C334D23C03DFFB4A5C204BA4351F827" box="[740,850,1930,1953]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03DFFB4A5C204BA4351F827" box="[740,850,1930,1953]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
. Instead of being moderately broad and short, as in
|
||
<taxonomicName id="4C334D23C03DFFB4A7B1049F4106F840" box="[151,261,1967,1990]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03DFFB4A7B1049F4106F840" box="[151,261,1967,1990]" italics="true" pageId="2" pageNumber="465">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, the process in
|
||
<taxonomicName id="4C334D23C03DFFB4A6EB049F425AF840" box="[461,601,1967,1990]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03DFFB4A6EB049F425AF840" box="[461,601,1967,1990]" italics="true" pageId="2" pageNumber="465">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
is narrow and forming a pair of long spines touching at middle and projecting posteriorly, so that the metacoxae are narrowly separated.
|
||
</paragraph>
|
||
<caption id="DF4C6628C03CFFB5A7B1078A41C5FACD" httpUri="https://zenodo.org/record/263215/files/figure.png" pageId="3" pageNumber="466" targetBox="[151,1436,181,1189]" targetPageId="3">
|
||
<paragraph id="8B8C36A0C03CFFB5A7B1078A41C5FACD" blockId="3.[151,1437,1210,1355]" pageId="3" pageNumber="466">
|
||
<emphasis id="B947EAB2C03CFFB5A7B1078A4157FB49" bold="true" box="[151,340,1210,1233]" pageId="3" pageNumber="466">FIGURES 10–12.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C334D23C03CFFB5A558078B441CFB56" authority="Franz." authorityName="Franz." box="[638,1055,1210,1232]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03CFFB5A558078B42E1FB56" box="[638,738,1211,1232]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03CFFB5A5D4078B4373FB56" box="[754,880,1211,1232]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03CFFB5A4A6078A43D4FB56" box="[896,983,1210,1232]" italics="true" pageId="3" pageNumber="466">kubienai</emphasis>
|
||
Franz.
|
||
</taxonomicName>
|
||
Head, prothorax and anterior portion of pterothorax in ventral view (10); right antennal club in ventral view (11); pterothorax in ventral view (12). Abbreviations: gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, notosternal suture; pcr, procoxal rest; ptp, posterior tentorial pit; smn, submentum; v3, metaventrite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C36A0C03CFFB5A7E106424554FA71" blockId="3.[151,1437,1394,2031]" pageId="3" pageNumber="466">
|
||
The elytra (
|
||
<figureCitation id="13082A25C03CFFB5A66C06424189FA0D" box="[330,394,1394,1419]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="3" pageNumber="466">Fig. 1</figureCitation>
|
||
) are oval, elongate, each with two barely noticeable asetose rudiments of basal foveae. This character differs from conditions known in most subgenera of
|
||
<taxonomicName id="4C334D23C03CFFB5A47B06A743C8FA28" box="[861,971,1431,1454]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A47B06A743C8FA28" box="[861,971,1431,1454]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, which have large and deep basal elytral foveae. However, the foveae are either entirely reduced or visible only as barely discernible rudiments in
|
||
<taxonomicName id="4C334D23C03CFFB5A7B106EE4147FA71" box="[151,324,1502,1527]" class="Insecta" family="Staphylinidae" genus="Rhomboconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A7B106EE4147FA71" box="[151,324,1502,1527]" italics="true" pageId="3" pageNumber="466">Rhomboconnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D23C03CFFB5A65C06EE4217FA71" box="[378,532,1502,1527]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A65C06EE4217FA71" box="[378,532,1502,1527]" italics="true" pageId="3" pageNumber="466">Filonapochus</emphasis>
|
||
</taxonomicName>
|
||
, or they can be very small (
|
||
<taxonomicName id="4C334D23C03CFFB5A46A06EF43EBFA70" box="[844,1000,1503,1526]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A46A06EF43EBFA70" box="[844,1000,1503,1526]" italics="true" pageId="3" pageNumber="466">Heteroconnus</emphasis>
|
||
</taxonomicName>
|
||
and some species of
|
||
<taxonomicName id="4C334D23C03CFFB5A3F106EE4544FA71" box="[1239,1351,1502,1527]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A3F106EE4544FA71" box="[1239,1351,1502,1527]" italics="true" pageId="3" pageNumber="466">Napochus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03CFFB5A7E1053241C7F92C" blockId="3.[151,1437,1394,2031]" pageId="3" pageNumber="466">
|
||
The aedeagus (
|
||
<figureCitation id="13082A25C03CFFB5A648053241D2F99D" box="[366,465,1538,1563]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="3" pageNumber="466">Figs 2–3</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C334D23C03CFFB5A6D80534437BF99D" box="[510,888,1538,1563]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03CFFB5A6D80534426FF99D" box="[510,620,1540,1563]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03CFFB5A55A0534430BF99D" box="[636,776,1540,1563]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03CFFB5A43E0532437BF99D" box="[792,888,1538,1563]" italics="true" pageId="3" pageNumber="466">kubienai</emphasis>
|
||
</taxonomicName>
|
||
is symmetrical, with free and slender parameres, sub-basally located dorsal orifice and subapically located ventral ostium, with particularly long apical projection of the dorsal wall.
|
||
<collectingCountry id="F3247630C03CFFB5A675057B4171F9E5" box="[339,370,1611,1635]" name="American Samoa" pageId="3" pageNumber="466">As</collectingCountry>
|
||
the aedeagus within
|
||
<taxonomicName id="4C334D23C03CFFB5A54D057C42DAF9E5" box="[619,729,1612,1635]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A54D057C42DAF9E5" box="[619,729,1612,1635]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
(and other genera of Glandulariini) is extremely diverse, the copulatory organ of
|
||
<taxonomicName id="4C334D23C03CFFB5A6AD055F4311F901" box="[395,786,1646,1671]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03CFFB5A6AD055F41FAF900" box="[395,505,1647,1670]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03CFFB5A536055F429FF900" box="[528,668,1647,1670]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03CFFB5A594055E4311F901" box="[690,786,1646,1671]" italics="true" pageId="3" pageNumber="466">kubienai</emphasis>
|
||
</taxonomicName>
|
||
does not provide any particular subgeneric or generic diagnostic character states.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C329652BC03CFFB5A7E10586435DF869" pageId="3" pageNumber="466" type="discussion">
|
||
<paragraph id="8B8C36A0C03CFFB5A7E10586435DF869" blockId="3.[151,1437,1394,2031]" pageId="3" pageNumber="466">
|
||
<emphasis id="B947EAB2C03CFFB5A7E10586415CF949" bold="true" box="[199,351,1718,1743]" pageId="3" pageNumber="466">Conclusions.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C334D23C03CFFB5A5E70587434EF948" box="[705,845,1719,1742]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A5E70587434EF948" box="[705,845,1719,1742]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="4C334D23C03CFFB5A35E058744E5F948" box="[1144,1254,1719,1742]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A35E058744E5F948" box="[1144,1254,1719,1742]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, especially the subgenus
|
||
<taxonomicName id="4C334D23C03CFFB5A62A05EA417FF975" box="[268,380,1754,1779]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A62A05EA417FF975" box="[268,380,1754,1779]" italics="true" pageId="3" pageNumber="466">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, but the narrow metaventral intercoxal process with a pair of posteriorly directed spines excludes this taxon from
|
||
<taxonomicName id="4C334D23C03CFFB5A69105CF4226F890" box="[439,549,1791,1814]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A69105CF4226F890" box="[439,549,1791,1814]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
. The broad vs. narrow metaventral intercoxal process was previously adopted as a distinction between
|
||
<taxonomicName id="4C334D23C03CFFB5A69104144226F8BD" box="[439,549,1828,1851]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A69104144226F8BD" box="[439,549,1828,1851]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D23C03CFFB5A54504124375F8BD" authority="Broun, 1983" authorityName="Broun" authorityYear="1983" box="[611,886,1826,1851]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A545041242DAF8BD" box="[611,729,1826,1851]" italics="true" pageId="3" pageNumber="466">Sciacharis</emphasis>
|
||
Broun, 1983
|
||
</taxonomicName>
|
||
, (
|
||
<bibRefCitation id="EFA24B51C03CFFB5A4AA04124462F8BD" author="Jaloszynski" box="[908,1121,1826,1851]" pageId="3" pageNumber="466" refString="Jaloszynski, P. (2015 c) Taxonomy of ' Euconnus complex'. Part V. Review of type species of Euconnus subgenera Tetramelus, Paratetramelus and Heteroconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4040 (3), 280 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 4040.3.2" type="journal article" year="2015" yearSuffix="c">Jałoszyński 2015c</bibRefCitation>
|
||
), genera that are similar in many other structures. However, in
|
||
<taxonomicName id="4C334D23C03CFFB5A50E0476429DF8D9" box="[552,670,1862,1887]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A50E0476429DF8D9" box="[552,670,1862,1887]" italics="true" pageId="3" pageNumber="466">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
the metacoxae are contiguous and the metaventral process lacks the pair of spines, found in
|
||
<taxonomicName id="4C334D23C03CFFB5A6BB045C422AF805" box="[413,553,1900,1923]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A6BB045C422AF805" box="[413,553,1900,1923]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
. This structure can be found in many genera of Glandulariini, but none of them has the body form so much
|
||
<taxonomicName id="4C334D23C03CFFB5A6F604BE4243F821" box="[464,576,1934,1959]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A6F604BE4243F821" box="[464,576,1934,1959]" italics="true" pageId="3" pageNumber="466">Napochus</emphasis>
|
||
</taxonomicName>
|
||
-like as
|
||
<taxonomicName id="4C334D23C03CFFB5A5BC04BF4416F821" box="[666,1045,1934,1959]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="kubienai" subGenus="Napoconnus">
|
||
<emphasis id="B947EAB2C03CFFB5A5BC04BF430BF820" box="[666,776,1935,1958]" italics="true" pageId="3" pageNumber="466">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03CFFB5A43F04BF43A6F820" box="[793,933,1935,1958]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03CFFB5A49304BE4416F821" box="[949,1045,1934,1959]" italics="true" pageId="3" pageNumber="466">kubienai</emphasis>
|
||
</taxonomicName>
|
||
and other species currently placed in
|
||
<taxonomicName id="4C334D23C03CFFB5A79E04844147F84D" box="[184,324,1972,1995]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A79E04844147F84D" box="[184,324,1972,1995]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
; also none of them has the antennal club composed of enlarged three terminal antennomeres. Consequently,
|
||
<taxonomicName id="4C334D23C03CFFB5A61B04E741CAF868" box="[317,457,2007,2030]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03CFFB5A61B04E741CAF868" box="[317,457,2007,2030]" italics="true" pageId="3" pageNumber="466">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
is here elevated to the generic rank.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C329652BC03BFFB2A7E103A7446FFE49" pageId="4" pageNumber="467" type="diagnosis">
|
||
<paragraph id="8B8C36A0C03BFFB2A7E103A7446FFE49" blockId="4.[151,1437,151,464]" pageId="4" pageNumber="467">
|
||
<emphasis id="B947EAB2C03BFFB2A7E103A743DDFF36" bold="true" box="[199,990,151,176]" pageId="4" pageNumber="467">
|
||
Emended diagnosis of the genus
|
||
<taxonomicName id="4C334D23C03BFFB2A55903A84312FF29" box="[639,785,152,175]" class="Insecta" family="Staphylinidae" genus="Napoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A55903A84312FF29" bold="true" box="[639,785,152,175]" italics="true" pageId="4" pageNumber="467">Napoconnus</emphasis>
|
||
</taxonomicName>
|
||
, status revised.
|
||
</emphasis>
|
||
Body
|
||
<taxonomicName id="4C334D23C03BFFB2A31B03A744AEFF36" box="[1085,1197,151,176]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A31B03A744AEFF36" box="[1085,1197,151,176]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
|
||
</taxonomicName>
|
||
-like, i.e., the head subpentagonal with long tempora, pronotum subtrapezoidal, broadest near base and strongly narrowing anteriorly, with lateral margins nearly straight, and oval elytra; head and pronotum with thick bristles. Antenna with distinct club composed of antennomeres IX–XI; hypostomal ridges short, strongly bent mesally and running nearly parallel to posterior margins of cardines; pronotum with faint antebasal impression and one pair of barely discernible lateral pits, lacking sublateral carinae; hypomeral ridges complete; prosternum lacking prosternal process or carina; mesoventral intercoxal process keel-like, anteriorly connected with anterior ridge of mesoventrite; metaventral intercoxal process narrowly separating metacoxae, with a pair of posteriorly directed, pointed spines adjacent at middle; each elytron with two rudimentary basal foveae; aedeagus with free parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |