123 lines
15 KiB
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123 lines
15 KiB
XML
<document id="04C6488D55E78BC71D21DE3E40565586" ID-CLB-Dataset="32715" ID-DOI="10.11646/zootaxa.4290.2.2" ID-GBIF-Dataset="4469947c-31f9-46e6-b963-03ae79d84554" ID-ISSN="1175-5326" ID-Zenodo-Dep="829009" ID-ZooBank="79FBA585-7662-4DE1-9BAB-37DFDA2BABBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110470243" checkinUser="plazi" docAuthor="Marshall, Stephen A." docDate="2017" docId="0399A674FFDAA005FF0AFC40AABC8BEB" docLanguage="en" docName="zootaxa.4290.2.2.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paramimegralla zinzala Marshall, 2017, new species" docType="treatment" docVersion="7" lastPageNumber="280" masterDocId="FFA0DE0CFFF8A021FF9DFF89A94D8B13" masterDocTitle="Micropezidae (Insecta, Diptera, Acalyptratae) of Madagascar and a revision of the genus Paramimegralla Hennig" masterLastPageNumber="280" masterPageNumber="244" pageNumber="278" updateTime="1698465370965" updateUser="ExternalLinkService">
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<mods:title id="9834EF260C0409B61A2B156132314B8E">Micropezidae (Insecta, Diptera, Acalyptratae) of Madagascar and a revision of the genus Paramimegralla Hennig</mods:title>
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<mods:name id="2240E2CBC377BE75E209D8AB471232AC" type="personal">
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<mods:namePart id="5E620A4E2C973A9AFC718D7B11FD152E">Marshall, Stephen A.</mods:namePart>
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<mods:title id="50C036E4CD12328CCB953CF2C91FD4C9">Zootaxa</mods:title>
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<mods:date id="220A7CF4F2469EB421DA61DEEE76C4EA">2017</mods:date>
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<mods:number id="06FDDA1A41B01BB6B1126C8431EA8006">4290</mods:number>
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<mods:identifier id="4D0873015CE95B27D771A44B622917A2" type="DOI">10.11646/zootaxa.4290.2.2</mods:identifier>
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<treatment id="0399A674FFDAA005FF0AFC40AABC8BEB" ID-DOI="http://doi.org/10.5281/zenodo.6033459" ID-GBIF-Taxon="132549261" ID-Zenodo-Dep="6033459" LSID="urn:lsid:plazi:treatment:0399A674FFDAA005FF0AFC40AABC8BEB" httpUri="http://treatment.plazi.org/id/0399A674FFDAA005FF0AFC40AABC8BEB" lastPageId="36" lastPageNumber="280" pageId="34" pageNumber="278">
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<subSubSection id="C32A44E9FFDAA003FF0AFC40A87D8F16" pageId="34" pageNumber="278" type="nomenclature">
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<paragraph id="8B8F1762FFDAA003FF0AFC40AB0288F0" blockId="34.[151,591,969,1029]" box="[151,591,969,995]" pageId="34" pageNumber="278">
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<heading id="D0C7A00EFFDAA003FF0AFC40AB0288F0" bold="true" box="[151,591,969,995]" fontSize="11" level="1" pageId="34" pageNumber="278" reason="1">
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<emphasis id="B944CB70FFDAA003FF0AFC40AB0288F0" bold="true" box="[151,591,969,995]" pageId="34" pageNumber="278">
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<taxonomicName id="4C306CE1FFDAA003FF0AFC40A8F588F0" ID-CoL="9HVWV" box="[151,440,969,995]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="278" phylum="Arthropoda" rank="species" species="zinzala" status="sp. nov.">
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<emphasis id="B944CB70FFDAA003FF0AFC40A8F588F0" bold="true" box="[151,440,969,995]" italics="true" pageId="34" pageNumber="278">Paramimegralla zinzala</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277760BFFDAA003FE22FC40AB0288F0" box="[447,591,969,995]" pageId="34" pageNumber="278" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F1762FFDAA003FF0AFC65A87D8F16" blockId="34.[151,591,969,1029]" box="[151,304,1004,1029]" pageId="34" pageNumber="278">
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<figureCitation id="130B0BE7FFDAA003FF0AFC65A9B78F16" box="[151,250,1004,1029]" captionStart="FIGURES 106 – 112" captionStartId="35.[151,264,1826,1849]" captionTargetBox="[151,1436,286,1805]" captionTargetId="figure@35.[151,1436,286,1805]" captionTargetPageId="35" captionText="FIGURES 106 – 112. Paramimegralla zinzala sp. nov., Male. 106. 107. Living fly along trail at Ranomafana; 108. Head, dorsal; 109. Head, lateral; 110. Epandrium and associated structures; left lateral 111. Abdominal segments 5 ‾ 8, lateral; 112. Phallus detail, lateral." httpUri="https://zenodo.org/record/829045/files/figure.png" pageId="34" pageNumber="278">Figs 106</figureCitation>
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̄112
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFDAA003FF0AFBA4AC5F8CE2" pageId="34" pageNumber="278" type="description">
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<paragraph id="8B8F1762FFDAA003FF0AFBA4ACCD8F55" blockId="34.[151,1437,1069,2033]" box="[151,1408,1069,1094]" pageId="34" pageNumber="278">
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<emphasis id="B944CB70FFDAA003FF0AFBA4A8648F55" bold="true" box="[151,297,1069,1094]" pageId="34" pageNumber="278">Description.</emphasis>
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Length
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<quantity id="4CC8BA87FFDAA003FE18FBA4A8958F56" box="[389,472,1069,1094]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="34" pageNumber="278" unit="mm" value="16.0">16 mm</quantity>
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. Ground colour mostly black, legs and lower head mostly yellow as detailed below.
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</paragraph>
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<paragraph id="8B8F1762FFDAA003FF5AFBD8A82B8E51" blockId="34.[151,1437,1069,2033]" pageId="34" pageNumber="278">Head with frons broad, ridged longitudinally; epicephalon shining; frontal vitta velvety black pruinose, strongly tapering anteriorly, broad and gently tapered posteriorly to a rounded apex ringed by a silvery pollinose area. Back of head strikingly depressed; occipital suture shining brown, expanded posteriorly, back of head dark, ventral third of head, including face, clypeus, palpus, gena, and lower third of postocciput, yellow to reddish. Head chaetotaxy complete, with 2 frontal setae. Parafacial silvery dorsally and ventrally, black at middle. Palpus broad. Clypeus bare, shorter than face, with a prominent peak at middle of upper margin. Length of first flagellomere subequal to width.</paragraph>
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<paragraph id="8B8F1762FFDAA003FF5AFAC4AA838D2D" blockId="34.[151,1437,1069,2033]" pageId="34" pageNumber="278">Thorax with scutellum silvery, scutum mostly densely silver microsetulose, central and lateral strips golden microsetulose. Proepisternum with short, inconspicuous yellow ventral setae. Posterior anepimeron and anterior anepisternum bare and shining with shining area continuing behind procoxae, pleuron otherwise silvery microsetulose. Legs mostly yellow, mid femur with a long but incomplete distal black ring, apex of mid femur and base of mid tibia black. Fore femur with anterior surface black on basal half. Fore tibia and basal half of fore tarsomere one dark brown to black, fore tarsomeres 1̄3 otherwise bright white, tarsomeres 4̄5 black. Hind tarsomere one white. Hind femur slightly flattened, hind tibia not sulcate.</paragraph>
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<paragraph id="8B8F1762FFDAA003FF5AF9C0ABA88D9B" blockId="34.[151,1437,1069,2033]" pageId="34" pageNumber="278">
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Wing uniformly darkly infuscated except for a transverse preapical pale band. Anal cell bare, CuA2 sinuate and distinctly proximal to dm̄cu, distal angle about 45
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<emphasis id="B944CB70FFDAA003FD4EF9F9AB938D9A" bold="true" box="[723,734,1648,1673]" pageId="34" pageNumber="278">°</emphasis>
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.
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</paragraph>
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<paragraph id="8B8F1762FFDAA003FF5AF91DAC5F8CE2" blockId="34.[151,1437,1069,2033]" pageId="34" pageNumber="278">
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Abdominal syntergite 1+2 slightly humped, T1 entirely silvery, T2 and T3 extensively silvery around margins; pleural membrane damaged on unique
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<typeStatus id="548BA9C0FFDAA003FDD5F930AB368DC2" box="[584,635,1721,1745]" pageId="34" pageNumber="278">type</typeStatus>
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, but apparently dark at base only. Male genital fork (S5) with very short distally tapered arms with inner distal margins densely packed with short, stout setae. Unlike all congeners except
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<taxonomicName id="4C306CE1FFDAA003FF0AF888A8308C0B" box="[151,381,1792,1816]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="278" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="B944CB70FFDAA003FF0AF888A8308C0B" box="[151,381,1792,1816]" italics="true" pageId="34" pageNumber="278">P. madagascariensis</emphasis>
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</taxonomicName>
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, the phallic plate extends far beyond the apex of the phallapodeme, with the basal part of the phallus forked and correspondingly extended posteriorly beyond the point of entry of the ejaculatory duct; extended part of phallic plate about one tenth as long as distiphallus beyond the phallapodeme. Distiphallus with basal part only slightly wider than distal part, ending in a simple phallic bulb with a spherical proximal part and a broad dorsal distal band; distal part of distiphallus separated from phallic bulb by a dorsal notch, straight, ending in a weakly differentiated but apically cupped glans. Blade of ejaculatory apodeme smaller than epandrium but much larger than sperm pump, ejaculatory apodeme including sperm pump subequal in length to epandrium.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFDBA002FF5AFF1EABB68BC6" pageId="35" pageNumber="279" type="materials_examined">
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<paragraph id="8B8F1762FFDBA002FF5AFF1EABB68BC6" blockId="35.[151,1436,151,213]" pageId="35" pageNumber="279">
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<emphasis id="B944CB70FFDBA002FF5AFF1EA8348BA3" bold="true" box="[199,377,151,176]" pageId="35" pageNumber="279">Type material.</emphasis>
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Holotype (♂, CAS, sequence MYCRO195̄15) Ranomafana National Park, Sahamalaotra, 21°14.44’ 47°23.67’,
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<date id="FF8E31A2FFDBA002FE16FF35A8B38BC6" box="[395,510,188,213]" pageId="35" pageNumber="279" value="2014-10-16">
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16.x.
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<quantity id="4CC8BA87FFDBA002FE23FF35A8B38BC6" box="[446,510,188,213]" metricMagnitude="6" metricUnit="m" metricValue="2.01465" pageId="35" pageNumber="279" unit="m" value="2014650.0">2014</quantity>
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</date>
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, 650m, S.A. Marshall.
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</paragraph>
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</subSubSection>
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<caption id="DF4F47EAFFDBA002FF0AF8ABA83E8C65" httpUri="https://zenodo.org/record/829045/files/figure.png" pageId="35" pageNumber="279" targetBox="[151,1436,286,1805]" targetPageId="35">
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<paragraph id="8B8F1762FFDBA002FF0AF8ABA83E8C65" blockId="35.[151,1436,1826,1910]" pageId="35" pageNumber="279">
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<emphasis id="B944CB70FFDBA002FF0AF8ABAB988C2B" bold="true" box="[151,725,1826,1849]" pageId="35" pageNumber="279">
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FIGURES 106–112.
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<taxonomicName id="4C306CE1FFDBA002FEE7F8AAAB3A8C2A" box="[378,631,1827,1849]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="279" phylum="Arthropoda" rank="species" species="zinzala" status="sp. nov.">
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<emphasis id="B944CB70FFDBA002FEE7F8AAAB3A8C2A" bold="true" box="[378,631,1827,1849]" italics="true" pageId="35" pageNumber="279">Paramimegralla zinzala</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277760BFFDBA002FD1CF8AAAB988C2B" box="[641,725,1827,1848]" pageId="35" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Male. 106. 107. Living fly along trail at Ranomafana; 108. Head, dorsal; 109. Head, lateral; 110. Epandrium and associated structures; left lateral 111. Abdominal segments 5‾8, lateral; 112. Phallus detail, lateral.
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</paragraph>
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</caption>
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<subSubSection id="C32A44E9FFDBA005FF5AF82AAA348BC6" lastPageId="36" lastPageNumber="280" pageId="35" pageNumber="279" type="discussion">
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<paragraph id="8B8F1762FFDBA005FF5AF82AAA348BC6" blockId="35.[151,1436,1955,2016]" lastBlockId="36.[151,1436,151,248]" lastPageId="36" lastPageNumber="280" pageId="35" pageNumber="279">
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<emphasis id="B944CB70FFDBA002FF5AF82AA8028CAF" bold="true" box="[199,335,1955,1980]" pageId="35" pageNumber="279">Comments.</emphasis>
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<taxonomicName id="4C306CE1FFDBA002FECAF82AAB258CA8" box="[343,616,1955,1979]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="279" phylum="Arthropoda" rank="species" species="zinzala">
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<emphasis id="B944CB70FFDBA002FECAF82AAB258CA8" box="[343,616,1955,1979]" italics="true" pageId="35" pageNumber="279">Paramimegralla zinzala</emphasis>
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</taxonomicName>
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is similar to, and obviously closely related to,
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<taxonomicName id="4C306CE1FFDBA002FBE6F82DAC2C8CA8" box="[1147,1377,1955,1979]" class="Insecta" family="Micropezidae" genus="Paramimegralla" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="279" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="B944CB70FFDBA002FBE6F82DAC2C8CA8" box="[1147,1377,1955,1979]" italics="true" pageId="35" pageNumber="279">P. madagascariensis</emphasis>
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</taxonomicName>
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. The dark wing with a distal clear band, the unusually short genital fork, tubular distiphallus apex, extended phallic plate and several other characters are unique to this group of two species, which differ in details of the distal distiphallus, length of phallic plate, leg colour, head colour and wing venation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A44E9FFDCA005FF5AFF56AABC8BEB" box="[199,1009,223,248]" pageId="36" pageNumber="280" type="etymology">
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<paragraph id="8B8F1762FFDCA005FF5AFF56AABC8BEB" blockId="36.[151,1436,151,248]" box="[199,1009,223,248]" pageId="36" pageNumber="280">
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<emphasis id="B944CB70FFDCA005FF5AFF56A80B8BEB" bold="true" box="[199,326,223,248]" pageId="36" pageNumber="280">Etymology</emphasis>
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. The species name is from the Latin for a kind of gnat or fly.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |