treatments-xml/data/03/93/91/0393914C2A67873EC282FD3A6AB9F9B2.xml

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<mods:title id="815788B48430E99D9EAA9BFD5839581C">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<taxonomicName id="4C3A5BD92A67873DC282FD3A6BE5F98F" ID-CoL="BDMR4" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[151,701,690,717]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="formosa" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A67873DC282FD3A69A1F98E" bold="true" box="[151,249,691,717]" italics="true" pageId="69" pageNumber="70">Stegana</emphasis>
(
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) 
<emphasis id="B94EFC482A67873DC380FD3B68A0F98F" bold="true" box="[405,504,690,716]" italics="true" pageId="69" pageNumber="70">formosa</emphasis>
Zhang &amp; Chen
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, 
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<paragraph id="8B85205A2A67873DC282FD5E683CF9B2" blockId="69.[151,808,690,753]" box="[151,356,727,753]" pageId="69" pageNumber="70">
(
<figureCitation id="13013CDF2A67873DC28AFD5E686BF9B2" box="[159,307,727,753]" captionStart="FIGURE 8" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[182,1418,198,1919]" captionTargetId="figure-19@31.[169,1418,181,1919]" captionTargetPageId="31" captionText="FIGURE 8. A, B. Stegana (Steganina) dendrophila Zhang &amp; Chen, sp. nov.; C, D. S. (S.) flabella Li &amp; Chen, sp. nov.; E, F. S. (S.) flavipes Li &amp; Chen, sp. nov.; G, H. S. (S.) formosa Zhang &amp; Chen, sp. nov.; I, J. S. (S.) fusca Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705683" httpUri="https://zenodo.org/record/7705683/files/figure.png" pageId="69" pageNumber="70">Figs. 8G, 8H</figureCitation>
, 
<figureCitation id="13013CDF2A67873DC32AFD5E6804F9B2" box="[319,348,727,753]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="69" pageNumber="70">39</figureCitation>
)
</paragraph>
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<subSubSection id="C32073D12A67873DC282FC97684BF8E6" pageId="69" pageNumber="70" type="diagnosis">
<paragraph id="8B85205A2A67873DC282FC97684BF8E6" blockId="69.[151,1437,798,933]" pageId="69" pageNumber="70">
<emphasis id="B94EFC482A67873DC282FC97684AF87B" bold="true" box="[151,274,798,824]" pageId="69" pageNumber="70">Diagnosis.</emphasis>
This species is similar to 
<emphasis id="B94EFC482A67873DC03BFC966B64F87A" box="[558,572,799,825]" italics="true" pageId="69" pageNumber="70">S</emphasis>
. (
<emphasis id="B94EFC482A67873DC047FC966B38F87A" box="[594,608,799,825]" italics="true" pageId="69" pageNumber="70">S</emphasis>
.) 
<emphasis id="B94EFC482A67873DC062FC966A58F87A" box="[631,768,799,825]" italics="true" pageId="69" pageNumber="70">clavispinula</emphasis>
Chen &amp; Chen, 
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the shape of male terminalia (as figs. 
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Chen &amp; Chen 2009, 
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), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth (
<figureCitation id="13013CDF2A67873DC2F5FCEE681EF8C2" box="[224,326,871,897]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="69" pageNumber="70">Fig. 39B</figureCitation>
); the pregonite small, with 3 sensilla (
<figureCitation id="13013CDF2A67873DC0FBFCEE6A2DF8C2" box="[750,885,871,897]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="69" pageNumber="70">Fig. 39C, D</figureCitation>
); the aedeagus distally broadened in ventral view (
<figureCitation id="13013CDF2A67873DC28AFC02685EF8E6" box="[159,262,907,933]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="69" pageNumber="70">Fig. 39C</figureCitation>
).
</paragraph>
</subSubSection>
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<emphasis id="B94EFC482A67873DC282F9416879FDA3" bold="true" box="[151,289,1736,1760]" pageId="69" pageNumber="70">FIGURE 39.</emphasis>
<taxonomicName id="4C3A5BD92A67873DC33CF9416BAEFDA3" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" box="[297,758,1736,1760]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="formosa" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A67873DC33CF9416823FD9C" box="[297,379,1736,1759]" italics="true" pageId="69" pageNumber="70">Stegana</emphasis>
(
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) 
<emphasis id="B94EFC482A67873DC015F9416B0BFDA3" box="[512,595,1736,1760]" italics="true" pageId="69" pageNumber="70">formosa</emphasis>
Zhang &amp; Chen
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, 
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<taxonomicNameLabel id="A27D41332A67873DC114F9416A0BFDA3" box="[769,851,1736,1760]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view.
</paragraph>
</caption>
<subSubSection id="C32073D12A67873EC2D2F8DF6A1DF97E" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="description">
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<emphasis id="B94EFC482A67873DC2D2F8DF6801FC33" bold="true" box="[199,345,1878,1904]" pageId="69" pageNumber="70">Description.</emphasis>
Male: First flagellomere brown. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally (
<figureCitation id="13013CDF2A67873DC317F8F36804FCD7" box="[258,348,1914,1940]" captionStart="FIGURE 8" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[182,1418,198,1919]" captionTargetId="figure-19@31.[169,1418,181,1919]" captionTargetPageId="31" captionText="FIGURE 8. A, B. Stegana (Steganina) dendrophila Zhang &amp; Chen, sp. nov.; C, D. S. (S.) flabella Li &amp; Chen, sp. nov.; E, F. S. (S.) flavipes Li &amp; Chen, sp. nov.; G, H. S. (S.) formosa Zhang &amp; Chen, sp. nov.; I, J. S. (S.) fusca Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705683" httpUri="https://zenodo.org/record/7705683/files/figure.png" pageId="69" pageNumber="70">Fig. 8G</figureCitation>
). Katepisternum yellowish (
<figureCitation id="13013CDF2A67873DC082F8F36BA9FCD7" box="[663,753,1914,1940]" captionStart="FIGURE 8" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[182,1418,198,1919]" captionTargetId="figure-19@31.[169,1418,181,1919]" captionTargetPageId="31" captionText="FIGURE 8. A, B. Stegana (Steganina) dendrophila Zhang &amp; Chen, sp. nov.; C, D. S. (S.) flabella Li &amp; Chen, sp. nov.; E, F. S. (S.) flavipes Li &amp; Chen, sp. nov.; G, H. S. (S.) formosa Zhang &amp; Chen, sp. nov.; I, J. S. (S.) fusca Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705683" httpUri="https://zenodo.org/record/7705683/files/figure.png" pageId="69" pageNumber="70">Fig. 8H</figureCitation>
). Scutellum dark brown (
<figureCitation id="13013CDF2A67873DC61EF8F36D3DFCD7" box="[1035,1125,1914,1940]" captionStart="FIGURE 8" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[182,1418,198,1919]" captionTargetId="figure-19@31.[169,1418,181,1919]" captionTargetPageId="31" captionText="FIGURE 8. A, B. Stegana (Steganina) dendrophila Zhang &amp; Chen, sp. nov.; C, D. S. (S.) flabella Li &amp; Chen, sp. nov.; E, F. S. (S.) flavipes Li &amp; Chen, sp. nov.; G, H. S. (S.) formosa Zhang &amp; Chen, sp. nov.; I, J. S. (S.) fusca Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705683" httpUri="https://zenodo.org/record/7705683/files/figure.png" pageId="69" pageNumber="70">Fig. 8G</figureCitation>
). Halter mostly brown (Fig. Epandrium with 
<emphasis id="B94EFC482A64873EC340FF1E6837FBF3" box="[341,367,151,176]" italics="true" pageId="70" pageNumber="71">ca</emphasis>
. 26 setae on dorsal to posterior portions per side (
<figureCitation id="13013CDF2A64873EC1BCFF1E6D4CFBF2" box="[937,1044,151,177]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39A</figureCitation>
). Surstylus with numerous minute, serrated processes on outer surface and setulae along distal margin (
<figureCitation id="13013CDF2A64873EC1B4FF326D74FB96" box="[929,1068,187,213]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39A, B</figureCitation>
). Cercus with much pubescence (
<figureCitation id="13013CDF2A64873EC28AFF566853FBBA" box="[159,267,223,249]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39A</figureCitation>
). Hypandrium expanded anteromedially (
<figureCitation id="13013CDF2A64873EC0F7FF566A36FBBA" box="[738,878,223,249]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39C, D</figureCitation>
). Aedeagal sheath distally slightly dorsad curved and with several minute, serrated processes (
<figureCitation id="13013CDF2A64873EC08BFE8A6A75FA5E" box="[670,813,259,285]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39C, D</figureCitation>
). Aedeagus with numerous minute, serrated processes distally (
<figureCitation id="13013CDF2A64873EC2ECFEAE68DAFA02" box="[249,386,295,321]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39C, D</figureCitation>
). Subepandrial sclerite slightly vaulted, rounded distally (
<figureCitation id="13013CDF2A64873EC61DFEAE6D36FA02" box="[1032,1134,295,321]" captionStart="FIGURE 39" captionStartId="69.[151,250,1736,1760]" captionTargetBox="[157,1428,967,1702]" captionTargetId="figure-275@69.[151,1436,957,1713]" captionTargetPageId="69" captionText="FIGURE 39. Stegana (Steganina) formosa Zhang &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; E. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705762" httpUri="https://zenodo.org/record/7705762/files/figure.png" pageId="70" pageNumber="71">Fig. 39E</figureCitation>
).
</paragraph>
<paragraph id="8B85205A2A64873EC2D2FEC26A1DF97E" blockId="70.[151,1437,151,753]" pageId="70" pageNumber="71">
<emphasis id="B94EFC482A64873EC2D2FEC26B55FA27" bold="true" box="[199,525,330,357]" pageId="70" pageNumber="71">Measurements and indices.</emphasis>
BL = 
<quantity id="4CC28DBF2A64873EC042FEC36BE4FA26" box="[599,700,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="70" pageNumber="71" unit="mm" value="2.45">2.45 mm</quantity>
in 
<typeStatus id="54819EF82A64873EC0F7FEC26A1CFA26" box="[738,836,331,357]" pageId="70" pageNumber="71" type="holotype">holotype</typeStatus>
(
<quantity id="4CC28DBF2A64873EC146FEC36ABBFA26" box="[851,995,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="6.8834" metricValueMax="7.493" metricValueMin="6.2738" pageId="70" pageNumber="71" unit="in" value="2.71" valueMax="2.95" valueMin="2.47">2.47–2.95 in</quantity>
3♂ 
<typeStatus id="54819EF82A64873EC600FEC26DDDFA26" box="[1045,1157,331,357]" pageId="70" pageNumber="71" type="paratype">paratypes</typeStatus>
), ThL = 
<quantity id="4CC28DBF2A64873EC6F2FEC26C14FA26" box="[1255,1356,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="70" pageNumber="71" unit="mm" value="1.13">1.13 mm</quantity>
(1.17– 1.36), WL = 
<quantity id="4CC28DBF2A64873EC33EFEE668C8FACA" box="[299,400,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="70" pageNumber="71" unit="mm" value="1.79">1.79 mm</quantity>
(1.91–2.31), WW = 
<quantity id="4CC28DBF2A64873EC06BFEE66BBBFACA" box="[638,739,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="70" pageNumber="71" unit="mm" value="0.91">0.91 mm</quantity>
(0.87–1.05), arb = 5/4 (6/4–7/5), avd = 0.74 (0.71–0.93), adf = 0.91 (0.93–1.16), flw = 1.83 (1.64–1.89), FW/HW = 0.37 (0.36–0.40), ch/o = 0.29 (0.26–0.30), prorb = 1.24 (0.92–1.25), rcorb = 0.77 (0.66–0.79), vb = 0.68 (0.59–0.72), dcl = 0.79 (0.69–0.81), presctl = 0.71 (0.57–0.62), sctl = 1.62 (1.34–1.96), sterno = 1.13 (0.72–0.96), orbito = 1.73 (1.68–2.00), dcp = 0.34 (0.36–0.44), sctlp = 1.35 (1.67–1.84), C = 2.58 (1.72–1.97), 4c = 1.00 (1.15–1.24), 4v = 1.71 (1.51–2.00), 5x = 1.32 (1.34–1.58), ac = 7.67 (9.89–10.75), M = 0.59 (0.61–0.66), C3F = 0.61 (0.60–0.62).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A64873EC2D2FDCE688EF9EA" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="8B85205A2A64873EC2D2FDCE688EF9EA" blockId="70.[151,1437,151,753]" pageId="70" pageNumber="71">
<emphasis id="B94EFC482A64873EC2D2FDCE6826F923" bold="true" box="[199,382,582,609]" pageId="70" pageNumber="71">Type materials.</emphasis>
<materialsCitation id="3B522A072A64873EC396FDCE6A6FF9C6" ID-GBIF-Occurrence="4097216376" collectingDate="2012-10-30" collectionCode="SCAU" collectorName="H. W. Chen" country="China" county="Taiwan" elevation="500" latitude="22.764444" location="Chipen" longLatPrecision="21" longitude="121.113335" municipality="Taitung" pageId="70" pageNumber="71" specimenCount="1" typeStatus="holotype">
<typeStatus id="54819EF82A64873EC396FDCE68B3F922" box="[387,491,583,609]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED2BB89F2A64873EC004FDCE6B38F922" box="[529,608,583,609]" pageId="70" pageNumber="71">SCAU</collectionCode>
, No. 123317, DNA #1490), 
<collectingCountry id="F32D60CA2A64873EC186FDCE6AA8F922" box="[915,1008,583,609]" name="China" pageId="70" pageNumber="71">CHINA</collectingCountry>
: 
<location id="8EE576812A64873EC1EDFDCE6D14F922" LSID="urn:lsid:plazi:treatment:0393914C2A67873EC282FD3A6AB9F9B2:8EE576812A64873EC1EDFDCE6D14F922" box="[1016,1100,583,609]" country="China" county="Taiwan" latitude="22.764444" longLatPrecision="21" longitude="121.113335" municipality="Taitung" name="Chipen" pageId="70" pageNumber="71">Chipen</location>
, 
<collectingMunicipality id="6BE1BA202A64873EC641FDCE6DF4F922" box="[1108,1196,583,609]" pageId="70" pageNumber="71">Taitung</collectingMunicipality>
, 
<collectingCounty id="62E458D62A64873EC6A0FDCE6C52F922" box="[1205,1290,583,609]" pageId="70" pageNumber="71">Taiwan</collectingCounty>
, 
<geoCoordinate id="EE0E469D2A64873EC706FDCF6CC0F922" box="[1299,1432,582,609]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="70" pageNumber="71" precision="15" seconds="52" value="22.764444">22°45'52''N</geoCoordinate>
, 
<geoCoordinate id="EE0E469D2A64873EC282FDE2687FF9C6" box="[151,295,619,645]" degrees="121" direction="east" minutes="06" orientation="longitude" pageId="70" pageNumber="71" precision="15" seconds="48" value="121.113335">121°06'48''E</geoCoordinate>
, alt. 
<quantity id="4CC28DBF2A64873EC34CFDE368C4F9C6" box="[345,412,618,645]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">
<elevation id="0017C7692A64873EC34CFDE368C4F9C6" box="[345,412,618,645]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500m</elevation>
</quantity>
, 
<date id="FF84069A2A64873EC3B0FDE26B40F9C6" box="[421,536,619,645]" pageId="70" pageNumber="71" value="2012-10-30">
<collectingDate id="EFC0FF722A64873EC3B0FDE26B40F9C6" box="[421,536,619,645]" pageId="70" pageNumber="71" value="2012-10-30">30.x.2012</collectingDate>
</date>
, ex tree trunk, 
<collectorName id="26CF458C2A64873EC0ADFDE26A6FF9C6" box="[696,823,619,645]" pageId="70" pageNumber="71">H.W. Chen</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B522A072A64873EC12AFDE2688EF9EA" ID-GBIF-Occurrence="4097216492" collectingDate="2012-10-30" collectionCode="SCAU" collectorName="H. W. Chen" country="China" county="Taiwan" elevation="500" latitude="22.764444" location="Chipen" longLatPrecision="21" longitude="121.113335" municipality="Taitung" pageId="70" pageNumber="71" specimenCount="1" typeStatus="paratype">
<typeStatus id="54819EF82A64873EC12AFDE26AE8F9C6" box="[831,944,619,645]" pageId="70" pageNumber="71" type="paratype">Paratypes</typeStatus>
: 3♂ (
<collectionCode id="ED2BB89F2A64873EC1FCFDE26D60F9C6" box="[1001,1080,619,645]" pageId="70" pageNumber="71">SCAU</collectionCode>
, Nos. 123318–20, DNA #2737), other data same as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A64873EC2D2FD3B6B46F98E" box="[199,542,690,717]" pageId="70" pageNumber="71" type="distribution">
<paragraph id="8B85205A2A64873EC2D2FD3B6B46F98E" blockId="70.[151,1437,151,753]" box="[199,542,690,717]" pageId="70" pageNumber="71">
<emphasis id="B94EFC482A64873EC2D2FD3B6839F98F" bold="true" box="[199,353,690,716]" pageId="70" pageNumber="71">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A64873EC372FD3A68F2F98E" box="[359,426,691,717]" name="China" pageId="70" pageNumber="71">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A64873EC3ACFD3A6B42F98E" box="[441,538,691,717]" country="China" name="Taiwan)" pageId="70" pageNumber="71">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A64873EC2D2FD5E6AB9F9B2" box="[199,993,727,753]" pageId="70" pageNumber="71" type="etymology">
<paragraph id="8B85205A2A64873EC2D2FD5E6AB9F9B2" blockId="70.[151,1437,151,753]" box="[199,993,727,753]" pageId="70" pageNumber="71">
<emphasis id="B94EFC482A64873EC2D2FD5E6816F9B2" bold="true" box="[199,334,727,753]" pageId="70" pageNumber="71">Etymology.</emphasis>
From the name of the 
<typeStatus id="54819EF82A64873EC058FD5E6B25F9B2" box="[589,637,727,753]" pageId="70" pageNumber="71">type</typeStatus>
locality &quot; 
<collectingCountry id="F32D60CA2A64873EC0F2FD5E6A08F9B2" box="[743,848,727,753]" name="Taiwan" pageId="70" pageNumber="71">Formosa</collectingCountry>
&quot; (= Taiwan).
</paragraph>
</subSubSection>
</treatment>
</document>