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<document id="49192DF47622A063ACE3DB51A5D3AEAD" ID-DOI="10.5281/zenodo.5386711" ID-ISSN="2345-7600" ID-Zenodo-Dep="5386711" ID-ZooBank="BAA493D3-8195-417B-ABD8-A36D3FD90533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303402995" checkinUser="valdenar" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2015" docId="039387F0C662FF97ADB9FC990B45FBA4" docLanguage="en" docName="63rbz564-582.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Lepidothelphusa flavochela Grinang &amp; Ng 2015, new species" docType="treatment" docUuid="2B94C97A-7093-4DB8-983C-13765B6923D8" docUuidSource="ZooBank" docVersion="5" lastPageNumber="568" masterDocId="FFAAFF88C666FF93AD69FFE40E06FFB1" masterDocTitle="Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo" masterLastPageNumber="582" masterPageNumber="564" pageNumber="568" updateTime="1699221322658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D8448A5D521A83046AF022357D39DDCD">Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo</mods:title>
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<mods:namePart id="0AAA96B78063E1C8052489946040FE6B">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="4D80782922D48AD1ACF226766BD446F0">Ng, Peter K. L.</mods:namePart>
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<mods:title id="221668B1C57F0C29CB3D698762ED1E59">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="7EE0FF63996A74CFF5C3379EB5F58838">2015</mods:date>
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<mods:number id="07AD76BE0EEEEE24D3A4D90E3E05853B">2015-12-18</mods:number>
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<treatment id="039387F0C662FF97ADB9FC990B45FBA4" ID-DOI="http://doi.org/10.5281/zenodo.5467450" ID-Zenodo-Dep="5467450" LSID="urn:lsid:zoobank.org:act:2B94C97A-7093-4DB8-983C-13765B6923D8" httpUri="http://treatment.plazi.org/id/039387F0C662FF97ADB9FC990B45FBA4" lastPageNumber="568" pageId="4" pageNumber="568">
<subSubSection id="C320656DC662FF97ADB9FC990C92FC24" box="[208,660,893,917]" pageId="4" pageNumber="568" type="nomenclature">
<paragraph id="8B8536E6C662FF97ADB9FC990C92FC24" blockId="4.[208,660,893,949]" box="[208,660,893,917]" pageId="4" pageNumber="568">
<heading id="D0CD818AC662FF97ADB9FC990C92FC24" bold="true" box="[208,660,893,917]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="568" reason="2">
<emphasis id="B94EEAF4C662FF97ADB9FC990C92FC24" bold="true" box="[208,660,893,917]" pageId="4" pageNumber="568">
<taxonomicName id="4C3A4D65C662FF97ADB9FC990FF9FC24" ID-CoL="3TDTG" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[208,511,893,917]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="flavochela" status="sp. nov.">
<emphasis id="B94EEAF4C662FF97ADB9FC990FF9FC24" bold="true" box="[208,511,893,917]" italics="true" pageId="4" pageNumber="568">Lepidothelphusa flavochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC662FF97AF67FC990C92FC24" box="[526,660,893,917]" pageId="4" pageNumber="568" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C320656DC662FF97AC71FC790C4CFC04" box="[280,586,925,949]" pageId="4" pageNumber="568" type="description">
<paragraph id="8B8536E6C662FF97AC71FC790C4CFC04" blockId="4.[208,660,893,949]" box="[280,586,925,949]" pageId="4" pageNumber="568">
(
<figureCitation id="13012A63C662FF97AC49FC790F91FC04" box="[288,407,925,949]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Figs. 3AI</figureCitation>
,
<figureCitation id="13012A63C662FF97ACCDFC790FE3FC04" box="[420,485,925,949]" captionStart="Fig" captionStartId="6.[112,147,1068,1090]" captionTargetBox="[117,1453,190,1052]" captionTargetId="figure-12@6.[91,1473,171,1063]" captionTargetPageId="6" captionText="Fig. 4. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386727" httpUri="https://zenodo.org/record/5386727/files/figure.png" pageId="4" pageNumber="568">4AE</figureCitation>
,
<figureCitation id="13012A63C662FF97AC9BFC790C44FC04" box="[498,578,925,949]" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.0038290), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.0040205), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="4" pageNumber="568">13C, D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320656DC662FF97AD18FC390ED0FB24" pageId="4" pageNumber="568" type="materials_examined">
<paragraph id="8B8536E6C662FF97AD18FC390ED0FB24" blockId="4.[113,753,989,1173]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AD18FC390F57FC44" bold="true" box="[113,337,989,1013]" pageId="4" pageNumber="568">Material examined.</emphasis>
<materialsCitation id="3B523CBBC662FF97AC30FC390EACFBE4" collectingDate="2013-02-15" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" elevation="194" location="Sarawak" pageId="4" pageNumber="568" specimenCode="ZRC 2015.0604" specimenCount="1" typeStatus="holotype">
<typeStatus id="54818844C662FF97AC30FC390FC6FC44" box="[345,448,989,1013]" pageId="4" pageNumber="568" type="holotype">Holotype</typeStatus>
: male (8.1 ×
<quantity id="4CC29B03C662FF97AF39FC390CA2FC44" box="[592,676,989,1013]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="568" unit="mm" value="7.2">7.2 mm</quantity>
) (
<specimenCode id="DB9C9E9DC662FF97AFD2FC390EE1FBA4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="568">ZRC 2015.0604</specimenCode>
),
<elevation id="0017D1D5C662FF97AD92FC190FF2FBA4" box="[251,500,1021,1045]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="4" pageNumber="568" unit="m" value="194.0">
<quantity id="4CC29B03C662FF97AD92FC190F44FBA4" box="[251,322,1021,1045]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="4" pageNumber="568" unit="m" value="194.0">194 m</quantity>
above sea level
</elevation>
, Pagar Besi, Kampung Gumbang, Bau,
<collectingCountry id="F32D7676C662FF97AC49FBF90F79FB84" box="[288,383,1053,1077]" name="Malaysia" pageId="4" pageNumber="568">Sarawak</collectingCountry>
, coll.
<collectorName id="26CF5330C662FF97ACD7FBFA0C20FB84" box="[446,550,1053,1077]" pageId="4" pageNumber="568">J Grinang</collectorName>
et al.,
<date id="FF841026C662FF97AF06FBF90EACFBE4" pageId="4" pageNumber="568" value="2013-02-15">
<collectingDate id="EFC0E9CEC662FF97AF06FBF90EACFBE4" pageId="4" pageNumber="568" value="2013-02-15">15 February 2013</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54818844C662FF97ADDCFBDA0F27FBE7" box="[181,289,1086,1110]" pageId="4" pageNumber="568" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3CFD6FC662FF97AC45FBD90F79FBE4" box="[300,383,1085,1109]" pageId="4" pageNumber="568" type="male">4 males</specimenCount>
(largest 8.5 ×
<quantity id="4CC29B03C662FF97AF77FBD90C72FBE4" box="[542,628,1085,1109]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="568" unit="mm" value="7.5">7.5 mm</quantity>
),
<materialsCitation id="3B523CBBC662FF97AFE1FBD90ED4FB24" collectingDate="2013-02-15" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" elevation="194" location="Sarawak" pageId="4" pageNumber="568" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<specimenCount id="9D3CFD6FC662FF97AFE1FBD90CF6FBE4" box="[648,752,1085,1109]" pageId="4" pageNumber="568" type="female">5 females</specimenCount>
(largest 9.9 ×
<quantity id="4CC29B03C662FF97AC66FBB90F60FBC4" box="[271,358,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="568" unit="mm" value="8.8">8.8 mm</quantity>
) (SBC.C.00392401), data same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC662FF97AD18FB590ED6F7A4" pageId="4" pageNumber="568" type="diagnosis">
<paragraph id="8B8536E6C662FF97AD18FB590C1CF8E4" blockId="4.[113,753,1213,1878]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AD18FB590EE0FB64" bold="true" box="[113,230,1213,1237]" pageId="4" pageNumber="568">Diagnosis.</emphasis>
Carapace quadrate, surface relatively flat, rugose, anterolateral margins smooth; antero- and posterolateral regions slightly rugose; epibranchial tooth undiscernible; infra-, supra-orbital margins confluent gradually, external orbital tooth with outer margin concave, smooth; serrated projection on frontal region distinct; epigastric and postorbital cristae distinct, relatively low, margins serrated; cervical and H-grooves relatively shallow, not confluent (
<figureCitation id="13012A63C662FF97AF27FA790CE5FA04" box="[590,739,1437,1461]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3A, B, D</figureCitation>
); ischium of third maxilliped rhomboidal, much longer than broad, sulcus undiscernible (
<figureCitation id="13012A63C662FF97ACC8FA390FF7FA44" box="[417,497,1501,1525]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3F</figureCitation>
); inner margin of merus of chelipeds with relatively lower serrated projection (
<figureCitation id="13012A63C662FF97AFACFA190E8CF984" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3I</figureCitation>
); carpus of chelipeds smooth, inner angle with broad, low triangular tooth (
<figureCitation id="13012A63C662FF97AC4FF9D90F7DF9E4" box="[294,379,1597,1621]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3A</figureCitation>
); fingers of adult major male chela gaping (
<figureCitation id="13012A63C662FF97ADA0F9B90F46F9C4" box="[201,320,1629,1653]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3G, H</figureCitation>
); ambulatory legs relatively long, merus of fourth ambulatory legs subequal to length of carapace (
<figureCitation id="13012A63C662FF97AFAFF9990EB4F904" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3A, E</figureCitation>
); male abdomen triangular, somite 6 width and length subequal, subequal to length of telson (
<figureCitation id="13012A63C662FF97AF49F9590C70F964" box="[544,630,1725,1749]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3C</figureCitation>
); G1 stout, sinuous in ventral view, straight in dorsal view; terminal segment straight in dorsal view, curved in ventral view (
<figureCitation id="13012A63C662FF97AFACF9190EF3F884" captionStart="Fig" captionStartId="6.[112,147,1068,1090]" captionTargetBox="[117,1453,190,1052]" captionTargetId="figure-12@6.[91,1473,171,1063]" captionTargetPageId="6" captionText="Fig. 4. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386727" httpUri="https://zenodo.org/record/5386727/files/figure.png" pageId="4" pageNumber="568">Fig. 4A, B, D, E</figureCitation>
); G2 with relatively long distal segment, about half length of basal segment (
<figureCitation id="13012A63C662FF97ACDEF8D90C0BF8E4" box="[439,525,1853,1877]" captionStart="Fig" captionStartId="6.[112,147,1068,1090]" captionTargetBox="[117,1453,190,1052]" captionTargetId="figure-12@6.[91,1473,171,1063]" captionTargetPageId="6" captionText="Fig. 4. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386727" httpUri="https://zenodo.org/record/5386727/files/figure.png" pageId="4" pageNumber="568">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C662FF97AD18F8990ED6F7A4" blockId="4.[113,753,1917,2069]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AD18F8990ECEF824" bold="true" box="[113,200,1917,1941]" pageId="4" pageNumber="568">Colour.</emphasis>
In life, males have uniformly yellow chela, creamwhite ambulatory legs and the anterior part of the carapace is yellowish-orange with the posterior part pale orange to yellowish-white. Females are dark brown overall (
<figureCitation id="13012A63C662FF97AFACF8390EC5F7A4" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.0038290), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.0040205), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="4" pageNumber="568">Fig. 13C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320656DC662FF97AE44FF590AFDFEA4" pageId="4" pageNumber="568" type="etymology">
<paragraph id="8B8536E6C662FF97AE44FF590AFDFEA4" blockId="4.[813,1452,189,277]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AE44FF590DB7FF64" bold="true" box="[813,945,189,213]" pageId="4" pageNumber="568">Etymology.</emphasis>
The name is derived from the Latin “flavo” combined with chela, alluding to the entire yellowish chela. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC662FF97AE44FED90B45FBA4" pageId="4" pageNumber="568" type="discussion">
<paragraph id="8B8536E6C662FF97AE44FED90B92FC04" blockId="4.[813,1454,317,950]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AE44FED90D9DFEE4" bold="true" box="[813,923,317,341]" pageId="4" pageNumber="568">Remarks.</emphasis>
<taxonomicName id="4C3A4D65C662FF97AEC8FED90AC1FEE4" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[929,1223,317,341]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="flavochela" status="sp. nov.">
<emphasis id="B94EEAF4C662FF97AEC8FED90AC1FEE4" box="[929,1223,317,341]" italics="true" pageId="4" pageNumber="568">Lepidothelphusa flavochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC662FF97A9BAFEDA0B50FEE4" box="[1235,1366,317,341]" pageId="4" pageNumber="568" rank="species">new species</taxonomicNameLabel>
, is most easily distinguished from
<taxonomicName id="4C3A4D65C662FF97A923FEBA0AC3FEC4" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1098,1221,350,373]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" sensu="stricto" species="cognettii">
<emphasis id="B94EEAF4C662FF97A923FEBA0AC3FEC4" box="[1098,1221,350,373]" italics="true" pageId="4" pageNumber="568">L. cognettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D578FC662FF97A9A5FEBA0B0EFEC4" box="[1228,1288,350,373]" pageId="4" pageNumber="568" sensu="stricto">s. str.</taxonomicNameLabel>
and congeners in having poorly developed epigastric and postorbital cristae (
<figureCitation id="13012A63C662FF97AE5CFE790DAEFE04" box="[821,936,413,437]" captionStart="Fig" captionStartId="5.[136,171,2006,2028]" captionTargetBox="[256,1353,189,1991]" captionTargetId="figure-90@5.[256,1354,189,1991]" captionTargetPageId="5" captionText="Fig. 3. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, outer view of right chela; H, outer view of left chela; I, left cheliped showing serrated inner margin of merus." figureDoi="http://doi.org/10.5281/zenodo.5386723" httpUri="https://zenodo.org/record/5386723/files/figure.png" pageId="4" pageNumber="568">Fig. 3B, D</figureCitation>
versus
<figureCitation id="13012A63C662FF97AE94FE790A7AFE04" box="[1021,1148,413,437]" captionStart="Fig" captionStartId="2.[113,149,2006,2028]" captionTargetBox="[190,1374,189,1991]" captionTargetId="figure-93@2.[190,1374,189,1991]" captionTargetPageId="2" captionText="Fig. 1. Lepidothelphusa cognettii (Nobili, 1903), male (12.2 × 10.2 mm) (ZRC 2015.0601), Batu Panggah Trail, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386715" httpUri="https://zenodo.org/record/5386715/files/figure.png" pageId="4" pageNumber="568">Figs. 1B, D</figureCitation>
,
<figureCitation id="13012A63C662FF97A9E0FE790ACAFE04" box="[1161,1228,413,437]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="4" pageNumber="568">5B, D</figureCitation>
,
<figureCitation id="13012A63C662FF97A9B0FE7A0B1AFE04" box="[1241,1308,414,437]" captionStart="Fig" captionStartId="9.[136,171,2006,2028]" captionTargetBox="[235,1376,189,1991]" captionTargetId="figure-91@9.[235,1376,189,1991]" captionTargetPageId="9" captionText="Fig. 7. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386737" httpUri="https://zenodo.org/record/5386737/files/figure.png" pageId="4" pageNumber="568">7B, D</figureCitation>
,
<figureCitation id="13012A63C662FF97A840FE790B6DFE04" box="[1321,1387,413,437]" captionStart="Fig" captionStartId="11.[136,171,2006,2028]" captionTargetBox="[218,1391,189,1991]" captionTargetId="figure-88@11.[218,1393,189,1991]" captionTargetPageId="11" captionText="Fig. 9. Lepidothelphusa padawan, new species, holotype male (13.6 × 12.5 mm) (ZRC 2015.0608), Kampung Sentah, Padawan, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386743" httpUri="https://zenodo.org/record/5386743/files/figure.png" pageId="4" pageNumber="568">9B, D</figureCitation>
,
<figureCitation id="13012A63C662FF97A811FE790D45FE64" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="4" pageNumber="568">11B, D</figureCitation>
). Its coloration in life is closest to
<taxonomicName id="4C3A4D65C662FF97A9D5FE5A0B13FE64" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1212,1301,445,469]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="limau" status="sp. nov.">
<emphasis id="B94EEAF4C662FF97A9D5FE5A0B13FE64" box="[1212,1301,445,469]" italics="true" pageId="4" pageNumber="568">L. limau</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC662FF97A84DFE5A0BAEFE64" box="[1316,1448,445,469]" pageId="4" pageNumber="568" rank="species">new species</taxonomicNameLabel>
, especially with the dorsal surface of adults having the anterior half orange and posterior part lighter; and the legs cream-white. The chelae are also similar in colour except that in
<taxonomicName id="4C3A4D65C662FF97AE10FDDA0A05FDE4" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[889,1027,573,597]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="flavochela">
<emphasis id="B94EEAF4C662FF97AE10FDDA0A05FDE4" box="[889,1027,573,597]" italics="true" pageId="4" pageNumber="568">L. flavochela</emphasis>
</taxonomicName>
, even the fingers are yellow (
<figureCitation id="13012A63C662FF97A82AFDD90BA5FDE4" box="[1347,1443,573,597]" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.0038290), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.0040205), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="4" pageNumber="568">Fig. 13C</figureCitation>
) while in
<taxonomicName id="4C3A4D65C662FF97AEFCFDBA0DF5FDC4" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[917,1011,605,629]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="limau">
<emphasis id="B94EEAF4C662FF97AEFCFDBA0DF5FDC4" box="[917,1011,605,629]" italics="true" pageId="4" pageNumber="568">L. limau</emphasis>
</taxonomicName>
, the fingers are white (
<figureCitation id="13012A63C662FF97A86AFDB90B6CFDC4" box="[1283,1386,605,629]" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.0038290), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.0040205), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="4" pageNumber="568">Fig. 13E</figureCitation>
). The yellow chela of
<taxonomicName id="4C3A4D65C662FF97AE88FD9A0A69FD24" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[993,1135,637,661]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="flavochela">
<emphasis id="B94EEAF4C662FF97AE88FD9A0A69FD24" box="[993,1135,637,661]" italics="true" pageId="4" pageNumber="568">L. flavochela</emphasis>
</taxonomicName>
(
<figureCitation id="13012A63C662FF97A9E9FD990B00FD24" box="[1152,1286,637,661]" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.0038290), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.0040205), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="4" pageNumber="568">Fig. 13C, D</figureCitation>
) is a character shared by
<taxonomicName id="4C3A4D65C662FF97AEC9FD7A0DDAFD04" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[928,988,669,693]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="loi">
<emphasis id="B94EEAF4C662FF97AEC9FD7A0DDAFD04" box="[928,988,669,693]" italics="true" pageId="4" pageNumber="568">L. loi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D65C662FF97A97FFD7A0A84FD04" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1046,1154,670,693]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="sangon">
<emphasis id="B94EEAF4C662FF97A97FFD7A0A84FD04" box="[1046,1154,670,693]" italics="true" pageId="4" pageNumber="568">L. sangon</emphasis>
</taxonomicName>
but these species have the carapace a more uniform reddish-brown to orange and the legs are reddish-brown (
<figureCitation id="13012A63C662FF97A954FD390B12FD44" box="[1085,1300,733,757]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.004404), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="4" pageNumber="568">Figs. 14A, B, E, F</figureCitation>
). The G1s of these species are quite different. The terminal segment of the G1 of
<taxonomicName id="4C3A4D65C662FF97AEF1FCFA0A24FC84" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[920,1058,797,821]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="flavochela">
<emphasis id="B94EEAF4C662FF97AEF1FCFA0A24FC84" box="[920,1058,797,821]" italics="true" pageId="4" pageNumber="568">L. flavochela</emphasis>
</taxonomicName>
(
<figureCitation id="13012A63C662FF97A959FCF90AE6FC84" box="[1072,1248,797,821]" captionStart="Fig" captionStartId="6.[112,147,1068,1090]" captionTargetBox="[117,1453,190,1052]" captionTargetId="figure-12@6.[91,1473,171,1063]" captionTargetPageId="6" captionText="Fig. 4. Lepidothelphusa flavochela, new species, holotype male (8.1 × 7.2 mm) (ZRC 2015.0604), Pagar Besi, Kampung Gumbang, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386727" httpUri="https://zenodo.org/record/5386727/files/figure.png" pageId="4" pageNumber="568">Fig. 4A, B, D, E</figureCitation>
) is proportionately longer than that of
<taxonomicName id="4C3A4D65C662FF97A96DFCDA0A66FCE4" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1028,1120,829,853]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="limau">
<emphasis id="B94EEAF4C662FF97A96DFCDA0A66FCE4" box="[1028,1120,829,853]" italics="true" pageId="4" pageNumber="568">L. limau</emphasis>
</taxonomicName>
(
<figureCitation id="13012A63C662FF97A918FCD90B28FCE4" box="[1137,1326,829,853]" captionStart="Fig" captionStartId="8.[113,148,958,980]" captionTargetBox="[114,1452,188,944]" captionTargetId="figure-12@8.[100,1462,177,951]" captionTargetPageId="8" captionText="Fig. 6. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386733" httpUri="https://zenodo.org/record/5386733/files/figure.png" pageId="4" pageNumber="568">Fig. 6A, B, D, E</figureCitation>
) and
<taxonomicName id="4C3A4D65C662FF97A819FCDA0BADFCE4" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1392,1451,829,853]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="loi">
<emphasis id="B94EEAF4C662FF97A819FCDA0BADFCE4" box="[1392,1451,829,853]" italics="true" pageId="4" pageNumber="568">L. loi</emphasis>
</taxonomicName>
(
<figureCitation id="13012A63C662FF97AE5CFCB90DECFCC4" box="[821,1002,861,885]" captionStart="Fig" captionStartId="10.[113,148,1287,1309]" captionTargetBox="[115,1450,189,1271]" captionTargetId="figure-12@10.[95,1468,178,1283]" captionTargetPageId="10" captionText="Fig. 8. Lepidothelphusa loi, new species, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386739" httpUri="https://zenodo.org/record/5386739/files/figure.png" pageId="4" pageNumber="568">Fig. 8A, B, D, E</figureCitation>
), while in
<taxonomicName id="4C3A4D65C662FF97A934FCBA0AC1FCC4" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1117,1223,862,885]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" species="sangon">
<emphasis id="B94EEAF4C662FF97A934FCBA0AC1FCC4" box="[1117,1223,862,885]" italics="true" pageId="4" pageNumber="568">L. sangon</emphasis>
</taxonomicName>
, it is much straighter with the distal opening more flared and the lateral margins of the subterminal segment sinuous (
<figureCitation id="13012A63C662FF97A9D6FC790B8EFC04" box="[1215,1416,925,949]" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="4" pageNumber="568">Fig. 12A, B, D, E</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C662FF97AE44FC390B45FBA4" blockId="4.[813,1452,989,1045]" pageId="4" pageNumber="568">
<emphasis id="B94EEAF4C662FF97AE44FC390D8DFC44" bold="true" box="[813,907,989,1013]" pageId="4" pageNumber="568">Habitat.</emphasis>
The habitat is similar to that of
<taxonomicName id="4C3A4D65C662FF97A986FC3A0B6CFC44" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1263,1386,990,1013]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="568" phylum="Arthropoda" rank="species" sensu="stricto" species="cognettii">
<emphasis id="B94EEAF4C662FF97A986FC3A0B6CFC44" box="[1263,1386,990,1013]" italics="true" pageId="4" pageNumber="568">L. cognettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D578FC662FF97A818FC3A0BAAFC44" box="[1393,1452,990,1013]" pageId="4" pageNumber="568" sensu="stricto">s. str.</taxonomicNameLabel>
except that it is less than
<quantity id="4CC29B03C662FF97A92FFC190A8AFBA4" box="[1094,1164,1021,1045]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="568" unit="m" value="200.0">200 m</quantity>
above sea level.
</paragraph>
</subSubSection>
</treatment>
</document>