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<document id="D9D29D31A0863235EC51B5955992CF31" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706805705" checkinUser="felipe" docAuthor="Hara, Marcos Ryotaro &amp; Pinto-Da-Rocha, Ricardo" docDate="2010" docId="039387E595654628FF65FE658E75FB6B" docLanguage="en" docName="zt02698p136.pdf" docOrigin="Zootaxa 2698" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eusarcus geometricus Hara &amp; Pinto-Da-Rocha 2010, sp. n." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFAAFF9D955B4668FFF2FFA78A56FF9F" masterDocTitle="Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698" masterLastPageNumber="136" masterPageNumber="1" pageNumber="63" updateTime="1699098874055" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7F067D625A763F035D5301DBE933E572" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3D55ACFCAB41A2161A879FDA52C3ADFC">Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698</mods:title>
</mods:titleInfo>
<mods:name id="5DFB000803F5C9C2207964754396F8D4" type="personal">
<mods:role id="BC2110586C3D23840D0E27335E59C16C">
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<mods:namePart id="8DF2074FAC52CB1FA4F07D7007B6D121">Hara, Marcos Ryotaro</mods:namePart>
</mods:name>
<mods:name id="31F86FBD832A2B29E2CF07CEF2E4B429" type="personal">
<mods:role id="4777D23A9193243E1E4ED5E5369516C1">
<mods:roleTerm id="606110C14EADC372B8E5E09C2DB6875D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E5C4BEDF31E49F1136E97362B3BE8E9">Pinto-Da-Rocha, Ricardo</mods:namePart>
</mods:name>
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<mods:title id="7CF5EACB0B8714DD93D4601E11C33195">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4CA7E4EC6F47753207AF82862CDBD052">
<mods:date id="066DFED0E9FBFC360E39A9F7CD4D3344">2010</mods:date>
<mods:detail id="D85D09278721966FFDE9C7E4A1F3068C" type="pubDate">
<mods:number id="7A6DAEC56361EA9EE5680382C7A520BC">2010-12-03</mods:number>
</mods:detail>
<mods:detail id="65C68120630E8DF5A0A801FB3DEC8F7A" type="volume">
<mods:number id="6BE45EFEB0AA1BDC808EF15455541B59">2698</mods:number>
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<mods:classification id="F6D8070CE1D4F031148E904BCA7F1C7C">journal article</mods:classification>
<mods:identifier id="719BE4028EF88E0F6F29DAA56EE81531" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="039387E595654628FF65FE658E75FB6B" ID-DOI="http://doi.org/10.5281/zenodo.5295597" ID-Zenodo-Dep="5295597" LSID="urn:lsid:plazi:treatment:039387E595654628FF65FE658E75FB6B" httpUri="http://treatment.plazi.org/id/039387E595654628FF65FE658E75FB6B" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63">
<subSubSection id="C320657895654656FF65FE658BBAFE43" box="[151,492,450,476]" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8B8536F395654656FF65FE658BBAFE43" blockId="62.[151,492,450,511]" box="[151,492,450,476]" pageId="62" pageNumber="63">
<heading id="D0CD819F95654656FF65FE658BBAFE43" bold="true" box="[151,492,450,476]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B94EEAE195654656FF65FE658BBAFE43" bold="true" box="[151,492,450,476]" pageId="62" pageNumber="63">
<taxonomicName id="4C3A4D7095654656FF65FE658BF4FE43" ID-CoL="7W5RK" authority="Hara &amp; Pinto-Da-Rocha, 2010" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[151,418,450,476]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="geometricus" status="sp. nov.">
<emphasis id="B94EEAE195654656FF65FE658BF4FE43" bold="true" box="[151,418,450,476]" italics="true" pageId="62" pageNumber="63">Eusarcus geometricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D579A95654656FE58FE658BBAFE43" box="[426,492,450,476]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320657895654656FF65FE428BC7FE60" box="[151,401,485,511]" pageId="62" pageNumber="63" type="description">
<paragraph id="8B8536F395654656FF65FE428BC7FE60" blockId="62.[151,492,450,511]" box="[151,401,485,511]" pageId="62" pageNumber="63">
(
<figureCitation id="13012A7695654656FF6DFE428B56FE60" box="[159,256,485,511]" captionStart="FIGURE 17" captionStartId="63.[151,255,1905,1929]" captionTargetBox="[209,1376,209,1879]" captionTargetId="figure-82@63.[204,1378,194,1892]" captionTargetPageId="63" captionText="FIGURE 17. Eusarcus geometricus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, right tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, idem, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="62" pageNumber="63">Figs. 17</figureCitation>
,
<figureCitation id="13012A7695654656FEFCFE428B0BFE60" box="[270,349,485,511]" captionStart="FIGURE 44" captionStartId="120.[151,255,1644,1668]" captionTargetBox="[172,1417,388,1612]" captionTargetId="figure-219@120.[165,1422,381,1623]" captionTargetPageId="120" captionText="FIGURE 44. Distal part of penis in dorsal and lateral view: A–B, Eusarcus geometricus sp. n.; C–D, E. caparaoensis sp. n.; E–F, E. insperatus. Arrow indicates the angular medio-ventral projection of the stylus. Scale bars, 0.1 mm." pageId="62" pageNumber="63">44A,B</figureCitation>
,
<figureCitation id="13012A7695654656FE98FE428BDEFE60" box="[362,392,485,511]" captionStart="FIGURE 51" captionStartId="127.[151,255,1031,1055]" captionTargetBox="[232,1362,180,1010]" captionTargetId="figure-89@127.[228,1362,165,1027]" captionTargetPageId="127" captionText="FIGURE 51. Localities of Eusarcus spp. in eastern Brazil, Uruguay, and northeastern Argentina." pageId="62" pageNumber="63">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320657895654656FF65FD888F5FFD25" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8B8536F395654656FF65FD888F5FFD25" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">
<emphasis id="B94EEAE195654656FF65FD888B06FDD6" bold="true" box="[151,336,559,585]" pageId="62" pageNumber="63">Type material:</emphasis>
<materialsCitation id="3B523CAE95654656FEABFD9788EBFDF0" collectingDate="2001-01-27" collectorName="Parque Nacional Serra dos Orgaos &amp; A. P. L. Giupponi &amp; R. L. C. Baptista" country="Brazil" location="Teresopolis" pageId="62" pageNumber="63" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<collectingCountry id="F32D766395654656FEABFD978B91FDD5" box="[345,455,560,586]" name="Brazil" pageId="62" pageNumber="63">BRAZIL</collectingCountry>
.
<collectingRegion id="49FEF81195654656FE26FD978828FDD5" box="[468,638,560,586]" country="Brazil" name="Rio de Janeiro" pageId="62" pageNumber="63">Rio de Janeiro</collectingRegion>
:
<location id="8EE5602895654656FD7DFD978940FDD5" LSID="urn:lsid:plazi:treatment:039387E595654628FF65FE658E75FB6B:8EE5602895654656FD7DFD978940FDD5" box="[655,790,560,586]" country="Brazil" name="Teresopolis" pageId="62" pageNumber="63" stateProvince="Rio de Janeiro">Teresópolis</location>
(
<collectorName id="26CF532595654656FCD4FD978E93FDD5" box="[806,1221,560,586]" pageId="62" pageNumber="63">Parque Nacional Serra dos Órgãos</collectorName>
),
<collectorName id="26CF532595654656FB28FD978FCAFDD5" box="[1242,1436,560,586]" pageId="62" pageNumber="63">A.P.L. Giupponi</collectorName>
&amp;
<collectorName id="26CF532595654656FF47FDF28B3DFDF0" box="[181,363,597,623]" pageId="62" pageNumber="63">R.L.C. Baptista</collectorName>
leg.,
<date id="FF84103395654656FE5EFDF2884BFDF0" box="[428,541,597,623]" pageId="62" pageNumber="63" value="2001-01-27">
<collectingDate id="EFC0E9DB95654656FE5EFDF2884BFDF0" box="[428,541,597,623]" pageId="62" pageNumber="63" value="2001-01-27">27.i.2001</collectingDate>
</date>
, ma
<typeStatus id="5481885195654656FDA6FDF288EBFDF0" box="[596,701,597,623]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="3B523CAE95654656FD3AFDF28F53FD25" collectingDate="1989-08-05" collectingDateMax="1998-11-02" collectingDateMin="1989-08-05" collectionCode="MZSP" collectorName="R. Pinto-da-Rocha &amp; A. B. Kury &amp; A. P. L. Giupponi &amp; A. Pires" country="Brazil" location="Teresopolis" pageId="62" pageNumber="63" specimenCode="MNRJ 4480, MHNC 6589, MNRJ 5736" specimenCount="4" stateProvince="Rio de Janeiro" typeStatus="paratype">
1 ma, 2 fe &amp; 1 im
<typeStatus id="5481885195654656FC6CFDF28E58FDF0" box="[926,1038,597,623]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DB9C9E8895654656FBEFFDF28EE5FDF0" box="[1053,1203,597,623]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="62" pageNumber="63">MNRJ 4480</specimenCode>
); idem, 1 ma &amp; 2 fe
<typeStatus id="5481885195654656FF65FDDD8B5FFD0B" box="[151,265,634,660]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED2BAE3695654656FEEBFDDD8B3BFD0B" box="[281,365,634,660]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="62" pageNumber="63" type="Museum">MZSP</collectionCode>
); idem,
<collectorName id="26CF532595654656FE3CFDDD88FDFD0B" box="[462,683,634,660]" pageId="62" pageNumber="63">R. Pinto-da-Rocha</collectorName>
&amp;
<collectorName id="26CF532595654656FD21FDDD8907FD0B" box="[723,849,634,660]" pageId="62" pageNumber="63">A.B. Kury</collectorName>
leg.,
<date id="FF84103395654656FC67FDDD8E4FFD0B" box="[917,1049,634,660]" pageId="62" pageNumber="63" value="1989-08-05">
<collectingDate id="EFC0E9DB95654656FC67FDDD8E4FFD0B" box="[917,1049,634,660]" pageId="62" pageNumber="63" value="1989-08-05">5.viii.1989</collectingDate>
</date>
, 1 ma &amp; 1 fe
<typeStatus id="5481885195654656FB4DFDDD8F67FD0B" box="[1215,1329,634,660]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DB9C9E8895654656FAB3FDDD8A80FD25" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="62" pageNumber="63">MHNC 6589</specimenCode>
); idem,
<collectorName id="26CF532595654656FEC6FD078BA5FD25" box="[308,499,672,698]" pageId="62" pageNumber="63">A.P.L. Giupponi</collectorName>
&amp;
<collectorName id="26CF532595654656FDEBFD078821FD25" box="[537,631,672,698]" pageId="62" pageNumber="63">A. Pires</collectorName>
leg.,
<date id="FF84103395654656FD4BFD078911FD25" box="[697,839,672,698]" pageId="62" pageNumber="63" value="1998-11-01" valueMax="1998-11-02" valueMin="1998-11-01">
<collectingDate id="EFC0E9DB95654656FD4BFD078911FD25" box="[697,839,672,698]" pageId="62" pageNumber="63" value="1998-11-01" valueMax="1998-11-02" valueMin="1998-11-01">1–2.xi.1998</collectingDate>
</date>
, 1 ma &amp; 1 fe
<typeStatus id="5481885195654656FC15FD078E01FD25" box="[999,1111,672,698]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DB9C9E8895654656FB94FD078EABFD25" box="[1126,1277,672,698]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="62" pageNumber="63">MNRJ 5736</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320657895654656FF34FD628B26FC7B" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8B8536F395654656FF34FD628B26FC7B" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">
<emphasis id="B94EEAE195654656FF34FD628B1EFD40" bold="true" box="[198,328,709,735]" pageId="62" pageNumber="63">Diagnosis:</emphasis>
<taxonomicName id="4C3A4D7095654656FEA4FD62885DFD41" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[342,523,709,735]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="geometricus">
<emphasis id="B94EEAE195654656FEA4FD628B3EFD41" box="[342,360,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FE8EFD62885DFD41" box="[380,523,709,734]" italics="true" pageId="62" pageNumber="63">geometricus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3A4D7095654656FD6EFD628957FD41" box="[668,769,709,735]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="incus">
<emphasis id="B94EEAE195654656FD6EFD6288F8FD41" box="[668,686,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FD31FD628957FD41" box="[707,769,709,734]" italics="true" pageId="62" pageNumber="63">incus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095654656FCE7FD6289C1FD41" authorityName="Mello-Leitao" authorityYear="1931" box="[789,919,709,735]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94EEAE195654656FCE7FD628971FD41" box="[789,807,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FCCEFD6289C1FD41" box="[828,919,709,734]" italics="true" pageId="62" pageNumber="63">pusillus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095654656FC5EFD628ED5FD41" box="[940,1155,709,735]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nigrimaculatus">
<emphasis id="B94EEAE195654656FC5EFD6289E8FD41" box="[940,958,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FC20FD628ED5FD41" box="[978,1155,709,734]" italics="true" pageId="62" pageNumber="63">nigrimaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095654656FB6AFD628F38FD41" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[1176,1390,709,735]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="acrophthalmus">
<emphasis id="B94EEAE195654656FB6AFD628EFCFD41" box="[1176,1194,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FB4DFD628F38FD41" box="[1215,1390,709,734]" italics="true" pageId="62" pageNumber="63">acrophthalmus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095654656FA71FD628AA4FC9C" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B94EEAE195654656FA71FD628FC3FD41" box="[1411,1429,709,734]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FF65FD4D8AA4FC9C" box="[151,242,746,771]" italics="true" pageId="62" pageNumber="63">alpinus</emphasis>
</taxonomicName>
e
<taxonomicName id="4C3A4D7095654656FEE0FD4D8BE2FC9C" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[274,436,746,772]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="didactylus">
<emphasis id="B94EEAE195654656FEE0FD4D8B72FC9C" box="[274,292,746,771]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FEC7FD4D8BE2FC9C" box="[309,436,746,771]" italics="true" pageId="62" pageNumber="63">didactylus</emphasis>
</taxonomicName>
because of the cylindrical prolateral median process on male trochanter IV and because forms of the generally large body size, and can be distinguished by: The ocularium with similar-sized tubercles; rectangular projection on the retrolateral margin of the prolateral apical apophysis of coxa IV and the shape of the cylindrical prolateral median process of male trochanter IV: Prolateral distal edge of apex with cylindrical projection, retrolateral distal edge of apex with conical blunt projection; basally with a small pyramidal apophysis. Penis without ventral process of glans (unusual within the genus) and similar to that of
<taxonomicName id="4C3A4D7095654656FF65FC6D8B3EFC7C" box="[151,360,970,996]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nigrimaculatus">
<emphasis id="B94EEAE195654656FF65FC6D8AFFFC7C" box="[151,169,970,995]" italics="true" pageId="62" pageNumber="63">E</emphasis>
.
<emphasis id="B94EEAE195654656FF45FC6D8B3EFC7C" box="[183,360,970,995]" italics="true" pageId="62" pageNumber="63">nigrimaculatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C320657895654656FF34FC4889E2FBB0" pageId="62" pageNumber="63" type="etymology">
<paragraph id="8B8536F395654656FF34FC4889E2FBB0" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">
<emphasis id="B94EEAE195654656FF34FC488B01FB96" bold="true" box="[198,343,1007,1033]" pageId="62" pageNumber="63">Etymology:</emphasis>
The name, an adjective, refers to the prolateral apophyses on the male trochanter IV that resemble geometrical forms, such as pyramids, cylinders and cones.
</paragraph>
</subSubSection>
<subSubSection id="C320657895654628FF37FB9D8EA2FB35" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63" type="description">
<paragraph id="8B8536F395654656FF37FB9D8B7BFAAB" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">
<emphasis id="B94EEAE195654656FF37FB9D8B36FBCB" bold="true" box="[197,352,1082,1108]" pageId="62" pageNumber="63">Description:</emphasis>
<emphasis id="B94EEAE195654656FE9AFB9D8BF2FBCC" box="[360,420,1082,1107]" italics="true" pageId="62" pageNumber="63">Male</emphasis>
(
<typeStatus id="5481885195654656FE47FB9D8877FBCB" box="[437,545,1082,1108]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
; MNRJ 4480): Dorsum (
<figureCitation id="13012A7695654656FCBAFB9D8984FBCB" box="[840,978,1082,1108]" captionStart="FIGURE 17" captionStartId="63.[151,255,1905,1929]" captionTargetBox="[209,1376,209,1879]" captionTargetId="figure-82@63.[204,1378,194,1892]" captionTargetPageId="63" captionText="FIGURE 17. Eusarcus geometricus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, right tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, idem, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="62" pageNumber="63">Fig. 17A,B</figureCitation>
): Measurements: SL 5.10; SMW 4.30; femur I 2.25; II 4.55; III 3.55; IV 4.45. Median paracheliceral projection larger than PAM. Ocularium far from anterior scutal margin; with 15 tubercles. Carapace with 20 scattered tubercles. Scutal area I with 24–25 scattered tubercles on each side; II with 42; III with 57 and a median spine long, almost straight, strongly pointing backwards, surpassing posterior margin of dorsal scutum; IV with 28 tubercles. Posterior margin of dorsal scutum and free tergites I–III with a row of 20, 16, 13, 11 tubercles, respectively. Anal operculum with 17 tubercles.
</paragraph>
<paragraph id="8B8536F395654656FF37FAE788B8FAE0" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">Venter: Coxa I with 25–26 tubercles, median and distal ones largest; II with 36–38, increasing in size distally; III–IV densely and irregularly tuberculate.</paragraph>
<paragraph id="8B8536F395654656FF37FA2D88D0FA3B" blockId="62.[151,1436,559,2004]" box="[197,646,1418,1444]" pageId="62" pageNumber="63">Chelicera: Segment I with 3 tubercles.</paragraph>
<paragraph id="8B8536F395654656FF37FA178E78F98B" blockId="62.[151,1436,559,2004]" pageId="62" pageNumber="63">Pedipalpus: Trochanter dorsally inflated, with 2 tubercles. Femur dorsally with 3 aligned tubercles, 6–7 apical small ones; ventrally with 1 large pointed basal tubercle, 2 small scattered ones. Tibial setation: Prolateral IiIi, retrolateral IiIi/iIi. Tarsal setation: Prolateral IIi, retrolateral IiIi.</paragraph>
<paragraph id="8B8536F395654628FF37F9878EECFE9D" blockId="62.[151,1436,559,2004]" lastBlockId="64.[151,1436,152,1268]" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63">
Legs (
<figureCitation id="13012A7695654656FEFCF9878BC5F9A5" box="[270,403,1568,1594]" captionStart="FIGURE 17" captionStartId="63.[151,255,1905,1929]" captionTargetBox="[209,1376,209,1879]" captionTargetId="figure-82@63.[204,1378,194,1892]" captionTargetPageId="63" captionText="FIGURE 17. Eusarcus geometricus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, right tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, idem, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm." pageId="62" pageNumber="63">Fig. 17C–I</figureCitation>
): Coxa II with 3 tubercles on retrolateral margin; IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, long, blunt, curved posteriad and slightly ventrad, with 1 rectangular projection on retrolateral margin. Trochanters I–IV with scattered tubercles; I–III with large retrolateral tubercles; I–II with 1 large ventro-basal tubercle; IV prolaterally with 1 cylindrical and flattened median process, prolateral edge of its apex with cylindrical projection; retrolateral edge of apex with conical, blunt projection; basally with 1 small pyramidal apophysis; retrolaterally with 1 large pointed median tubercle, 1 large apical tubercle. Femur I with reduced PDS, RDS, two ventral irregular rows of slightly enlarged tubercles; II–III with reduced PDS, large RDS; II with two ventral rows of tubercles increasing in size distally; III sinuous, ventrally with two irregular rows of tubercles increasing in size distally, prolateral ones enlarged, 1 blunt prolateral apical spine, 1 large retrolateral apical tubercle; IV sinuous, with medium PDS, large RDS; prolateral and retrolateral rows of enlarged, pointed tubercles; ventrally with 2 rows of tubercles, prolateral ones enlarged and increasing in size distally, 2 apical spines. Tibia III with 2 irregular ventral rows of pointed tubercles increasing in size subapically (3 large); IV with 2 lateral apical spines, prolateral one smaller, ventrally with 2 rows of enlarged tubercles increasing in size distally, 2 apical spines. Metatarsi III– IV with tubercles decreasing in size and amount distally. Tarsal segmentation: 6, 8–9, 6, 6.
</paragraph>
<caption id="DF45667B95644657FF65F8D68B00F877" pageId="63" pageNumber="64" startId="63.[151,255,1905,1929]" targetBox="[209,1376,209,1879]" targetPageId="63">
<paragraph id="8B8536F395644657FF65F8D68B00F877" blockId="63.[151,1438,1905,2024]" pageId="63" pageNumber="64">
<emphasis id="B94EEAE195644657FF65F8D68B70F816" bold="true" box="[151,294,1905,1929]" pageId="63" pageNumber="64">FIGURE 17.</emphasis>
<taxonomicName id="4C3A4D7095644657FEDEF8D58841F816" authority="Hara &amp; Pinto-Da-Rocha, 2010" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[300,535,1906,1929]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="geometricus" status="sp. nov.">
<emphasis id="B94EEAE195644657FEDEF8D58841F816" box="[300,535,1906,1929]" italics="true" pageId="63" pageNumber="64">Eusarcus geometricus</emphasis>
</taxonomicName>
<emphasis id="B94EEAE195644657FDECF8D6880CF816" bold="true" box="[542,602,1905,1929]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="A27D579A95644657FDECF8D6880CF816" box="[542,602,1905,1929]" pageId="63" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right femur III, ventral view; E, right tibia III, ventral view; F, right trochanter and femur IV, dorsal view; G, idem, ventral view; H, idem, prolateral view; I, right tibia IV, ventral view. Scale bars, 1 mm.
</paragraph>
</caption>
<paragraph id="8B8536F3951B4628FF37FEB7882CFE20" blockId="64.[151,1436,152,1268]" pageId="64" pageNumber="65">
Penis (
<figureCitation id="13012A76951B4628FEE4FEB78BF4FEB5" box="[278,418,272,298]" captionStart="FIGURE 44" captionStartId="120.[151,255,1644,1668]" captionTargetBox="[172,1417,388,1612]" captionTargetId="figure-219@120.[165,1422,381,1623]" captionTargetPageId="120" captionText="FIGURE 44. Distal part of penis in dorsal and lateral view: A–B, Eusarcus geometricus sp. n.; C–D, E. caparaoensis sp. n.; E–F, E. insperatus. Arrow indicates the angular medio-ventral projection of the stylus. Scale bars, 0.1 mm." pageId="64" pageNumber="65">Fig. 44A,B</figureCitation>
): Stylus smooth, apex swollen, curved ventrad; without medio-ventral trichomes and angular medio-ventral projection. Ventral process of glans absent. Ventral plate covering only basal third of glans; with 2 pairs of short, straight distal setae, 1 dorsal pair of small distal setae, 1 pair of short, straight median setae, 2 pairs of ventral subapical tubercles; 3–4 pairs of long, straight basal setae, basalmost pair far from others and placed on trunk of penis.
</paragraph>
<paragraph id="8B8536F3951B4628FF37FE6D880EFD95" blockId="64.[151,1436,152,1268]" pageId="64" pageNumber="65">Coloration: Generally dark brown; brown on grooves, pedipalps, chelicerae, median portions of femora, tibiae, metatarsi. Light brown on tarsi.</paragraph>
<paragraph id="8B8536F3951B4628FF37FDB28F46FC70" blockId="64.[151,1436,152,1268]" pageId="64" pageNumber="65">
<emphasis id="B94EEAE1951B4628FF37FDB28B4BFDB1" box="[197,285,533,558]" italics="true" pageId="64" pageNumber="65">Female</emphasis>
(
<typeStatus id="54818851951B4628FEDFFDB28BCEFDB0" box="[301,408,533,559]" pageId="64" pageNumber="65" type="paratype">paratype</typeStatus>
; MNRJ 4480): Dorsum: Measurements: SL 4.75; SMW 3.60; femur I 1.85; II 3.95; III 3.10; IV 3.90. Posterior margin of dorsal scutum with 10 scattered tubercles and an irregular row of 21 tubercles. Free tergites I–III with an irregular row of 20, 13, 14 tubercles, respectively. Pedipalpus: Tibial setation: Prolateral IiIi, retrolateral iiIi. Legs: Coxa IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, short, blunt, curved backwards, surpassing posterior margin of this segment. Trochanter III with scattered tubercles, retrolateral ones enlarged, 1 retrolateral apical standing out; IV with scattered tubercles, with 1 large pointed retrolateral apical tubercle. Femur I ventrally with rows of similarsized tubercles; III ventrally with two irregular rows of tubercles slightly increasing in size distally, prolateral ones enlarged, 1 small, blunt prolateral apical spine, 1 large retrolateral apical tubercle; IV with prolateral and retrolateral rows of similar-sized tubercles; ventrally with 2 rows of tubercles, prolateral ones enlarged and increasing in size distally, 2 small blunt apical spines. Patella IV ventrally with enlarged tubercles; tibia III only with rows of similar-sized tubercles; IV with two irregular rows of enlarged tubercles slightly increasing in size distally, 2 large apical tubercles. Metatarsus III unarmed. Tarsal segmentation: 6, 7–8, 6, 6.
</paragraph>
<paragraph id="8B8536F3951B4628FF37FC5D883FFBC0" blockId="64.[151,1436,152,1268]" pageId="64" pageNumber="65">
<emphasis id="B94EEAE1951B4628FF37FC5D8BF2FB8C" box="[197,420,1018,1043]" italics="true" pageId="64" pageNumber="65">Variation in males</emphasis>
(n=5): Measurements: SL 4.30–5.20; SMW 3.50–4.40; femur I 1.90–2.25; II 4.00– 4.55; III 3.00–3.55; IV 3.90–4.45. Pedipalpus: Tibial setation: Retrolateral IiIi/IIi/iIi. Femur I reduced or small RDS; II–III with reduced or small PDS.
</paragraph>
<paragraph id="8B8536F3951B4628FF37FBCC8EA2FB35" blockId="64.[151,1436,152,1268]" pageId="64" pageNumber="65">
<emphasis id="B94EEAE1951B4628FF37FBCC8BE3FB1B" box="[197,437,1131,1156]" italics="true" pageId="64" pageNumber="65">Variation in females</emphasis>
(n=6): Measurements: SL 4.70–5.10; SMW 3.60–4.00; femur I 1.70–2.00; II 3.40– 4.20; III 2.50–3.30; IV 3.30–4.30. Pedipalpus: Tibial setation: Prolateral IiIi, retrolateral iiIi/iIi.
</paragraph>
</subSubSection>
<subSubSection id="C3206578951B4628FF37FB128E75FB6B" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8B8536F3951B4628FF37FB128EE5FB50" blockId="64.[151,1436,152,1268]" box="[197,1203,1205,1231]" pageId="64" pageNumber="65">
<materialsCitation id="3B523CAE951B4628FF37FB128EE6FB50" box="[197,1200,1205,1231]" collectorName="Parque Nacional Serra dos Orgaos" country="Brazil" location="Rio de Janeiro" pageId="64" pageNumber="65" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="B94EEAE1951B4628FF37FB128B38FB50" bold="true" box="[197,366,1205,1231]" pageId="64" pageNumber="65">
<typeStatus id="54818851951B4628FF37FB128B54FB50" box="[197,258,1205,1231]" pageId="64" pageNumber="65">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F32D7663951B4628FE84FB128B96FB50" box="[374,448,1205,1231]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
,
<collectingRegion id="49FEF811951B4628FE3EFB12882EFB50" box="[460,632,1205,1231]" country="Brazil" name="Rio de Janeiro" pageId="64" pageNumber="65">Rio de Janeiro</collectingRegion>
, Teresópolis,
<collectorName id="26CF5325951B4628FCEBFB128EE6FB50" box="[793,1200,1205,1231]" pageId="64" pageNumber="65">Parque Nacional Serra dos Órgãos</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536F3951B4628FF37FB7D8E75FB6B" blockId="64.[151,1436,152,1268]" box="[197,1059,1242,1268]" pageId="64" pageNumber="65">
<emphasis id="B94EEAE1951B4628FF37FB7D885FFB6B" bold="true" box="[197,521,1242,1268]" pageId="64" pageNumber="65">Geographical distribution</emphasis>
(
<figureCitation id="13012A76951B4628FDE8FB7D8827FB6B" box="[538,625,1242,1268]" captionStart="FIGURE 51" captionStartId="127.[151,255,1031,1055]" captionTargetBox="[232,1362,180,1010]" captionTargetId="figure-89@127.[228,1362,165,1027]" captionTargetPageId="127" captionText="FIGURE 51. Localities of Eusarcus spp. in eastern Brazil, Uruguay, and northeastern Argentina." pageId="64" pageNumber="65">Fig. 51</figureCitation>
): Known only from the
<typeStatus id="54818851951B4628FC79FB7D89EBFB6B" box="[907,957,1242,1268]" pageId="64" pageNumber="65">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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