304 lines
56 KiB
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304 lines
56 KiB
XML
<document id="1ADF2DA4960F815A030A87FF1B2E743B" ID-DOI="10.11646/zootaxa.5486.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13210005" ID-ZooBank="C7F7E7D5-6F6E-49C8-AF31-9B769EC8B56C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722788280951" checkinUser="plazi" docAuthor="Orr, Albert G. W., Dow, Rory A. & Steinhoff, Philip O. M." docDate="2024" docId="0393879EFFD7FFC5DBF74CA7FCBB7F3C" docLanguage="en" docName="zootaxa.5486.2.3.1.pdf" docOrigin="Zootaxa 5486 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sundacypha petiolata" docType="treatment" docVersion="2" lastPageNumber="317" masterDocId="FFAAFFE6FFD8FFD5DB60480DFF897815" masterDocTitle="Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family." masterLastPageNumber="337" masterPageNumber="301" pageNumber="316" updateTime="1722788985873" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ADB5DA92CBED79AC83DC3ED512C81B79">Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family.</mods:title>
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<mods:roleTerm id="E5F272010BBC1C7B21566A452704BECC">Author</mods:roleTerm>
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<mods:namePart id="60FB18FA8C05451C3EA293545913A7E2">Orr, Albert G. W.</mods:namePart>
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<mods:affiliation id="CD4713A99A7F37ECF7F4B57468D944AB">Environmental Futures Research Institute, Griffith University, Nathan, Australia.</mods:affiliation>
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<mods:namePart id="7DA71999762A83677479CEE9EE8C9087">Dow, Rory A.</mods:namePart>
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<mods:affiliation id="B8D1C6296E855C17D05E9AB4A4B23B6F">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands. & Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia.</mods:affiliation>
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<mods:roleTerm id="27B8B7C14FDEDD66B858D1ACA4B2BAC1">Author</mods:roleTerm>
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<mods:namePart id="D736A0ABFAEBBB9745B9A39983811E99">Steinhoff, Philip O. M.</mods:namePart>
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<mods:affiliation id="595B17040FA4BFB8791941267135B3D3">Höxter, Germany.</mods:affiliation>
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<mods:typeOfResource id="01EB894ED974F1B08459F887532F8684">text</mods:typeOfResource>
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<mods:title id="05B55B7B558CBCCCE8B14693431A0955">Zootaxa</mods:title>
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<mods:part id="F1EE40DDD59DA767D8FC7699F71A23CF">
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<mods:date id="004110F3771684A39CF24F5061A68099">2024</mods:date>
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<mods:number id="BEC1A4242C5EA9DE13A4F1C2454CB98D">2024-07-26</mods:number>
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<mods:number id="FCC0B9ED6135E38057809A18741855FF">5486</mods:number>
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<mods:url id="2D8BF19739DB3E68983CE9AFA02048B4">http://dx.doi.org/10.11646/zootaxa.5486.3.1</mods:url>
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<treatment id="0393879EFFD7FFC5DBF74CA7FCBB7F3C" LSID="urn:lsid:plazi:treatment:0393879EFFD7FFC5DBF74CA7FCBB7F3C" httpUri="http://treatment.plazi.org/id/0393879EFFD7FFC5DBF74CA7FCBB7F3C" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
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<paragraph id="8B853688FFD7FFDADBF74CA7FE7E7CD1" blockId="15.[151,906,1194,1257]" box="[151,503,1194,1221]" pageId="15" pageNumber="316">
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<heading id="D0CD81E4FFD7FFDADBF74CA7FE7E7CD1" bold="true" box="[151,503,1194,1221]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
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<taxonomicName id="4C3A4D0BFFD7FFDADBF74CA7FE7E7CD1" ID-CoL="53G6J" authority="(Selys)" baseAuthorityName="Selys" box="[151,503,1194,1221]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="petiolata">
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<emphasis id="B94EEA9AFFD7FFDADBF74CA7FE7E7CD1" bold="true" box="[151,503,1194,1221]" pageId="15" pageNumber="316">
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<emphasis id="B94EEA9AFFD7FFDADBF74CA7FE157CD0" bold="true" box="[151,412,1194,1221]" italics="true" pageId="15" pageNumber="316">Sundacypha petiolata</emphasis>
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(Selys)
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</emphasis>
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<subSubSection id="C3206503FFD7FFDADBF74CC2FC037CFC" box="[151,906,1230,1257]" pageId="15" pageNumber="316" type="description">
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<paragraph id="8B853688FFD7FFDADBF74CC2FC037CFC" blockId="15.[151,906,1194,1257]" box="[151,906,1230,1257]" pageId="15" pageNumber="316">
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(
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<figureCitation id="13012A0DFFD7FFDADBFF4CC2FF7D7CFC" box="[159,244,1231,1257]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="15" pageNumber="316">Figs 4c</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADA604CC3FE947CFC" box="[256,285,1230,1257]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="15" pageNumber="316">5c</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADA494CC2FECF7CFC" box="[297,326,1231,1257]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="15" pageNumber="316">7c</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADA314CC2FEE47CFC" box="[337,365,1231,1257]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="15" pageNumber="316">8c</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADA194CC2FE1E7CFC" box="[377,407,1231,1257]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="15" pageNumber="316">9c</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADAC24CC2FE447CFC" box="[418,461,1231,1257]" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="15" pageNumber="316">11b</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDADAB94CC2FD967CFC" box="[473,543,1231,1257]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="15" pageNumber="316">12c, g</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDAD94B4CC2FDFB7CFC" box="[555,626,1231,1257]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="15" pageNumber="316">14c, d</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDAD91E4CC3FD4D7CFC" box="[638,708,1230,1257]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="15" pageNumber="316">15c, d</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDAD9B04CC2FC9D7CFC" box="[720,788,1231,1257]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="15" pageNumber="316">16b, f</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDAD87F4CC2FCC57CFC" box="[799,844,1231,1257]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="15" pageNumber="316">17b</figureCitation>
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,
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<figureCitation id="13012A0DFFD7FFDAD8374CC2FC087CFC" box="[855,897,1231,1257]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="15" pageNumber="316">18c</figureCitation>
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)
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</paragraph>
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<emphasis id="B94EEA9AFFD7FFDADBF74D1BFE0D7D25" bold="true" box="[151,388,1302,1328]" pageId="15" pageNumber="316">Material examined.</emphasis>
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<materialsCitation id="3B523CD5FFD7FFDADAEE4D1AFDF07D24" box="[398,633,1303,1329]" country="Malaysia" location="MALAYSIA" pageId="15" pageNumber="316" specimenCount="1" specimenCount-male="1">
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<collectingCountry id="F32D7618FFD7FFDADAEE4D1AFDAB7D24" box="[398,546,1303,1329]" name="Malaysia" pageId="15" pageNumber="316">MALAYSIA</collectingCountry>
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F
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,
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<materialsCitation id="3B523CD5FFD7FFDAD9E74D1AFD237D6C" collectingDate="2014-03-26" collectingDateMax="2014-03-27" collectingDateMin="2014-03-26" collectionCode="RMNH" collectorName="Camp C & S. G. Butler." country="Malaysia" county="Planted Forest Project" latitude="2.7171" location="Bintulu Division" longLatPrecision="7" longitude="113.3463" municipality="Planted Forest" pageId="15" pageNumber="316" specimenCount="1" specimenCount-female="1" stateProvince="Sarawak">
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F (
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.5008329,
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<collectionCode id="ED2BAE4DFFD7FFDAD8D14D1BFB807D24" box="[945,1033,1302,1329]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="316" type="Museum">RMNH</collectionCode>
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.5008330), stream in kerangas near
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<collectorName id="26CF535EFFD7FFDADBF74D36FF7E7D40" box="[151,247,1339,1365]" pageId="15" pageNumber="316">Camp C</collectorName>
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<collectingMunicipality id="6BE1ACF2FFD7FFDADA644D36FE2F7D40" box="[260,422,1339,1365]" pageId="15" pageNumber="316">Planted Forest</collectingMunicipality>
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Project
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<collectingRegion id="49FEF86AFFD7FFDAD9B94D36FCB57D40" box="[729,828,1339,1365]" country="Malaysia" name="Sarawak" pageId="15" pageNumber="316">Sarawak</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE0E504FFFD7FFDAD8294D36FC247D40" box="[841,941,1339,1365]" degrees="2.7171" direction="north" orientation="latitude" pageId="15" pageNumber="316" precision="5" value="2.7171">2.7171N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0E504FFFD7FFDAD8D94D36FBBC7D40" box="[953,1077,1339,1365]" degrees="113.3463" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="5" value="113.3463">113.3463E</geoCoordinate>
|
||
,
|
||
<date id="FF841048FFD7FFDADF224D36FB347D40" box="[1090,1213,1339,1365]" pageId="15" pageNumber="316" value="2014-03-26">
|
||
<collectingDate id="EFC0E9A0FFD7FFDADF224D36FB347D40" box="[1090,1213,1339,1365]" pageId="15" pageNumber="316" value="2014-03-26">26 iii 2014</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED2BAE4DFFD7FFDADFAD4D37FAAF7D40" box="[1229,1318,1338,1365]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="316" type="Museum">RMNH</collectionCode>
|
||
.5008329),
|
||
<date id="FF841048FFD7FFDADBF74D52FE987D6C" box="[151,273,1375,1401]" pageId="15" pageNumber="316" value="2014-03-27">
|
||
<collectingDate id="EFC0E9A0FFD7FFDADBF74D52FE987D6C" box="[151,273,1375,1401]" pageId="15" pageNumber="316" value="2014-03-27">27 iii 2014</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED2BAE4DFFD7FFDADA7F4D53FEF17D6C" box="[287,376,1374,1401]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="316" type="Museum">RMNH</collectionCode>
|
||
.5008330), leg.
|
||
<collectorName id="26CF535EFFD7FFDAD9454D52FD237D6C" box="[549,682,1375,1401]" pageId="15" pageNumber="316">S.G. Butler.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD7FFDADBA74D8FFEC77E38" blockId="15.[151,1437,1302,2013]" pageId="15" pageNumber="316">
|
||
<emphasis id="B94EEA9AFFD7FFDADBA74D8FFEA67D89" bold="true" box="[199,303,1410,1436]" pageId="15" pageNumber="316">Habitus:</emphasis>
|
||
similar to
|
||
<taxonomicName id="4C3A4D0BFFD7FFDADAD44D8EFDB77D88" box="[436,574,1411,1437]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDADAD44D8EFDB77D88" box="[436,574,1411,1437]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but obviously smaller (
|
||
<figureCitation id="13012A0DFFD7FFDAD8374D8EFC247D88" box="[855,941,1411,1437]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="15" pageNumber="316">Fig. 4c</figureCitation>
|
||
). Significantly paler mottling on head and body in specimens examined which have predominantly pale yellowish ground colour. Both head and antennae relatively longer, especially the latter; prothorax lower in profile with anterior tubercles much reduced; prothorax legs proportionally slightly longer, caudal spikes proportionally longer but thinner and more evenly tapered to a point from base.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFD7FFC5DBA74E3BFEE679E0" blockId="15.[151,1437,1302,2013]" lastBlockId="16.[151,1437,151,1401]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
|
||
<emphasis id="B94EEA9AFFD7FFDADBA74E3BFE877E44" bold="true" box="[199,270,1590,1617]" pageId="15" pageNumber="316">
|
||
<emphasis id="B94EEA9AFFD7FFDADBA74E3BFE8C7E44" bold="true" box="[199,261,1590,1617]" italics="true" pageId="15" pageNumber="316">Head</emphasis>
|
||
:
|
||
</emphasis>
|
||
in dorsal view as illustrated (
|
||
<figureCitation id="13012A0DFFD7FFDAD9044E3AFD3E7E44" box="[612,695,1590,1617]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="15" pageNumber="316">Fig. 5c</figureCitation>
|
||
), similar to
|
||
<taxonomicName id="4C3A4D0BFFD7FFDAD8234E3AFC427E44" box="[835,971,1591,1617]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDAD8234E3AFC427E44" box="[835,971,1591,1617]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but slightly more elongate and antennae proportionally longer (1.63 times maximum head width versus
|
||
<quantity id="4CC29B6DFFD7FFDAD8114E57FC487E60" box="[881,961,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.8608" pageId="15" pageNumber="316" unit="in" value="1.52">1.52 in</quantity>
|
||
<taxonomicName id="4C3A4D0BFFD7FFDAD8AB4E56FBDD7E60" box="[971,1108,1627,1653]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDAD8AB4E56FBDD7E60" box="[971,1108,1627,1653]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
) and markings weaker, with subtle differences but tip of pedicel with similar dark coloration; scape with longer and mainly longer straighter setae but a few recurved short setae so that wet specimens may appear sparsely warty. Labrum as in
|
||
<taxonomicName id="4C3A4D0BFFD7FFDADE754EAEFA147EA8" box="[1301,1437,1699,1725]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDADE754EAEFA147EA8" box="[1301,1437,1699,1725]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but more strongly curved, accommodating long mandibular incisor lobes; ‘moustache’ with 9–10 strong setae. Ventrally, genae bear a cluster of subocular spines shorter and less numerous than in
|
||
<taxonomicName id="4C3A4D0BFFD7FFDADF3C4EE6FB6A7F10" box="[1116,1251,1771,1797]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDADF3C4EE6FB6A7F10" box="[1116,1251,1771,1797]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
, all main spines except first appear rounded in ventral view (
|
||
<figureCitation id="13012A0DFFD7FFDAD9F04F02FD697F3C" box="[656,736,1807,1833]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="15" pageNumber="316">Fig. 7c</figureCitation>
|
||
right) but these are upturned and acuminate as seen in lateral view (
|
||
<figureCitation id="13012A0DFFD7FFDADBBD4F3EFEA57F58" box="[221,300,1843,1869]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="15" pageNumber="316">Fig. 7c</figureCitation>
|
||
left). Head, especially seen in lateral view longer and shallower than previous two species, with other differences as shown (
|
||
<figureCitation id="13012A0DFFD7FFDADAF94F5AFE647F64" box="[409,493,1879,1905]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="15" pageNumber="316">Fig. 7c</figureCitation>
|
||
). Mask very long and thin, articulation resting between posterior part of coxae of mesothorax (
|
||
<figureCitation id="13012A0DFFD7FFDADA494F76FEF37F80" box="[297,378,1915,1941]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="15" pageNumber="316">Fig. 8c</figureCitation>
|
||
); prementum narrow basally, gradually flared anteriorly with margins of anterior half straight; lateral margins with sparse fine setae on distal 2/3
|
||
<superScript id="7C4F9BC0FFD7FFDAD9AC4F93FD537FB9" attach="left" box="[716,730,1950,1964]" fontSize="6" pageId="15" pageNumber="316">rd</superScript>
|
||
(
|
||
<figureCitation id="13012A0DFFD7FFDAD9864F92FCBF7FAC" box="[742,822,1951,1977]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="15" pageNumber="316">Fig. 9c</figureCitation>
|
||
) seated in distinctly raised sockets (
|
||
<figureCitation id="13012A0DFFD7FFDADFAE4F92FAA67FAC" box="[1230,1327,1951,1977]" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="15" pageNumber="316">Fig. 11b</figureCitation>
|
||
); anterior median lobe as in
|
||
<taxonomicName id="4C3A4D0BFFD7FFDADA074FCEFE677FC8" box="[359,494,1987,2013]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFD7FFDADA074FCEFE677FC8" box="[359,494,1987,2013]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
but only slightly convex; median cleft very short with tinted area marking point of closure (
|
||
<figureCitation id="13012A0DFFC8FFC5DB97489AFEDE78A4" box="[247,343,151,177]" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="16" pageNumber="317">Fig. 11b</figureCitation>
|
||
); lobe of labial palp as in previous species, but with both palpal lobe and movable hook relatively slightly longer relative to anterior breadth of prementum as indicated (
|
||
<figureCitation id="13012A0DFFC8FFC5D8CD48B6FBCE78C0" box="[941,1095,187,213]" captionStart="FIGURE 10" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[167,1433,182,1916]" captionTargetId="figure-22@18.[155,1433,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Details of anterior prementum and labial palp Heliocypha biseriata: (a) anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below); (b) labial palp and anterior margin of prementum, frontal view; (c) labial palp, interior view. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210025" httpUri="https://zenodo.org/record/13210025/files/figure.png" pageId="16" pageNumber="317">Figs 10a, b,c</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DF3448B6FB0978C0" box="[1108,1152,187,213]" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="16" pageNumber="317">11b</figureCitation>
|
||
). Maxillae as in previous species (
|
||
<figureCitation id="13012A0DFFC8FFC5DB9648D2FEDD78EC" box="[246,340,223,249]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="16" pageNumber="317">Fig. 12c</figureCitation>
|
||
), but with basal pair of secondary denticles on distal tooth well separated as in
|
||
<taxonomicName id="4C3A4D0BFFC8FFC5DFD948D2FAC578EC" box="[1209,1356,223,249]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="fenestrella">
|
||
<emphasis id="B94EEA9AFFC8FFC5DFD948D2FAC578EC" box="[1209,1356,223,249]" italics="true" pageId="16" pageNumber="317">A. fenestrella</emphasis>
|
||
</taxonomicName>
|
||
and all relatively shorter and lighter than in that species (
|
||
<figureCitation id="13012A0DFFC8FFC5D9D8490EFC917908" box="[696,792,259,285]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(a–d) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (e–h) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="16" pageNumber="317">
|
||
Fig.
|
||
<quantity id="4CC29B6DFFC8FFC5D98B490EFC917908" box="[747,792,259,285]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="16" pageNumber="317" unit="g" value="12.0">12g</quantity>
|
||
</figureCitation>
|
||
). Mandibles (
|
||
<figureCitation id="13012A0DFFC8FFC5D8CF490EFBA57908" box="[943,1068,259,285]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="16" pageNumber="317">Figs 14c, d</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DF56490FFBF07908" box="[1078,1145,258,285]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="16" pageNumber="317">15c, d</figureCitation>
|
||
)—both left and right with five distinct teeth on incisor visible in internal view, dorsal one small; right with small supplementary tooth basal to strong ventral tooth; right molar crest a low elongate shallow ridge without obvious dentition and lightly sclerotised (
|
||
<figureCitation id="13012A0DFFC8FFC5DBFF4962FE8D799C" box="[159,260,367,393]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="16" pageNumber="317">Figs 14c</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DA724963FEB6799C" box="[274,319,366,393]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="16" pageNumber="317">15d</figureCitation>
|
||
); left incisor (
|
||
<figureCitation id="13012A0DFFC8FFC5DA814962FDC1799C" box="[481,584,367,393]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="16" pageNumber="317">Figs 14d</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5D9364963FDF6799C" box="[598,639,366,393]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="16" pageNumber="317">15c</figureCitation>
|
||
) with five well developed teeth, the strongest ventrally; molar crest a strong flat dorso-ventral blade with eight minute denticles along a nearly straight distal margin, with end denticles markedly heavier and longer, especially ventrally (
|
||
<figureCitation id="13012A0DFFC8FFC5D9AD49BAFCBB79C4" box="[717,818,439,465]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="16" pageNumber="317">Figs 14d</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5D85D49BBFCE179C4" box="[829,872,438,465]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R & L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="16" pageNumber="317">15c</figureCitation>
|
||
). A strong outer basal seta on both mandibles as in other three species.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFC8FFC5DBA749F3FC517AE4" blockId="16.[151,1437,151,1401]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBA749F3FEAF7A0C" bold="true" box="[199,294,510,537]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBA749F3FE947A0C" bold="true" box="[199,285,510,537]" italics="true" pageId="16" pageNumber="317">Thorax</emphasis>
|
||
:
|
||
</emphasis>
|
||
anterior tubercles on prothorax less developed and more centrally placed than in other two species (
|
||
<figureCitation id="13012A0DFFC8FFC5DE0F49F2FF3D7A28" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="16" pageNumber="317">Fig. 5c</figureCitation>
|
||
, arrowed), and anterior and posterior flanges defining saddle lower. Otherwise similar. Marking (not illustrated on body) lightly mottled but always with broad pale band between two anterior tubercles on prothorax. Wing sheaths slightly spatulate, not completely obscuring abdomen at sides, forewing reaching to near end of S5, hindwing to 2/3 along S6 (
|
||
<figureCitation id="13012A0DFFC8FFC5DA544A82FE0E7ABC" box="[308,391,655,681]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="16" pageNumber="317">Fig. 4c</figureCitation>
|
||
). Legs long and thin, metafemur reaching base of S8, clearly proportionally longer than in
|
||
<taxonomicName id="4C3A4D0BFFC8FFC5DEE14A82FF727AD8" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="biseriata">
|
||
<emphasis id="B94EEA9AFFC8FFC5DEE14A82FF727AD8" italics="true" pageId="16" pageNumber="317">H. biseriata</emphasis>
|
||
</taxonomicName>
|
||
which they resemble in all other respects including presence of two rows of pectinate setae on the venter of the tarsus. Unlike the previous two species the tibiae have no markings.
|
||
</paragraph>
|
||
<paragraph id="8B853688FFC8FFC5DBA74AF7FE0E7D18" blockId="16.[151,1437,151,1401]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBA74AF7FEB67B00" bold="true" box="[199,319,762,789]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBA74AF7FEBC7B00" bold="true" box="[199,309,762,789]" italics="true" pageId="16" pageNumber="317">Abdomen</emphasis>
|
||
:
|
||
</emphasis>
|
||
squat, mainly mottled brown with well-defined pale narrow middorsal stripe (not figured). Margins of S3–7 distinctly crimped forming shallow lateral keels; venter of S6–8 with irregular sparse row of tiny submarginal spines; ventrolateral margin of S9 with 5–7 strong, curved spines and scattering of tiny marginal and submarginal spines along venter (
|
||
<figureCitation id="13012A0DFFC8FFC5DA1D4B6AFE6A7B94" box="[381,483,871,897]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Figs 16b</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DA8E4B6AFD927B94" box="[494,539,871,897]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="16" pageNumber="317">17b</figureCitation>
|
||
); S10 with 3–6 strong ventrolateral spines (
|
||
<figureCitation id="13012A0DFFC8FFC5DF614B6AFBEF7B94" box="[1025,1126,871,897]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Figs 16b</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DF114B6AFB157B94" box="[1137,1180,871,897]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="16" pageNumber="317">17b</figureCitation>
|
||
) and cluster of smaller spines on venter and ventrally on tergite (
|
||
<figureCitation id="13012A0DFFC8FFC5D93F4B86FD327BB0" box="[607,699,907,933]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Fig. 16f</figureCitation>
|
||
); hind-margin S10 with 3–4 strong rearward pointing ventrolateral spines (
|
||
<figureCitation id="13012A0DFFC8FFC5DB8E4BA2FEE77BDC" box="[238,366,943,969]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Figs 16b, f</figureCitation>
|
||
,
|
||
<figureCitation id="13012A0DFFC8FFC5DA1B4BA2FE2F7BDC" box="[379,422,943,969]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="16" pageNumber="317">17b</figureCitation>
|
||
) and small spines along all of hind margin, being sparsest ventrally. Male gonapophyses (
|
||
<figureCitation id="13012A0DFFC8FFC5DBFF4BDEFE8B7BF8" box="[159,258,979,1005]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="16" pageNumber="317">Fig. 17b</figureCitation>
|
||
) short and conical, well separated, arising near posterior margin S9 and just reaching margin. Outer female gonapophyses (
|
||
<figureCitation id="13012A0DFFC8FFC5DAF84BFAFD9A7C04" box="[408,531,1015,1041]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Figs 16b, f</figureCitation>
|
||
) arising from base of S9 and extending well beyond hind margin of S10 to about level of epiproct tip (
|
||
<figureCitation id="13012A0DFFC8FFC5DAE24C16FE567C20" box="[386,479,1051,1077]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Fig. 16f</figureCitation>
|
||
), ventral margin with series of sparse heavy spines to about level of hind margin S10, thence continuing almost directly backward to a sharp point about level with epiproct tip (
|
||
<figureCitation id="13012A0DFFC8FFC5DF184C32FB5C7C4C" box="[1144,1237,1087,1113]" captionStart="FIGURE 16" captionStartId="24.[151,250,1964,1988]" captionTargetBox="[158,1439,262,1931]" captionTargetId="figure-44@24.[149,1439,237,1940]" captionTargetPageId="24" captionText="FIGURE 16. Female gonapophyses and terminal abdominal segments in ventral view: (a) H. biseriata, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) H. biseriata, (e) L. hyalina, (f) S. petiolata. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210037" httpUri="https://zenodo.org/record/13210037/files/figure.png" pageId="16" pageNumber="317">Fig. 16f</figureCitation>
|
||
); inner apophyses smooth, curved smoothly and strongly upward to a rounded point extending well beyond outer apophyses and epiproct. Epiproct, short, subconical, not obviously different from other species studied. Caudal spikes about 65% body length, of medium breadth at their base and tapered almost uniformly to a point (
|
||
<figureCitation id="13012A0DFFC8FFC5DF3E4CA6FB377CD0" box="[1118,1214,1195,1221]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="16" pageNumber="317">Fig. 18c</figureCitation>
|
||
); bearing numerous backward facing strong sharp curved spines, heavy setae, and also long fine setae too matted to assess accurately, but apparently dense.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206503FFC8FFC5DBA74D1AFCE17D6C" pageId="16" pageNumber="317" type="description">
|
||
<paragraph id="8B853688FFC8FFC5DBA74D1AFCE17D6C" blockId="16.[151,1437,151,1401]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBA74D1AFE4E7D25" bold="true" box="[199,455,1302,1329]" pageId="16" pageNumber="317">Measurements (mm):</emphasis>
|
||
body length, 9.9; lateral caudal spikes 6.4; head width 2.6; prementum 2.65; mandibles 0.75; palpal lobe (to tip of dorsal branch) 0.58; movable hook 0.52 maxillary palp 0.57; galeolacinia to distal tooth 1.2; antennae 1.70, 0.74, 0.80, 0.49, 0.24, 0.15, 0.11 (total 4.23).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206503FFC8FFC5DBF74DC6FCBB7F3C" pageId="16" pageNumber="317" type="discussion">
|
||
<paragraph id="8B853688FFC8FFC5DBF74DC6FE807DF0" blockId="16.[151,265,1483,1509]" box="[151,265,1483,1509]" pageId="16" pageNumber="317">
|
||
<emphasis id="B94EEA9AFFC8FFC5DBF74DC6FE807DF0" bold="true" box="[151,265,1483,1509]" pageId="16" pageNumber="317">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B853688FFC8FFC5DBF74E1EFCBB7F3C" blockId="16.[151,1437,1555,1833]" pageId="16" pageNumber="317">
|
||
This species exhibits several distinctive characters when compared with other genera studied: the mask is elongate, as is the head, although the latter is not very obvious. The antennae are long in proportion to head width. The anterior tubercles on the prothorax are reduced and the caudal spikes are very long thin, spiny and distinctively tapered. The posterior segments of the abdomen bear considerably more small spines on the venter and ventral margins than in the other species and the specimens examined were much paler than other genera, but it is unclear if this is a reliable character. Only one other species
|
||
<taxonomicName id="4C3A4D0BFFC8FFC5D9724ECAFDF17EF5" box="[530,632,1735,1761]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="striata">
|
||
<emphasis id="B94EEA9AFFC8FFC5D9724ECAFDF17EF5" box="[530,632,1735,1761]" italics="true" pageId="16" pageNumber="317">S. striata</emphasis>
|
||
</taxonomicName>
|
||
is known from the genus, closely related to
|
||
<taxonomicName id="4C3A4D0BFFC8FFC5DF0A4ECAFB627EF4" box="[1130,1259,1735,1761]" class="Insecta" family="Chlorocyphidae" genus="Sundacypha" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="petiolata">
|
||
<emphasis id="B94EEA9AFFC8FFC5DF0A4ECAFB627EF4" box="[1130,1259,1735,1761]" italics="true" pageId="16" pageNumber="317">S. petiolata</emphasis>
|
||
</taxonomicName>
|
||
. It seems likely that its larvae when discovered will be very similar, but it is possible differences occur especially in the shape of the prementum, to judge from known variation in other genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |