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<document id="C271FA2893834C91776933F1DA08B4F9" ID-DOI="10.11646/zootaxa.5486.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13210005" ID-ZooBank="C7F7E7D5-6F6E-49C8-AF31-9B769EC8B56C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1722788280951" checkinUser="plazi" docAuthor="Orr, Albert G. W., Dow, Rory A. &amp; Steinhoff, Philip O. M." docDate="2024" docId="0393879EFFD6FFDADBF7498AFE297C4C" docLanguage="en" docName="zootaxa.5486.2.3.1.pdf" docOrigin="Zootaxa 5486 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aristocypha fenestrella" docType="treatment" docVersion="3" lastPageNumber="316" masterDocId="FFAAFFE6FFD8FFD5DB60480DFF897815" masterDocTitle="Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family." masterLastPageNumber="337" masterPageNumber="301" pageNumber="315" updateTime="1726085199428" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="645F78FE0620B192AB391DB7653A5E45">Descriptions of larvae of four mainly DNA barcode-matched species of chlorocyphids from south-east Asia (Odonata: Chlorocyphidae) with notes on the generic and species level larval identification of Oriental region members of the family.</mods:title>
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<mods:namePart id="8F8897105117024FAA0B62440CA9E513">Orr, Albert G. W.</mods:namePart>
<mods:affiliation id="017A4C5BCC3E01156C16111E125923D4">Environmental Futures Research Institute, Griffith University, Nathan, Australia.</mods:affiliation>
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<mods:namePart id="432ADFB1BED8C05D3F33B80B11D4C061">Dow, Rory A.</mods:namePart>
<mods:affiliation id="966A436CCA5F6AF26BCE085C8504D789">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands. &amp; Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia.</mods:affiliation>
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<mods:namePart id="6EC9AA55D0AA44223FBE789AA32553DB">Steinhoff, Philip O. M.</mods:namePart>
<mods:affiliation id="F23FF13C53D71A0C530E31AB047DEA5C">Höxter, Germany.</mods:affiliation>
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<mods:title id="4CBE7EE84CFD96903BD77E7548CCA7D4">Zootaxa</mods:title>
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<mods:date id="FD3F40175EDDE722E7FABC4FD2D2A90D">2024</mods:date>
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<treatment id="0393879EFFD6FFDADBF7498AFE297C4C" LSID="urn:lsid:plazi:treatment:0393879EFFD6FFDADBF7498AFE297C4C" httpUri="http://treatment.plazi.org/id/0393879EFFD6FFDADBF7498AFE297C4C" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
<subSubSection id="C3206503FFD6FFDBDBF7498AFDBA79B7" box="[151,563,391,418]" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="8B853688FFD6FFDBDBF7498AFDBA79B7" blockId="14.[151,787,391,455]" box="[151,563,391,418]" pageId="14" pageNumber="315">
<heading id="D0CD81E4FFD6FFDBDBF7498AFDBA79B7" bold="true" box="[151,563,391,418]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<taxonomicName id="4C3A4D0BFFD6FFDBDBF7498AFDBA79B7" ID-CoL="GPJ8" authority="(Rambur)" baseAuthorityName="Rambur" box="[151,563,391,418]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD6FFDBDBF7498AFDBA79B7" bold="true" box="[151,563,391,418]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBF7498AFE2679B7" bold="true" box="[151,431,391,418]" italics="true" pageId="14" pageNumber="315">Aristocypha fenestrella</emphasis>
(Rambur)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3206503FFD6FFDBDBF749A1FC9A79D3" box="[151,787,428,455]" pageId="14" pageNumber="315" type="description">
<paragraph id="8B853688FFD6FFDBDBF749A1FC9A79D3" blockId="14.[151,787,391,455]" box="[151,787,428,455]" pageId="14" pageNumber="315">
(
<figureCitation id="13012A0DFFD6FFDBDBFF49A1FF7F79D3" box="[159,246,428,454]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="14" pageNumber="315">Figs 4b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDA6249A1FEA979D2" box="[258,288,428,455]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="14" pageNumber="315">5b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDA4C49A1FEC179D3" box="[300,328,428,454]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="14" pageNumber="315">6c</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDA3449A1FEFA79D3" box="[340,371,428,454]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="14" pageNumber="315">7b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDA1E49A1FE1579D3" box="[382,412,428,454]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="14" pageNumber="315">8b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDAC849A1FE4E79D3" box="[424,455,428,454]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="14" pageNumber="315">9b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDAB249A1FE7379D3" box="[466,506,428,454]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(ad) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (eh) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="14" pageNumber="315">12f</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD96649A1FDC579D3" box="[518,588,428,454]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">14a, b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD93849A1FD1779D3" box="[600,670,428,455]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="14" pageNumber="315">15a, b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD9CA49A1FD5C79D3" box="[682,725,428,454]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="14" pageNumber="315">17a</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD98149A1FC8379D3" box="[737,778,428,454]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="14" pageNumber="315">18a</figureCitation>
)
</paragraph>
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<subSubSection id="C3206503FFD6FFDBDBF749F9FCAB7A27" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="8B853688FFD6FFDBDBF749F9FCAB7A27" blockId="14.[151,1437,500,2039]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBF749F9FE0B7A1B" bold="true" box="[151,386,500,526]" pageId="14" pageNumber="315">Material examined.</emphasis>
<materialsCitation id="3B523CD5FFD6FFDBDAE949F9FCAB7A27" collectingDate="2008-09-19" collectionCode="RMNH, INS" collectorName="C. Y. Choong." country="Malaysia" latitude="4.613" location="Cameron Highlands" longLatPrecision="78" longitude="101.404" municipality="Tributary of Sungai Telom" pageId="14" pageNumber="315" specimenCount="1" specimenCount-female="1" stateProvince="Pahang">
<collectingCountry id="F32D7618FFD6FFDBDAE949F9FD947A1B" box="[393,541,500,526]" name="Malaysia" pageId="14" pageNumber="315">MALAYSIA</collectingCountry>
,
<specimenCount id="9D3CFD01FFD6FFDBD94A49F9FDDD7A1A" box="[554,596,500,527]" count="1" pageId="14" pageNumber="315" type="female">1 ♀</specimenCount>
F (
<collectionCode id="ED2BAE4DFFD6FFDBD91C49F9FD5F7A1A" box="[636,726,500,527]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED2BAE4DFFD6FFDBD9B949F9FC8E7A1A" box="[729,775,500,527]" pageId="14" pageNumber="315">INS</collectionCode>
.500054),
<collectingMunicipality id="6BE1ACF2FFD6FFDBD81849F9FB2E7A1B" box="[888,1191,500,526]" pageId="14" pageNumber="315">Tributary of Sungai Telom</collectingMunicipality>
,
<location id="8EE56053FFD6FFDBDFD449F9FA117A1B" LSID="urn:lsid:plazi:treatment:0393879EFFD6FFDADBF7498AFE297C4C:8EE56053FFD6FFDBDFD449F9FA117A1B" box="[1204,1432,500,526]" country="Malaysia" latitude="4.613" longLatPrecision="78" longitude="101.404" municipality="Tributary of Sungai Telom" name="Cameron Highlands" pageId="14" pageNumber="315" stateProvince="Pahang">Cameron Highlands</location>
,
<collectingRegion id="49FEF86AFFD6FFDBDBF74A15FF657A27" box="[151,236,536,562]" country="Malaysia" name="Pahang" pageId="14" pageNumber="315">Pahang</collectingRegion>
,
<geoCoordinate id="EE0E504FFFD6FFDBDB984A15FEC77A27" box="[248,334,536,562]" degrees="4.613" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="55" value="4.613">4.613N</geoCoordinate>
,
<geoCoordinate id="EE0E504FFFD6FFDBDA394A15FE417A27" box="[345,456,536,562]" degrees="101.404" direction="east" orientation="longitude" pageId="14" pageNumber="315" precision="55" value="101.404">101.404E</geoCoordinate>
,
<date id="FF841048FFD6FFDBDAB34A15FDC77A27" box="[467,590,536,562]" pageId="14" pageNumber="315" value="2008-09-19">
<collectingDate id="EFC0E9A0FFD6FFDBDAB34A15FDC77A27" box="[467,590,536,562]" pageId="14" pageNumber="315" value="2008-09-19">19 ix 2008</collectingDate>
</date>
, leg.
<collectorName id="26CF535EFFD6FFDBD9E94A15FCAB7A27" box="[649,802,536,562]" pageId="14" pageNumber="315">C.Y. Choong.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3206503FFD6FFDBDBA74A31FC4F7AF2" pageId="14" pageNumber="315" type="biology_ecology">
<paragraph id="8B853688FFD6FFDBDBA74A31FC4F7AF2" blockId="14.[151,1437,500,2039]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74A31FEA67A43" bold="true" box="[199,303,572,598]" pageId="14" pageNumber="315">Habitus:</emphasis>
very similar to
<taxonomicName id="4C3A4D0BFFD6FFDBDAB74A30FDD27A43" box="[471,603,572,598]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDAB74A30FDD27A43" box="[471,603,572,598]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
with the following differences: overall slightly larger with long, thin banded legs relatively slightly shorter (
<figureCitation id="13012A0DFFD6FFDBDA924A6DFDCF7A6F" box="[498,582,608,634]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="14" pageNumber="315">Fig. 4b</figureCitation>
); very similar mottled pattern on the body darker in specimen examined (not figured); sagittate head, antennae and very similar to
<taxonomicName id="4C3A4D0BFFD6FFDBD9874A88FCE57A8B" box="[743,876,644,670]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD9874A88FCE57A8B" box="[743,876,644,670]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
; abdomen obviously narrower and more elongate, bearing distinct lateral keels on most segments; caudal spikes relatively shorter and less tapered than in all other species studied, with numerous short rearward pointing spines and setae.
</paragraph>
</subSubSection>
<subSubSection id="C3206503FFD6FFDADBA74AE2FDC679B8" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315" type="description">
<paragraph id="8B853688FFD6FFDBDBA74AE2FCA87E53" blockId="14.[151,1437,500,2039]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74AE2FE877B1F" bold="true" box="[199,270,751,778]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74AE2FE8C7B1F" bold="true" box="[199,261,751,778]" italics="true" pageId="14" pageNumber="315">Head</emphasis>
:
</emphasis>
in dorsal view as illustrated (
<figureCitation id="13012A0DFFD6FFDBD9374AFDFD207B1E" box="[599,681,752,779]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="14" pageNumber="315">Fig. 5b</figureCitation>
), almost identical to
<taxonomicName id="4C3A4D0BFFD6FFDBD8F24AFCFB917B1F" box="[914,1048,752,778]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD8F24AFCFB917B1F" box="[914,1048,752,778]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
except antennae relatively slightly shorter (1.44 times maximum head width versus
<quantity id="4CC29B6DFFD6FFDBD9BD4B19FCB97B3B" box="[733,816,788,815]" metricMagnitude="-2" metricUnit="m" metricValue="3.8608" pageId="14" pageNumber="315" unit="in" value="1.52">1.52 in</quantity>
<taxonomicName id="4C3A4D0BFFD6FFDBD85C4B18FC4E7B3B" box="[828,967,788,814]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD85C4B18FC4E7B3B" box="[828,967,788,814]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
) and markings stronger and with slight differences but tip of pedicel with similar dark coloration; scape of similar length but with longer and mainly straighter setae so that even wet specimens appear smooth, rather than warty (
<figureCitation id="13012A0DFFD6FFDBDF6E4B51FBE87B63" box="[1038,1121,860,886]" captionStart="FIGURE 6" captionStartId="9.[151,250,1855,1879]" captionTargetBox="[153,1429,242,1805]" captionTargetId="figure-18@9.[151,1436,181,1831]" captionTargetPageId="9" captionText="FIGURE 6. (a) Frontal view of labrum of Heliocypha biseriata; (b, c) detail of scape showing short recurved and longer, mainly straighter setae on H. biseriata and Aristocypha fenestrella respectively; (d) Fore-tarsus in H. biseriata and (e) detail of pectinate setae on ventral side of same. Scale bars as indicated." figureDoi="http://doi.org/10.5281/zenodo.13210017" httpUri="https://zenodo.org/record/13210017/files/figure.png" pageId="14" pageNumber="315">Fig. 6c</figureCitation>
). Labrum as in
<taxonomicName id="4C3A4D0BFFD6FFDBDE754B50FA147B63" box="[1301,1437,860,886]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDE754B50FA147B63" box="[1301,1437,860,886]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
except moustache with 14 setae. Ventrally, genae bear a cluster of subocular spines longer and less numerous than in
<taxonomicName id="4C3A4D0BFFD6FFDBDBD44BA8FEB27BAB" box="[180,315,932,958]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDBD44BA8FEB27BAB" box="[180,315,932,958]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
, with anterior strong three distinctive, being longer, more curved and sharply acuminate; remaining spines differ as shown (
<figureCitation id="13012A0DFFD6FFDBDACC4BC5FD8A7BF7" box="[428,515,968,994]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="14" pageNumber="315">Fig. 7b</figureCitation>
). Proportions and other features of head seen in lateral view very similar to
<taxonomicName id="4C3A4D0BFFD6FFDBDEE14BC4FF727C13" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDEE14BC4FF727C13" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
with small differences as shown (
<figureCitation id="13012A0DFFD6FFDBD9E04BE1FD667C13" box="[640,751,1004,1031]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[228,1386,186,1919]" captionTargetId="figure-22@10.[155,1433,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Left lateral view of head (left) and ventral view of genae showing spines (right): (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bars left side, 1 mm; right side 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210019" httpUri="https://zenodo.org/record/13210019/files/figure.png" pageId="14" pageNumber="315">Fig. 7a, b</figureCitation>
), some of which may relate to subtle differences in angle of view, although ventral spur at margin of gena and occiput is clearly directed forward more strongly and the post occipital dorsal tubercle set further back in
<taxonomicName id="4C3A4D0BFFD6FFDBD91D4C39FC9B7C5B" box="[637,786,1076,1102]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD6FFDBD91D4C39FC9B7C5B" box="[637,786,1076,1102]" italics="true" pageId="14" pageNumber="315">A. fenestrella</emphasis>
</taxonomicName>
. Mask long and thin, articulation resting between anterior part of coxae of mesothorax (
<figureCitation id="13012A0DFFD6FFDBDABD4C55FDA77C67" box="[477,558,1112,1138]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="14" pageNumber="315">Fig. 8b</figureCitation>
) as in
<taxonomicName id="4C3A4D0BFFD6FFDBD9164C54FD757C67" box="[630,764,1112,1138]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD9164C54FD757C67" box="[630,764,1112,1138]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
; prementum narrow basally, gradually flared anteriorly with margins of anterior half weakly convex, slightly more so than in
<taxonomicName id="4C3A4D0BFFD6FFDBD80F4C70FC7F7C83" box="[879,1014,1148,1174]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD80F4C70FC7F7C83" box="[879,1014,1148,1174]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
; similarly lateral margins with sparse fine setae on distal 2/3
<superScript id="7C4F9BC0FFD6FFDBDAF24CADFE297CBB" attach="left" box="[402,416,1184,1198]" fontSize="6" pageId="14" pageNumber="315">rd</superScript>
(
<figureCitation id="13012A0DFFD6FFDBDACC4CADFE747CAF" box="[428,509,1184,1210]" captionStart="FIGURE 9" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[244,1341,217,1810]" captionTargetId="figure-18@17.[155,1433,181,1919]" captionTargetPageId="17" captionText="FIGURE 9. Prementum in dorsal view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina, (e) Libellago lineata. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210023" httpUri="https://zenodo.org/record/13210023/files/figure.png" pageId="14" pageNumber="315">Fig. 9b</figureCitation>
) seated in slightly raised sockets (
<figureCitation id="13012A0DFFD6FFDBD81B4CADFC507CAF" box="[891,985,1184,1210]" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="14" pageNumber="315">Fig. 11a</figureCitation>
); anterior median lobe as in
<taxonomicName id="4C3A4D0BFFD6FFDBDE774CACFA147CAF" box="[1303,1437,1184,1210]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDE774CACFA147CAF" box="[1303,1437,1184,1210]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
but slightly more produced; median cleft almost completely closed with tinted groove marking line of closure (
<figureCitation id="13012A0DFFD6FFDBDE0F4CC9FF487D17" captionStart="FIGURE 11" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-18@19.[155,1433,181,1919]" captionTargetPageId="19" captionText="FIGURE 11. Details of anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below): (a) Aristocypha fenestrella, (b) Sundacypha petiolata, (c) Libellago hyalina. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210027" httpUri="https://zenodo.org/record/13210027/files/figure.png" pageId="14" pageNumber="315">Fig. 11a</figureCitation>
); lobe of labial palp bifid with shorter process on each branch as in
<taxonomicName id="4C3A4D0BFFD6FFDBD8CC4CE4FBBB7D17" box="[940,1074,1256,1282]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD8CC4CE4FBBB7D17" box="[940,1074,1256,1282]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
with slight differences in degree of torsion of distal lobe and form of processes as indicated (
<figureCitation id="13012A0DFFD6FFDBD8294D01FC287D33" box="[841,929,1292,1318]" captionStart="FIGURE 8" captionStartId="11.[151,250,1675,1699]" captionTargetBox="[160,1432,215,1639]" captionTargetId="figure-18@11.[151,1436,181,1652]" captionTargetPageId="11" captionText="FIGURE 8. Ventral view of head with mask at rest and anterior two thoracic segments in ventral view: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210021" httpUri="https://zenodo.org/record/13210021/files/figure.png" pageId="14" pageNumber="315">Figs 8a</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD8CE4D01FB9A7D33" box="[942,1043,1292,1318]" captionStart="FIGURE 10" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[167,1433,182,1916]" captionTargetId="figure-22@18.[155,1433,181,1919]" captionTargetPageId="18" captionText="FIGURE 10. Details of anterior prementum and labial palp Heliocypha biseriata: (a) anterior prementum and labial palp in dorsal view (base of movable hook dotted to reveal details of palpal lobe below); (b) labial palp and anterior margin of prementum, frontal view; (c) labial palp, interior view. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210025" httpUri="https://zenodo.org/record/13210025/files/figure.png" pageId="14" pageNumber="315">10a, b, c</figureCitation>
); movable hook long and strongly curved inward. Maxillae general form and arrangement of setae as in
<taxonomicName id="4C3A4D0BFFD6FFDBD8D24D3CFBB07D5F" box="[946,1081,1328,1354]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBD8D24D3CFBB07D5F" box="[946,1081,1328,1354]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
; apical tooth with row of three inner denticles (
<figureCitation id="13012A0DFFD6FFDBDA294D59FE2F7D7B" box="[329,422,1364,1390]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[205,1382,181,1764]" captionTargetId="figure-22@20.[205,1382,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Details of maxillae—(ad) Right maxillae: Heliocypha biseriata, (a) ventral view, (b) oral view; (c) Sundacypha petiolata distal detail, oral view; (d) L. hyalina oral view. (eh) Left maxillae, oral view, details of terminal strong spine with three shorter basal to median supplementary spines: (e) Heliocypha biseriata, (f) Aristocypha fenestrella, (g) Sundacypha petiolata, (h) Libellago hyalina. Scale bars: above 1 mm; below 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13210029" httpUri="https://zenodo.org/record/13210029/files/figure.png" pageId="14" pageNumber="315">Fig. 12f</figureCitation>
), all well separated, longest one apicad, all finer and longer than in
<taxonomicName id="4C3A4D0BFFD6FFDBDFF34D58FA907D7B" box="[1171,1305,1364,1390]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDFF34D58FA907D7B" box="[1171,1305,1364,1390]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
. Mandibles (
<figureCitation id="13012A0DFFD6FFDBDBFF4D75FE977D87" box="[159,286,1400,1426]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">Figs 14a, b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDA4A4D75FEE77D87" box="[298,366,1400,1427]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="14" pageNumber="315">15a, b</figureCitation>
)—both left and right with five distinct teeth on incisor visible in internal view; right with small supplementary tooth basal to strong ventral tooth; right molar crest a low swelling with two strong, sharp spines, the ventral being longer, heavily sclerotised (
<figureCitation id="13012A0DFFD6FFDBD9064DCDFD4C7DCF" box="[614,709,1472,1498]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">Fig. 14a</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBD9B04DCDFD747DCE" box="[720,765,1472,1499]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="14" pageNumber="315">15b</figureCitation>
); left incisor (
<figureCitation id="13012A0DFFD6FFDBD8F94DCDFC767DCF" box="[921,1023,1472,1498]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">Figs 14b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDF6A4DCDFBBD7DCE" box="[1034,1076,1472,1499]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="14" pageNumber="315">15a</figureCitation>
) with five well developed teeth, the strongest ventrally; molar crest a strong flat dorso-ventral blade with 13 minute denticles along a convex distal margin, with end denticles markedly heavier and longer, especially ventrally (
<figureCitation id="13012A0DFFD6FFDBDF664E05FBE57E37" box="[1030,1132,1544,1570]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">Figs 14b</figureCitation>
,
<figureCitation id="13012A0DFFD6FFDBDF184E05FB2D7E36" box="[1144,1188,1544,1571]" captionStart="FIGURE 15" captionStartId="23.[151,250,1548,1572]" captionTargetBox="[198,1381,260,1486]" captionTargetId="figure-18@23.[164,1423,181,1524]" captionTargetPageId="23" captionText="FIGURE 15. Mandibles in interior view: (a) Aristocypha fenestrella L, (b) A. fenestrella R (rotated slightly outward to expose part of venter); (c) Sundacypha petiolata L, (d) S. petiolata R; (e) Libellago hyalina L, (f) L. hyalina R. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210035" httpUri="https://zenodo.org/record/13210035/files/figure.png" pageId="14" pageNumber="315">15a</figureCitation>
). Strong basal seta on both mandibles as in
<taxonomicName id="4C3A4D0BFFD6FFDBDAE54E20FD827E53" box="[389,523,1580,1606]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDAE54E20FD827E53" box="[389,523,1580,1606]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
(not visible in
<figureCitation id="13012A0DFFD6FFDBD9D54E21FC9D7E53" box="[693,788,1580,1606]" captionStart="FIGURE 14" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[202,1392,181,1919]" captionTargetId="figure-22@22.[193,1394,181,1919]" captionTargetPageId="22" captionText="FIGURE 14. Mandibles in ventral view: (a) Aristocypha fenestrella R, (b) A. fenestrella L; (c) Sundacypha petiolata R, (d) S. petiolata L; (e) Libellago hyalina R, (f) L. hyalina L. R &amp; L signify right and left below, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210033" httpUri="https://zenodo.org/record/13210033/files/figure.png" pageId="14" pageNumber="315">Fig. 14a</figureCitation>
).
</paragraph>
<paragraph id="8B853688FFD6FFDBDBA74E42FD1A7F0A" blockId="14.[151,1437,500,2039]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74E42FEAF7E7F" bold="true" box="[199,294,1615,1642]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74E42FE947E7F" bold="true" box="[199,285,1615,1642]" italics="true" pageId="14" pageNumber="315">Thorax</emphasis>
:
</emphasis>
prothorax saddle-shaped with anterior and posterior flanges and tubercles as in
<taxonomicName id="4C3A4D0BFFD6FFDBDFC64E5CFAA47E7F" box="[1190,1325,1616,1642]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDFC64E5CFAA47E7F" box="[1190,1325,1616,1642]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
(
<figureCitation id="13012A0DFFD6FFDBDE5B4E5DFA067E7E" box="[1339,1423,1616,1643]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[206,1300,192,1855]" captionTargetId="figure-22@8.[196,1391,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Detail of heads, antennae and prothoraces of: (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210015" httpUri="https://zenodo.org/record/13210015/files/figure.png" pageId="14" pageNumber="315">Fig. 5b</figureCitation>
). Marking (not illustrated on body) pale and mottled but always with broad pale band pale between two anterior tubercles on prothorax. Wing sheaths slightly spatulate, not completely obscuring abdomen at sides, forewing reaching to end of S4, hindwing to 2/3 along S5 (
<figureCitation id="13012A0DFFD6FFDBD9DF4EB1FC9A7EC3" box="[703,787,1724,1750]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[302,1377,197,1855]" captionTargetId="figure-18@7.[172,1416,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Habitus of (a) Heliocypha biseriata, (b) Aristocypha fenestrella, (c) Sundacypha petiolata, (d) Libellago hyalina. Markings shown only on legs and antennae. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210013" httpUri="https://zenodo.org/record/13210013/files/figure.png" pageId="14" pageNumber="315">Fig. 4b</figureCitation>
). Legs long and thin, metafemur reaching midpoint of S6, clearly shorter than in
<taxonomicName id="4C3A4D0BFFD6FFDBDAF44EECFD927EEF" box="[404,539,1760,1786]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDAF44EECFD927EEF" box="[404,539,1760,1786]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
which they resemble in all other respects including the presence of two rows of pectinate setae on the underside of the tarsus.
</paragraph>
<paragraph id="8B853688FFD6FFDADBA74F2AFD1D7908" blockId="14.[151,1437,500,2039]" lastBlockId="15.[151,1436,151,429]" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74F2AFEB67F57" bold="true" box="[199,319,1831,1858]" pageId="14" pageNumber="315">
<emphasis id="B94EEA9AFFD6FFDBDBA74F2AFEBC7F57" bold="true" box="[199,309,1831,1858]" italics="true" pageId="14" pageNumber="315">Abdomen</emphasis>
:
</emphasis>
squat, mainly dark with well-defined pale narrow middorsal stripe (not figured). Margins of S28 distinctly crimped forming shallow lateral keels; dorsal surface similar to
<taxonomicName id="4C3A4D0BFFD6FFDBDF544F40FB377F73" box="[1076,1214,1868,1894]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD6FFDBDF544F40FB377F73" box="[1076,1214,1868,1894]" italics="true" pageId="14" pageNumber="315">H. biseriata</emphasis>
</taxonomicName>
with sparser setae; ventrolateral margin of S9 with ca 4 strong, curved spines intermingled with 45 smaller spines and scattering of tiny marginal and submarginal spines along venter; S10 hind-margin with 4 strong rearward pointing ventrolateral spines (
<figureCitation id="13012A0DFFD6FFDBDB8D4FB5FECE7FC7" box="[237,327,1976,2002]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="14" pageNumber="315">Fig.17a</figureCitation>
). Male gonapophyses (
<figureCitation id="13012A0DFFD6FFDBD9294FB5FD2F7FC7" box="[585,678,1976,2002]" captionStart="FIGURE 17" captionStartId="25.[151,250,1625,1649]" captionTargetBox="[155,1421,190,1546]" captionTargetId="figure-18@25.[151,1436,181,1601]" captionTargetPageId="25" captionText="FIGURE 17. Male gonapophyses and terminal abdominal segments in ventral view: (a) A. fenestrella, (b) S. petiolata, (c) L. hyalina, and in lateral view: (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210039" httpUri="https://zenodo.org/record/13210039/files/figure.png" pageId="14" pageNumber="315">Fig. 17a</figureCitation>
) short and conical, well separated, arising near posterior margin S9 and just reaching margin hind margin. Female gonapophyses unknown. Epiproct, short, subconical, not obviously different from
<taxonomicName id="4C3A4D0BFFD7FFDADA23489AFE4778A4" box="[323,462,151,177]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD7FFDADA23489AFE4778A4" box="[323,462,151,177]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
</taxonomicName>
. Caudal spikes about 40% body length, broad for most of their length and tapered abruptly to a point subapically (
<figureCitation id="13012A0DFFD7FFDADA9C48B6FDD478C0" box="[508,605,187,213]" captionStart="FIGURE 18" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[191,1381,197,1917]" captionTargetId="figure-18@27.[189,1399,181,1919]" captionTargetPageId="27" captionText="FIGURE 18. Caudal spikes scaled in proportion to overall body length: (a) H. biseriata, (b) A. fenstrella, (c) S. petiolata, (d) L. hyalina. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210041" httpUri="https://zenodo.org/record/13210041/files/figure.png" pageId="15" pageNumber="316">Fig. 18b</figureCitation>
); bearing numerous backward facing strong sharp spines on outer margin, these rather sparse on inner margin; heavy setae, and also long fine setae too entangled to assess accurately, but appear to be less profuse than in
<taxonomicName id="4C3A4D0BFFD7FFDAD967490EFD047908" box="[519,653,259,285]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD7FFDAD967490EFD047908" box="[519,653,259,285]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B853688FFD7FFDADBA7492AFDC679B8" blockId="15.[151,1436,151,429]" pageId="15" pageNumber="316">
<emphasis id="B94EEA9AFFD7FFDADBA7492AFE617955" bold="true" box="[199,488,294,321]" pageId="15" pageNumber="316">Measurements (in mm):</emphasis>
body length excluding antennae, caudal appendages and spikes 12.4; lateral caudal gills 5.3; head width 2.81; prementum 2.45; palpal lobe (to tip of dorsal branch) 0.80; movable hook 0.68; maxillae base to distal hook of galeolacinea (1.26), maxillary palp (0.55); mandibles long axis (0.86); antenna 1.70, 0.65, 0.70, 0.50, 0.29, 0.16, 0.07 (total 4.07).
</paragraph>
</subSubSection>
<subSubSection id="C3206503FFD7FFDADBF749F2FE297C4C" pageId="15" pageNumber="316" type="discussion">
<paragraph id="8B853688FFD7FFDADBF749F2FE807A0C" blockId="15.[151,265,511,537]" box="[151,265,511,537]" pageId="15" pageNumber="316">
<emphasis id="B94EEA9AFFD7FFDADBF749F2FE807A0C" bold="true" box="[151,265,511,537]" pageId="15" pageNumber="316">Remarks</emphasis>
</paragraph>
<paragraph id="8B853688FFD7FFDADBF74A4AFE297C4C" blockId="15.[151,1437,583,1113]" pageId="15" pageNumber="316">
The principal diagnostic differences between
<taxonomicName id="4C3A4D0BFFD7FFDAD9F24A4AFCAF7A74" box="[658,806,583,609]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDAD9F24A4AFCAF7A74" box="[658,806,583,609]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0BFFD7FFDAD83B4A4AFC687A74" box="[859,993,583,609]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD7FFDAD83B4A4AFC687A74" box="[859,993,583,609]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
</taxonomicName>
are the longer subocular anterior spines on the genae and the relatively shorter caudal spikes, tapering to a point abruptly near apex in the former. The length of the spikes in proportion to body length is ca 40% in
<taxonomicName id="4C3A4D0BFFD7FFDAD8664A82FC127ABC" box="[774,923,655,681]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDAD8664A82FC127ABC" box="[774,923,655,681]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
versus ca 55% in
<taxonomicName id="4C3A4D0BFFD7FFDADF0A4A82FB787ABC" box="[1130,1265,655,681]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="biseriata">
<emphasis id="B94EEA9AFFD7FFDADF0A4A82FB787ABC" box="[1130,1265,655,681]" italics="true" pageId="15" pageNumber="316">H. biseriata</emphasis>
</taxonomicName>
. The legs in
<taxonomicName id="4C3A4D0BFFD7FFDADEE44A82FE857AD8" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDADEE44A82FE857AD8" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
are proportionally shorter. There are also clear differences in the armature of the maxillae and dentition of the mandibles and the setae on the scape of the antenna are mainly long and do not create a warty appearance when wet. The larvae of two other species of
<taxonomicName id="4C3A4D0BFFD7FFDAD9044AF6FD627B00" box="[612,747,763,789]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9AFFD7FFDAD9044AF6FD627B00" box="[612,747,763,789]" italics="true" pageId="15" pageNumber="316">Aristocypha</emphasis>
</taxonomicName>
have been described in some detail.
<taxonomicName id="4C3A4D0BFFD7FFDADFF84AF6FA157B00" box="[1176,1436,763,789]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="trifasciata">
<emphasis id="B94EEA9AFFD7FFDADFF84AF6FA157B00" box="[1176,1436,763,789]" italics="true" pageId="15" pageNumber="316">Aristocypha trifasciata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0BFFD7FFDADBA84B12FE127B2C" box="[200,411,799,825]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis id="B94EEA9AFFD7FFDADBA84B12FE127B2C" box="[200,411,799,825]" italics="true" pageId="15" pageNumber="316">A. quadrimaculata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAB4B79FFD7FFDADAA74B12FD527B2C" author="Kumar, A. &amp; Prasad, M." box="[455,731,799,825]" pageId="15" pageNumber="316" pagination="547 - 554" refId="ref15440" refString="Kumar, A. &amp; Prasad, M. (1977) On the larvae of Rhinocypha (Odonata: Chlorocyphidae) from Garhwal hills. Oriental Insects, 11, 547 - 554. https: // doi. org / 10.1080 / 00305316.1977.11090927" type="journal article" year="1977">Kumar &amp; Prasad (1977)</bibRefCitation>
. Both species differ from
<taxonomicName id="4C3A4D0BFFD7FFDADF634B12FB0D7B2C" box="[1027,1156,799,825]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9AFFD7FFDADF634B12FB0D7B2C" box="[1027,1156,799,825]" italics="true" pageId="15" pageNumber="316">Heliocypha</emphasis>
</taxonomicName>
in possessing long setae on the scape without a warty appearance and the caudal spikes are proportionally shorter (ca 40% body length), and are not tapered until near the apex, as in
<taxonomicName id="4C3A4D0BFFD7FFDAD9FE4B6AFCBA7B94" box="[670,819,871,897]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDAD9FE4B6AFCBA7B94" box="[670,819,871,897]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
. In other respects consistent genus specific characters are not evident. For example in
<taxonomicName id="4C3A4D0BFFD7FFDADA9D4B86FD067BB0" box="[509,655,907,933]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="trifasciata">
<emphasis id="B94EEA9AFFD7FFDADA9D4B86FD067BB0" box="[509,655,907,933]" italics="true" pageId="15" pageNumber="316">A. trifasciata</emphasis>
</taxonomicName>
the subocular spines on the genae are small and scattered, whereas in
<taxonomicName id="4C3A4D0BFFD7FFDADBF74BA2FEE37BDC" box="[151,362,943,969]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis id="B94EEA9AFFD7FFDADBF74BA2FEE37BDC" box="[151,362,943,969]" italics="true" pageId="15" pageNumber="316">A. quadrimaculata</emphasis>
</taxonomicName>
they are very similar to those found in
<taxonomicName id="4C3A4D0BFFD7FFDAD84B4BA2FC497BDC" box="[811,960,943,969]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDAD84B4BA2FC497BDC" box="[811,960,943,969]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
. Conversely, in
<taxonomicName id="4C3A4D0BFFD7FFDADF194BA2FA857BDC" box="[1145,1292,943,969]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="trifasciata">
<emphasis id="B94EEA9AFFD7FFDADF194BA2FA857BDC" box="[1145,1292,943,969]" italics="true" pageId="15" pageNumber="316">A. trifasciata</emphasis>
</taxonomicName>
, the legs are relatively shorter than in
<taxonomicName id="4C3A4D0BFFD7FFDADADA4BDEFDD97BF8" box="[442,592,979,1005]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDADADA4BDEFDD97BF8" box="[442,592,979,1005]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C3A4D0BFFD7FFDAD9844BDEFC307BF8" box="[740,953,979,1005]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="quadrimaculata">
<emphasis id="B94EEA9AFFD7FFDAD9844BDEFC307BF8" box="[740,953,979,1005]" italics="true" pageId="15" pageNumber="316">A. quadrimaculata</emphasis>
</taxonomicName>
they are much longer. In both species the prementum, while of similar general shape to
<taxonomicName id="4C3A4D0BFFD7FFDAD9C14BFAFCBF7C04" box="[673,822,1015,1041]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="fenestrella">
<emphasis id="B94EEA9AFFD7FFDAD9C14BFAFCBF7C04" box="[673,822,1015,1041]" italics="true" pageId="15" pageNumber="316">A. fenestrella</emphasis>
</taxonomicName>
, is much wider anteriorly and the median cleft is well defined. Hence only the terminal shape of the caudal spikes consistently separates
<taxonomicName id="4C3A4D0BFFD7FFDADF374C16FB517C20" box="[1111,1240,1051,1077]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9AFFD7FFDADF374C16FB517C20" box="[1111,1240,1051,1077]" italics="true" pageId="15" pageNumber="316">Heliocypha</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D0BFFD7FFDADE754C16FA157C20" box="[1301,1436,1051,1077]" class="Insecta" family="Chlorocyphidae" genus="Aristocypha" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEA9AFFD7FFDADE754C16FA157C20" box="[1301,1436,1051,1077]" italics="true" pageId="15" pageNumber="316">Aristocypha</emphasis>
</taxonomicName>
based on known larvae.
</paragraph>
</subSubSection>
</treatment>
</document>