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<document id="3EEA4E0A4FF161440FAB655102A5682E" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FFBBA921A7D8B3E6FA9EFEBC" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole uranus Salata &amp; Fisher 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="54" updateTime="1698850390174" updateUser="ExternalLinkService">
<mods:mods id="306D2F0BFA964F841FA715DF27A57F0A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E072DEB3685A6873865CD4F1A75A34E">
<mods:title id="485231D97BAF03BE059DBBD2D7DD191E">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
</mods:titleInfo>
<mods:name id="FAB79386E60F83B86AA67958C07B7770" type="personal">
<mods:role id="632C5BF2074CD9C6E1F57A1C7DF4B1D0">
<mods:roleTerm id="6F528BF49152375CDF28945D959B6C31">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="014B552B48AC61D117A7D99E7D82B354">Salata, Sebastian</mods:namePart>
</mods:name>
<mods:name id="8AD271249932E5F6DF0ABA485E23307A" type="personal">
<mods:role id="A466D9A720F2F9787FAAA4D5F8265348">
<mods:roleTerm id="658BC373666002035B0593E7DAF5991F">Author</mods:roleTerm>
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<mods:namePart id="1A6EE2F9B82DEA81A8E648E636220D51">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="727AB0BAE669670DDF73431314D159E0" type="ORCID">0000-0002-4653-3270</mods:nameIdentifier>
<mods:nameIdentifier id="6A9B996A1DB008662991DD1CBF9B44C5" type="email">bfisher@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="054AC405AE9D87F0D4455DB656BB41EA">text</mods:typeOfResource>
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<mods:titleInfo id="A20D2EEFE1C7096CEDF4F8B6C471E4E6">
<mods:title id="BA3A80E2EA9EF9D078F287322846A288">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EBA45D9E35BA61FBF45F83C4B9EFD982">
<mods:date id="FEA76D5AF2D3D16C8D48CEDDA6275AD6">2020</mods:date>
<mods:detail id="45E5C668B5B157EA44A2770E6114F686" type="pubDate">
<mods:number id="C0AD695786E4A083713895E6036018EF">2020-08-31</mods:number>
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<mods:detail id="806E26131E8476B3B2E013D72A513769" type="volume">
<mods:number id="34912C90664C1103E4EDFBC1B3F08AA5">4843</mods:number>
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<mods:number id="BC61E7FB93D165E59E9F54F6D729BAA6">1</mods:number>
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<mods:classification id="B1D35F26F9F2F20BD1DD0C757BA0D7DD">journal article</mods:classification>
<mods:identifier id="7349B83960F52BE1E843C90A5441BAB9" type="CLB-Dataset">8556</mods:identifier>
<mods:identifier id="9C9D350B4F0D1F5BFE4797DD7F60C368" type="DOI">10.11646/zootaxa.4843.1.1</mods:identifier>
<mods:identifier id="2E3A41C6661005407165CDB632DD8490" type="GBIF-Dataset">a2aef684-44f1-429d-9dd5-dd394539fa36</mods:identifier>
<mods:identifier id="6D95D89D233244C8F8C8FD2D9C395677" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="51785A7540E8F4E35EF36B7896DFA906" type="Zenodo-Dep">4405729</mods:identifier>
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</mods:mods>
<treatment id="03938025FFBBA921A7D8B3E6FA9EFEBC" ID-DOI="http://doi.org/10.5281/zenodo.4488409" ID-GBIF-Taxon="177202114" ID-Zenodo-Dep="4488409" LSID="urn:lsid:plazi:treatment:03938025FFBBA921A7D8B3E6FA9EFEBC" httpUri="http://treatment.plazi.org/id/03938025FFBBA921A7D8B3E6FA9EFEBC" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<subSubSection id="C32062B8FFBBA923A7D8B3E6FE02F9F4" box="[151,452,1518,1545]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8B853133FFBBA923A7D8B3E6FE02F9F4" blockId="53.[151,452,1518,1581]" box="[151,452,1518,1545]" pageId="53" pageNumber="54">
<heading id="D0CD865FFFBBA923A7D8B3E6FE02F9F4" bold="true" box="[151,452,1518,1545]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
<emphasis id="B94EED21FFBBA923A7D8B3E6FE02F9F4" bold="true" box="[151,452,1518,1545]" pageId="53" pageNumber="54">
<taxonomicName id="4C3A4AB0FFBBA923A7D8B3E6FE98F9F4" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,350,1518,1545]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="uranus" status="sp. nov.">
<emphasis id="B94EED21FFBBA923A7D8B3E6FE98F9F4" bold="true" box="[151,350,1518,1545]" italics="true" pageId="53" pageNumber="54">Pheidole uranus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D505AFFBBA923A629B3E7FE02F9F4" box="[358,452,1519,1545]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBBA923A7D8B01BFE4CF9D0" box="[151,394,1555,1581]" pageId="53" pageNumber="54" type="description">
<paragraph id="8B853133FFBBA923A7D8B01BFE4CF9D0" blockId="53.[151,452,1518,1581]" box="[151,394,1555,1581]" pageId="53" pageNumber="54">
<figureCitation id="13012DB6FFBBA923A7D8B01BFF2FF9D0" box="[151,233,1555,1581]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="53" pageNumber="54">Figs 24</figureCitation>
A–F, 27P, 28P
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBBA923A7D8B053FC92F8B0" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8B853133FFBBA923A7D8B053FB81F940" blockId="53.[151,1436,1627,2013]" pageId="53" pageNumber="54">
<materialsCitation id="3B523B6EFFBBA923A7D8B053FF30F940" ID-GBIF-Occurrence="3027954664" accessionNumber="BLF36505" collectingDate="2015-02-26" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="1125" latitude="-24.13894" location="Anosyenne Mts" longLatPrecision="1" longitude="47.06804" municipality="Anosy Region" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="54818F91FFBBA923A7D8B053FEE9F988" box="[151,303,1627,1653]" pageId="53" pageNumber="54" type="holotype">HOLOTYPE</typeStatus>
: 1s.,
<collectingCountry id="F32D71A3FFBBA923A63FB053FE3FF988" box="[368,505,1627,1653]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FFBBA923A548B053FD9FF988" box="[519,601,1627,1653]" country="Madagascar" name="Toliara" pageId="53" pageNumber="54">Toliara</collectingRegion>
,
<collectingMunicipality id="6BE1AB49FFBBA923A529B053FCC8F988" box="[614,782,1627,1653]" pageId="53" pageNumber="54">Anosy Region</collectingMunicipality>
,
<location id="8EE567E8FFBBA923A453B053FC15F988" LSID="urn:lsid:plazi:treatment:03938025FFBBA921A7D8B3E6FA9EFEBC:8EE567E8FFBBA923A453B053FC15F988" box="[796,979,1627,1653]" country="Madagascar" latitude="-24.13894" longLatPrecision="1" longitude="47.06804" municipality="Anosy Region" name="Anosyenne Mts" pageId="53" pageNumber="54" stateProvince="Toliara">Anosyenne Mts</location>
,
<locationDeviation id="9A372811FFBBA923A4ADB053FADDF988" box="[994,1307,1627,1653]" location="Manantenina" pageId="53" pageNumber="54">
<quantity id="4CC29CD6FFBBA923A4ADB053FB84F988" box="[994,1090,1627,1653]" metricMagnitude="4" metricUnit="m" metricValue="3.12" pageId="53" pageNumber="54" unit="km" value="31.2">31.2 km</quantity>
NW Manantenina
</locationDeviation>
,
<geoCoordinate id="EE0E57F4FFBBA923A265B053FA5AF988" box="[1322,1436,1627,1653]" degrees="24.13894" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-24.13894">-24.13894</geoCoordinate>
<geoCoordinate id="EE0E57F4FFBBA923A7D8B077FEC5F964" box="[151,259,1663,1689]" degrees="47.06804" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="47.06804">47.06804</geoCoordinate>
,
<quantity id="4CC29CD6FFBBA923A65DB076FEADF964" box="[274,363,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="53" pageNumber="54" unit="m" value="1125.0">
<elevation id="0017D600FFBBA923A65DB076FEADF964" box="[274,363,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="53" pageNumber="54" unit="m" value="1125.0">1125 m</elevation>
</quantity>
,
<date id="FF8417F3FFBBA923A635B076FDC8F964" box="[378,526,1662,1689]" pageId="53" pageNumber="54" value="2015-02-26">
<collectingDate id="EFC0EE1BFFBBA923A635B076FDC8F964" box="[378,526,1662,1689]" pageId="53" pageNumber="54" value="2015-02-26">26-Feb-2015</collectingDate>
</date>
, rainforest, ex root mat,
<collectorName id="26CF54E5FFBBA923A47EB077FC78F964" box="[817,958,1663,1689]" pageId="53" pageNumber="54">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFBBA923A486B077FC3BF964" box="[969,1021,1663,1689]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. leg. BLF36505, CASENT0704257 (
<collectionCode id="ED2BA9F6FFBBA923A7D0B0ABFF28F940" box="[159,238,1699,1725]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFFBBA923A64EB0ABFB85F940" ID-GBIF-Occurrence="3027954425" accessionNumber="BLF36505" box="[257,1091,1699,1725]" collectingDate="2015-02-26" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="1125" latitude="-24.13894" location="Anosyenne Mts" longLatPrecision="1" longitude="47.06804" municipality="Anosy Region" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="54818F91FFBBA923A64EB0ABFE66F940" box="[257,416,1699,1725]" pageId="53" pageNumber="54" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3CFABAFFBBA923A6E5B0ABFE15F940" box="[426,467,1699,1725]" pageId="53" pageNumber="54" type="generic">1w.</specimenCount>
, the same data as holotype, CASENT0923260 (
<collectionCode id="ED2BA9F6FFBBA923A4A4B0ABFBFCF940" box="[1003,1082,1699,1725]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FFBBA923A788B0CEFB44F8D4" blockId="53.[151,1436,1627,2013]" pageId="53" pageNumber="54">
<emphasis id="B94EED21FFBBA923A788B0CEFE47F91D" bold="true" box="[199,385,1734,1760]" pageId="53" pageNumber="54">Other material.</emphasis>
<materialsCitation id="3B523B6EFFBBA923A6C7B0CFFD0DF91C" ID-GBIF-Occurrence="3027954498" box="[392,715,1735,1761]" country="Madagascar" location="Toliara" pageId="53" pageNumber="54" specimenCount="4" stateProvince="Toliara">
<collectingCountry id="F32D71A3FFBBA923A6C7B0CFFDD7F91C" box="[392,529,1735,1761]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
.
<collectingRegion id="49FEFFD1FFBBA923A554B0CFFDA8F91C" box="[539,622,1735,1761]" country="Madagascar" name="Toliara" pageId="53" pageNumber="54">Toliara</collectingRegion>
:
<specimenCount id="9D3CFABAFFBBA923A537B0CFFD64F91C" box="[632,674,1735,1761]" pageId="53" pageNumber="54" type="generic">4w.</specimenCount>
, 2s
</materialsCitation>
.,
<materialsCitation id="3B523B6EFFBBA923A594B0CFFC55F8F8" ID-GBIF-Occurrence="3027954730" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="1" latitude="-24.13894" location="Anosyenne Mts" longLatPrecision="1" longitude="47.06804" municipality="Anosy Region" pageId="53" pageNumber="54" specimenCount="2" stateProvince="Toliara">
<specimenCount id="9D3CFABAFFBBA923A594B0CFFCC1F91C" box="[731,775,1735,1761]" pageId="53" pageNumber="54" type="generic">
<quantity id="4CC29CD6FFBBA923A594B0CFFCC5F91C" box="[731,771,1735,1761]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="m" value="1.0">
<elevation id="0017D600FFBBA923A594B0CFFCC5F91C" box="[731,771,1735,1761]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="m" value="1.0">1m</elevation>
</quantity>
.
</specimenCount>
,
<specimenCount id="9D3CFABAFFBBA923A45BB0CFFCFEF91C" box="[788,824,1735,1761]" pageId="53" pageNumber="54" type="generic">1q.</specimenCount>
,
<collectingMunicipality id="6BE1AB49FFBBA923A40CB0CFFC21F91C" box="[835,999,1735,1761]" pageId="53" pageNumber="54">Anosy Region</collectingMunicipality>
,
<location id="8EE567E8FFBBA923A4BFB0CFFB65F91C" LSID="urn:lsid:plazi:treatment:03938025FFBBA921A7D8B3E6FA9EFEBC:8EE567E8FFBBA923A4BFB0CFFB65F91C" box="[1008,1187,1735,1761]" country="Madagascar" latitude="-24.13894" longLatPrecision="1" longitude="47.06804" municipality="Anosy Region" name="Anosyenne Mts" pageId="53" pageNumber="54" stateProvince="Toliara">Anosyenne Mts</location>
,
<locationDeviation id="9A372811FFBBA923A3E1B0CFFF18F8F8" location="Manantenina" pageId="53" pageNumber="54">
<quantity id="4CC29CD6FFBBA923A3E1B0CFFACCF91C" box="[1198,1290,1735,1761]" metricMagnitude="4" metricUnit="m" metricValue="3.12" pageId="53" pageNumber="54" unit="km" value="31.2">31.2 km</quantity>
NW Manantenina
</locationDeviation>
,
<geoCoordinate id="EE0E57F4FFBBA923A7A6B0E3FE9DF8F8" box="[233,347,1771,1797]" degrees="24.13894" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-24.13894">-24.13894</geoCoordinate>
<geoCoordinate id="EE0E57F4FFBBA923A62DB0E3FE08F8F8" box="[354,462,1771,1797]" degrees="47.06804" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="47.06804">47.06804</geoCoordinate>
,
<quantity id="4CC29CD6FFBBA923A696B0E2FDE9F8F8" box="[473,559,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="53" pageNumber="54" unit="m" value="1125.0">
<elevation id="0017D600FFBBA923A696B0E2FDE9F8F8" box="[473,559,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="53" pageNumber="54" unit="m" value="1125.0">1125 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FFBBA923A575B0E3FD79F8F8" box="[570,703,1771,1797]" pageId="53" pageNumber="54">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFBBA923A589B0E3FD31F8F8" box="[710,759,1771,1797]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFBBA923A473B0E3FC4DF8F8" box="[828,907,1771,1797]" pageId="53" pageNumber="54">CASC</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D3CFABAFFBBA923A4D0B0E3FC0EF8F8" box="[927,968,1771,1797]" pageId="53" pageNumber="54" type="generic">2w.</specimenCount>
, 1s.,
<specimenCount id="9D3CFABAFFBBA923A34EB0E3FBEBF8F8" box="[1025,1069,1771,1797]" pageId="53" pageNumber="54" type="generic">
<quantity id="4CC29CD6FFBBA923A34EB0E3FBEFF8F8" box="[1025,1065,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="m" value="1.0">1m</quantity>
<materialsCitation id="3B523B6EFFBBA923A366B0E3FBEBF8F8" ID-GBIF-Occurrence="3027954754" box="[1065,1069,1771,1797]" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="1315" latitude="-24.13632" location="Anosyenne Mts" longLatPrecision="1" longitude="47.05485" municipality="Anosy Region" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Toliara">.</materialsCitation>
</specimenCount>
,
<collectingMunicipality id="6BE1AB49FFBBA923A377B0E3FB1AF8F8" box="[1080,1244,1771,1797]" pageId="53" pageNumber="54">Anosy Region</collectingMunicipality>
,
<location id="8EE567E8FFBBA923A3AAB0E3FA5EF8F8" LSID="urn:lsid:plazi:treatment:03938025FFBBA921A7D8B3E6FA9EFEBC:8EE567E8FFBBA923A3AAB0E3FA5EF8F8" box="[1253,1432,1771,1797]" country="Madagascar" latitude="-24.13632" longLatPrecision="1" longitude="47.05485" municipality="Anosy Region" name="Anosyenne Mts" pageId="53" pageNumber="54" stateProvince="Toliara">Anosyenne Mts</location>
,
<locationDeviation id="9A372811FFBBA923A7D8B107FE02F8D4" box="[151,452,1807,1833]" location="Manantenina" pageId="53" pageNumber="54">
<quantity id="4CC29CD6FFBBA923A7D8B107FF35F8D4" box="[151,243,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="3.12" pageId="53" pageNumber="54" unit="km" value="31.2">31.2 km</quantity>
NW Manantenina
</locationDeviation>
,
<geoCoordinate id="EE0E57F4FFBBA923A680B107FD87F8D4" box="[463,577,1807,1833]" degrees="24.13632" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-24.13632">-24.13632</geoCoordinate>
<geoCoordinate id="EE0E57F4FFBBA923A506B106FD73F8D4" box="[585,693,1806,1833]" degrees="47.05485" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="47.05485">47.05485</geoCoordinate>
,
<quantity id="4CC29CD6FFBBA923A58FB106FCD1F8D4" box="[704,791,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.315" pageId="53" pageNumber="54" unit="m" value="1315.0">
<elevation id="0017D600FFBBA923A58FB106FCD1F8D4" box="[704,791,1806,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.315" pageId="53" pageNumber="54" unit="m" value="1315.0">1315 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FFBBA923A46DB107FC6EF8D4" box="[802,936,1807,1833]" pageId="53" pageNumber="54">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFBBA923A4E0B107FC26F8D4" box="[943,992,1807,1833]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFBBA923A369B107FBB3F8D4" box="[1062,1141,1807,1833]" pageId="53" pageNumber="54">CASC</collectionCode>
).
</paragraph>
<paragraph id="8B853133FFBBA923A788B13AFC92F8B0" blockId="53.[151,1436,1627,2013]" box="[199,852,1842,1869]" pageId="53" pageNumber="54">
<emphasis id="B94EED21FFBBA923A788B13AFE60F8B0" bold="true" box="[199,422,1842,1869]" pageId="53" pageNumber="54">Geographic range.</emphasis>
<collectingCountry id="F32D71A3FFBBA923A6E2B13BFDF0F8B0" box="[429,566,1843,1869]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FFBBA923A50FB13BFD54F8B0" box="[576,658,1843,1869]" country="Madagascar" name="Toliara" pageId="53" pageNumber="54">Toliara</collectingRegion>
, Anosyenne Mts.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFBBA920A788B15EFBECFE08" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="8B853133FFBBA920A788B15EFBECFE08" blockId="53.[151,1436,1627,2013]" lastBlockId="54.[151,1437,151,2013]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<emphasis id="B94EED21FFBBA923A788B15EFDC5F88C" bold="true" box="[199,515,1878,1905]" pageId="53" pageNumber="54">Diagnosis. Major workers.</emphasis>
Head in full-face view sub-rectangular, not widening posteriorly, with lateral margins slightly convex; side of head with dense, long, erect pilosity; medial frons with moderately sparse and thick rugae; interspaces between rugae smooth; frons laterally with dense rugae; interspaces between rugae reticulate; occipital lobes mostly smooth, posteriorly with sparse and weakly developed rugae; scape, when laid back, slightly exceeding the midlength of head; inner hypostomal tooth distinct, moderate and bulge-like, with top arching posteroventrally; outer hypostomal tooth bulge-like, high, and wide, with top arching posteroventrally; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum smoothly descending towards propodeum; mesonotal process very distinct and tubercle-like; mesosoma rugoreticulate; promesonotal dorsum with reduced rugoreticulae and few additional transverse and thick rugae; gaster indistinctly shagreened at base of first tergite; body brown.
<emphasis id="B94EED21FFB8A920A691B742FD5AFE98" bold="true" box="[478,668,330,357]" pageId="54" pageNumber="55">Minor workers.</emphasis>
Occiput slightly elongated; nuchal collar indistinct; head sculpture mostly smooth; frons with weakly developed and sparse puncta; scape, when laid back, exceeding the posterior head margin by three-fifths of its length; promesonotum low, moderately long, and arched; promesonotal groove distinct; metanotal groove distinct; propodeal spines small and with wide base; mesosoma mostly smooth; pronotum, mesonotum, and propodeum with weakly developed and sparse puncta; body brown.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFB8A920A788B7F6FE47F91C" pageId="54" pageNumber="55" type="description">
<paragraph id="8B853133FFB8A920A788B7F6FB75FD78" blockId="54.[151,1437,151,2013]" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B7F6FDD8FDE4" bold="true" box="[199,542,510,537]" pageId="54" pageNumber="55">Description. Major workers.</emphasis>
Measurements (n=4): HL: 2.08–2.16 (2.12); HW: 2.09–2.14 (2.11); SL: 1.19– 1.24 (1.21); EL: 0.21–0.25 (0.23); WL: 1.82–1.91 (1.86); PSL: 0.33–0.37 (0.35); MTL: 1.38–1.41 (1.4); PNW: 0.81–0.97 (0.86); PTW: 0.26–0.29 (0.27); PPW: 0.69–0.76 (0.72); CI: 99.0–103.3 (100.4); SI: 56.5–59.0 (57.5); PSLI: 15.6–17.5 (16.6); PPI: 35.6–38.5 (37.3); PNI: 38.8–45.4 (40.8); MTI: 65.8–67.3 (66.5).
</paragraph>
<paragraph id="8B853133FFB8A920A788B487FBC1FAF0" blockId="54.[151,1437,151,2013]" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B487FEC8FD54" bold="true" box="[199,270,655,681]" pageId="54" pageNumber="55">Head.</emphasis>
In full-face view sub-rectangular, not widening posteriorly, with lateral sides slightly convex (
<figureCitation id="13012DB6FFB8A920A266B487FA56FD54" box="[1321,1424,655,681]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24B</figureCitation>
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth visible. Side of head with dense, long, and erect pilosity; head dorsum with relatively dense, long, decumbent to erect pilosity. Medial frons with moderately sparse and thick rugae; interspaces between rugae smooth. Frons laterally with dense rugae; interspaces between rugae reticulate. Occipital lobes mostly smooth; posteriorly with sparse and weakly developed rugae. Gena with dense and thick costulae; interspaces between costulae distinctly rugulate. Sides posterolateral from eyes with dense and thinner network of rugoreticulae. Center of clypeus shiny and smooth, lateral sides with distinct rugulae; median notch present, moderately wide and shallow; median longitudinal carina absent; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity appressed and subdecumbent (
<figureCitation id="13012DB6FFB8A920A55CB5DBFD5BFC10" box="[531,669,979,1005]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24B, D</figureCitation>
). Inner hypostomal tooth distinct, moderate and bulge-like, with top arching posteroventrally; outer hypostomal tooth bulge-like, high, and wide, with top arching posteroventrally; median tooth absent (
<figureCitation id="13012DB6FFB8A920A6DFB213FE31FBC8" box="[400,503,1051,1077]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="54" pageNumber="55">Fig. 27P</figureCitation>
).
<emphasis id="B94EED21FFB8A920A542B213FD54FBC8" bold="true" box="[525,658,1051,1077]" pageId="54" pageNumber="55">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum smoothly descending towards propodeum; mesonotal process very distinct and tubercle-like; promesonotal groove absent; metanotal groove indistinct; propodeal spines long with narrow base and acute top; humeral tubercle weakly produced (
<figureCitation id="13012DB6FFB8A920A569B28FFD4BFB5C" box="[550,653,1159,1185]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24D</figureCitation>
). Surface shiny and rugoreticulate; promesonotal dorsum with reduced rugoreticulae and few additional transverse, thick rugae. Pilosity moderately dense, moderately long, and erect (
<figureCitation id="13012DB6FFB8A920A220B2A3FF21FB14" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24D, F</figureCitation>
).
<emphasis id="B94EED21FFB8A920A7B5B2C6FE89FB15" bold="true" box="[250,335,1230,1256]" pageId="54" pageNumber="55">Gaster.</emphasis>
Shiny and indistinctly shagreened at base of first tergite; pilosity moderately sparse, long, and erect (
<figureCitation id="13012DB6FFB8A920A7EFB2FBFEE3FAF0" box="[160,293,1267,1293]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24D, F</figureCitation>
).
<emphasis id="B94EED21FFB8A920A676B2FAFE44FAF1" bold="true" box="[313,386,1266,1292]" pageId="54" pageNumber="55">Color.</emphasis>
Brown, legs and antennae slightly brighter (
<figureCitation id="13012DB6FFB8A920A43BB2FBFC3CFAF0" box="[884,1018,1267,1293]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24D, F</figureCitation>
).
</paragraph>
<paragraph id="8B853133FFB8A920A788B31EFB1BFA63" blockId="54.[151,1437,151,2013]" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B31EFDD8FACC" bold="true" box="[199,542,1302,1329]" pageId="54" pageNumber="55">Description. Minor workers.</emphasis>
Measurements (n=5): HL: 0.79–0.95 (0.87); HW: 0.57–0.67 (0.62); SL: 1.11– 1.19 (1.15); EL: 0.15–0.17 (0.16); WL: 1.13–1.32 (1.2); PSL: 0.1–0.14 (0.12); MTL: 0.95–1.09 (1.04); PNW: 0.41– 0.51 (0.46); PTW: 0.09–0.13 (0.11); PPW: 0.17–0.22 (0.19); CI: 132.0–143.4 (139.3); SI: 178.8–193.7 (185.7); PSLI: 12.8–15.5 (14.2); PPI: 50.6–70.1 (57.4); PNI: 70.7–77.0 (73.5); MTI: 164.0–177.1 (167.3).
</paragraph>
<paragraph id="8B853133FFB8A920A788B3AFFE47F91C" blockId="54.[151,1437,151,2013]" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B3AFFEC8FA3C" bold="true" box="[199,270,1447,1473]" pageId="54" pageNumber="55">Head.</emphasis>
In full-face view oval, posterior region slightly elongated; nuchal collar weakly developed (
<figureCitation id="13012DB6FFB8A920A269B3AFFA56FA3C" box="[1318,1424,1447,1473]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24A</figureCitation>
). Pilosity sparse, moderately long and subdecumbent. Sculpture shiny and mostly smooth; frons with weakly developed and sparse puncta. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by three-fifths of its length; pilosity dense, subdecumbent to erect (
<figureCitation id="13012DB6FFB8A920A660B03FFE71F9AC" box="[303,439,1591,1617]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24A, C</figureCitation>
).
<emphasis id="B94EED21FFB8A920A684B03FFD96F9AC" bold="true" box="[459,592,1591,1617]" pageId="54" pageNumber="55">Mesosoma.</emphasis>
In lateral view, promesonotum low, moderately long, and arched; promesonotal groove distinct; metanotal groove distinct; propodeal spines small and triangular, with wide base (
<figureCitation id="13012DB6FFB8A920A267B053FA56F988" box="[1320,1424,1627,1653]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24C</figureCitation>
). Sculpture mostly smooth; pronotum, mesonotum, and propodeum with weakly developed and sparse puncta. Pilosity very sparse, short, and erect (
<figureCitation id="13012DB6FFB8A920A544B0ABFD55F940" box="[523,659,1699,1725]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24C, E</figureCitation>
).
<emphasis id="B94EED21FFB8A920A5E7B0AAFD3BF941" bold="true" box="[680,765,1698,1724]" pageId="54" pageNumber="55">Gaster.</emphasis>
With sparse and erect pilosity (
<figureCitation id="13012DB6FFB8A920A328B0ABFB29F940" box="[1127,1263,1699,1725]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24C, E</figureCitation>
).
<emphasis id="B94EED21FFB8A920A24BB0AAFA8BF941" bold="true" box="[1284,1357,1698,1724]" pageId="54" pageNumber="55">Color.</emphasis>
Bright brown (
<figureCitation id="13012DB6FFB8A920A7A1B0CFFEB2F91C" box="[238,372,1735,1761]" captionStart="FIGURE 24" captionStartId="55.[151,250,1834,1858]" captionTargetBox="[151,1436,345,1811]" captionTargetId="figure-96@55.[151,1436,345,1811]" captionTargetPageId="55" captionText="FIGURE 24. Pheidole uranus, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0235036) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0704257)." figureDoi="http://doi.org/10.5281/zenodo.4405795" httpUri="https://zenodo.org/record/4405795/files/figure.png" pageId="54" pageNumber="55">Fig. 24C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFB8A920A788B0E2FEABF8D4" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="8B853133FFB8A920A788B0E2FEABF8D4" blockId="54.[151,1437,151,2013]" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B0E2FEE1F8F9" bold="true" box="[199,295,1770,1796]" pageId="54" pageNumber="55">Biology.</emphasis>
The species was collected between
<quantity id="4CC29CD6FFB8A920A5F5B0E2FC92F8F8" box="[698,852,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.22" metricValueMax="1.315" metricValueMin="1.125" pageId="54" pageNumber="55" unit="m" value="1220.0" valueMax="1315.0" valueMin="1125.0">1125–1315 m</quantity>
elevation, in rainforest. Nests were located in rotten logs and root mats.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFB8A921A788B13AFED3FEE0" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8B853133FFB8A921A788B13AFED3FEE0" blockId="54.[151,1437,151,2013]" lastBlockId="55.[151,1437,151,321]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis id="B94EED21FFB8A920A788B13AFE89F8B1" bold="true" box="[199,335,1842,1868]" pageId="54" pageNumber="55">Comments.</emphasis>
<taxonomicName id="4C3A4AB0FFB8A920A617B13BFDD4F8B1" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[344,530,1843,1869]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FFB8A920A617B13BFDD4F8B1" box="[344,530,1843,1869]" italics="true" pageId="54" pageNumber="55">Pheidole uranus</emphasis>
</taxonomicName>
is known only from the Anosyenne Mts. in
<collectingRegion id="49FEFFD1FFB8A920A343B13BFB99F8B0" box="[1036,1119,1843,1869]" country="Madagascar" name="Toliara" pageId="54" pageNumber="55">Toliara</collectingRegion>
. Its major workers are most similar to two species known from northern parts of the island:
<taxonomicName id="4C3A4AB0FFB8A920A428B15FFC38F88C" authorityName="Forel" authorityYear="1892" box="[871,1022,1879,1905]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFB8A920A428B15FFC38F88C" box="[871,1022,1879,1905]" italics="true" pageId="54" pageNumber="55">P. madecassa</emphasis>
</taxonomicName>
and sympatric
<emphasis id="B94EED21FFB8A920A3E3B15FFACCF88D" box="[1196,1290,1879,1904]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="4C3A4AB0FFB8A920A3E3B15FFAC1F88D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1196,1287,1879,1904]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="maro">P. maro</taxonomicName>
.
</emphasis>
Majors of
<taxonomicName id="4C3A4AB0FFB8A920A2C7B15FFF23F869" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FFB8A920A2C7B15FFF23F869" italics="true" pageId="54" pageNumber="55">P. uranus</emphasis>
</taxonomicName>
can be easily separated based on presence of dense rugae with reticulate interspaces on lateral frons, sparse and weakly developed rugae on posterior occipital lobes, dense pilosity on side of head, and darker body coloration. Majors of
<taxonomicName id="4C3A4AB0FFB8A920A645B1CBFE66F820" authorityName="Forel" authorityYear="1892" box="[266,416,1987,2013]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFB8A920A645B1CBFE66F820" box="[266,416,1987,2013]" italics="true" pageId="54" pageNumber="55">P. madecassa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FFB8A920A69AB1CBFDECF821" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[469,554,1987,2012]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FFB8A920A69AB1CBFDECF821" box="[469,554,1987,2012]" italics="true" pageId="54" pageNumber="55">P. maro</emphasis>
</taxonomicName>
have lateral frons with weaker sculpture, posterior occipital lobes with reduced or absent sculpture, sparser pilosity on side of head, and orange to brownish orange body. Minors of
<taxonomicName id="4C3A4AB0FFB9A921A246B69FFAB5FF4D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1289,1395,151,176]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FFB9A921A246B69FFAB5FF4D" box="[1289,1395,151,176]" italics="true" pageId="55" pageNumber="56">P. uranus</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="4C3A4AB0FFB9A921A605B6B3FE11FF28" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[330,471,187,213]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFB9A921A605B6B3FE11FF28" box="[330,471,187,213]" italics="true" pageId="55" pageNumber="56">P. ovalinoda</emphasis>
</taxonomicName>
and can be separated based on more elongated posterior portion of head, and mostly smooth mesosoma. Minors of
<taxonomicName id="4C3A4AB0FFB9A921A6BFB6D7FDBBFF04" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[496,637,223,249]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FFB9A921A6BFB6D7FDBBFF04" box="[496,637,223,249]" italics="true" pageId="55" pageNumber="56">P. ovalinoda</emphasis>
</taxonomicName>
have less elongated posterior portion of head, and partially sculptured mesosoma.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFB9A921A788B72FFA9EFEBC" box="[199,1368,295,321]" pageId="55" pageNumber="56" type="etymology">
<paragraph id="8B853133FFB9A921A788B72FFA9EFEBC" blockId="55.[151,1437,151,321]" box="[199,1368,295,321]" pageId="55" pageNumber="56">
<emphasis id="B94EED21FFB9A921A788B72FFE88FEBC" bold="true" box="[199,334,295,321]" pageId="55" pageNumber="56">Etymology.</emphasis>
Named after the Greek god of the sky, in reference to the high elevation of the
<typeStatus id="54818F91FFB9A921A388B72FFB31FEBC" box="[1223,1271,295,321]" pageId="55" pageNumber="56">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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