293 lines
40 KiB
XML
293 lines
40 KiB
XML
<document id="4C65BBACC06A6B4224D33E3C4794CCCB" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="03938025FF80A906A7D8B462FD95FC34" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole atsirakambiaty Salata & Fisher 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="15" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:mods id="F662F1AAB3E3279BFA9D09E139F74564" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="72B902B06C28A26143FA0B4C6154032A">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:name id="93192D411FC5FA5EBDA2744D2B345019" type="personal">
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<mods:roleTerm id="C394EA25E871421F4E5301CDD4F32202">Author</mods:roleTerm>
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<mods:namePart id="CBEF77126757C5A04753D7FDE95E0CDB">Salata, Sebastian</mods:namePart>
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<mods:name id="C7820969607487D9373B5ED291A4039D" type="personal">
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<mods:roleTerm id="F45C99084741DBFE82CAC65FB2794516">Author</mods:roleTerm>
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<mods:namePart id="E8D2A9159D5FBE1BF227207453228091">Fisher, Brian L.</mods:namePart>
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<mods:nameIdentifier id="CE2DEB22FD262A5C36711B94A713F0DD" type="ORCID">0000-0002-4653-3270</mods:nameIdentifier>
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<mods:nameIdentifier id="4B2FED0F1126D2BE8580DBF52C29C661" type="email">bfisher@calacademy.org</mods:nameIdentifier>
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<mods:typeOfResource id="5EE944C7A001A5F728E4C4BCBEB98EB8">text</mods:typeOfResource>
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<mods:title id="F75DF45CD35F6B226B7B8237C37ACCBE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7B6B1AB2C0E9071F28DF8AD0372641A4">
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<mods:date id="E6DE7F323F3E52A1567A2B642411F020">2020</mods:date>
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<mods:number id="ADA3E1C65C4A7D5ED30E4755A754C14C">2020-08-31</mods:number>
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<mods:detail id="6A8B2B99C61E44853E04F638DE08D5A9" type="volume">
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<mods:number id="D845CB3A3E53C009861979EFA0470DBA">4843</mods:number>
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<mods:detail id="95BAA23BCE2D50290839F79F9ED4F28E" type="issue">
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<mods:identifier id="3260BF365CCF271C68AB02CD3757155F" type="CLB-Dataset">8556</mods:identifier>
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<mods:identifier id="7F785E83418497CBFB5B373EF2737EEC" type="DOI">10.11646/zootaxa.4843.1.1</mods:identifier>
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<mods:identifier id="FE2D3DB13F9AAB95A708AEFE496535BD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8B5956A7036F85317835B66A6646FF22" type="Zenodo-Dep">4405729</mods:identifier>
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<treatment id="03938025FF80A906A7D8B462FD95FC34" ID-DOI="http://doi.org/10.5281/zenodo.4488379" ID-GBIF-Taxon="177202129" ID-Zenodo-Dep="4488379" LSID="urn:lsid:plazi:treatment:03938025FF80A906A7D8B462FD95FC34" httpUri="http://treatment.plazi.org/id/03938025FF80A906A7D8B462FD95FC34" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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<subSubSection id="C32062B8FF80A918A7D8B462FDE7FD78" box="[151,545,618,645]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8B853133FF80A918A7D8B462FDE7FD78" blockId="14.[151,545,618,681]" box="[151,545,618,645]" pageId="14" pageNumber="15">
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<heading id="D0CD865FFF80A918A7D8B462FDE7FD78" bold="true" box="[151,545,618,645]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
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<emphasis id="B94EED21FF80A918A7D8B462FDE7FD78" bold="true" box="[151,545,618,645]" pageId="14" pageNumber="15">
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<taxonomicName id="4C3A4AB0FF80A918A7D8B462FE7DFD78" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" box="[151,443,618,645]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="atsirakambiaty" status="sp. nov.">
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<emphasis id="B94EED21FF80A918A7D8B462FE7DFD78" bold="true" box="[151,443,618,645]" italics="true" pageId="14" pageNumber="15">Pheidole atsirakambiaty</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27D505AFF80A918A68CB463FDE7FD78" box="[451,545,619,645]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C32062B8FF80A918A7D8B487FE55FD54" box="[151,403,655,681]" pageId="14" pageNumber="15" type="description">
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<paragraph id="8B853133FF80A918A7D8B487FE55FD54" blockId="14.[151,545,618,681]" box="[151,403,655,681]" pageId="14" pageNumber="15">
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<figureCitation id="13012DB6FF80A918A7D8B487FF2EFD54" box="[151,232,655,681]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="14" pageNumber="15">Figs 10</figureCitation>
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A–F, 27B, 28B
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32062B8FF80A918A7D8B4DFFC2CFC10" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8B853133FF80A918A7D8B4DFFF2BFC7C" blockId="14.[151,1437,726,2013]" pageId="14" pageNumber="15">
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<materialsCitation id="3B523B6EFF80A918A7D8B4DFFE13FCC4" ID-GBIF-Occurrence="3027954757" accessionNumber="BLF07274" collectingDate="2003-01-22" collectionCode="CASC" country="Madagascar" elevation="1550" latitude="-20.59333" location="Foret d'Atsirakambiaty" longLatPrecision="1" longitude="46.56333" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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||
<typeStatus id="54818F91FF80A918A7D8B4DFFEE9FD0C" box="[151,303,727,753]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
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||
: 1s.,
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||
<collectingCountry id="F32D71A3FF80A918A63BB4DFFE3BFD0C" box="[372,509,727,753]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
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,
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||
<collectingRegion id="49FEFFD1FF80A918A540B4DFFD67FD0C" box="[527,673,727,753]" country="Madagascar" name="Fianarantsoa" pageId="14" pageNumber="15">Fianarantsoa</collectingRegion>
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||
,
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||
<location id="8EE567E8FF80A918A5FDB4DFFC06FD0C" LSID="urn:lsid:plazi:treatment:03938025FF80A906A7D8B462FD95FC34:8EE567E8FF80A918A5FDB4DFFC06FD0C" box="[690,960,727,753]" country="Madagascar" latitude="-20.59333" longLatPrecision="1" longitude="46.56333" name="Foret d'Atsirakambiaty" pageId="14" pageNumber="15" stateProvince="Fianarantsoa">Forêt d’Atsirakambiaty</location>
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,
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||
<locationDeviation id="9A372811FF80A918A49EB4DFFADFFD0C" box="[977,1305,726,753]" location="Itremo" pageId="14" pageNumber="15">
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<quantity id="4CC29CD6FF80A918A49EB4DFFBE0FD0C" box="[977,1062,727,753]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="15" unit="km" value="7.6">7.6 km</quantity>
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||
285° WNW Itremo
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||
</locationDeviation>
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,
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<geoCoordinate id="EE0E57F4FF80A918A265B4DEFA5AFD0C" box="[1322,1436,726,753]" degrees="20.59333" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-20.59333">-20.59333</geoCoordinate>
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<geoCoordinate id="EE0E57F4FF80A918A7D8B4F2FEC5FCE8" box="[151,259,762,789]" degrees="46.56333" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="46.56333">46.56333</geoCoordinate>
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,
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<quantity id="4CC29CD6FF80A918A65EB4F2FEAAFCE8" box="[273,364,762,789]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="m" value="1550.0">
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<elevation id="0017D600FF80A918A65EB4F2FEAAFCE8" box="[273,364,762,789]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="m" value="1550.0">1550 m</elevation>
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</quantity>
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,
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<date id="FF8417F3FF80A918A634B4F3FDCFFCE8" box="[379,521,763,789]" pageId="14" pageNumber="15" value="2003-01-22">
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<collectingDate id="EFC0EE1BFF80A918A634B4F3FDCFFCE8" box="[379,521,763,789]" pageId="14" pageNumber="15" value="2003-01-22">22-Jan-2003</collectingDate>
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</date>
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||
, montane rainforest, ex rotten log/termite mound, Fisher
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||
<emphasis id="B94EED21FF80A918A3ECB4F3FB1EFCE8" box="[1187,1240,763,789]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
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||
. leg. BLF07274, CASENT0491660 (
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<collectionCode id="ED2BA9F6FF80A918A631B517FE0BFCC4" box="[382,461,799,825]" pageId="14" pageNumber="15">CASC</collectionCode>
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)
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||
</materialsCitation>
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||
.
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||
<typeStatus id="54818F91FF80A918A6A6B517FD4EFCC4" box="[489,648,799,825]" pageId="14" pageNumber="15" type="paratype">PARATYPES</typeStatus>
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||
:
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||
<specimenCount id="9D3CFABAFF80A918A5D4B517FD03FCC4" box="[667,709,799,825]" pageId="14" pageNumber="15" type="generic">6w.</specimenCount>
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||
, 2s.,
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||
<materialsCitation id="3B523B6EFF80A918A44EB517FF2FFC7C" ID-GBIF-Occurrence="3027954500" accessionNumber="BLF07274" collectingDate="2003-01-22" collectionCode="CASC, MHNG, PBZT" country="Madagascar" elevation="1" latitude="-20.59333" location="Foret d'Atsirakambiaty" longLatPrecision="1" longitude="46.56333" pageId="14" pageNumber="15" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
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||
<specimenCount id="9D3CFABAFF80A918A44EB517FCE8FCC4" box="[769,814,799,825]" pageId="14" pageNumber="15" type="generic">
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||
<quantity id="4CC29CD6FF80A918A44EB517FCECFCC4" box="[769,810,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="1.0">
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||
<elevation id="0017D600FF80A918A44EB517FCECFCC4" box="[769,810,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="1.0">1m</elevation>
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||
</quantity>
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||
.
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||
</specimenCount>
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,
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<specimenCount id="9D3CFABAFF80A918A47BB517FC90FCC4" box="[820,854,799,825]" pageId="14" pageNumber="15" type="generic">1q.</specimenCount>
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||
, the same data as holotype, CASENT0491661, CASENT0491664, CASENT0491665, CASENT0491666, CASENT0872258, CASENT0491662 (
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||
<collectionCode id="ED2BA9F6FF80A918A3A8B54BFAF2FCA0" box="[1255,1332,835,861]" pageId="14" pageNumber="15">CASC</collectionCode>
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,
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||
<collectionCode id="ED2BA9F6FF80A918A20FB54BFA50FCA0" box="[1344,1430,835,861]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="15" type="Museum">MHNG</collectionCode>
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||
,
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<collectionCode id="ED2BA9F6FF80A918A7D8B56FFF27FC7C" box="[151,225,871,897]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="14" pageNumber="15">PBZT</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B853133FF80A918A788B582FCC6FC34" blockId="14.[151,1437,726,2013]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EED21FF80A918A788B582FE42FC59" bold="true" box="[199,388,906,932]" pageId="14" pageNumber="15">Other material.</emphasis>
|
||
<materialsCitation id="3B523B6EFF80A918A6C2B583FD3AFC34" ID-GBIF-Occurrence="3027954317" collectionCode="CASC" country="Madagascar" elevation="1550" latitude="-20.59333" location="Foret d'Atsirakambiaty" longLatPrecision="1" longitude="46.56333" pageId="14" pageNumber="15" specimenCount="11" stateProvince="Fianarantsoa">
|
||
<collectingCountry id="F32D71A3FF80A918A6C2B583FDD0FC58" box="[397,534,907,933]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
|
||
.
|
||
<collectingRegion id="49FEFFD1FF80A918A56CB583FD71FC58" box="[547,695,907,933]" country="Madagascar" name="Fianarantsoa" pageId="14" pageNumber="15">Fianarantsoa</collectingRegion>
|
||
:
|
||
<specimenCount id="9D3CFABAFF80A918A58CB583FD3CFC58" box="[707,762,907,933]" pageId="14" pageNumber="15" type="generic">11w.</specimenCount>
|
||
, 3s.,
|
||
<location id="8EE567E8FF80A918A477B583FB87FC58" LSID="urn:lsid:plazi:treatment:03938025FF80A906A7D8B462FD95FC34:8EE567E8FF80A918A477B583FB87FC58" box="[824,1089,907,933]" country="Madagascar" latitude="-20.59333" longLatPrecision="1" longitude="46.56333" name="Foret d'Atsirakambiaty" pageId="14" pageNumber="15" stateProvince="Fianarantsoa">Forêt d’Atsirakambiaty</location>
|
||
,
|
||
<locationDeviation id="9A372811FF80A918A300B583FA40FC58" box="[1103,1414,906,933]" location="Itremo" pageId="14" pageNumber="15">
|
||
<quantity id="4CC29CD6FF80A918A300B583FB59FC58" box="[1103,1183,907,933]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="15" unit="km" value="7.6">7.6 km</quantity>
|
||
285° WNW Itremo
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE0E57F4FF80A918A2DCB583FEC6FC34" degrees="20.59333" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-20.59333">- 20.59333</geoCoordinate>
|
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<geoCoordinate id="EE0E57F4FF80A918A648B5A6FEB5FC34" box="[263,371,942,969]" degrees="46.56333" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="46.56333">46.56333</geoCoordinate>
|
||
,
|
||
<quantity id="4CC29CD6FF80A918A631B5A6FE13FC34" box="[382,469,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="m" value="1550.0">
|
||
<elevation id="0017D600FF80A918A631B5A6FE13FC34" box="[382,469,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="m" value="1550.0">1550 m</elevation>
|
||
</quantity>
|
||
, Fisher
|
||
<emphasis id="B94EED21FF80A918A561B5A7FD99FC34" box="[558,607,943,969]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
|
||
. leg. (
|
||
<collectionCode id="ED2BA9F6FF80A918A5EAB5A7FD32FC34" box="[677,756,943,969]" pageId="14" pageNumber="15">CASC</collectionCode>
|
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)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B853133FF80A918A788B5DAFC2CFC10" blockId="14.[151,1437,726,2013]" box="[199,1002,978,1005]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EED21FF80A918A788B5DAFE60FC10" bold="true" box="[199,422,978,1005]" pageId="14" pageNumber="15">Geographic range.</emphasis>
|
||
<collectingCountry id="F32D71A3FF80A918A6E2B5DBFDF0FC10" box="[429,566,979,1005]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
|
||
,
|
||
<collectingRegion id="49FEFFD1FF80A918A50EB5DBFD15FC10" box="[577,723,979,1005]" country="Madagascar" name="Fianarantsoa" pageId="14" pageNumber="15">Fianarantsoa</collectingRegion>
|
||
, Forêt d’Atsirakambiaty.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32062B8FF80A918A788B5FEFA4BFA3C" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="8B853133FF80A918A788B5FEFA4BFA3C" blockId="14.[151,1437,726,2013]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EED21FF80A918A788B5FEFDCEFBEC" bold="true" box="[199,520,1014,1041]" pageId="14" pageNumber="15">Diagnosis. Major workers.</emphasis>
|
||
Head in full-face view sub-rectangular, not widening posteriorly, lateral margin relatively straight; sides of head with moderately dense, moderately long, suberect pilosity; frons with moderately thick and moderately dense costulae that arch posterolaterally; interspaces between costulae smooth to indistinctly rugopunctate; occipital lobes with thin and sparse rugulae that arch posterolaterally; interspaces between rugulae sparsely rugopunctate; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal tooth absent; outer hypostomal tooth lobe-like, high, and wide, with top arching posterolaterally; median tooth present; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process tubercle-like; promesonotum with thick, dense and transverse rugae; interspaces between rugae distinctly punctate; katepisternum and propodeum with sparse rugae; interspaces between rugae rugopunctate; gaster shagreened; body black.
|
||
<emphasis id="B94EED21FF80A918A682B332FD41FAA8" bold="true" box="[461,647,1338,1365]" pageId="14" pageNumber="15">Minor workers.</emphasis>
|
||
Occiput slightly elongated; nuchal collar distinct; head mostly smooth; frons rugopunctate; scape, when laid back, exceeding posterior head margin by half its length; promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines moderately small with wide base; mesosoma punctate; promesonotal dorsum with sparser puncta and transverse, thin rugae; body black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32062B8FF80A906A788B3C2FB6AFDC0" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8B853133FF80A918A788B3C2FB74F9AF" blockId="14.[151,1437,726,2013]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EED21FF80A918A788B3C2FDE6FA18" bold="true" box="[199,544,1482,1509]" pageId="14" pageNumber="15">Description. Major workers.</emphasis>
|
||
Measurements (n=7): HL: 1.88–2.1 (1.99); HW: 1.75–1.97 (1.86); SL: 1.21– 1.25 (1.24); EL: 0.23–0.29 (0.26); WL: 1.62–1.74 (1.69); PSL: 0.23–0.32 (0.28); MTL: 1.36–1.49 (1.42); PNW: 0.59–0.74 (0.64); PTW: 0.19–0.31 (0.23); PPW: 0.52–0.73 (0.6); CI: 103.4–110.8 (107.0); SI: 61.5–71.5 (66.7); PSLI: 11.2–15.9 (14.0); PPI: 32.2–42.7 (39.3); PNI: 32.6–37.6 (34.5); MTI: 72.1–79.2 (76.1).
|
||
</paragraph>
|
||
<paragraph id="8B853133FF80A919A788B053FB9FFE50" blockId="14.[151,1437,726,2013]" lastBlockId="15.[151,1437,151,429]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EED21FF80A918A788B053FEC8F988" bold="true" box="[199,270,1627,1653]" pageId="14" pageNumber="15">Head.</emphasis>
|
||
In full-face view sub-rectangular, not widening posteriorly, lateral margin relatively straight (
|
||
<figureCitation id="13012DB6FF80A918A266B053FA49F988" box="[1321,1423,1627,1653]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="14" pageNumber="15">Fig. 10B</figureCitation>
|
||
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth not visible. Side of the head with moderately dense, moderately long, suberect pilosity; head dorsum with relatively dense, long, decumbent to erect pilosity. Frons with moderately thick and moderately dense costulae that arch posterolaterally; interspaces between costulae smooth to indistinctly rugopunctate. Occipital lobe with thin and sparse rugulae that arch posterolaterally; interspaces between rugulae sparsely rugopunctate. Gena with dense and thick costulae; interspaces between costulae indistinctly rugopunctate. Sides posterolateral from eyes with dense but thin rugocostulae; interspaces between rugocostulae rugopunctate. Center of clypeus shiny and indistinctly punctate, lateral sides with distinct rugulae; median notch present, moderately wide and shallow; median and lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (
|
||
<figureCitation id="13012DB6FF80A918A6E7B1CBFDF3F820" box="[424,565,1987,2013]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="14" pageNumber="15">Fig. 10B, D</figureCitation>
|
||
). Inner hypostomal tooth absent; outer hypostomal tooth lobe-like, high and wide, with top arching posterolaterally; median tooth present (
|
||
<figureCitation id="13012DB6FF81A919A408B69FFC6BFF4C" box="[839,941,151,177]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="15" pageNumber="16">Fig. 27B</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF81A919A4F0B69FFB82FF4C" bold="true" box="[959,1092,151,177]" pageId="15" pageNumber="16">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tuberclelike; promesonotal groove absent; metanotal groove present; propodeal spines moderate with wide base and acute top; humeral tubercle weakly produced (
|
||
<figureCitation id="13012DB6FF81A919A512B70BFD03FEE0" box="[605,709,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16">Fig. 10D</figureCitation>
|
||
). Surface shiny; promesonotum with thick, dense, and transverse rugae; interspaces between rugae distinctly punctate; katepisternum and propodeum with sparse rugae; interspaces between rugae rugopunctate. Pilosity moderately dense, long, and erect (
|
||
<figureCitation id="13012DB6FF81A919A49BB743FB9BFE98" box="[980,1117,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16">Fig. 10D, F</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF81A919A33DB742FB0CFE99" bold="true" box="[1138,1226,330,356]" pageId="15" pageNumber="16">Petiole.</emphasis>
|
||
In rear view node dorsoventrally convex (
|
||
<figureCitation id="13012DB6FF81A919A6D0B767FDE2FE74" box="[415,548,367,393]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16">Fig. 10D, F</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF81A919A577B766FD4BFE75" bold="true" box="[568,653,366,392]" pageId="15" pageNumber="16">Gaster.</emphasis>
|
||
Dull and shagreened; pilosity moderately sparse, long, and erect (
|
||
<figureCitation id="13012DB6FF81A919A220B767FF2EFE50" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16">Fig. 10D, F</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF81A919A7B3B79AFE83FE51" bold="true" box="[252,325,402,428]" pageId="15" pageNumber="16">Color.</emphasis>
|
||
Black, antennae and legs brown, mandibles ochraceous (
|
||
<figureCitation id="13012DB6FF81A919A489B79BFB8AFE50" box="[966,1100,403,429]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16">Fig. 10D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4561BBFF81A919A7D8B1ACFABCF81D" ID-DOI="http://doi.org/10.5281/zenodo.4405757" ID-Zenodo-Dep="4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1956,1980]" targetBox="[151,1436,453,1933]" targetPageId="15">
|
||
<paragraph id="8B853133FF81A919A7D8B1ACFABCF81D" blockId="15.[151,1436,1956,2017]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94EED21FF81A919A7D8B1ACFEE0F841" bold="true" box="[151,294,1956,1980]" pageId="15" pageNumber="16">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C3A4AB0FF81A919A67CB1ACFDF7F841" authorityName="Salata & Fisher" authorityYear="2020" box="[307,561,1956,1980]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="atsirakambiaty">
|
||
<emphasis id="B94EED21FF81A919A67CB1ACFDF7F841" box="[307,561,1956,1980]" italics="true" pageId="15" pageNumber="16">Pheidole atsirakambiaty</emphasis>
|
||
</taxonomicName>
|
||
, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B853133FF9EA906A788B69EFB1BFEE0" blockId="16.[151,1437,150,969]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EED21FF9EA906A788B69EFDDAFF4C" bold="true" box="[199,540,150,177]" pageId="16" pageNumber="17">Description. Minor workers.</emphasis>
|
||
Measurements (n=10): HL: 0.85–0.95 (0.91); HW: 0.59–0.71 (0.65); SL: 1.13– 1.29 (1.23); EL: 0.16–0.2 (0.18); WL: 1.16–1.29 (1.23); PSL: 0.12–0.15 (0.13); MTL: 1.07–1.21 (1.13); PNW: 0.41–0.47 (0.44); PTW: 0.1–0.14 (0.12); PPW: 0.17–0.22 (0.2); CI: 133.4–143.6 (139.1); SI: 180.3–196.8 (189.1); PSLI: 13.0–16.5 (14.8); PPI: 47.7–68.3 (59.2); PNI: 64.1–72.1 (67.9); MTI: 163.4–183.0 (173.8).
|
||
</paragraph>
|
||
<paragraph id="8B853133FF9EA906A788B72FFB6AFDC0" blockId="16.[151,1437,150,969]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EED21FF9EA906A788B72FFEC8FEBC" bold="true" box="[199,270,295,321]" pageId="16" pageNumber="17">Head.</emphasis>
|
||
In full-face view oval, occiput slightly elongated; nuchal collar distinct (
|
||
<figureCitation id="13012DB6FF9EA906A31BB72FFB78FEBC" box="[1108,1214,295,321]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10A</figureCitation>
|
||
). Pilosity relatively dense, moderately long, subdecumbent to erect. Sculpture shining and smooth; frons rugopunctate. Clypeus with median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, exceeding posterior head margin by half its length; pilosity dense, subdecumbent to erect (
|
||
<figureCitation id="13012DB6FF9EA906A49EB79BFB9EFE50" box="[977,1112,403,429]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10A, C</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF9EA906A324B79BFB36FE50" bold="true" box="[1131,1264,403,429]" pageId="16" pageNumber="17">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum low, long and arched; promesonotal groove present; metanotal groove distinct; propodeal spines moderately small, triangular and with wide base (
|
||
<figureCitation id="13012DB6FF9EA906A584B7D3FCF2FE08" box="[715,820,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10C</figureCitation>
|
||
). Surface punctate; promesonotal dorsum with sparser puncta and thin, transverse rugae. Pilosity very sparse, long, and erect (
|
||
<figureCitation id="13012DB6FF9EA906A4FAB7F7FBFAFDE4" box="[949,1084,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10C, E</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF9EA906A300B7F6FB63FDE5" bold="true" box="[1103,1189,510,536]" pageId="16" pageNumber="17">Gaster.</emphasis>
|
||
With sparse and erect pilosity (
|
||
<figureCitation id="13012DB6FF9EA906A7B4B42BFE47FDC0" box="[251,385,547,573]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10C, E</figureCitation>
|
||
).
|
||
<emphasis id="B94EED21FF9EA906A6DBB42AFE1BFDC1" bold="true" box="[404,477,546,572]" pageId="16" pageNumber="17">Color.</emphasis>
|
||
Black, with legs, mandibles, and antennae brown (
|
||
<figureCitation id="13012DB6FF9EA906A356B42BFB59FDC0" box="[1049,1183,547,573]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="16" pageNumber="17">Fig. 10C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32062B8FF9EA906A788B44EFCE1FD78" pageId="16" pageNumber="17" type="biology_ecology">
|
||
<paragraph id="8B853133FF9EA906A788B44EFCE1FD78" blockId="16.[151,1437,150,969]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EED21FF9EA906A788B44EFEE1FD9D" bold="true" box="[199,295,582,608]" pageId="16" pageNumber="17">Biology.</emphasis>
|
||
The specimens were collected at
|
||
<quantity id="4CC29CD6FF9EA906A5E7B44EFD38FD9C" box="[680,766,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="16" pageNumber="17" unit="m" value="1550.0">1550 m</quantity>
|
||
elevation, in montane rainforest and grassland. Nests were located in rotten logs on termite mounds and under stones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32062B8FF9EA906A788B486FD45FC58" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8B853133FF9EA906A788B486FD45FC58" blockId="16.[151,1437,150,969]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EED21FF9EA906A788B486FE89FD55" bold="true" box="[199,335,654,680]" pageId="16" pageNumber="17">Comments.</emphasis>
|
||
Considering the ants known from northern
|
||
<collectingCountry id="F32D71A3FF9EA906A40CB487FC0AFD54" box="[835,972,655,681]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3A4AB0FF9EA906A496B487FB2AFD54" authorityName="Salata & Fisher" authorityYear="2020" box="[985,1260,655,681]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="atsirakambiaty">
|
||
<emphasis id="B94EED21FF9EA906A496B487FB2AFD54" box="[985,1260,655,681]" italics="true" pageId="16" pageNumber="17">Pheidole atsirakambiaty</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C3A4AB0FF9EA906A7F9B4BBFEE2FD31" authorityName="Salata & Fisher" authorityYear="2020" box="[182,292,691,716]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="voreios">
|
||
<emphasis id="B94EED21FF9EA906A7F9B4BBFEE2FD31" box="[182,292,691,716]" italics="true" pageId="16" pageNumber="17">P. voreios</emphasis>
|
||
</taxonomicName>
|
||
and parapatric
|
||
<taxonomicName id="4C3A4AB0FF9EA906A698B4BBFD96FD30" authorityName="Forel" authorityYear="1891" box="[471,592,691,717]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bessonii">
|
||
<emphasis id="B94EED21FF9EA906A698B4BBFD96FD30" box="[471,592,691,717]" italics="true" pageId="16" pageNumber="17">P. bessonii</emphasis>
|
||
</taxonomicName>
|
||
. Its majors can be separated based on the black body and the presence of moderately thick, moderately dense costulae that arch posterolaterally, and rugulate frons and occipital lobes with smooth to indistinctly rugopunctate interspaces between rugulae. Majors of
|
||
<taxonomicName id="4C3A4AB0FF9EA906A4A0B4F3FB9AFCE9" authorityName="Salata & Fisher" authorityYear="2020" box="[1007,1116,763,788]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="voreios">
|
||
<emphasis id="B94EED21FF9EA906A4A0B4F3FB9AFCE9" box="[1007,1116,763,788]" italics="true" pageId="16" pageNumber="17">P. voreios</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4AB0FF9EA906A3DBB4F3FACAFCE8" authorityName="Forel" authorityYear="1891" box="[1172,1292,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bessonii">
|
||
<emphasis id="B94EED21FF9EA906A3DBB4F3FACAFCE8" box="[1172,1292,763,789]" italics="true" pageId="16" pageNumber="17">P. bessonii</emphasis>
|
||
</taxonomicName>
|
||
have a body that is brown to blackish brown, their head sculpture is never arches uniformly posterolaterally, and interspaces between rugocostulae are smooth to punctate. Minors can be distinguished based on their black body and distinctly punctate mesosoma. In contrast, minors of
|
||
<taxonomicName id="4C3A4AB0FF9EA906A537B56FFD22FC7D" authorityName="Salata & Fisher" authorityYear="2020" box="[632,740,871,896]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="voreios">
|
||
<emphasis id="B94EED21FF9EA906A537B56FFD22FC7D" box="[632,740,871,896]" italics="true" pageId="16" pageNumber="17">P. voreios</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A4AB0FF9EA906A454B56FFC54FC7C" authorityName="Forel" authorityYear="1891" box="[795,914,871,897]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bessonii">
|
||
<emphasis id="B94EED21FF9EA906A454B56FFC54FC7C" box="[795,914,871,897]" italics="true" pageId="16" pageNumber="17">P. bessonii</emphasis>
|
||
</taxonomicName>
|
||
have a body that is bright brown to brown and their mesosoma is never distinctly punctate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32062B8FF9EA906A788B5A7FD95FC34" box="[199,595,943,969]" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="8B853133FF9EA906A788B5A7FD95FC34" blockId="16.[151,1437,150,969]" box="[199,595,943,969]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EED21FF9EA906A788B5A7FE88FC34" bold="true" box="[199,334,943,969]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
From the
|
||
<typeStatus id="54818F91FF9EA906A68DB5A7FE34FC34" box="[450,498,943,969]" pageId="16" pageNumber="17">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |